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<document id="4B88B282EEFB44BA263E5FE2670AFB2C" ID-CLB-Dataset="45896" ID-DOI="10.5281/zenodo.207142" ID-GBIF-Dataset="1f0ce477-206a-4ba6-912e-2822d97d6ff8" ID-ISSN="1175-5326" ID-Zenodo-Dep="207142" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460414129179" checkinUser="plazi" docAuthor="Heard, Richard W. &amp; Morales-Núñez, Andrés G." docDate="2011" docId="038587E8FFFC51102593FF74FCDBFD89" docLanguage="en" docName="zt03111p063.pdf" docOrigin="Zootaxa 3111" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Akanthoparapseudes Heard &amp; Morales-Núñez, 2011, new genus" docType="treatment" docVersion="10" lastPageNumber="52" masterDocId="FFBCFF90FFFE51142504FFE3FFCCFF9E" masterDocTitle="Akanthoparapseudes alfaroae, a new genus and species of apseudomorphan tanaidacean (Crustacea: Malacostraca: Peracarida) from Puerto Rican coastal waters" masterLastPageNumber="63" masterPageNumber="49" pageNumber="51" updateTime="1698328350723" updateUser="ExternalLinkService">
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<mods:title id="9E0BDDC97DA13351C18966F7243D02FF">Akanthoparapseudes alfaroae, a new genus and species of apseudomorphan tanaidacean (Crustacea: Malacostraca: Peracarida) from Puerto Rican coastal waters</mods:title>
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<mods:namePart id="32AE54D3E8273D003C6A7AE50CB99018">Heard, Richard W.</mods:namePart>
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<mods:namePart id="B34848BE583380B2B4A05D8B88D4D4DE">Morales-Núñez, Andrés G.</mods:namePart>
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<mods:date id="38490F8F24C0CF19554B45A09EB88135">2011</mods:date>
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<treatment id="038587E8FFFC51102593FF74FCDBFD89" ID-DOI="http://doi.org/10.5281/zenodo.6191496" ID-GBIF-Taxon="119424507" ID-Zenodo-Dep="6191496" LSID="urn:lsid:plazi:treatment:038587E8FFFC51102593FF74FCDBFD89" httpUri="http://treatment.plazi.org/id/038587E8FFFC51102593FF74FCDBFD89" lastPageId="4" lastPageNumber="52" pageId="2" pageNumber="51">
<subSubSection id="C3366575FFFC51162593FF74FDE9FF2F" box="[151,549,151,177]" pageId="2" pageNumber="51" type="nomenclature">
<paragraph id="8B9336FEFFFC51162593FF74FDE9FF2F" blockId="2.[151,549,151,177]" box="[151,549,151,177]" pageId="2" pageNumber="51">
<heading id="D0DB8192FFFC51162593FF74FDE9FF2F" bold="true" box="[151,549,151,177]" fontSize="11" level="1" pageId="2" pageNumber="51" reason="1">
<emphasis id="B958EAECFFFC51162593FF74FDE9FF2F" bold="true" box="[151,549,151,177]" pageId="2" pageNumber="51">
<emphasis id="B958EAECFFFC51162593FF74FE50FF2F" bold="true" box="[151,412,151,177]" italics="true" pageId="2" pageNumber="51">
<taxonomicName id="4C2C4D7DFFFC51162593FF74FE5AFF2F" ID-CoL="84L7Y" box="[151,406,151,177]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">Akanthoparapseudes</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26B5797FFFC511624A0FF74FDE9FF2F" box="[420,549,151,177]" pageId="2" pageNumber="51" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3366575FFFC51162593FF3DFCC9FD39" pageId="2" pageNumber="51" type="diagnosis">
<paragraph id="8B9336FEFFFC51162593FF3DFC43FD1C" blockId="2.[151,1437,222,1110]" pageId="2" pageNumber="51">
<emphasis id="B958EAECFFFC51162593FF3DFEC6FF69" bold="true" box="[151,266,222,247]" pageId="2" pageNumber="51">Diagnosis</emphasis>
(Subadult). Apseudomorpha,
<taxonomicName id="4C2C4D7DFFFC51162764FF3DFCC9FF68" box="[608,773,222,246]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="family">Parapseudidae</taxonomicName>
,
<taxonomicName id="4C2C4D7DFFFC51162615FF3DFC69FF68" box="[785,933,222,246]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="tribe" tribe="Parapseudini">Parapseudini</taxonomicName>
. Carapace without spinous processes. Triangular, apically-subacute rostrum. Epistomal spine present. Pereonites 26 with posterolateral margins armed with small, but distinct spiniform seta. Left mandible with tridentate
<emphasis id="B958EAECFFFC51162659FEC5FBC8FEA0" box="[861,1028,294,318]" italics="true" pageId="2" pageNumber="51">lacinia mobilis</emphasis>
, palp with article 1 having cluster of 10 or more simple distally-attenuated setae. Eye lobe distinct; eyes appearing poorly developed, lacking distinct pigment (in preserved specimens), ommatidia (about 8) small, poorly-formed. Maxilliped having distolateral margin of article 2 with strongly-developed spiniform seta. Pereopod 1 with strongly-developed spiniform setae on anterior margin of basis, coxa with reduced process present, not visible in dorsal view. Pereopod 2 with spine on anterior margin of basis; pereopods 26 with ischium bearing one, occasionally two narrow, acute spines on the anterior margin. Crenulate, rod-like, spiniform setae present on the merus, carpus and/or propodus of pereopods 2 6. Pereopod 4 with dactylus, not greatly reduced, but with some adjacent setae about equal or greater in length. Uropodal peduncle with subdistal and distal setae only. Pereopod 6 having long simple setae along anterior margin. First pleonite lacking distinct transverse row of small setulate setae.
</paragraph>
<paragraph id="8B9336FEFFFC511625C1FD6DFCC9FD39" blockId="2.[151,1437,222,1110]" box="[197,773,654,679]" pageId="2" pageNumber="51">
<emphasis id="B958EAECFFFC511625C1FD6DFEAEFD39" bold="true" box="[197,354,654,679]" pageId="2" pageNumber="51">Type-species.</emphasis>
<taxonomicName id="4C2C4D7DFFFC5116246DFD6DFD75FD38" box="[361,697,654,678]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="alfaroae" status="sp. nov.">
<emphasis id="B958EAECFFFC5116246DFD6DFD75FD38" box="[361,697,654,678]" italics="true" pageId="2" pageNumber="51">Akanthoparapseudes alfaroae</emphasis>
</taxonomicName>
,
<emphasis id="B958EAECFFFC511627C3FD6CFCC9FD39" bold="true" box="[711,773,655,679]" pageId="2" pageNumber="51">
<taxonomicNameLabel id="A26B5797FFFC511627C3FD6CFCC9FD39" box="[711,773,655,679]" pageId="2" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3366575FFFC511625C1FD50FECEFD70" pageId="2" pageNumber="51" type="etymology">
<paragraph id="8B9336FEFFFC511625C1FD50FECEFD70" blockId="2.[151,1437,222,1110]" pageId="2" pageNumber="51">
<emphasis id="B958EAECFFFC511625C1FD50FE87FD55" bold="true" box="[197,331,691,715]" pageId="2" pageNumber="51">Etymology.</emphasis>
<emphasis id="B958EAECFFFC51162451FD50FE7EFD55" box="[341,434,691,715]" italics="true" pageId="2" pageNumber="51">Akantho</emphasis>
, from the Greek = spiny, plus the generic name
<taxonomicName id="4C2C4D7DFFFC511626E8FD50FBB1FD55" box="[1004,1149,691,715]" class="Malacostraca" family="Parapseudidae" genus="Parapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC511626E8FD50FBB1FD55" box="[1004,1149,691,715]" italics="true" pageId="2" pageNumber="51">Parapseudes</emphasis>
</taxonomicName>
, from the Greek; gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3366575FFFC511625C1FD18FE10FBC8" pageId="2" pageNumber="51" type="discussion">
<paragraph id="8B9336FEFFFC511625C1FD18FE10FBC8" blockId="2.[151,1437,222,1110]" pageId="2" pageNumber="51">
<emphasis id="B958EAECFFFC511625C1FD18FEF5FC8D" bold="true" box="[197,313,763,787]" pageId="2" pageNumber="51">Remarks.</emphasis>
Although
<taxonomicName id="4C2C4D7DFFFC511624C1FD18FD7CFC8D" box="[453,688,763,787]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B958EAECFFFC511624C1FD18FD7CFC8D" box="[453,688,763,787]" italics="true" pageId="2" pageNumber="51">Akanthoparapseudes</emphasis>
</taxonomicName>
,
<emphasis id="B958EAECFFFC511627CCFD18FCE5FC8D" bold="true" box="[712,809,763,787]" pageId="2" pageNumber="51">
<taxonomicNameLabel id="A26B5797FFFC511627CCFD18FCEEFC8D" box="[712,802,763,787]" pageId="2" pageNumber="51" rank="genus">n. gen.</taxonomicNameLabel>
,
</emphasis>
also appears to share several characters with the
<taxonomicName id="4C2C4D7DFFFC51162593FCFDFED2FCA8" box="[151,286,798,822]" class="Malacostraca" family="Apseudidae" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="family">Apseudidae</taxonomicName>
, with reservations we refer it to the
<taxonomicName id="4C2C4D7DFFFC511627C2FCFDFCA7FCA8" box="[710,875,798,822]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="family">Parapseudidae</taxonomicName>
. Previously 22 genera have been referred to this family (see
<bibRefCitation id="EFBD4B0FFFFC5116241FFCA1FE07FCC4" author="Anderson" box="[283,459,834,858]" pageId="2" pageNumber="51" refString="Anderson, G. 2011, May 11. Tanaidacea Taxa and Literature http: // percarida. usm. edu /&gt;." type="book chapter" year="2011">Anderson 2011</bibRefCitation>
). Of these genera,
<taxonomicName id="4C2C4D7DFFFC51162798FCA0FC4BFCC5" box="[668,903,835,859]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC51162798FCA0FC4BFCC5" box="[668,903,835,859]" italics="true" pageId="2" pageNumber="51">Akanthoparapseudes</emphasis>
</taxonomicName>
appears, at least superficially, to have affinities with
<taxonomicName id="4C2C4D7DFFFC511625D4FC85FE2EFCE0" authority="Gutu, 2006" authorityName="Gutu" authorityYear="2006" box="[208,482,870,895]" class="Malacostraca" family="Parapseudidae" genus="Brachylicoa" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC511625D4FC85FE95FCE0" box="[208,345,870,894]" italics="true" pageId="2" pageNumber="51">Brachylicoa</emphasis>
<bibRefCitation id="EFBD4B0FFFFC51162464FC84FE2EFCE0" author="Gutu" box="[352,482,870,895]" pageId="2" pageNumber="51" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C4D7DFFFC511624E9FC85FC0DFCE0" authority="Larsen and Shimomura, 2006" authorityName="Larsen and Shimomura" authorityYear="2006" box="[493,961,870,894]" class="Malacostraca" family="Parapseudidae" genus="Longipedis" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC511624E9FC85FDA6FCE0" box="[493,618,870,894]" italics="true" pageId="2" pageNumber="51">Longipedis</emphasis>
Larsen and Shimomura, 2006
</taxonomicName>
, and
<taxonomicName id="4C2C4D7DFFFC511626F8FC85FAC5FCE0" authority="Gutu, 2006" authorityName="Gutu" authorityYear="2006" box="[1020,1289,870,895]" class="Malacostraca" family="Parapseudidae" genus="Podictenius" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC511626F8FC85FB4CFCE0" box="[1020,1152,870,894]" italics="true" pageId="2" pageNumber="51">Podictenius</emphasis>
<bibRefCitation id="EFBD4B0FFFFC5116218CFC84FAC5FCE0" author="Gutu" box="[1160,1289,870,895]" pageId="2" pageNumber="51" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu, 2006</bibRefCitation>
</taxonomicName>
, all of which lack a distinct dorsal row of setae on the first pleonite. The new genus can be distinguished from these and from the other members of the family by a combination of characters, including having (1) pereopod 1 with a strongly developed spiniform seta on the anterior margin of the basis, (2) long, distally blunt, rod-like setae on the pereopods, and (3) pereopods 26 with the anterior margin of the ischium armed with one, occasionally two, spiniform setae. The following key may be used to distinguish
<taxonomicName id="4C2C4D7DFFFC5116267AFBF8FBA5FBAD" box="[894,1129,1051,1075]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC5116267AFBF8FBA5FBAD" box="[894,1129,1051,1075]" italics="true" pageId="2" pageNumber="51">Akanthoparapseudes</emphasis>
</taxonomicName>
further from
<taxonomicName id="4C2C4D7DFFFC51162008FBF8FA59FBAD" box="[1292,1429,1051,1075]" class="Malacostraca" family="Parapseudidae" genus="Brachylicoa" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC51162008FBF8FA59FBAD" box="[1292,1429,1051,1075]" italics="true" pageId="2" pageNumber="51">Brachylicoa</emphasis>
</taxonomicName>
,
<emphasis id="B958EAECFFFC51162593FBDDFEEEFBC8" box="[151,290,1086,1110]" italics="true" pageId="2" pageNumber="51">
<taxonomicName id="4C2C4D7DFFFC51162593FBDDFED2FBC8" box="[151,286,1086,1110]" class="Malacostraca" family="Parapseudidae" genus="Podictenius" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Podictenius</taxonomicName>
,
</emphasis>
and
<emphasis id="B958EAECFFFC5116245CFBDDFE10FBC8" box="[344,476,1086,1110]" italics="true" pageId="2" pageNumber="51">
<taxonomicName id="4C2C4D7DFFFC5116245CFBDDFE14FBC8" box="[344,472,1086,1110]" class="Malacostraca" family="Parapseudidae" genus="Longipedis" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Longipedis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3366575FFFC51172593FB49FCCCFF4A" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51" type="key">
<paragraph id="8B9336FEFFFC51162593FB49FD17FB79" blockId="2.[151,1436,1194,1255]" pageId="2" pageNumber="51">
<emphasis id="B958EAECFFFC51162593FB49FC38FB5A" bold="true" box="[151,1012,1194,1220]" pageId="2" pageNumber="51">
Key to parapseudid genera within the Tribe
<taxonomicName id="4C2C4D7DFFFC511627CBFB49FCB9FB5A" ID-CoL="3d865ae1-1061-4e6f-baad-3931f0cb6683" box="[719,885,1194,1220]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="tribe" tribe="Parapseudini">Parapseudini</taxonomicName>
<emphasis id="B958EAECFFFC5116267BFB49FC0FFB5A" bold="true" box="[895,963,1194,1220]" italics="true" pageId="2" pageNumber="51">sensu</emphasis>
Gu
</emphasis>
ţ
<emphasis id="B958EAECFFFC511626F9FB49FD17FB79" bold="true" pageId="2" pageNumber="51">u lacking a distinct, dense transverse row of setae dorsally on the first pleonite</emphasis>
</paragraph>
<key id="FFBD61CFFFFC51162593FAF7FA50F938" pageId="2" pageNumber="51">
<keyStep id="30D82E5BFFFC51162593FAF7FA50FA1F" pageId="2" pageNumber="51">
<paragraph id="8B9336FEFFFC51162593FAF7FA5AFAD8" blockId="2.[151,1436,1300,1702]" pageId="2" pageNumber="51">
<keyLead id="30DD95CBFFFC51162593FAF7FA5AFAD8" pageId="2" pageNumber="51">
1. Pereopod 4 with dactylus greatly reduced, shorter than adjacent setae. Often with diffuse transverse row of setae on the first pleonite. Usually only four pairs of pleopods present.......................................
<taxonomicName id="4C2C4D7DFFFC511621ABFAD2FA5AFAD8" ID-CoL="84RQM" authority="Sars, 1882" authorityName="Sars" authorityYear="1882" box="[1199,1430,1329,1350]" class="Malacostraca" family="Parapseudidae" genus="Parapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC511621ABFAD2FAE7FAD8" box="[1199,1323,1329,1350]" italics="true" pageId="2" pageNumber="51">Parapseudes</emphasis>
Sars, 1882
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9336FEFFFC51162593FAADFA50FA1F" blockId="2.[151,1436,1300,1702]" pageId="2" pageNumber="51">
<keyLead id="30DD95CBFFFC51162593FAADFA50FA1F" pageId="2" pageNumber="51">- Pereopod 4 with dactylus not greatly reduced, not distinctly shorter than adjacent setae. Five pairs of pleopods present. No obvious dorsal transverse row of setae on first pleonite........................................................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D82E5BFFFC51162593FA6AFA50FA25" pageId="2" pageNumber="51">
<paragraph id="8B9336FEFFFC51162593FA6AFA57FA00" blockId="2.[151,1436,1300,1702]" box="[151,1435,1417,1438]" pageId="2" pageNumber="51">
<keyLead id="30DD95CBFFFC51162593FA6AFA57FA00" box="[151,1435,1417,1438]" pageId="2" pageNumber="51">
2. Mandibular palp with article 1 bearing single long seta........................
<taxonomicName id="4C2C4D7DFFFC51162114FA6AFA57FA00" ID-CoL="84R6S" authority="Larsen and Shimomura, 2000" authorityName="Larsen and Shimomura" authorityYear="2000" box="[1040,1435,1417,1438]" class="Malacostraca" family="Parapseudidae" genus="Longipedis" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC51162114FA6AFBB6FA00" box="[1040,1146,1417,1438]" italics="true" pageId="2" pageNumber="51">Longipedis</emphasis>
Larsen and Shimomura, 2000
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9336FEFFFC51162593FA45FA50FA25" blockId="2.[151,1436,1300,1702]" box="[151,1436,1446,1467]" pageId="2" pageNumber="51">
<keyLead id="30DD95CBFFFC51162593FA45FA50FA25" box="[151,1436,1446,1467]" pageId="2" pageNumber="51">- Mandibular palp with article 1 having 515 simple setae....................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D82E5BFFFC51162593FA27FA5AF9AF" pageId="2" pageNumber="51">
<paragraph id="8B9336FEFFFC51162593FA27FA50FA6B" blockId="2.[151,1436,1300,1702]" pageId="2" pageNumber="51">
<keyLead id="30DD95CBFFFC51162593FA27FA50FA6B" pageId="2" pageNumber="51">
3. Palp of maxilliped with tip of outer margin of article 1 armed with well-developed spiniform seta. Pereopod 2 with anterior margin of basis armed with distinct spine...........................................
<taxonomicName id="4C2C4D7DFFFC51162187FA02FA80FA68" ID-CoL="84L7Y" box="[1155,1356,1505,1526]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B958EAECFFFC51162187FA02FA80FA68" box="[1155,1356,1505,1526]" italics="true" pageId="2" pageNumber="51">Akanthoparapseudes</emphasis>
</taxonomicName>
,
<emphasis id="B958EAECFFFC5116205CFA03FA50FA6B" bold="true" box="[1368,1436,1504,1525]" pageId="2" pageNumber="51">
<taxonomicNameLabel id="A26B5797FFFC5116205CFA03FA50FA6B" box="[1368,1436,1504,1525]" pageId="2" pageNumber="51" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9336FEFFFC51162593FA1DFA5AF9AF" blockId="2.[151,1436,1300,1702]" pageId="2" pageNumber="51">
<keyLead id="30DD95CBFFFC51162593FA1DFA5AF9AF" pageId="2" pageNumber="51">- Palp of maxilliped with article 1 having distal corner of outer margin armed with minute spiniform seta. Pereopod 2 with anterior margin of basis lacking distinct spine................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D82E5BFFFC51162593F9DAFA50F938" pageId="2" pageNumber="51">
<paragraph id="8B9336FEFFFC51162593F9DAFA50F9F5" blockId="2.[151,1436,1300,1702]" pageId="2" pageNumber="51">
<keyLead id="30DD95CBFFFC51162593F9DAFA50F9F5" pageId="2" pageNumber="51">
4. Pereopod 1 with anterior margin of basis armed with row of 512 spiniform setae often increasing in length distally. Mandibular palp with first article bearing five or six simple setae.....................................
<taxonomicName id="4C2C4D7DFFFC511621BDF9B5FA50F9F5" ID-CoL="84SNG" authority="Gutu, 2006" authorityName="Gutu" authorityYear="2006" box="[1209,1436,1622,1643]" class="Malacostraca" family="Parapseudidae" genus="Podictenius" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC511621BDF9B5FAE6F9F5" box="[1209,1322,1622,1643]" italics="true" pageId="2" pageNumber="51">Podictenius</emphasis>
<bibRefCitation id="EFBD4B0FFFFC51162034F9B5FA50F9F5" author="Gutu" box="[1328,1436,1622,1643]" pageId="2" pageNumber="51" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu, 2006</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9336FEFFFC51162593F997FA50F938" blockId="2.[151,1436,1300,1702]" pageId="2" pageNumber="51">
<keyLead id="30DD95CBFFFC51162593F997FA50F938" pageId="2" pageNumber="51">
- Pereopod 1 with anterior margin of basis lacking spiniform setae. Mandibular palp with first article bearing cluster (about 15) of simple setae.....................................................................
<taxonomicName id="4C2C4D7DFFFC511621B0F972FA50F938" ID-CoL="84MKY" authority="Gutu, 2006" authorityName="Gutu" authorityYear="2006" box="[1204,1436,1681,1702]" class="Malacostraca" family="Parapseudidae" genus="Brachylicoa" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC511621B0F972FAE5F938" box="[1204,1321,1681,1702]" italics="true" pageId="2" pageNumber="51">Brachylicoa</emphasis>
<bibRefCitation id="EFBD4B0FFFFC51162034F972FA50F938" author="Gutu" box="[1328,1436,1681,1702]" pageId="2" pageNumber="51" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu, 2006</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8B9336FEFFFC511725C1F931FCCCFF4A" blockId="2.[151,1436,1746,2022]" lastBlockId="3.[151,1437,151,2012]" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
Though the presence on the pereopods of spines on the anterior margin of the ischium is quite pronounced in the new genus; it is not unique and occurs in other apseudomorphans genera, including the apseudid genus,
<taxonomicName id="4C2C4D7DFFFC51162593F8F8FEC3F8AD" ID-CoL="84MQM" box="[151,271,1819,1843]" class="Malacostraca" family="Apseudidae" genus="Bunakenia" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC51162593F8F8FEC3F8AD" box="[151,271,1819,1843]" italics="true" pageId="2" pageNumber="51">Bunakenia</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFBD4B0FFFFC51162448F8F8FE0BF8AC" author="Gutu" box="[332,455,1818,1843]" pageId="2" pageNumber="51" refString="Gutu, M. (1995) Bunakenia indonesiana, new genus and species of Apseudomorpha (Crustacea, Tanaidacea), from the south of the Sulawesi Sea. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 35, 7 - 16." type="journal article" year="1995">Guţu 1995</bibRefCitation>
,
<bibRefCitation id="EFBD4B0FFFFC511624D7F8F9FDC2F8AC" author="Gutu" box="[467,526,1818,1842]" pageId="2" pageNumber="51" refString="Gutu, M. (1996) Description of Bunakenia tanzaniana n. sp. (Crustacea, Tanaidacea) from the East African marine waters. Revue Roumaine de Biologie (Serie de Biologie Animale), 41, 17 - 23." type="journal article" year="1996">1996</bibRefCitation>
;
<bibRefCitation id="EFBD4B0FFFFC5116271EF8F9FD79F8AC" author="Bamber" box="[538,693,1818,1842]" pageId="2" pageNumber="51" refString="Bamber, R. N (2005) The tanaidaceans (Arthropoda: Crustacea: Peracarida: Tanaidacea) of Esperance, Western Australia, Australia. Pages 613 - 728, In: Wells, F. E., D. I. Walker, &amp; G. A. Kendrick (eds.), The Marine Flora and Fauna of Esperance, Western Australia. Western Australian Museum, Perth." type="book chapter" year="2005">Bamber 2005</bibRefCitation>
) and the parapseudid genera,
<taxonomicName id="4C2C4D7DFFFC5116210CF8F8FB5DF8AD" ID-CoL="84MKY" box="[1032,1169,1819,1843]" class="Malacostraca" family="Parapseudidae" genus="Brachylicoa" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC5116210CF8F8FB5DF8AD" box="[1032,1169,1819,1843]" italics="true" pageId="2" pageNumber="51">Brachylicoa</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFBD4B0FFFFC511621CAF8F8FA87F8AC" author="Gutu" box="[1230,1355,1818,1843]" pageId="2" pageNumber="51" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu 2006</bibRefCitation>
);
<taxonomicName id="4C2C4D7DFFFC5116205BF8F8FE95F8C8" ID-CoL="84TDJ" authority="Gutu, 1995" authorityName="Gutu" authorityYear="1995" class="Malacostraca" family="Parapseudidae" genus="Saltipedis" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC5116205BF8F8FF1EF8C8" italics="true" pageId="2" pageNumber="51">Saltipedis</emphasis>
<bibRefCitation id="EFBD4B0FFFFC511625DEF8DCFE95F8C8" author="Gutu" box="[218,345,1854,1879]" pageId="2" pageNumber="51" refString="Gutu, M. (1995) Bunakenia indonesiana, new genus and species of Apseudomorpha (Crustacea, Tanaidacea), from the south of the Sulawesi Sea. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 35, 7 - 16." type="journal article" year="1995">Guţu, 1995</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFBD4B0FFFFC51162493F8DCFDC3F8C8" author="Gutu" box="[407,527,1854,1879]" pageId="2" pageNumber="51" refString="Gutu, M. (1998) Description of three new species of Tanaidacea (Crustacea) from the Tanzanian Coasts. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 40, 179 - 209." type="journal article" year="1998">Guţu 1998</bibRefCitation>
;
<bibRefCitation id="EFBD4B0FFFFC51162725F8DDFC9AF8C8" author="Morales-Nunez" box="[545,854,1854,1879]" pageId="2" pageNumber="51" refString="Morales-Nunez, A. G., Heard, R. W. &amp; Alfaro. M. (2010) Saltipedis (Spinosaltipedis) puertoricensis, a new subgenus and species of apseudomorphan (Crustacea: Tanaidacea: Parapseudidae) from coastal waters off Culebra Island, Puerto Rico, with keys and taxonomic observations. Zootaxa, 2587, 25 - 41." type="journal article" year="2010">
Morales-Nuñez
<emphasis id="B958EAECFFFC511627D3F8A3FCDDF8C8" box="[727,785,1854,1879]" italics="true" pageId="2" pageNumber="51">et al.</emphasis>
2010
</bibRefCitation>
);
<taxonomicName id="4C2C4D7DFFFC5116266EF8DDFB57F8C8" ID-CoL="84QC7" authority="Edgar, 1997" authorityName="Edgar" authorityYear="1997" box="[874,1179,1854,1878]" class="Malacostraca" family="Parapseudidae" genus="Gutuapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC5116266EF8DDFB57F8C8" box="[874,1179,1854,1878]" italics="true" pageId="2" pageNumber="51">Gutuapseudes Edgar, 1997</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFBD4B0FFFFC511621DDF8DCFEF0F8E4" author="Gutu" pageId="2" pageNumber="51" refString="Gutu, M. &amp; Angsupanich. S. (2004 b) Two new parapseudid species and some first records of Tanaidacea (Crustacea: Peracarida) from Thailand. Travaux du Museum Nationa l d´Histoire naturelle &quot; Grigore Antipa &quot;, 47, 75 - 87." type="journal article" year="2004" yearSuffix="b">Guţu and Angsupanich 2004b</bibRefCitation>
); and
<taxonomicName id="4C2C4D7DFFFC51162495F880FDFBF8E5" ID-CoL="84TZ8" box="[401,567,1891,1915]" class="Malacostraca" family="Parapseudidae" genus="Swireapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC51162495F880FDFBF8E5" box="[401,567,1891,1915]" italics="true" pageId="2" pageNumber="51">Swireapseudes</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFBD4B0FFFFC51162787F880FC49F8E4" author="Gutu" box="[643,901,1890,1915]" pageId="2" pageNumber="51" refString="Gutu, M. &amp; Iliffe, T. M. (2008) A new species and the first description of the male belonging to the genus Swireapseudes Bamber, from the submarine caves of the Eleuthera Island (Crustacea: Tanaidacea: Apseudomorpha). Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 51, 7 - 16." type="journal article" year="2008">Guţu and Iliffe 2008</bibRefCitation>
). It should be noted that within the genus
<taxonomicName id="4C2C4D7DFFFC51162593F865FEE4F800" ID-CoL="84RQM" box="[151,296,1926,1950]" class="Malacostraca" family="Parapseudidae" genus="Parapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC51162593F865FEE4F800" box="[151,296,1926,1950]" italics="true" pageId="2" pageNumber="51">Parapseudes</emphasis>
</taxonomicName>
, the anterior margin of the ischium on some pereopods bears a long simple seta; in some species this seta can extend well-past the merus (see
<bibRefCitation id="EFBD4B0FFFFC5116275AF848FD15F85C" author="Lang" box="[606,729,1962,1987]" pageId="2" pageNumber="51" refString="Lang, K. (1966) Taxonomische und phylogenetische Untesuchungen uber die Tanaidaceen, 2 Die Gattung Parapseudes G. O Sars. Arkiv for Zoologi, Series, 2, (18), 549 - 566." type="journal article" year="1966">Lang 1966</bibRefCitation>
,
<bibRefCitation id="EFBD4B0FFFFC511627E1F849FCBDF85C" author="Shiino" box="[741,881,1962,1986]" pageId="2" pageNumber="51" refString="Shiino, S. M. (1952) A new genus and two new species of the order Tanaidacea found at Seto. Publications of the Seto Marine Biological Laboratory, 2, 53 - 68." type="journal article" year="1952">Shiino 1952</bibRefCitation>
); these long setae may be unique to
<taxonomicName id="4C2C4D7DFFFC51162000F848FA59F85D" ID-CoL="84RQM" box="[1284,1429,1963,1987]" class="Malacostraca" family="Parapseudidae" genus="Parapseudes" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFC51162000F848FA59F85D" box="[1284,1429,1963,1987]" italics="true" pageId="2" pageNumber="51">Parapseudes</emphasis>
</taxonomicName>
, and possibly homologous with the ischial spines discussed above. Also, in at least in one apseudid species,
<taxonomicName id="4C2C4D7DFFFC51172044F82DFE08FF31" authority="Gutu, 2006" authorityName="Gutu" authorityYear="2006" class="Malacostraca" family="Apseudidae" genus="Androgynella" higherTaxonomySource="CoL" kingdom="Animalia" lastPageId="3" lastPageNumber="52" order="Tanaidacea" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="fecunda">
<emphasis id="B958EAECFFFC51172044F82DFEF1FF2E" italics="true" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">Androgynella fecunda</emphasis>
<bibRefCitation id="EFBD4B0FFFFD51172440FF7BFE08FF31" author="Gutu" box="[324,452,151,176]" pageId="3" pageNumber="52" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu, 2006</bibRefCitation>
</taxonomicName>
, a plumose seta is present on the anterior face of the ischium of pereopod 6 (see
<bibRefCitation id="EFBD4B0FFFFD51172060FF7BFF18FF4A" author="Gutu" pageId="3" pageNumber="52" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu 2006</bibRefCitation>
); this may be an independently-derived character.
</paragraph>
</subSubSection>
<subSubSection id="C3366575FFFD511025C1FF03FCDBFD89" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52" type="reference_group">
<paragraph id="8B9336FEFFFD511725C1FF03FABFFDC1" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="52">
<taxonomicName id="4C2C4D7DFFFD511725C1FF03FE7CFF66" ID-CoL="84L7Y" box="[197,432,224,248]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511725C1FF03FE7CFF66" box="[197,432,224,248]" italics="true" pageId="3" pageNumber="52">Akanthoparapseudes</emphasis>
</taxonomicName>
also appears to have some affinities with the family
<taxonomicName id="4C2C4D7DFFFD51172109FF3CFB59FF69" ID-CoL="84JW5" box="[1037,1173,223,247]" class="Malacostraca" family="Apseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Apseudidae</taxonomicName>
; however, pending further study we tentatively refer it to the family
<taxonomicName id="4C2C4D7DFFFD5117279EFEE7FCF3FE82" ID-CoL="84KM3" box="[666,831,260,284]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Parapseudidae</taxonomicName>
, within the Tribe
<taxonomicName id="4C2C4D7DFFFD51172107FEE7FAD2FE82" ID-CoL="3d865ae1-1061-4e6f-baad-3931f0cb6683" authority="Gutu, 2008" authorityName="Gutu" authorityYear="2008" box="[1027,1310,260,284]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Parapseudini">Parapseudini Guţu, 2008</taxonomicName>
. This Tribe was characterized by Guţu (2008) as having the pereonites distinctly broader and longer than those of the pleon, which is compressed (i.e. usually less than half the length of thorax) with the pleonites being distinctly wider than long). The
<taxonomicName id="4C2C4D7DFFFD51172415FE8CFE6EFE19" ID-CoL="3d865ae1-1061-4e6f-baad-3931f0cb6683" box="[273,418,367,391]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Parapseudini">Parapseudini</taxonomicName>
also contains tube- or burrow-dwelling genera such as
<taxonomicName id="4C2C4D7DFFFD5117210CFE93FF03FE32" ID-CoL="84NXS" authority="Bacescu and Gutu, 1974" authorityName="Bacescu and Gutu" authorityYear="1974" class="Malacostraca" family="Parapseudidae" genus="Halmyrapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD5117210CFE93FB08FE16" box="[1032,1220,368,392]" italics="true" pageId="3" pageNumber="52">Halmyrapseudes</emphasis>
<bibRefCitation id="EFBD4B0FFFFD511721CEFE93FF03FE32" author="Bacescu" pageId="3" pageNumber="52" refString="Bacescu, M. &amp; Gutu., M. (1974) Halmyrapseudes cubanensis n. g., n. sp. and Halmyrapseudes Bahamensis n. sp., brackishwater species of Tanaidacea (Crustacea). Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 15, 91 - 101." type="journal article" year="1974">Bačescu and Guţu, 1974</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2C4D7DFFFD51172400FE77FD7AFE32" ID-CoL="84N8X" authority="Bacescu and Gutu, 1975" authorityName="Bacescu and Gutu" authorityYear="1975" box="[260,694,404,428]" class="Malacostraca" family="Parapseudidae" genus="Discapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD51172400FE77FE52FE32" box="[260,414,404,428]" italics="true" pageId="3" pageNumber="52">Discapseudes</emphasis>
<bibRefCitation id="EFBD4B0FFFFD511724A1FE77FD7AFE32" author="Bacescu" box="[421,694,404,428]" pageId="3" pageNumber="52" refString="Bacescu, M., &amp; Gutu., M. (1975) A new genus (Discapseudes n. g.) and three new species of Apseudidae (Crustacea, Tanaidacea) from the northeastern coast of South America. Zoologische Mededelingen, 49 (11), 95 - 113." type="journal article" year="1975">Bačescu and Guţu, 1975</bibRefCitation>
</taxonomicName>
that have a distinct transverse row of tightly spaced, short setulate setae present on the dorsoproximal surface of at least the first pleonite.
<taxonomicName id="4C2C4D7DFFFD511726ABFE5BFB56FE4E" ID-CoL="84L7Y" box="[943,1178,440,464]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511726ABFE5BFB56FE4E" box="[943,1178,440,464]" italics="true" pageId="3" pageNumber="52">Akanthoparapseudes</emphasis>
</taxonomicName>
, along with three other genera,
<taxonomicName id="4C2C4D7DFFFD511725F5FE3FFDCBFE6A" ID-CoL="84MKY" authority="Gutu, 2006" authorityName="Gutu" authorityYear="2006" box="[241,519,476,500]" class="Malacostraca" family="Parapseudidae" genus="Brachylicoa" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511725F5FE3FFEB6FE6A" box="[241,378,476,500]" italics="true" pageId="3" pageNumber="52">Brachylicoa</emphasis>
<bibRefCitation id="EFBD4B0FFFFD51172480FE3FFDCBFE6A" author="Gutu" box="[388,519,476,500]" pageId="3" pageNumber="52" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu, 2006</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C2C4D7DFFFD51172710FE3FFCE9FE6A" ID-CoL="84SNG" authority="Gutu, 2006" authorityName="Gutu" authorityYear="2006" box="[532,805,476,500]" class="Malacostraca" family="Parapseudidae" genus="Podictenius" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD51172710FE3FFD54FE6A" box="[532,664,476,500]" italics="true" pageId="3" pageNumber="52">Podictenius</emphasis>
<bibRefCitation id="EFBD4B0FFFFD511727A6FE3FFCE9FE6A" author="Gutu" box="[674,805,476,500]" pageId="3" pageNumber="52" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu, 2006</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C2C4D7DFFFD51172660FE3FFC2DFE6A" ID-CoL="84R6S" box="[868,993,476,500]" class="Malacostraca" family="Parapseudidae" genus="Longipedis" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD51172660FE3FFC2DFE6A" box="[868,993,476,500]" italics="true" pageId="3" pageNumber="52">Longipedis</emphasis>
</taxonomicName>
are characterized by lacking a distinct dense row of setulate setae. Within the genus
<taxonomicName id="4C2C4D7DFFFD511727A4FDE3FC7CFD89" ID-CoL="84RQM" authority="Sars, 1882" authorityName="Sars" authorityYear="1882" box="[672,944,511,536]" class="Malacostraca" family="Parapseudidae" genus="Parapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511727A4FDE3FCFDFD86" box="[672,817,512,536]" italics="true" pageId="3" pageNumber="52">Parapseudes</emphasis>
Sars, 1882
</taxonomicName>
some species appear to have a weak transverse row of setae (e.g.,
<taxonomicName id="4C2C4D7DFFFD511724A9FDC6FD3FFDA2" ID-CoL="75QZY" authority="Lang 1966" authorityName="Lang" authorityYear="1966" box="[429,755,548,573]" class="Malacostraca" family="Parapseudidae" genus="Parapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B958EAECFFFD511724A9FDC6FDA2FDA2" box="[429,622,548,572]" italics="true" pageId="3" pageNumber="52">P. latifrons sensu</emphasis>
<bibRefCitation id="EFBD4B0FFFFD51172772FDC6FD3FFDA2" author="Lang" box="[630,755,548,573]" pageId="3" pageNumber="52" refString="Lang, K. (1966) Taxonomische und phylogenetische Untesuchungen uber die Tanaidaceen, 2 Die Gattung Parapseudes G. O Sars. Arkiv for Zoologi, Series, 2, (18), 549 - 566." type="journal article" year="1966">Lang 1966</bibRefCitation>
</taxonomicName>
), but they are simple and not setulose as in the known tubedwelling genera and others only have a few diffuse setae (e. g.
<taxonomicName id="4C2C4D7DFFFD5117265CFDAAFBBEFDC1" ID-CoL="4D9ZG" authority="Miller, 1940" authorityName="Miller" authorityYear="1940" box="[856,1138,583,608]" class="Malacostraca" family="Parapseudidae" genus="Parapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="B958EAECFFFD5117265CFDAAFC12FDFE" box="[856,990,584,608]" italics="true" pageId="3" pageNumber="52">P. neglectus</emphasis>
Miller, 1940
</taxonomicName>
, R.
<collectingCountry id="F33B766EFFFD5117219AFDA4FB29FDC1" box="[1182,1253,583,607]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="52">Heard</collectingCountry>
, per. obser.).
</paragraph>
<paragraph id="8B9336FEFFFD511725C1FD8FFCFFFC72" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="52">
As exemplified by
<taxonomicName id="4C2C4D7DFFFD5117249FFD8FFDE0FD1A" ID-CoL="84RQM" box="[411,556,620,644]" class="Malacostraca" family="Parapseudidae" genus="Parapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD5117249FFD8FFDE0FD1A" box="[411,556,620,644]" italics="true" pageId="3" pageNumber="52">Parapseudes</emphasis>
</taxonomicName>
, the overall presence or absence of the transverse row of setae on the first pleonite may not be as an important phylogenetic character as previously thought. Whether the other genera assigned to the
<taxonomicName id="4C2C4D7DFFFD511725D9FD57FEA2FD52" ID-CoL="3d865ae1-1061-4e6f-baad-3931f0cb6683" box="[221,366,692,716]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Parapseudini">Parapseudini</taxonomicName>
(e. g.,
<taxonomicName id="4C2C4D7DFFFD511724BFFD57FC8CFD52" ID-CoL="84P8J" authority="Bamber et al., 1996" authorityName="Bamber et al." authorityYear="1996" box="[443,832,692,716]" class="Malacostraca" family="Parapseudidae" genus="Ctenapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511724BFFD57FD9BFD52" box="[443,599,692,716]" italics="true" pageId="3" pageNumber="52">Ctenapseudes</emphasis>
Bamber
<emphasis id="B958EAECFFFD511727B9FD56FD31FD52" box="[701,765,692,716]" italics="true" pageId="3" pageNumber="52">et al.,</emphasis>
1996
</taxonomicName>
;
<taxonomicName id="4C2C4D7DFFFD5117264FFD57FA5BFD52" ID-CoL="84SXM" authority="Larsen &amp; Hansknecht, 2004" authorityName="Larsen &amp; Hansknecht" authorityYear="2004" box="[843,1431,692,716]" class="Malacostraca" family="Parapseudidae" genus="Pseudohalmyrapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD5117264FFD57FB9FFD52" box="[843,1107,692,716]" italics="true" pageId="3" pageNumber="52">Pseudohalmyrapseudes</emphasis>
<bibRefCitation id="EFBD4B0FFFFD5117215EFD56FA5BFD52" author="Larsen" box="[1114,1431,692,716]" pageId="3" pageNumber="52" refString="Heard, R. W., Hansknecht, T. &amp; Larsen, K. (2004) An illustrated identification guide to Tanaidacea (Crustacea: Peracarida) occurring in depths of less than 200 m. &quot; Final Supplemental Report, Florida Department of Environmental Protection, Tallahassee, Florida, 163 p. http: // www. dep. state. fl. us / labs / cgi-bin / sbio / keys. asp" type="book" year="2004">Larsen &amp; Hansknecht, 2004</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C2C4D7DFFFD51172593FD3BFE40FD71" ID-CoL="84TDJ" authority="Gutu, 1995" authorityName="Gutu" authorityYear="1995" box="[151,396,727,752]" class="Malacostraca" family="Parapseudidae" genus="Saltipedis" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD51172593FD3BFEC9FD6E" box="[151,261,728,752]" italics="true" pageId="3" pageNumber="52">Saltipedis</emphasis>
<bibRefCitation id="EFBD4B0FFFFD51172408FD3BFE40FD71" author="Gutu" box="[268,396,727,752]" pageId="3" pageNumber="52" refString="Gutu, M. (1995) Bunakenia indonesiana, new genus and species of Apseudomorpha (Crustacea, Tanaidacea), from the south of the Sulawesi Sea. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 35, 7 - 16." type="journal article" year="1995">Guţu, 1995</bibRefCitation>
</taxonomicName>
) having a dense transverse row of finely setulate setae on the first pleonite are tube, burrow, or&quot; “nest” builders remains to be discovered. Notwithstanding this, for the species attributed to the
<taxonomicName id="4C2C4D7DFFFD51172001FD1FFA55FC8A" ID-CoL="3d865ae1-1061-4e6f-baad-3931f0cb6683" box="[1285,1433,764,788]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Parapseudini">Parapseudini</taxonomicName>
, this condition appears to be an important and distinctive character, which may be associated with a tube, burrow, or “nest” dwelling as exemplified by the genera
<taxonomicName id="4C2C4D7DFFFD5117279AFCA7FC96FCC2" ID-CoL="84NXS" box="[670,858,836,860]" class="Malacostraca" family="Parapseudidae" genus="Halmyrapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD5117279AFCA7FC96FCC2" box="[670,858,836,860]" italics="true" pageId="3" pageNumber="52">Halmyrapseudes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D7DFFFD5117266EFCA7FBC8FCC2" ID-CoL="84N8X" box="[874,1028,836,860]" class="Malacostraca" family="Parapseudidae" genus="Discapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD5117266EFCA7FBC8FCC2" box="[874,1028,836,860]" italics="true" pageId="3" pageNumber="52">Discapseudes</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2C4D7DFFFD51172147FCA7FB30FCC2" ID-CoL="84SW4" box="[1091,1276,836,860]" class="Malacostraca" family="Parapseudidae" genus="Pseudoapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD51172147FCA7FB30FCC2" box="[1091,1276,836,860]" italics="true" pageId="3" pageNumber="52">Pseudoapseudes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBD4B0FFFFD51172008FCA7FEDDFCE1" author="Bacescu" pageId="3" pageNumber="52" refString="Bacescu, M. &amp; Gutu., M. (1974) Halmyrapseudes cubanensis n. g., n. sp. and Halmyrapseudes Bahamensis n. sp., brackishwater species of Tanaidacea (Crustacea). Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 15, 91 - 101." type="journal article" year="1974">Bačescu and Guţu 1974</bibRefCitation>
,
<bibRefCitation id="EFBD4B0FFFFD51172418FC84FE9BFCE1" author="Bacescu" box="[284,343,871,895]" pageId="3" pageNumber="52" refString="Bacescu, M., &amp; Gutu., M. (1975) A new genus (Discapseudes n. g.) and three new species of Apseudidae (Crustacea, Tanaidacea) from the northeastern coast of South America. Zoologische Mededelingen, 49 (11), 95 - 113." type="journal article" year="1975">1975</bibRefCitation>
; R.
<collectingCountry id="F33B766EFFFD51172481FC84FE01FCE1" box="[389,461,871,895]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="52">Heard</collectingCountry>
, per. obser.). Based on observations of living specimens of
<taxonomicName id="4C2C4D7DFFFD5117216CFC8BFE04FC3A" ID-CoL="6KYMX" authority="Bacescu and Gutu 1974" authorityName="Bacescu and Gutu" authorityYear="1974" class="Malacostraca" family="Parapseudidae" genus="Halmyrapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="bahamensis">
<emphasis id="B958EAECFFFD5117216CFC8BFF79FC3A" italics="true" pageId="3" pageNumber="52">Halmyrapseudes bahamensis</emphasis>
<bibRefCitation id="EFBD4B0FFFFD511725B8FC6FFE04FC3A" author="Bacescu" box="[188,456,908,932]" pageId="3" pageNumber="52" refString="Bacescu, M. &amp; Gutu., M. (1974) Halmyrapseudes cubanensis n. g., n. sp. and Halmyrapseudes Bahamensis n. sp., brackishwater species of Tanaidacea (Crustacea). Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 15, 91 - 101." type="journal article" year="1974">Bačescu and Guţu 1974</bibRefCitation>
</taxonomicName>
from southern Florida, the transverse dorsal rows of setulate setae on the pleon appear to filter out silt and other fine particles. This might be a mechanism to prevent fouling of the pleopods and, possibly, to retain potential food particles (R.
<collectingCountry id="F33B766EFFFD51172759FC37FD68FC72" box="[605,676,980,1004]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="52">Heard</collectingCountry>
, per. obser.).
</paragraph>
<paragraph id="8B9336FEFFFD511725C1FC14FD31FBE2" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="52">
Such setae appear to be absent in members of the Tribe
<taxonomicName id="4C2C4D7DFFFD51172632FC14FBCFFB91" ID-CoL="7dd5ba3f-0e2f-457e-9a2b-36bd38a26ad8" box="[822,1027,1015,1039]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Pakistanapseudini">Pakistanapseudini</taxonomicName>
, which, beside the distinctive sexual dimorphism of the male antenna and body form mentioned previously, appear to have no tube- or burrow-dwelling representatives. In our opinion the
<taxonomicName id="4C2C4D7DFFFD5117271BFBDCFD25FBC9" ID-CoL="7dd5ba3f-0e2f-457e-9a2b-36bd38a26ad8" box="[543,745,1087,1111]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Pakistanapseudini">Pakistanapseudini</taxonomicName>
appear to form a more natural group than the more heterogeneous genera currently placed within the
<taxonomicName id="4C2C4D7DFFFD51172761FB87FD35FBE2" ID-CoL="3d865ae1-1061-4e6f-baad-3931f0cb6683" box="[613,761,1124,1148]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Parapseudini">Parapseudini</taxonomicName>
.
</paragraph>
<paragraph id="8B9336FEFFFD511725C1FB6BFED1FAAE" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="52">
A small, reduced coxal process or lobe is present on the first pereopod of
<taxonomicName id="4C2C4D7DFFFD511726F2FB6BFB2DFB3E" ID-CoL="84L7Y" box="[1014,1249,1160,1184]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511726F2FB6BFB2DFB3E" box="[1014,1249,1160,1184]" italics="true" pageId="3" pageNumber="52">Akanthoparapseudes</emphasis>
</taxonomicName>
, but it is not visible from the dorsal aspect (
<figureCitation id="13172A7BFFFD511724D9FB4FFDEEFB5A" box="[477,546,1196,1220]" captionStart="FIGURE 2" captionStartId="5.[151,250,1911,1934]" captionTargetBox="[205,1390,322,1893]" captionTargetId="figure@5.[197,1390,309,1904]" captionTargetPageId="5" captionText="FIGURE 2. Akanthoparapseudes alfaroae n. gen., n. sp., subadult female holotype. A, body, dorsal view; B, enlargement of posterolateral margin of pereonite 2; C, enlargement of posterolateral margin of pereonite 3: D, E. enlargement of lateral margins of first and fifth pleonites, repectively, 1; F, enlargement of pleotelson; G, enlargement of uropod; H, lateral view of cephalothorax and pereonite 1. Scale line: A = 1.0 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/207144/files/figure.png" pageId="3" pageNumber="52">Fig. 2</figureCitation>
H). A similar more reduced coxal process has been reported for a few other parapseudids (e.g.,
<taxonomicName id="4C2C4D7DFFFD51172473FB33FC7FFB76" ID-CoL="72MQ5" authority="Larsen and Shimomura, 2006" authorityName="Larsen and Shimomura" authorityYear="2006" box="[375,947,1232,1256]" class="Malacostraca" family="Parapseudidae" genus="Longipedis" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B958EAECFFFD51172473FB33FD82FB76" box="[375,590,1232,1256]" italics="true" pageId="3" pageNumber="52">Longipedis fragilis</emphasis>
Larsen and Shimomura, 2006
</taxonomicName>
;
<taxonomicName id="4C2C4D7DFFFD511726C7FB33FA5BFB76" ID-CoL="79FPR" authority="Morales-Nunez" authorityName="Morales-Nunez" box="[963,1431,1232,1256]" class="Malacostraca" family="Parapseudidae" genus="Saltipedis" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="puertoricensis">
<emphasis id="B958EAECFFFD511726C7FB33FB10FB76" box="[963,1244,1232,1256]" italics="true" pageId="3" pageNumber="52">Saltipedis puertoricensis</emphasis>
Morales-Núñez
</taxonomicName>
,
<collectingCountry id="F33B766EFFFD51172593FB17FF12FA92" box="[151,222,1268,1292]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="52">Heard</collectingCountry>
, and Alfaro, 2010). It is possible, however, that this structure may have been overlooked in other parapseudid taxa.
</paragraph>
<paragraph id="8B9336FEFFFD511725C3FADFFD99F922" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="52">
<taxonomicName id="4C2C4D7DFFFD511725C3FADFFE7EFACA" ID-CoL="84L7Y" box="[199,434,1340,1364]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511725C3FADFFE7EFACA" box="[199,434,1340,1364]" italics="true" pageId="3" pageNumber="52">Akanthoparapseudes</emphasis>
</taxonomicName>
appears to be related to “Parapseudid? species A”
<emphasis id="B958EAECFFFD5117210CFADEFB8AFACA" box="[1032,1094,1341,1364]" italics="true" pageId="3" pageNumber="52">sensu</emphasis>
<bibRefCitation id="EFBD4B0FFFFD51172154FADFFAE3FACA" box="[1104,1327,1340,1364]" pageId="3" pageNumber="52" refString="Heard, R. W., Hansknecht, T. &amp; Larsen, K. (2004) An illustrated identification guide to Tanaidacea (Crustacea: Peracarida) occurring in depths of less than 200 m. &quot; Final Supplemental Report, Florida Department of Environmental Protection, Tallahassee, Florida, 163 p. http: // www. dep. state. fl. us / labs / cgi-bin / sbio / keys. asp" type="book">
<collectingCountry id="F33B766EFFFD51172154FADFFB59FACA" box="[1104,1173,1340,1364]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="52">Heard</collectingCountry>
<emphasis id="B958EAECFFFD5117219BFADEFB17FACA" box="[1183,1243,1340,1364]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
(2004)
</bibRefCitation>
from the eastern Gulf of
<collectingCountry id="F33B766EFFFD5117244FFA83FE68FAE6" box="[331,420,1376,1400]" name="Mexico" pageId="3" pageNumber="52">Mexico</collectingCountry>
. Except for the lack of a strongly developed and dorsally visible, coxal spine or process,
<taxonomicName id="4C2C4D7DFFFD51172593FA67FE4EFA02" ID-CoL="84L7Y" box="[151,386,1412,1436]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD51172593FA67FE4EFA02" box="[151,386,1412,1436]" italics="true" pageId="3" pageNumber="52">Akanthoparapseudes</emphasis>
</taxonomicName>
and “Parapseudid? sp. A” appear to share several characters with some members of the apseudid genus
<taxonomicName id="4C2C4D7DFFFD5117244BFA4BFD99FA21" ID-CoL="84MQM" authority="Gutu, 1995" authorityName="Gutu" authorityYear="1995" box="[335,597,1447,1472]" class="Malacostraca" family="Apseudidae" genus="Bunakenia" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD5117244BFA4BFE0BFA5E" box="[335,455,1448,1472]" italics="true" pageId="3" pageNumber="52">Bunakenia</emphasis>
<bibRefCitation id="EFBD4B0FFFFD511724D5FA4BFD99FA21" author="Gutu" box="[465,597,1447,1472]" pageId="3" pageNumber="52" refString="Gutu, M. (1995) Bunakenia indonesiana, new genus and species of Apseudomorpha (Crustacea, Tanaidacea), from the south of the Sulawesi Sea. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 35, 7 - 16." type="journal article" year="1995">Guţu, 1995</bibRefCitation>
</taxonomicName>
, especially those species attributed to the subgenus
<taxonomicName id="4C2C4D7DFFFD511721B4FA4BFF1EFA7A" authority="Gutu, 1998" authorityName="Gutu" authorityYear="1998" class="Malacostraca" family="Apseudidae" genus="Extensibasella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511721B4FA4BFA98FA5E" box="[1200,1364,1448,1472]" italics="true" pageId="3" pageNumber="52">Extensibasella</emphasis>
<bibRefCitation id="EFBD4B0FFFFD51172059FA4BFF1EFA7A" author="Gutu" pageId="3" pageNumber="52" refString="Gutu, M. (1998) Description of three new species of Tanaidacea (Crustacea) from the Tanzanian Coasts. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 40, 179 - 209." type="journal article" year="1998">Guţu, 1998</bibRefCitation>
</taxonomicName>
. The possible affinity of
<taxonomicName id="4C2C4D7DFFFD511724E2FA2FFD92FA7A" ID-CoL="84MQM" box="[486,606,1484,1508]" class="Malacostraca" family="Apseudidae" genus="Bunakenia" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511724E2FA2FFD92FA7A" box="[486,606,1484,1508]" italics="true" pageId="3" pageNumber="52">Bunakenia</emphasis>
</taxonomicName>
with the family
<taxonomicName id="4C2C4D7DFFFD51172611FA2FFC7BFA7A" ID-CoL="84KM3" box="[789,951,1484,1508]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Parapseudidae</taxonomicName>
also has been broached previously by
<bibRefCitation id="EFBD4B0FFFFD51172060FA2FFF29F999" author="Gutu" pageId="3" pageNumber="52" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">Guţu (2006)</bibRefCitation>
. These morphological similarities are most event in the mouth parts (e.g., tridentate or bidentate
<emphasis id="B958EAECFFFD5117204AFA13FF26F9B2" italics="true" pageId="3" pageNumber="52">lacinia mobilis</emphasis>
), setation and shape of the maxilliped, and the setation of the pereopods (e.g., presence of spiniform setae on the anterior margin of the ischium for pereopods 26,), and propodal setae on pereopod 4 extending to or beyond the dactylus. Since members of the genus
<taxonomicName id="4C2C4D7DFFFD511727CDF9BFFC8DF9EA" ID-CoL="84MQM" box="[713,833,1628,1652]" class="Malacostraca" family="Apseudidae" genus="Bunakenia" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD511727CDF9BFFC8DF9EA" box="[713,833,1628,1652]" italics="true" pageId="3" pageNumber="52">Bunakenia</emphasis>
</taxonomicName>
appear to share some distinctive homologous characters with them,
<taxonomicName id="4C2C4D7DFFFD51172448F963FDFBF906" ID-CoL="84L7Y" box="[332,567,1664,1688]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD51172448F963FDFBF906" box="[332,567,1664,1688]" italics="true" pageId="3" pageNumber="52">Akanthoparapseudes</emphasis>
</taxonomicName>
and “Parapseudid? species A” may represent transitional links between the families
<taxonomicName id="4C2C4D7DFFFD511725FCF947FEB0F922" ID-CoL="84JW5" box="[248,380,1700,1724]" class="Malacostraca" family="Apseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Apseudidae</taxonomicName>
and
<taxonomicName id="4C2C4D7DFFFD511724B7F947FD99F922" ID-CoL="84KM3" box="[435,597,1700,1724]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Parapseudidae</taxonomicName>
</paragraph>
<paragraph id="8B9336FEFFFD511025C1F924FCDBFD89" blockId="3.[151,1437,151,2012]" lastBlockId="4.[151,1437,151,535]" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
Some of the systematic problems exhibited by several apseudomorphan families are exemplified by overlapping characters within the
<taxonomicName id="4C2C4D7DFFFD511724C7F90FFDA9F89A" ID-CoL="84KM3" box="[451,613,1772,1796]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Parapseudidae</taxonomicName>
and
<taxonomicName id="4C2C4D7DFFFD51172798F90FFCE8F89A" ID-CoL="84JW5" box="[668,804,1772,1796]" class="Malacostraca" family="Apseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Apseudidae</taxonomicName>
. As indicated by
<bibRefCitation id="EFBD4B0FFFFD511726ECF90FFBB9F89A" author="Gutu" box="[1000,1141,1772,1796]" pageId="3" pageNumber="52" refString="Gutu, M. (2007) Contribution to the knowledge of the Indo-West-Pacific Apseudomorpha (Crustacea: Tanaidacea). Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 50, 47 - 86." type="journal article" year="2007">Guţu (2007)</bibRefCitation>
, the designation of a variety of apseudomophan genera during the past 20 years, has eroded and diluted some of the basic differences between these two families. This is especially true for some of the genera discussed in this paper. M. Guţu (per. comm., 2011) considers the most consistent family character for the
<taxonomicName id="4C2C4D7DFFFD511726C4F8B4FBAEF8F1" ID-CoL="84KM3" box="[960,1122,1879,1903]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Parapseudidae</taxonomicName>
to be the extension of the propodal setae on pereopod 4 to or beyond the tip of the dactylus. Except for the small deep-water family
<taxonomicName id="4C2C4D7DFFFD5117204AF89FFED7F826" ID-CoL="84K8C" class="Malacostraca" family="Gigantapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Gigantapseudidae</taxonomicName>
and the apseudid genus
<taxonomicName id="4C2C4D7DFFFD51172729F843FD69F826" ID-CoL="84MQM" box="[557,677,1952,1976]" class="Malacostraca" family="Apseudidae" genus="Bunakenia" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAECFFFD51172729F843FD69F826" box="[557,677,1952,1976]" italics="true" pageId="3" pageNumber="52">Bunakenia</emphasis>
</taxonomicName>
, this character appears to hold for all of the presently known parapseudid species (see
<bibRefCitation id="EFBD4B0FFFFD51172471F827FDA7F842" author="Gutu" box="[373,619,1988,2012]" pageId="3" pageNumber="52" refString="Gutu, M., &amp; Heard., R. W. (2002) A new genus and four new species of parapseudid and sphyrapid apseudomorphans (Crustacea: Tanaidacea) from the Caribbean Sea and the Gulf of Mexico. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 44, 69 - 92." type="journal article" year="2002">
Guţu and
<collectingCountry id="F33B766EFFFD511724E0F827FDE5F842" box="[484,553,1988,2012]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="52">Heard</collectingCountry>
2002
</bibRefCitation>
;
<bibRefCitation id="EFBD4B0FFFFD51172773F827FD3DF842" author="Gutu" box="[631,753,1988,2012]" pageId="3" pageNumber="52" refString="Gutu, M. (1998) Description of three new species of Tanaidacea (Crustacea) from the Tanzanian Coasts. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 40, 179 - 209." type="journal article" year="1998">Guţu 1998</bibRefCitation>
,
<bibRefCitation id="EFBD4B0FFFFD511727F9F827FCF4F842" author="Gutu" box="[765,824,1988,2012]" pageId="3" pageNumber="52" refString="Gutu, M. (2006) New Apseudomorph Taxa of the World Ocean: Crustacea, Tanaidacea. Curtea Veche, Bucharest, Romania. 307 pp." type="book" year="2006">2006</bibRefCitation>
;
<bibRefCitation id="EFBD4B0FFFFD51172640F827FB51F842" author="Gutu" box="[836,1181,1988,2012]" pageId="3" pageNumber="52" refString="Gutu, M., &amp; Angsupanich, S. (2004 a) Description of two new genera and species of Tanaidacea (Crustacea) from shallow waters of the Andaman Sea, Thailand. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 46, 29 - 44." type="journal article" year="2004" yearSuffix="a">Guţu and Angsupanich 2004 a</bibRefCitation>
,
<bibRefCitation id="EFBD4B0FFFFD511721ADF827FB76F842" author="Gutu" box="[1193,1210,1988,2012]" pageId="3" pageNumber="52" refString="Gutu, M. &amp; Angsupanich. S. (2004 b) Two new parapseudid species and some first records of Tanaidacea (Crustacea: Peracarida) from Thailand. Travaux du Museum Nationa l d´Histoire naturelle &quot; Grigore Antipa &quot;, 47, 75 - 87." type="journal article" year="2004" yearSuffix="b">b</bibRefCitation>
). The occurrence of a reduced coxal process of the first pereopod on some parapseudids (e.g.,
<emphasis id="B958EAECFFFA511026C8FF7BFA8DFF2E" box="[972,1345,152,176]" italics="true" pageId="4" pageNumber="53">
<taxonomicName id="4C2C4D7DFFFA511026C8FF7BFB87FF2E" ID-CoL="84R6S" box="[972,1099,152,176]" class="Malacostraca" family="Parapseudidae" genus="Longipedis" kingdom="Animalia" order="Tanaidacea" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">Longipedis</taxonomicName>
,
<taxonomicName id="4C2C4D7DFFFA51102152FF7BFA8DFF2E" ID-CoL="84L7Y" box="[1110,1345,152,176]" class="Malacostraca" family="Parapseudidae" genus="Akanthoparapseudes" kingdom="Animalia" order="Tanaidacea" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">Akanthoparapseudes</taxonomicName>
</emphasis>
), weakens this once definitive morphological maker for the separation of the two families making it difficult to identify unique delineating characters to distinguish them. As mentioned by
<bibRefCitation id="EFBD4B0FFFFA511026A7FF3CFB83FF69" author="Bamber" box="[931,1103,223,247]" pageId="4" pageNumber="53" refString="Bamber, R. N. (1998) Tanaidaceans (Crustacea, Peracarida) from the Southeast of the South China Sea. Asian Marine Biology, 15, 169 - 197." type="journal article" year="1998">Bamber (1998)</bibRefCitation>
and
<bibRefCitation id="EFBD4B0FFFFA5110218EFF03FA54FF69" author="Gutu" box="[1162,1432,223,248]" pageId="4" pageNumber="53" refString="Gutu, M., &amp; Heard., R. W. (2002) A new genus and four new species of parapseudid and sphyrapid apseudomorphans (Crustacea: Tanaidacea) from the Caribbean Sea and the Gulf of Mexico. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 44, 69 - 92." type="journal article" year="2002">
Guţu and
<collectingCountry id="F33B766EFFFA511021F9FF3CFA8EFF69" box="[1277,1346,223,247]" name="Heard Island and McDonald Islands" pageId="4" pageNumber="53">Heard</collectingCountry>
(2002)
</bibRefCitation>
, another factor complicating apseudomorphan systematics, is the presence of a distinctive leaf-shaped spine on the inner distal margin of the maxillipedal endite in some parapseudid genera (e.g.,
<emphasis id="B958EAECFFFA5110210BFECBFA9FFEDE" box="[1039,1363,296,320]" italics="true" pageId="4" pageNumber="53">
<taxonomicName id="4C2C4D7DFFFA5110210BFECBFB17FEDE" ID-CoL="84RHB" box="[1039,1243,296,320]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">Pakistanapseudes</taxonomicName>
,
<taxonomicName id="4C2C4D7DFFFA511021E1FECBFA9FFEDE" ID-CoL="84TDJ" box="[1253,1363,296,320]" class="Malacostraca" family="Parapseudidae" genus="Saltipedis" kingdom="Animalia" order="Tanaidacea" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">Saltipedis</taxonomicName>
</emphasis>
). This “leaf-shaped spine appears to be similar to those found in the same location on members of the apseudid subfamily
<taxonomicName id="4C2C4D7DFFFA51102593FE8CFE0BFE19" ID-CoL="85GKQ" authority="Sieg, 1983" authorityName="Sieg" authorityYear="1983" box="[151,455,367,391]" class="Malacostraca" family="Apseudidae" kingdom="Animalia" order="Tanaidacea" pageId="4" pageNumber="53" phylum="Arthropoda" rank="subFamily" subFamily="Leviapseudinae">Leviapseudinae Sieg, 1983</taxonomicName>
and family
<taxonomicName id="4C2C4D7DFFFA51102755FE8CFC4BFE19" ID-CoL="84KXL" authority="Gutu, 1972" authorityName="Gutu" authorityYear="1972" box="[593,903,367,392]" class="Malacostraca" family="Whiteleggiidae" kingdom="Animalia" order="Tanaidacea" pageId="4" pageNumber="53" phylum="Arthropoda" rank="family">Whiteleggiidae Guţu, 1972</taxonomicName>
; whether this condition is due to homoplasy or has systematic significance remains to be seen. Based our present limited knowledge, an adequate understanding of the present systematic status of the family
<taxonomicName id="4C2C4D7DFFFA5110278AFE54FCFCFE51" ID-CoL="84KM3" box="[654,816,439,463]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="4" pageNumber="53" phylum="Arthropoda" rank="family">Parapseudidae</taxonomicName>
is beyond the scope of this study. Its resolution will require detailed morphological studies with robust systematic analyses (e.g., cladistics), coupled with new information derived from comparative molecular genetic studies.
</paragraph>
</subSubSection>
</treatment>
</document>