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<document id="95D768193FC772BA9F63AF35D966FD26" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE3B0FF2BFF75F9D5BAA4FE98" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madagaster simplissima Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="187" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="184" updateTime="1698480684898" updateUser="ExternalLinkService">
<mods:mods id="F2CB8CD0844252FB9222609AFA3C85C7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A326CFBEF8A4BD34F93C5D192F4ED804">
<mods:title id="242718562CF569D89FD2DBE2D5AD37A1">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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<mods:namePart id="33BC34FA48E549357BA3E29B9F3F94BC">Perkins, Philip D.</mods:namePart>
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<mods:title id="BB62A25270E754FF84C7E5D69A56C8FC">Zootaxa</mods:title>
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<mods:part id="09BE462EC61C9F282A193C9B14B334B0">
<mods:date id="D243F64148C92FCD9F5FC8BC0636DF64">2017</mods:date>
<mods:detail id="34ACEC23672DAB012543B8F785A61A45" type="pubDate">
<mods:number id="F386F5EA1D0BA7D9D5F5616E0EE3E778">2017-11-03</mods:number>
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<mods:classification id="B0496B99A3C98802B890D147D5683041">journal article</mods:classification>
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<mods:identifier id="4ACC09EAB411082CDB122323B55ACF54" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<treatment id="038587BBE3B0FF2BFF75F9D5BAA4FE98" ID-DOI="http://doi.org/10.5281/zenodo.5323732" ID-GBIF-Taxon="137233387" ID-Zenodo-Dep="5323732" LSID="urn:lsid:plazi:treatment:038587BBE3B0FF2BFF75F9D5BAA4FE98" httpUri="http://treatment.plazi.org/id/038587BBE3B0FF2BFF75F9D5BAA4FE98" lastPageId="186" lastPageNumber="187" pageId="183" pageNumber="184">
<subSubSection id="C3366526E3B0FF26FF75F9D5BB98F92E" pageId="183" pageNumber="184" type="nomenclature">
<paragraph id="8B9336ADE3B0FF26FF75F9D5BB66F90F" blockId="183.[151,676,1648,1708]" box="[151,602,1648,1674]" pageId="183" pageNumber="184">
<heading id="D0DB81C1E3B0FF26FF75F9D5BB66F90F" bold="true" box="[151,602,1648,1674]" fontSize="11" level="1" pageId="183" pageNumber="184" reason="1">
<emphasis id="B958EABFE3B0FF26FF75F9D5BB66F90F" bold="true" box="[151,602,1648,1674]" pageId="183" pageNumber="184">
<taxonomicName id="4C2C4D2EE3B0FF26FF75F9D5B886F90F" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,442,1648,1674]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="simplissima" status="sp. nov.">
<emphasis id="B958EABFE3B0FF26FF75F9D5B886F90F" bold="true" box="[151,442,1648,1674]" italics="true" pageId="183" pageNumber="184">Madagaster simplissima</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B57C4E3B0FF26FE28F9D5BB66F90F" box="[458,602,1648,1674]" pageId="183" pageNumber="184" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE3B0FF26FF75F936BB98F92E" blockId="183.[151,676,1648,1708]" box="[151,676,1683,1708]" pageId="183" pageNumber="184">
<figureCitation id="13172A28E3B0FF26FF75F936B9C3F929" box="[151,255,1683,1708]" captionStart="FIGURE 176" captionStartId="184.[151,250,1520,1542]" captionText="FIGURE 176. Madagaster simplissima, holotype, habitus dorsal, lateral, and ventral." pageId="183" pageNumber="184">Figs. 176</figureCitation>
(habitus), 178 (aedeagus), 301 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B0FF26FF75F97EBB26F8BE" pageId="183" pageNumber="184" type="materials_examined">
<paragraph id="8B9336ADE3B0FF26FF75F97EBB26F8BE" blockId="183.[151,1437,1755,2032]" pageId="183" pageNumber="184">
<emphasis id="B958EABFE3B0FF26FF75F97EB882F971" bold="true" box="[151,446,1755,1780]" pageId="183" pageNumber="184">
<typeStatus id="5497880FE3B0FF26FF75F97EB9EEF971" box="[151,210,1755,1780]" pageId="183" pageNumber="184">Type</typeStatus>
Material.
<typeStatus id="5497880FE3B0FF26FEB3F97EB882F971" box="[337,446,1755,1780]" pageId="183" pageNumber="184" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingRegion id="49E8F84FE3B0FF26FDC2F97EBB83F971" box="[544,703,1755,1780]" country="Madagascar" name="Fianarantsoa" pageId="183" pageNumber="184">
<emphasis id="B958EABFE3B0FF26FDC2F97EBB83F971" bold="true" box="[544,703,1755,1780]" pageId="183" pageNumber="184">Fianarantsoa</emphasis>
</collectingRegion>
,
<quantity id="4CD49B48E3B0FF26FD2CF97EBA2AF971" box="[718,790,1755,1780]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="183" pageNumber="184" unit="km" value="32.0">32 km</quantity>
S Ambositra, in seepage over exposed granite cliff face, elev.
<quantity id="4CD49B48E3B0FF26FF30F95AB816F892" box="[210,298,1791,1815]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="183" pageNumber="184" unit="m" value="1400.0">1400 m</quantity>
,
<geoCoordinate id="EE18506AE3B0FF26FED4F95AB8A3F892" box="[310,415,1791,1815]" direction="south" orientation="latitude" pageId="183" pageNumber="184" precision="925" value="-20.716667">20° 43' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE3B0FF26FE4EF95ABB2AF892" box="[428,534,1791,1816]" direction="east" orientation="longitude" pageId="183" pageNumber="184" precision="925" value="47.266666">47° 16' E</geoCoordinate>
,
<date id="FF92106DE3B0FF26FDC1F95ABB9DF892" box="[547,673,1791,1816]" pageId="183" pageNumber="184" value="1990-03-25">25 iii 1990</date>
, W. E. Steiner, C. Kremen, V. Razafimahatratra (
<collectionCode id="ED3DAE68E3B0FF26FB2AF95ABC23F892" box="[1224,1311,1791,1815]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="183" pageNumber="184">USNM</collectionCode>
).
<typeStatus id="5497880FE3B0FF26FAD1F8A5BCA5F89D" box="[1331,1433,1792,1816]" pageId="183" pageNumber="184" type="paratype">Paratype</typeStatus>
: Same data as
<typeStatus id="5497880FE3B0FF26FED2F886B8AEF8B9" box="[304,402,1827,1852]" pageId="183" pageNumber="184" type="holotype">holotype</typeStatus>
(1
<collectionCode id="ED3DAE68E3B0FF26FE55F886BB31F8BE" box="[439,525,1827,1851]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="183" pageNumber="184">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B0FF29FF25F8E2B9C4FF7D" lastPageId="184" lastPageNumber="185" pageId="183" pageNumber="184" type="diagnosis">
<paragraph id="8B9336ADE3B0FF29FF25F8E2B9C4FF7D" blockId="183.[151,1437,1755,2032]" lastBlockId="184.[151,1436,151,248]" lastPageId="184" lastPageNumber="185" pageId="183" pageNumber="184">
<emphasis id="B958EABFE3B0FF26FF25F8E2B8EEF8E5" bold="true" box="[199,466,1863,1888]" pageId="183" pageNumber="184">Differential Diagnosis.</emphasis>
Differentiated from other members of
<emphasis id="B958EABFE3B0FF26FC61F8E2BD35F8E5" box="[899,1033,1863,1888]" italics="true" pageId="183" pageNumber="184">Madagaster</emphasis>
, except
<taxonomicName id="4C2C4D2EE3B0FF26FB85F8EDBDF1F8DA" box="[1127,1229,1863,1887]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B958EABFE3B0FF26FB85F8EDBDF1F8DA" box="[1127,1229,1863,1887]" italics="true" pageId="183" pageNumber="184">M. franzi</emphasis>
</taxonomicName>
, by the small size. Very similar in body size and dorsal habitus to
<taxonomicName id="4C2C4D2EE3B0FF26FD27F8C9BA0CF806" box="[709,816,1899,1923]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B958EABFE3B0FF26FD27F8C9BA0CF806" box="[709,816,1899,1923]" italics="true" pageId="183" pageNumber="184">M. franzi</emphasis>
</taxonomicName>
(
<figureCitation id="13172A28E3B0FF26FCA1F8CEBA8EF801" box="[835,946,1899,1924]" captionStart="FIGURE 175" captionStartId="183.[151,250,1551,1573]" captionText="FIGURE 175. Madagaster franzi, holotype, habitus dorsal, lateral, and ventral." pageId="183" pageNumber="184">Figs. 175</figureCitation>
,
<figureCitation id="13172A28E3B0FF26FC23F8CEBAD3F801" box="[961,1007,1899,1924]" captionStart="FIGURE 176" captionStartId="184.[151,250,1520,1542]" captionText="FIGURE 176. Madagaster simplissima, holotype, habitus dorsal, lateral, and ventral." pageId="183" pageNumber="184">176</figureCitation>
); reliable determinations will require careful examination of the male genitalia, in temporary microslide mounts (
<figureCitation id="13172A28E3B0FF26FC18F82ABD98F822" box="[1018,1188,1935,1960]" captionStart="FIGURES 177 – 178" captionStartId="185.[151,264,1725,1748]" captionText="FIGURES 177 – 178. Madagaster, aedeagi of holotypes." pageId="183" pageNumber="184">Figs. 177, 178</figureCitation>
). The aedeagus of
<taxonomicName id="4C2C4D2EE3B0FF26FA9FF835B827F849" authorityName="Perkins" authorityYear="2017" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="simplissima">
<emphasis id="B958EABFE3B0FF26FA9FF835B827F849" italics="true" pageId="183" pageNumber="184">M. simplissima</emphasis>
</taxonomicName>
is almost twice as long as the aedeagus of
<taxonomicName id="4C2C4D2EE3B0FF26FCEBF811BA4DF84E" box="[777,881,1971,1995]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B958EABFE3B0FF26FCEBF811BA4DF84E" box="[777,881,1971,1995]" italics="true" pageId="183" pageNumber="184">M. franzi</emphasis>
</taxonomicName>
; the gonopore bearing tubule is shorter and only slightly curved in
<taxonomicName id="4C2C4D2EE3B0FF26FE80F87DBB30F875" authorityName="Perkins" authorityYear="2017" box="[354,524,2007,2032]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="simplissima">
<emphasis id="B958EABFE3B0FF26FE80F87DBB30F875" box="[354,524,2007,2032]" italics="true" pageId="183" pageNumber="184">M. simplissima</emphasis>
</taxonomicName>
, whereas in
<taxonomicName id="4C2C4D2EE3B0FF26FD78F87DBA3CF86A" box="[666,768,2007,2031]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B958EABFE3B0FF26FD78F87DBA3CF86A" box="[666,768,2007,2031]" italics="true" pageId="183" pageNumber="184">M. franzi</emphasis>
</taxonomicName>
it is longer and sinuate; the tubule, in lateral view, emerges from the dorsal part of the apex of the main piece in
<taxonomicName id="4C2C4D2EE3BFFF29FCE4FF3CBA53FF35" box="[774,879,152,176]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B958EABFE3BFFF29FCE4FF3CBA53FF35" box="[774,879,152,176]" italics="true" pageId="184" pageNumber="185">M. franzi</emphasis>
</taxonomicName>
, but is more centrally located at the apex of
<taxonomicName id="4C2C4D2EE3BFFF29FA9FFF3CB827FF50" authorityName="Perkins" authorityYear="2017" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="simplissima">
<emphasis id="B958EABFE3BFFF29FA9FFF3CB827FF50" italics="true" pageId="184" pageNumber="185">M. simplissima</emphasis>
</taxonomicName>
. In both species the paramere setae are located on both the lateral and medial surfaces, not just at the margins.
</paragraph>
</subSubSection>
<caption id="DF536625E3BFFF29FF75FA55BAC6F983" box="[151,1018,1520,1542]" pageId="184" pageNumber="185">
<paragraph id="8B9336ADE3BFFF29FF75FA55BAC6F983" blockId="184.[151,1018,1520,1542]" box="[151,1018,1520,1542]" pageId="184" pageNumber="185">
<emphasis id="B958EABFE3BFFF29FF75FA55B810F980" bold="true" box="[151,300,1520,1542]" pageId="184" pageNumber="185">FIGURE 176.</emphasis>
<taxonomicName id="4C2C4D2EE3BFFF29FED0FA55BB16F983" authorityName="Perkins" authorityYear="2017" box="[306,554,1520,1542]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="simplissima">
<emphasis id="B958EABFE3BFFF29FED0FA55BB16F983" box="[306,554,1520,1542]" italics="true" pageId="184" pageNumber="185">Madagaster simplissima</emphasis>
</taxonomicName>
, holotype, habitus dorsal, lateral, and ventral.
</paragraph>
</caption>
<subSubSection id="C3366526E3BFFF2BFF25F988BB0AFF7D" lastPageId="186" lastPageNumber="187" pageId="184" pageNumber="185" type="description">
<paragraph id="8B9336ADE3BFFF29FF25F988B85AF90B" blockId="184.[151,1436,1581,2003]" pageId="184" pageNumber="185">
<emphasis id="B958EABFE3BFFF29FF25F988B86DF9C3" bold="true" box="[199,337,1581,1606]" pageId="184" pageNumber="185">Description</emphasis>
. Size:
<typeStatus id="5497880FE3BFFF29FE41F988BB39F9C3" box="[419,517,1581,1606]" pageId="184" pageNumber="185" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.61/0.68; head width 0.44; pronotum 0.36/0.59, PA 0.47, PB 0.50; elytra 1.02/0.68. Dorsum and venter dark brown to piceous, legs brown, ocelli light brown.
</paragraph>
<paragraph id="8B9336ADE3BFFF29FF25F93FBA26F8C6" blockId="184.[151,1436,1581,2003]" pageId="184" pageNumber="185">Clypeus and frons punctulate and, in places, microreticulate; setae very fine and sparse, recumbent Labrum bilobed, with deep V-shaped apicomedian notch, microreticulate and with sparse small setae. Labrum originating at anterior of very slightly overhanging margin of clypeus. Labroclypeal suture moderately deep. Short, deep sulcus in front of each ocellus. Frons with deep concavity between ocelli. Maxillary palpi ca. as along as antenna. Antenna with penultimate antennomere much larger than ultimate.</paragraph>
<paragraph id="8B9336ADE3BFFF29FF25F8EAB8C2F856" blockId="184.[151,1436,1581,2003]" pageId="184" pageNumber="185">Pronotum with small, moderately dense granules, each with short recumbent seta; disc with two narrowly separated midlongitudinal fovea, anterior ca. twice as long as posterior; each side with two admedian and two sublateral foveae; pronotum slightly cordiform, lateral margin rounded except emarginate near base; postocular emarginations moderately deep.</paragraph>
<caption id="DF536625E3BEFF28FF75F918BBEEF956" box="[151,722,1725,1748]" pageId="185" pageNumber="186">
<paragraph id="8B9336ADE3BEFF28FF75F918BBEEF956" blockId="185.[151,722,1725,1748]" box="[151,722,1725,1748]" pageId="185" pageNumber="186">
<emphasis id="B958EABFE3BEFF28FF75F918B851F956" bold="true" box="[151,365,1725,1748]" pageId="185" pageNumber="186">FIGURES 177–178.</emphasis>
<emphasis id="B958EABFE3BEFF28FE91F918B8D0F956" box="[371,492,1725,1747]" italics="true" pageId="185" pageNumber="186">Madagaster</emphasis>
, aedeagi of holotypes.
</paragraph>
</caption>
<paragraph id="8B9336ADE3BEFF28FF25F8A5B87AF8E5" blockId="185.[151,1436,1792,2033]" pageId="185" pageNumber="186">Elytral suture very weakly raised. Each elytron with four non-interrupted lines, each consisting of unilinear row of very slightly raised granules; areas between granular lines with four very irregular rows of granules. Elytral apices truncate.</paragraph>
<paragraph id="8B9336ADE3BEFF28FF25F8C8B9CBF848" blockId="185.[151,1436,1792,2033]" pageId="185" pageNumber="186">Mentum and postmentum densely micropunctulate, very sparsely pubescent; Genae micropunctulate, dull, concave behind cardo, without posterior ridge. Subocular groove for antenna narrow. Posterior margin of eye not carinate.</paragraph>
<paragraph id="8B9336ADE3BEFF2BFF25F87CBB0AFF7D" blockId="185.[151,1436,1792,2033]" lastBlockId="186.[151,1436,151,285]" lastPageId="186" lastPageNumber="187" pageId="185" pageNumber="186">
Prosternum without midlongitudinal carina. Mesoventrite without tubercle on intercoxal process. Metaventrite densely micropunctulate and with very short and very dense vestiture (plastron?); midlongitudinal depression moderately deep (
<figureCitation id="13172A28E3BDFF2BFE82FF19B8FFFF50" box="[352,451,188,213]" captionStart="FIGURE 176" captionStartId="184.[151,250,1520,1542]" captionText="FIGURE 176. Madagaster simplissima, holotype, habitus dorsal, lateral, and ventral." pageId="186" pageNumber="187">Fig. 176</figureCitation>
). First four abdominal ventrites and anterior margin of fifth with very dense, recumbent pubescence (plastron?). Legs simple.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3BDFF2BFF25FEA1BAA4FE98" box="[199,920,260,285]" pageId="186" pageNumber="187" type="etymology">
<paragraph id="8B9336ADE3BDFF2BFF25FEA1BAA4FE98" blockId="186.[151,1436,151,285]" box="[199,920,260,285]" pageId="186" pageNumber="187">
<emphasis id="B958EABFE3BDFF2BFF25FEA1B872FE98" bold="true" box="[199,334,260,285]" pageId="186" pageNumber="187">Etymology.</emphasis>
Named in reference to the very simple morphology.
</paragraph>
</subSubSection>
</treatment>
</document>
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