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<mods:title id="434DEF77A15A97CCF6440C1FD4EB58D1">A new deep-water goatfish of the genus Upeneus (Mullidae) from northern Australia and the Philippines, with a taxonomic account of U. subvittatus and remarks on U. mascareinsis</mods:title>
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<subSubSection id="3EF8FCBE820AFFAFFF5AFBB5FE49FBE5" box="[151,507,1047,1073]" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="765DAF35820AFFAFFF5AFBB5FE49FBE5" blockId="1.[151,507,1047,1073]" box="[151,507,1047,1073]" pageId="1" pageNumber="62">
<heading id="2D151859820AFFAFFF5AFBB5FE49FBE5" bold="true" box="[151,507,1047,1073]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="44967327820AFFAFFF5AFBB5FE49FBE5" bold="true" box="[151,507,1047,1073]" pageId="1" pageNumber="62">
Genus
<taxonomicName id="B1E2D4B6820AFFAFFF23FBB5FE49FBE5" ID-CoL="84ZF" ID-ENA="115430" authority="Cuvier, 1829" authorityName="Cuvier" authorityYear="1829" box="[238,507,1047,1073]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="44967327820AFFAFFF23FBB5FEEAFBE5" bold="true" box="[238,344,1047,1073]" italics="true" pageId="1" pageNumber="62">Upeneus</emphasis>
Cuvier, 1829
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE820AFFAFFF5AFBC2FE01FBA0" box="[151,435,1118,1141]" pageId="1" pageNumber="62" type="reference_group">
<paragraph id="765DAF35820AFFAFFF5AFBC2FE01FBA0" blockId="1.[151,1105,1118,1171]" box="[151,435,1118,1141]" pageId="1" pageNumber="62">
<taxonomicName id="B1E2D4B6820AFFAFFF5AFBC2FE1DFBA0" ID-CoL="84ZF" authority="Cuvier, 1829: 157" authorityName="Cuvier" authorityPageNumber="157" authorityYear="1829" box="[151,431,1118,1141]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="44967327820AFFAFFF5AFBC2FF42FBA1" box="[151,240,1120,1141]" italics="true" pageId="1" pageNumber="62">Upeneus</emphasis>
Cuvier, 1829: 157
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE820AFFAFFF5AFBDCFBE3FB47" box="[151,1105,1149,1171]" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="765DAF35820AFFAFFF5AFBDCFBE3FB47" blockId="1.[151,1105,1118,1171]" box="[151,1105,1149,1171]" pageId="1" pageNumber="62">
<typeStatus id="A9591197820AFFAFFF5AFBDCFF7AFB47" box="[151,200,1150,1171]" pageId="1" pageNumber="62">Type</typeStatus>
species
<taxonomicName id="B1E2D4B6820AFFAFFED2FBDFFDD0FB47" authority="Forsskal, 1775" authorityName="Forsskal" authorityYear="1775" box="[287,610,1149,1171]" class="Actinopterygii" family="Mullidae" genus="Mullus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="44967327820AFFAFFED2FBDFFE04FB47" box="[287,438,1149,1171]" italics="true" pageId="1" pageNumber="62">Mullus vittatus</emphasis>
(Forsskål, 1775)
</taxonomicName>
by subsequent designation of
<bibRefCitation id="1273D2C4820AFFAFFC58FBDCFBFFFB47" author="Desmarest" box="[917,1101,1149,1171]" pageId="1" pageNumber="62" refString="Desmarest, E. (1856) Reptiles et poissons. In: Chenu, J. G. (Ed.), Encyclopedie d'histoire naturelle; ou, Traite complet de cette science d'apres les travaux des naturalistes les plus eeminents de toutes les epoques (Paris), 19, 1 - 360 + 1 - 62." type="journal article" year="1856">Desmarest (1856)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE820AFFAFFF5AFB1DFB8CFA58" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="765DAF35820AFFAFFF5AFB1DFB8CFA58" blockId="1.[151,1436,1215,2033]" pageId="1" pageNumber="62">
<emphasis id="44967327820AFFAFFF5AFB1DFEA3FB0C" bold="true" box="[151,273,1215,1240]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
Dorsal fins VII or VIII + 9; anal fin I, 6; pectoral-fin rays 1217; principal caudal-fin rays 7 + 8 (median 13 branched); gill rakers 49 + 1324 = 1833; lateral-line scales 2839, lateral line complete; small scales present basally on second dorsal, anal and caudal fins; small teeth present on vomer, palatines and jaws, multiserial and villiform on jaws; body oblong, slightly compressed; barbel length in adults 47 times in SL; snout length 711 times in SL, subequal to postorbital length (710 times in SL); in fresh fish lateral body stripes and/or caudal-fin bars of differing colours, dark caudal-fin bars frequently retained on preserved fish.
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE820AFFAFFF0AFA35FC2DFA00" pageId="1" pageNumber="62" type="distribution">
<paragraph id="765DAF35820AFFAFFF0AFA35FC2DFA00" blockId="1.[151,1436,1215,2033]" pageId="1" pageNumber="62">
<emphasis id="44967327820AFFAFFF0AFA35FED2FA64" bold="true" box="[199,352,1431,1456]" pageId="1" pageNumber="62">Distribution.</emphasis>
In all major oceans, tropical to subtropical, only a single species in the Atlantic and two in the Mediterranean, both immigrants from the Red Sea (Ben
<bibRefCitation id="1273D2C4820AFFAFFCC2FA1EFC20FA00" author="Tuvia" box="[783,914,1468,1492]" pageId="1" pageNumber="62" refString="Ben-Tuvia, A. (1966) Red Sea fishes recently found in the Mediterranean. Copeia, 1966, 254 - 275. Boeseman, M. (1947) Revision of the fishes collected by Burger and Von Siebold in Japan. Zoologische Mededelingen, 28, i - vii + 1 - 242, pls. 1 - 5." type="journal article" year="1966">Tuvia 1966</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE820AFFACFF0AFA42FA27FEC8" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="discussion">
<paragraph id="765DAF35820AFFAFFF0AFA42FEADF88B" blockId="1.[151,1436,1215,2033]" pageId="1" pageNumber="62">
<emphasis id="44967327820AFFAFFF0AFA42FE89FA2C" bold="true" box="[199,315,1504,1528]" pageId="1" pageNumber="62">Remarks.</emphasis>
We recognize 30 species as valid. One single species,
<taxonomicName id="B1E2D4B6820AFFAFFC67FA43FBE3FA2C" box="[938,1105,1504,1528]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="filifer">
<emphasis id="44967327820AFFAFFC67FA43FBE3FA2C" box="[938,1105,1504,1528]" italics="true" pageId="1" pageNumber="62">Upeneus filifer</emphasis>
</taxonomicName>
, and four species groups can be distinguished based on number of dorsal spines and gill rakers, length of longest dorsal-fin spine, length of pelvic and pectoral fins, and presence or absence of caudal-fin bars and mid-lateral body stripes (
<bibRefCitation id="1273D2C4820AFFAFFAE9F985FEF9F9B0" author="Uiblein" pageId="1" pageNumber="62" refString="Uiblein F, &amp; Heemstra, P. C (2010) A taxonomic review of the Western Indian Ocean goatfishes of the genus Upeneus (Family Mullidae) with descriptions of four new species. Smithiana Bulletin, 11, 35 - 71." type="journal article" year="2010">Uiblein &amp; Heemstra 2010</bibRefCitation>
). Here,
<taxonomicName id="B1E2D4B6820AFFAFFE67F9EFFDCCF9B0" box="[426,638,1613,1636]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="stenopsis" status="sp. nov.">
<emphasis id="44967327820AFFAFFE67F9EFFDCCF9B0" box="[426,638,1613,1636]" italics="true" pageId="1" pageNumber="62">Upeneus stenopsis</emphasis>
</taxonomicName>
<emphasis id="44967327820AFFAFFD44F9EEFD7EF9B0" bold="true" box="[649,716,1612,1636]" pageId="1" pageNumber="62">
<taxonomicNameLabel id="5FA5CE5C820AFFAFFD44F9EEFD7EF9B0" box="[649,716,1612,1636]" pageId="1" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is added to the
<taxonomicName id="B1E2D4B6820AFFAFFC5EF9EFFC57F9B0" box="[915,997,1613,1636]" class="Actinopterygii" family="Mullidae" genus="Mullus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="44967327820AFFAFFC5EF9EFFC57F9B0" box="[915,997,1613,1636]" italics="true" pageId="1" pageNumber="62">vittatus</emphasis>
</taxonomicName>
group which includes
<taxonomicName id="B1E2D4B6820AFFAFFB38F9EFFE98F953" authority="Golani, 2001" authorityName="Golani" authorityYear="2001" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="davidaromi">
<emphasis id="44967327820AFFAFFB38F9EFFA2EF9B0" box="[1269,1436,1612,1636]" italics="true" pageId="1" pageNumber="62">U. davidaromi</emphasis>
<bibRefCitation id="1273D2C4820AFFAFFF5AF9CDFE98F953" author="Golani" box="[151,298,1647,1671]" pageId="1" pageNumber="62" refString="Golani, D. (2001) Upeneus davidaromi, a new deepwater goatfish (Osteichthyes, Mullidae) from the Red Sea. Israel Journal of Zoology, 47, 111 - 121." type="journal article" year="2001">Golani, 2001</bibRefCitation>
</taxonomicName>
(Red Sea),
<taxonomicName id="B1E2D4B6820AFFAFFE62F9D3FCE7F953" authority="Uiblein &amp; Heemstra, 2010" authorityName="Uiblein &amp; Heemstra" authorityYear="2010" box="[431,853,1647,1672]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="indicus">
<emphasis id="44967327820AFFAFFE62F9D3FD91F95C" box="[431,547,1648,1672]" italics="true" pageId="1" pageNumber="62">U. indicus</emphasis>
<bibRefCitation id="1273D2C4820AFFAFFDE6F9CDFCE7F953" author="Uiblein" box="[555,853,1647,1671]" pageId="1" pageNumber="62" refString="Uiblein F, &amp; Heemstra, P. C (2010) A taxonomic review of the Western Indian Ocean goatfishes of the genus Upeneus (Family Mullidae) with descriptions of four new species. Smithiana Bulletin, 11, 35 - 71." type="journal article" year="2010">Uiblein &amp; Heemstra, 2010</bibRefCitation>
</taxonomicName>
(SW
<collectingCountry id="0EF5EFA5820AFFAFFC5AF9CDFC67F953" box="[919,981,1647,1671]" name="India" pageId="1" pageNumber="62">India</collectingCountry>
),
<taxonomicName id="B1E2D4B6820AFFAFFC24F9D3FF7DF978" authority="Fourmanoir &amp; Gueze, 1967" authorityName="Fourmanoir &amp; Gueze" authorityYear="1967" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="mascareinsis">
<emphasis id="44967327820AFFAFFC24F9D3FB2FF95C" box="[1001,1181,1649,1672]" italics="true" pageId="1" pageNumber="62">U. mascareinsis</emphasis>
Fourmanoir &amp; Guézé, 1967
</taxonomicName>
(Mascarenes to
<collectingCountry id="0EF5EFA5820AFFAFFE5DF936FD97F978" box="[400,549,1684,1708]" name="Mozambique" pageId="1" pageNumber="62">Mozambique</collectingCountry>
),
<taxonomicName id="B1E2D4B6820AFFAFFDF3F937FC8BF978" authority="Poey, 1852" authorityName="Poey" authorityYear="1852" box="[574,825,1684,1709]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="parvus">
<emphasis id="44967327820AFFAFFDF3F937FD1DF978" box="[574,687,1685,1708]" italics="true" pageId="1" pageNumber="62">U. parvus</emphasis>
Poey, 1852
</taxonomicName>
(Western Central Atlantic),
<taxonomicName id="B1E2D4B6820AFFAFFB4EF937FEF8F91B" authority="Uiblein &amp; Heemstra, 2010" authorityName="Uiblein &amp; Heemstra" authorityYear="2010" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="suahelicus">
<emphasis id="44967327820AFFAFFB4EF937FAACF978" box="[1155,1310,1684,1708]" italics="true" pageId="1" pageNumber="62">U. suahelicus</emphasis>
<bibRefCitation id="1273D2C4820AFFAFFAE5F936FEF8F91B" author="Uiblein" pageId="1" pageNumber="62" refString="Uiblein F, &amp; Heemstra, P. C (2010) A taxonomic review of the Western Indian Ocean goatfishes of the genus Upeneus (Family Mullidae) with descriptions of four new species. Smithiana Bulletin, 11, 35 - 71." type="journal article" year="2010">Uiblein &amp; Heemstra, 2010</bibRefCitation>
</taxonomicName>
(Red Sea to
<collectingCountry id="0EF5EFA5820AFFAFFE13F915FDC6F91B" box="[478,628,1719,1743]" name="South Africa" pageId="1" pageNumber="62">South Africa</collectingCountry>
),
<taxonomicName id="B1E2D4B6820AFFAFFD45F91BFB36F91B" authority="Temminck &amp; Schlegel, 1843" authorityName="Temminck &amp; Schlegel" authorityYear="1843" box="[648,1156,1719,1744]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="subvittatus">
<emphasis id="44967327820AFFAFFD45F91BFC96F904" box="[648,804,1720,1744]" italics="true" pageId="1" pageNumber="62">U. subvittatus</emphasis>
(
<bibRefCitation id="1273D2C4820AFFAFFCF9F915FBCEF91B" author="Temminck" box="[820,1148,1719,1743]" pageId="1" pageNumber="62" refString="Temminck, C. J. &amp; Schlegel, H. (1843) Pisces. In: Fauna Japonica, sive descriptio animalium quae in itinere per Japoniam suscepto annis 1823 - 30 collegit, notis observationibus et adumbrationibus illustravit P. F. de Siebold. Fauna Japonica, sive descriptio animalium quae in itinere per Japoniam, Parts 2 - 4, 21 - 72." type="book chapter" year="1843">Temminck &amp; Schlegel, 1843</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="0EF5EFA5820AFFAFFB59F915FAB1F91B" box="[1172,1283,1719,1743]" name="Indonesia" pageId="1" pageNumber="62">Indonesia</collectingCountry>
to
<collectingCountry id="0EF5EFA5820AFFAFFAE5F915FAD8F91B" box="[1320,1386,1719,1743]" name="Japan" pageId="1" pageNumber="62">Japan</collectingCountry>
),
<taxonomicName id="B1E2D4B6820AFFAFFA4CF91BFDEBF920" authority="Uiblein &amp; Heemstra, 2010" authorityName="Uiblein &amp; Heemstra" authorityYear="2010" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="supravittatus">
<emphasis id="44967327820AFFAFFA4CF91BFE9BF920" italics="true" pageId="1" pageNumber="62">U. supravittatus</emphasis>
<bibRefCitation id="1273D2C4820AFFAFFEFDF97EFDEBF920" author="Uiblein" box="[304,601,1756,1780]" pageId="1" pageNumber="62" refString="Uiblein F, &amp; Heemstra, P. C (2010) A taxonomic review of the Western Indian Ocean goatfishes of the genus Upeneus (Family Mullidae) with descriptions of four new species. Smithiana Bulletin, 11, 35 - 71." type="journal article" year="2010">Uiblein &amp; Heemstra, 2010</bibRefCitation>
</taxonomicName>
(southern
<collectingCountry id="0EF5EFA5820AFFAFFD1DF97EFCB9F920" box="[720,779,1756,1780]" name="India" pageId="1" pageNumber="62">India</collectingCountry>
) and
<taxonomicName id="B1E2D4B6820AFFAFFC84F97FFAAAF920" authority="(Forsskal, 1775) (Indopacific)" authorityName="(Forsskal, 1775) (Indopacific" box="[841,1304,1756,1780]" class="Actinopterygii" family="Mullidae" genus="Mullus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="44967327820AFFAFFC84F97FFC0CF920" box="[841,958,1757,1780]" italics="true" pageId="1" pageNumber="62">U. vittatus</emphasis>
(Forsskål, 1775) (Indopacific)
</taxonomicName>
. This group can be distinguished by the following combination of characteristics: oblique dark bars on both caudal-fin lobes in fresh and preserved fish, 2533 total gill rakers, 1417 pectoral-fin rays, and pelvic-fin length 1.11.5 times in pectoral fin.
</paragraph>
<paragraph id="765DAF35820AFFACFF0AF8CEFA27FEC8" blockId="1.[151,1436,1215,2033]" lastBlockId="2.[151,1436,151,284]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
The three other species groups are the
<taxonomicName id="B1E2D4B6820AFFAFFD4AF8CFFD46F850" box="[647,756,1901,1924]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="44967327820AFFAFFD4AF8CFFD46F850" box="[647,756,1901,1924]" italics="true" pageId="1" pageNumber="62">japonicus</emphasis>
</taxonomicName>
group (which includes
<taxonomicName id="B1E2D4B6820AFFAFFBC8F8CFFAC1F850" authority="Lachner, 1954" authorityName="Lachner" authorityYear="1954" box="[1029,1395,1900,1924]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="asymmetricus">
<emphasis id="44967327820AFFAFFBC8F8CFFB76F850" box="[1029,1220,1901,1924]" italics="true" pageId="1" pageNumber="62">U. asymmetricus</emphasis>
Lachner, 1954
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820AFFAFFA4CF8CFFDB7F873" authority="Kim &amp; Nakaya, 2002" authorityName="Kim &amp; Nakaya" authorityYear="2002" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="australiae">
<emphasis id="44967327820AFFAFFA4CF8CFFEBBF87C" italics="true" pageId="1" pageNumber="62">U. australiae</emphasis>
Kim &amp; Nakaya, 2002
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820AFFAFFDDCF833FC29F873" authority="Randall &amp; Gueze, 1992" authorityName="Randall &amp; Gueze" authorityYear="1992" box="[529,923,1935,1960]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="francisi">
<emphasis id="44967327820AFFAFFDDCF833FD38F87C" box="[529,650,1936,1960]" italics="true" pageId="1" pageNumber="62">U. francisi</emphasis>
Randall &amp; Guézé, 1992
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820AFFAFFC64F833FB04F873" authority="Day, 1868" authorityName="Day" authorityYear="1868" box="[937,1206,1935,1960]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="guttatus">
<emphasis id="44967327820AFFAFFC64F833FB94F87C" box="[937,1062,1937,1960]" italics="true" pageId="1" pageNumber="62">U. guttatus</emphasis>
(Day, 1868)
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820AFFAFFB0FF833FE52F818" authority="Yamashita, Golani &amp; Motomura, 2011" authorityName="Yamashita, Golani &amp; Motomura" authorityYear="2011" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="itoui">
<emphasis id="44967327820AFFAFFB0FF833FAAAF87C" box="[1218,1304,1937,1960]" italics="true" pageId="1" pageNumber="62">U. itoui</emphasis>
<bibRefCitation id="1273D2C4820AFFAFFAEDF82DFE52F818" author="Yamashita" pageId="1" pageNumber="62" refString="Yamashita, Y., Golani, D. &amp; Motomura, H. (2011) A new species of Upeneus (Perciformes: Mullidae) from southern Japan. Zootaxa, 3107, 47 - 58." type="journal article" year="2011">Yamashita, Golani &amp; Motomura, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820AFFAFFE39F817FCD8F818" authority="Houttuyn, 1782" authorityName="Houttuyn" authorityYear="1782" box="[500,874,1972,1996]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="44967327820AFFAFFE39F817FD3FF818" box="[500,653,1973,1996]" italics="true" pageId="1" pageNumber="62">U. japonicus</emphasis>
(Houttuyn, 1782)
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820AFFAFFCB2F817FA86F818" authority="Ben-Tuvia &amp; Golani, 1989" authorityName="Ben-Tuvia &amp; Golani" authorityYear="1989" box="[895,1332,1972,1996]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="pori">
<emphasis id="44967327820AFFAFFCB2F817FC6BF818" box="[895,985,1973,1996]" italics="true" pageId="1" pageNumber="62">U. pori</emphasis>
Ben-Tuvia &amp; Golani, 1989
</taxonomicName>
, and
<taxonomicName id="B1E2D4B6820AFFAFFA4CF817FDDEF824" authority="Uiblein &amp; Heemstra 2011" authorityName="Uiblein &amp; Heemstra" authorityYear="2011" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="seychellensis">
<emphasis id="44967327820AFFAFFA4CF817FE9BF824" italics="true" pageId="1" pageNumber="62">U. seychellensis</emphasis>
<bibRefCitation id="1273D2C4820AFFAFFEFBF87AFDDEF824" author="Uiblein" box="[310,620,2008,2032]" pageId="1" pageNumber="62" refString="Uiblein, F. &amp; Heemstra, P. C. (2011) Description of a new goatfish species, Upeneus seychellensis sp. nov. (Mullidae), from the Seychelles Bank, with remarks on Upeneus guttatus geographic variation and a key for all Western Indian Ocean Upeneus species. Marine Biology Research, 7, 637 - 650." type="journal article" year="2011">Uiblein &amp; Heemstra 2011</bibRefCitation>
</taxonomicName>
), the
<taxonomicName id="B1E2D4B6820AFFAFFD79F87AFC8FF824" box="[692,829,2008,2032]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="moluccensis">
<emphasis id="44967327820AFFAFFD79F87AFC8FF824" box="[692,829,2008,2032]" italics="true" pageId="1" pageNumber="62">moluccensis</emphasis>
</taxonomicName>
group (
<taxonomicName id="B1E2D4B6820AFFAFFC6CF87BFB68F824" authority="Gunther, 1869" authorityName="Gunther" authorityYear="1869" box="[929,1242,2008,2032]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="doriae">
<emphasis id="44967327820AFFAFFC6CF87BFBA0F824" box="[929,1042,2008,2032]" italics="true" pageId="1" pageNumber="62">U. doriae</emphasis>
(Günther, 1869)
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820AFFACFB26F87BFEFAFF7B" authority="Bleeker, 1855" authorityName="Bleeker" authorityYear="1855" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" lastPageId="2" lastPageNumber="63" order="Perciformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="moluccensis">
<emphasis id="44967327820AFFAFFB26F87BFA2EF824" box="[1259,1436,2008,2032]" italics="true" pageId="1" pageNumber="62">U. moluccensis</emphasis>
(Bleeker, 1855)
</taxonomicName>
,
<taxonomicName id="B1E2D4B68209FFACFE98FF3BFCA8FF7B" authority="Cheng &amp; Wang, 1963" authorityName="Cheng &amp; Wang" authorityYear="1963" box="[341,794,151,176]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="quadrilineatus">
<emphasis id="449673278209FFACFE98FF3BFDA9FF64" box="[341,539,152,176]" italics="true" pageId="2" pageNumber="63">U. quadrilineatus</emphasis>
Cheng &amp; Wang, 1963
</taxonomicName>
,
<taxonomicName id="B1E2D4B68209FFACFCE4FF3BFBABFF7B" authority="Cuvier" authorityName="Cuvier" box="[809,1049,151,176]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="sulphureus">
<emphasis id="449673278209FFACFCE4FF3BFC74FF64" box="[809,966,152,176]" italics="true" pageId="2" pageNumber="63">U. sulphureus</emphasis>
Cuvier
</taxonomicName>
in Cuvier &amp; Valenciennes, 1829), and the
<taxonomicName id="B1E2D4B68209FFACFF22FF1EFEF0FF00" box="[239,322,188,212]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="tragula">
<emphasis id="449673278209FFACFF22FF1EFEF0FF00" box="[239,322,188,212]" italics="true" pageId="2" pageNumber="63">tragula</emphasis>
</taxonomicName>
group (
<taxonomicName id="B1E2D4B68209FFACFE54FF1FFDAAFF00" box="[409,536,188,212]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="luzonius">
<emphasis id="449673278209FFACFE54FF1FFDAAFF00" box="[409,536,188,212]" italics="true" pageId="2" pageNumber="63">U. luzonius</emphasis>
</taxonomicName>
<collectingCountry id="0EF5EFA58209FFACFDD2FF1EFDD8FF00" box="[543,618,188,212]" name="Jordan" pageId="2" pageNumber="63">Jordan</collectingCountry>
&amp; Seale, 1907,
<taxonomicName id="B1E2D4B68209FFACFCD0FF1FFB4DFF00" authority="Uiblein &amp; Heemstra, 2010" authorityName="Uiblein &amp; Heemstra" authorityYear="2010" box="[797,1279,188,212]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="margarethae">
<emphasis id="449673278209FFACFCD0FF1FFC7FFF00" box="[797,973,188,212]" italics="true" pageId="2" pageNumber="63">U. margarethae</emphasis>
<bibRefCitation id="1273D2C48209FFACFC19FF1EFB4DFF00" author="Uiblein" box="[980,1279,188,212]" pageId="2" pageNumber="63" refString="Uiblein F, &amp; Heemstra, P. C (2010) A taxonomic review of the Western Indian Ocean goatfishes of the genus Upeneus (Family Mullidae) with descriptions of four new species. Smithiana Bulletin, 11, 35 - 71." type="journal article" year="2010">Uiblein &amp; Heemstra, 2010</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B1E2D4B68209FFACFAC7FF1FFE60FF23" authority="Randall &amp; Kulbicki, 2006" authorityName="Randall &amp; Kulbicki" authorityYear="2006" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="mouthami">
<emphasis id="449673278209FFACFAC7FF1FFA2EFF00" box="[1290,1436,188,212]" italics="true" pageId="2" pageNumber="63">U. mouthami</emphasis>
Randall &amp; Kulbicki, 2006
</taxonomicName>
,
<taxonomicName id="B1E2D4B68209FFACFE28FF43FCF0FF23" authority="Lachner, 1954" authorityName="Lachner" authorityYear="1954" box="[485,834,223,248]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="oligospilus">
<emphasis id="449673278209FFACFE28FF43FD3BFF2C" box="[485,649,224,248]" italics="true" pageId="2" pageNumber="63">U. oligospilus</emphasis>
Lachner, 1954
</taxonomicName>
,
<taxonomicName id="B1E2D4B68209FFACFC98FF43FADEFF23" authority="Uiblein &amp; Heemstra, 2011" authorityName="Uiblein &amp; Heemstra" authorityYear="2011" box="[853,1388,223,248]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="randalli">
<emphasis id="449673278209FFACFC98FF43FBADFF2C" box="[853,1055,224,248]" italics="true" pageId="2" pageNumber="63">Upeneus randalli</emphasis>
<bibRefCitation id="1273D2C48209FFACFBE0FF7DFADEFF23" author="Uiblein" box="[1069,1388,223,247]" pageId="2" pageNumber="63" refString="Uiblein, F. &amp; Heemstra, P. C. (2011) Description of a new goatfish species, Upeneus seychellensis sp. nov. (Mullidae), from the Seychelles Bank, with remarks on Upeneus guttatus geographic variation and a key for all Western Indian Ocean Upeneus species. Marine Biology Research, 7, 637 - 650." type="journal article" year="2011">Uiblein &amp; Heemstra, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B1E2D4B68209FFACFA4DFF43FE0CFEC8" authority="Bleeker, 1855" authorityName="Bleeker" authorityYear="1855" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="sundaicus">
<emphasis id="449673278209FFACFA4DFF43FEB5FEC8" italics="true" pageId="2" pageNumber="63">U. sundaicus</emphasis>
(Bleeker, 1855)
</taxonomicName>
,
<taxonomicName id="B1E2D4B68209FFACFE07FEA7FD78FEC8" authority="Cuvier" authorityName="Cuvier" box="[458,714,260,284]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="taeniopterus">
<emphasis id="449673278209FFACFE07FEA7FDCAFEC8" box="[458,632,261,284]" italics="true" pageId="2" pageNumber="63">U. taeniopterus</emphasis>
Cuvier
</taxonomicName>
in Cuvier &amp; Valenciennes, 1829,
<taxonomicName id="B1E2D4B68209FFACFB89FEA7FA3BFEC8" authority="Richardson, 1846" authorityName="Richardson" authorityYear="1846" box="[1092,1417,260,284]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="tragula">
<emphasis id="449673278209FFACFB89FEA7FB0BFEC8" box="[1092,1209,260,284]" italics="true" pageId="2" pageNumber="63">U. tragula</emphasis>
Richardson, 1846
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>