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<document id="C1B40EADED9DEA7E15E5ED503AAD9A5B" ID-DOI="10.5281/zenodo.208421" ID-GBIF-Dataset="9fbffc4e-2741-47ec-9cf7-ccda0f18517f" ID-ISSN="1175-5326" ID-Zenodo-Dep="208421" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460553901718" checkinUser="plazi" docAuthor="Uiblein, Franz &amp; Mcgrouther, Mark" docDate="2012" docId="FE4B1E238209FFABFF5AFECDFC72FD50" docLanguage="en" docName="zt03550p070.pdf" docOrigin="Zootaxa 3550" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Upeneus stenopsis Uiblein &amp; Mcgrouther, 2012, n. sp." docType="treatment" docVersion="10" lastPageNumber="66" masterDocId="0272665B820BFFAEFFCDFFA2FFB2FFD4" masterDocTitle="A new deep-water goatfish of the genus Upeneus (Mullidae) from northern Australia and the Philippines, with a taxonomic account of U. subvittatus and remarks on U. mascareinsis" masterLastPageNumber="70" masterPageNumber="61" pageNumber="63" updateTime="1698318766311" updateUser="plazi">
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<mods:titleInfo id="6EF4646C1B8295553DFB65A05EFC2CD5">
<mods:title id="BAD4DCDB0DB22F9C926B9DD555B330B5">A new deep-water goatfish of the genus Upeneus (Mullidae) from northern Australia and the Philippines, with a taxonomic account of U. subvittatus and remarks on U. mascareinsis</mods:title>
</mods:titleInfo>
<mods:name id="FB47D3D43ADDED916971C42151CA23DD" type="personal">
<mods:role id="C5851863EFE679750B5043889F3B7C02">
<mods:roleTerm id="A58504575DD93E81CC6027E66B8CF1B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BE09247E47F38436CE48DEEB1E6C455A">Uiblein, Franz</mods:namePart>
</mods:name>
<mods:name id="10F4592380113D6624B65E1DB4D25D1D" type="personal">
<mods:role id="5383B3528FA55EBFA0AF697EF581AB84">
<mods:roleTerm id="4B3F93CBC69A6B08F309BADEA4E40D46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B1C89898ECDCC576ABBB15FBD59712A5">Mcgrouther, Mark</mods:namePart>
</mods:name>
<mods:typeOfResource id="86C9B6B8FE6B44AAD904E9EB1223F731">text</mods:typeOfResource>
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<mods:titleInfo id="464FA059DE4CE56DAA68C080290C5DD9">
<mods:title id="DB7FF0B3FEE00BC6218F642F473FFE39">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F6DC181BD132F90DFD9D59217F0255F4">
<mods:date id="3469576D4E719E9FB73F137F961CA86F">2012</mods:date>
<mods:detail id="351746244E06210848E0026C95BAFE59" type="volume">
<mods:number id="1AE1BA18E9EE06ACE8F74FCF94B66C29">3550</mods:number>
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<mods:classification id="573DC31DE1603564268A52AA471F9FB7">journal article</mods:classification>
<mods:identifier id="6AD015E4F9A07B5E8EE9B1E598C46BD9" type="DOI">10.5281/zenodo.208421</mods:identifier>
<mods:identifier id="6F551A729EA1C9CBD456A968999B910C" type="GBIF-Dataset">9fbffc4e-2741-47ec-9cf7-ccda0f18517f</mods:identifier>
<mods:identifier id="AE02AA42C99B24C5D5C685FFBE922223" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A1DB977F7AE41888183B89916EF72516" type="Zenodo-Dep">208421</mods:identifier>
</mods:mods>
<treatment id="FE4B1E238209FFABFF5AFECDFC72FD50" ID-DOI="http://doi.org/10.5281/zenodo.6168945" ID-GBIF-Taxon="119555292" ID-Zenodo-Dep="6168945" LSID="urn:lsid:zoobank.org:act:0FB24930-AE9D-4FAE-8BD0-83EF715A65F3" httpUri="http://treatment.plazi.org/id/FE4B1E238209FFABFF5AFECDFC72FD50" lastPageId="5" lastPageNumber="66" pageId="2" pageNumber="63">
<subSubSection id="3EF8FCBE8209FFACFF5AFECDFEC1FE1A" pageId="2" pageNumber="63" type="nomenclature">
<paragraph id="765DAF358209FFACFF5AFECDFE0CFE5D" blockId="2.[151,446,367,462]" box="[151,446,367,393]" pageId="2" pageNumber="63">
<heading id="2D1518598209FFACFF5AFECDFE0CFE5D" bold="true" box="[151,446,367,393]" fontSize="11" level="1" pageId="2" pageNumber="63" reason="1">
<emphasis id="449673278209FFACFF5AFECDFE0CFE5D" bold="true" box="[151,446,367,393]" pageId="2" pageNumber="63">
<taxonomicName id="B1E2D4B68209FFACFF5AFECDFEC7FE5D" ID-CoL="7DLSS" box="[151,373,367,393]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="stenopsis" status="sp. nov.">
<emphasis id="449673278209FFACFF5AFECDFEC7FE5D" bold="true" box="[151,373,367,393]" italics="true" pageId="2" pageNumber="63">Upeneus stenopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FA5CE5C8209FFACFEB1FECDFE0CFE5D" box="[380,446,367,393]" pageId="2" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="765DAF358209FFACFF5AFE30FEC8FE7E" blockId="2.[151,446,367,462]" box="[151,378,402,426]" pageId="2" pageNumber="63">Narrow-tail goatfish</paragraph>
<paragraph id="765DAF358209FFACFF5AFE14FEC1FE1A" blockId="2.[151,446,367,462]" box="[151,371,438,462]" pageId="2" pageNumber="63">
(
<figureCitation id="EED9B3B08209FFACFF52FE14FEB9FE1A" box="[159,267,438,462]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1061,1084]" captionStartId-1="4.[151,250,1916,1939]" captionTargetBox-0="[151,1435,193,1038]" captionTargetBox-1="[162,1424,342,1883]" captionTargetId-0="figure@3.[151,1436,193,1040]" captionTargetId-1="figure@4.[151,1436,337,1895]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. (a) Upeneus stenopsis, AMS I. 20918 - 017, 13.1 cm SL, NE Australia (D. F. Hoese); (b) U. davidaromi, SL 16.5 cm, Gulf of Aqaba (J. E. Randall), (c) U. mascareinsis, SAIAB 81951, SL 16.2 cm, Mozambique (P. C. Heemstra), (d) U. subvittatus, BPBM 32266, SL 20.5 cm, Moluccas, Indonesia (J. E. Randall); (e) U. subvittatus, MZB PB 389, SL 16.6 cm, Java, Indonesia (W. White); (f) U. vittatus, CSIRO KD 395, SL 14.1 cm, Bali, Indonesia (W. White). The photographs of U. stenopsis and U. vittatus are shown mirror-inverted in order to facilitate comparison." captionText-1="FIGURE 2. Relationships among morphometric and meristic characters in four closely related species of the vittatu s group, Upeneus davidaromi, U. mascareinsis, U. stenopsis n. sp., and U. subvittatus. Specimens &lt;80 mm SL are distinguished as juveniles by crossed symbols." httpUri-0="https://zenodo.org/record/208422/files/figure.png" httpUri-1="https://zenodo.org/record/208423/files/figure.png" pageId="2" pageNumber="63">Figs. 12</figureCitation>
; Table 1)
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE8209FFACFF5AFE5DFA2EFD3A" pageId="2" pageNumber="63" type="materials_examined">
<paragraph id="765DAF358209FFACFF5AFE5DFE63FDEE" blockId="2.[151,1437,510,2010]" pageId="2" pageNumber="63">
<emphasis id="449673278209FFACFF5AFE5DFEB9FDC3" bold="true" box="[151,267,511,535]" pageId="2" pageNumber="63">
<typeStatus id="A95911978209FFACFF5AFE5DFEB4FDC3" box="[151,262,511,535]" pageId="2" pageNumber="63" type="holotype">Holotype</typeStatus>
.
</emphasis>
AMS I.20918-017,
<quantity id="B11A02D08209FFACFE23FE5CFDFFFDC2" box="[494,589,510,534]" metricMagnitude="-1" metricUnit="m" metricValue="1.31" pageId="2" pageNumber="63" unit="mm" value="131.0">131 mm</quantity>
, NE
<collectingCountry id="0EF5EFA58209FFACFD48FE5CFD5DFDC2" box="[645,751,510,534]" name="Australia" pageId="2" pageNumber="63">Australia</collectingCountry>
, Queensland, Coral Sea, off Raine Island,
<geoCoordinate id="13D6C9F28209FFACFB06FE5CFA80FDC2" box="[1227,1330,510,534]" direction="south" orientation="latitude" pageId="2" pageNumber="63" precision="925" value="-11.583333">11° 35' S</geoCoordinate>
,
<geoCoordinate id="13D6C9F28209FFACFAF3FE5CFF19FDEE" direction="east" orientation="longitude" pageId="2" pageNumber="63" precision="925" value="144.03334">144° 02' E</geoCoordinate>
,
<quantity id="B11A02D08209FFACFF7BFD80FE85FDEE" box="[182,311,546,570]" metricMagnitude="2" metricUnit="m" metricValue="2.725" metricValueMax="2.75" metricValueMin="2.7" pageId="2" pageNumber="63" unit="m" value="272.5" valueMax="275.0" valueMin="270.0">270275 m</quantity>
, prawn trawl.
</paragraph>
<paragraph id="765DAF358209FFACFF0AFDE5FEEDFD1E" blockId="2.[151,1437,510,2010]" pageId="2" pageNumber="63">
<emphasis id="449673278209FFACFF0AFDE5FEF4FD8B" bold="true" box="[199,326,583,607]" pageId="2" pageNumber="63">
<typeStatus id="A95911978209FFACFF0AFDE5FEF3FD8B" box="[199,321,583,607]" pageId="2" pageNumber="63" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="60E464BC8209FFACFE83FDE4FE68FD8A" box="[334,474,582,606]" pageId="2" pageNumber="63" type="generic">3 specimens</specimenCount>
(
<quantity id="B11A02D08209FFACFE27FDE4FD30FD8A" box="[490,642,582,606]" metricMagnitude="-1" metricUnit="m" metricValue="1.065" metricValueMax="1.1199999999999999" metricValueMin="1.01" pageId="2" pageNumber="63" unit="mm" value="106.5" valueMax="112.0" valueMin="101.0">101112 mm</quantity>
) from northern
<collectingCountry id="0EF5EFA58209FFACFCF7FDE4FC13FD8A" box="[826,929,582,606]" name="Australia" pageId="2" pageNumber="63">Australia</collectingCountry>
and the
<collectingCountry id="0EF5EFA58209FFACFBCBFDE4FB3AFD8A" box="[1030,1160,582,606]" name="Philippines" pageId="2" pageNumber="63">Philippines</collectingCountry>
. AMS I.21628-001,
<quantity id="B11A02D08209FFACFABFFDE4FF77FD56" metricMagnitude="-1" metricUnit="m" metricValue="1.08" pageId="2" pageNumber="63" unit="mm" value="108.0">108 mm</quantity>
, Northern Territory, Timor Sea,
<geoCoordinate id="13D6C9F28209FFACFDFDFDC8FD2AFD56" box="[560,664,618,642]" direction="south" orientation="latitude" pageId="2" pageNumber="63" precision="925" value="-9.766666">09° 46' S</geoCoordinate>
,
<geoCoordinate id="13D6C9F28209FFACFD69FDC8FCAEFD56" box="[676,796,618,642]" direction="east" orientation="longitude" pageId="2" pageNumber="63" precision="925" value="128.95">128° 57' E</geoCoordinate>
,
<quantity id="B11A02D08209FFACFCE5FDC8FC1BFD56" box="[808,937,618,642]" metricMagnitude="2" metricUnit="m" metricValue="1.775" metricValueMax="1.9" metricValueMin="1.65" pageId="2" pageNumber="63" unit="m" value="177.5" valueMax="190.0" valueMin="165.0">165190 m</quantity>
, bottom trawl;
<collectionCode id="10F337F08209FFACFB98FDC9FB18FD56" LSID="urn:lsid:biocol.org:col:34988" box="[1109,1194,619,642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="63">MNHN</collectionCode>
2012-0212,
<quantity id="B11A02D08209FFACFAF4FDC8FA25FD56" box="[1337,1431,618,642]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="2" pageNumber="63" unit="mm" value="112.0">112 mm</quantity>
,
<collectingCountry id="0EF5EFA58209FFACFF5AFD2CFEAAFD72" box="[151,280,654,678]" name="Philippines" pageId="2" pageNumber="63">Philippines</collectingCountry>
, off Quezon Island,
<geoCoordinate id="13D6C9F28209FFACFE30FD2CFD3BFD72" box="[509,649,654,678]" direction="north" orientation="latitude" pageId="2" pageNumber="63" precision="15" value="13.983055">13°58'59'' N</geoCoordinate>
,
<geoCoordinate id="13D6C9F28209FFACFD58FD2CFC9FFD72" box="[661,813,654,678]" direction="east" orientation="longitude" pageId="2" pageNumber="63" precision="15" value="120.3">120°18'00'' E</geoCoordinate>
,
<quantity id="B11A02D08209FFACFCF5FD2CFC08FD72" box="[824,954,654,678]" metricMagnitude="2" metricUnit="m" metricValue="1.8649999999999998" metricValueMax="1.8699999999999999" metricValueMin="1.86" pageId="2" pageNumber="63" unit="m" value="186.5" valueMax="187.0" valueMin="186.0">186187 m</quantity>
, beam trawl;
<collectionCode id="10F337F08209FFACFB9EFD2DFB1BFD72" LSID="urn:lsid:biocol.org:col:34988" box="[1107,1193,655,678]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="63">MNHN</collectionCode>
1984-0802,
<quantity id="B11A02D08209FFACFAF5FD2CFA2AFD72" box="[1336,1432,654,678]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="2" pageNumber="63" unit="mm" value="101.0">101 mm</quantity>
, same station data.
</paragraph>
<paragraph id="765DAF358209FFACFF0AFD75FA2EFD3A" blockId="2.[151,1437,510,2010]" box="[199,1436,726,751]" pageId="2" pageNumber="63">
<emphasis id="449673278209FFACFF0AFD75FE8EFD3B" bold="true" box="[199,316,727,751]" pageId="2" pageNumber="63">Non-type.</emphasis>
<collectionCode id="10F337F08209FFACFE8EFD74FECBFD3A" LSID="urn:lsid:biocol.org:col:34700" box="[323,377,726,750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="2" pageNumber="63">CAS</collectionCode>
33367, 71 mm (juvenile),
<collectingCountry id="0EF5EFA58209FFACFD6CFD74FC90FD3A" box="[673,802,726,750]" name="Philippines" pageId="2" pageNumber="63">Philippines</collectingCountry>
, Quezon Island, Catanauan, Sandoval Point,
<quantity id="B11A02D08209FFACFADAFD74FA2AFD3A" box="[1303,1432,726,750]" metricMagnitude="2" metricUnit="m" metricValue="1.345" metricValueMax="1.42" metricValueMin="1.27" pageId="2" pageNumber="63" unit="m" value="134.5" valueMax="142.0" valueMin="127.0">
127
<specimenCount id="60E464BC8209FFACFA82FD74FA2AFD3A" box="[1359,1432,726,750]" pageId="2" pageNumber="63" type="generic">142 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE8209FFACFF0AFD58FEFCFB4A" pageId="2" pageNumber="63" type="diagnosis">
<paragraph id="765DAF358209FFACFF0AFD58FEFCFB4A" blockId="2.[151,1437,510,2010]" pageId="2" pageNumber="63">
<emphasis id="449673278209FFACFF0AFD58FE88FCC7" bold="true" box="[199,314,762,787]" pageId="2" pageNumber="63">Diagnosis</emphasis>
. Dorsal fins VIII + 9; pectoral fins 15 or 16; gill rakers 78 + 1820 = 2528; measurements in % SL: body depth at first dorsal-fin origin 2225; body depth at anus 1821; caudal-peduncle depth 7.47.8; maximum head depth 2223; head depth through eye 1819; head length 3334; orbit length 9.29.9; upper jaw length 1415; barbel length 2122; caudal-fin length 2628; anal-fin height 1516; pelvic-fin length 1819; pectoral-fin length 2325; first dorsal-fin height 2023; second dorsal-fin height 1416; total number of oblique caudal-fin bars 79, colour of bars changing from pale brown proximally to dark brown towards rear caudal-fin margin; 45 dark bars on upper caudal-fin lobe (including one bar close to rear end of lateral line), slightly increasing in width distally; 34 bars on lower caudal-fin lobe, of similar width as white interspaces between bars; first dorsal-fin tip black; caudal-fin bars and black dorsal-fin tip retained on preserved fish; barbels white; no lateral body stripes; head and body silvery laterally and reddish dorsally, belly and ventral side of peduncle white; preserved fish dorsally or entirely pale brown or brown, with ventral part of body and head covered by white or silvery pigment.
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE8209FFACFF0AFB08FCD2F80E" pageId="2" pageNumber="63" type="description">
<paragraph id="765DAF358209FFACFF0AFB08FC3AFAA2" blockId="2.[151,1437,510,2010]" pageId="2" pageNumber="63">
<emphasis id="449673278209FFACFF0AFB08FEE3FB17" bold="true" box="[199,337,1194,1219]" pageId="2" pageNumber="63">Description</emphasis>
. Measurements in % SL and counts are given in Table 1; morphometric data as ratios of SL for
<typeStatus id="A95911978209FFACFF5AFB6CFF4EFB32" box="[151,252,1230,1254]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
, data for
<typeStatus id="A95911978209FFACFEA3FB6DFE6BFB33" box="[366,473,1231,1255]" pageId="2" pageNumber="63" type="paratype">paratypes</typeStatus>
in brackets: body moderately deep, its depth at first dorsal-fin origin 4.1[3.94.5]; body depth at anal-fin origin 5.4[4.85.4]; head depth through eye 5.2[5.25.6]; head length 2.9[3.0], clearly longer than maximum depth of body and caudal-fin length (3.6[3.63.8]); height of second dorsal fin 7.0[6.16.3], clearly less than barbel length (4.6[4.75.0]); pectoral-fin length 4.0[4.14.4], longer than pelvic-fin length (5.3[5.25.7]); orbit length 10[1011], larger than caudal-peduncle depth (13[13]).
</paragraph>
<paragraph id="765DAF358209FFACFF0AFA21FE0EF90A" blockId="2.[151,1437,510,2010]" pageId="2" pageNumber="63">
<emphasis id="449673278209FFACFF0AFA21FEE5FA4F" box="[199,343,1411,1435]" italics="true" pageId="2" pageNumber="63">Fresh colour</emphasis>
(based on
<typeStatus id="A95911978209FFACFE1CFA20FD85FA4E" box="[465,567,1410,1434]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
;
<figureCitation id="EED9B3B08209FFACFD8FFA20FD2CFA4E" box="[578,670,1410,1434]" captionStart="FIGURE 1" captionStartId="3.[151,250,1061,1084]" captionTargetBox="[151,1435,193,1038]" captionTargetId="figure@3.[151,1436,193,1040]" captionTargetPageId="3" captionText="FIGURE 1. (a) Upeneus stenopsis, AMS I. 20918 - 017, 13.1 cm SL, NE Australia (D. F. Hoese); (b) U. davidaromi, SL 16.5 cm, Gulf of Aqaba (J. E. Randall), (c) U. mascareinsis, SAIAB 81951, SL 16.2 cm, Mozambique (P. C. Heemstra), (d) U. subvittatus, BPBM 32266, SL 20.5 cm, Moluccas, Indonesia (J. E. Randall); (e) U. subvittatus, MZB PB 389, SL 16.6 cm, Java, Indonesia (W. White); (f) U. vittatus, CSIRO KD 395, SL 14.1 cm, Bali, Indonesia (W. White). The photographs of U. stenopsis and U. vittatus are shown mirror-inverted in order to facilitate comparison." httpUri="https://zenodo.org/record/208422/files/figure.png" pageId="2" pageNumber="63">Figure 1</figureCitation>
a): Head and anterior part of body silvery laterally, the posterior and dorsal third of body as well as the dorsal head margin starting from upper jaw reddish; belly to ventral caudal peduncle white; no lateral body stripes; barbels white; both dorsal fins hyaline at base with white rays overlain by two horizontal pale red stripes and black tips (only the spiny dorsal-fin tip well visible in photograph of
<typeStatus id="A95911978209FFACFAE4FA4CFA22F9D2" box="[1321,1424,1518,1542]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
); paired fins and anal fin hyaline, caudal fin with 9 oblique bars (the two distal-most bars covering the fin tips); 5 bars (including one bar close to rear end of lateral line) on upper lobe, 4 bars on lower lobe, with hyaline-whitish interspaces between bars; the five anterior-most caudal-fin bars changing from pale brown proximally to dark brown at rear caudal-fin margin; the distal-most bars (including fin tip) on lower caudal-fin lobe entirely dark brown; the bars on the upper caudal-fin lobe slightly increasing in width distally, the interspaces of lesser width as interspaces on lower lobe.
</paragraph>
<paragraph id="765DAF358209FFACFF0AF949FCD2F80E" blockId="2.[151,1437,510,2010]" pageId="2" pageNumber="63">
<emphasis id="449673278209FFACFF0AF949FE3AF8D7" box="[199,392,1771,1795]" italics="true" pageId="2" pageNumber="63">Preserved colour</emphasis>
: Head and body uniformly brown in
<typeStatus id="A95911978209FFACFCE3F948FC22F8D6" box="[814,912,1770,1794]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
with white pigmentation patches behind jaws, on opercle, ventrally from belly to caudal peduncle, and some tiny white markings along lateral line at mid body; first dorsal-fin tip and caudal-fin bar pigmentation partly retained at or close to distal fin margins;
<typeStatus id="A95911978209FFACFB3AF891FAD0F89F" box="[1271,1378,1843,1867]" pageId="2" pageNumber="63" type="paratype">paratypes</typeStatus>
with large whitish or silvery patches on head below eye and/or surrounding the lower half of orbit, and on posterior opercle (reduced on preopercle); Body entirely whitish or silvery at ventral side, brown dorsally, with dark pigmentation on both dorsal-fin tips, and bars on lower caudal-fin lobe well retained, much weaker on upper lobe (upper caudal-fin lobe almost entirely broken in one specimen).
</paragraph>
</subSubSection>
<caption id="229DFFBD8208FFADFF5AFB87FC3FFB62" httpUri="https://zenodo.org/record/208422/files/figure.png" pageId="3" pageNumber="64" targetBox="[151,1435,193,1038]" targetPageId="3">
<paragraph id="765DAF358208FFADFF5AFB87FC3FFB62" blockId="3.[151,1436,1061,1207]" pageId="3" pageNumber="64">
<emphasis id="449673278208FFADFF5AFB87FEA0FBEF" bold="true" box="[151,274,1061,1084]" pageId="3" pageNumber="64">FIGURE 1.</emphasis>
(a)
<emphasis id="449673278208FFADFEF7FB85FE4EFBE8" box="[314,508,1063,1084]" italics="true" pageId="3" pageNumber="64">
<taxonomicName id="B1E2D4B68208FFADFEF7FB85FE45FBE8" box="[314,503,1063,1084]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="stenopsis">Upeneus stenopsis</taxonomicName>
,
</emphasis>
AMS I.20918-017, 13.1 cm SL, NE Australia (D. F. Hoese); (b)
<emphasis id="449673278208FFADFB45FB85FA93FBE8" box="[1160,1313,1062,1084]" italics="true" pageId="3" pageNumber="64">
<taxonomicName id="B1E2D4B68208FFADFB45FB85FAAFFBE8" box="[1160,1309,1062,1084]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="davidaromi">U. davidaromi</taxonomicName>
,
</emphasis>
SL16.5 cm, Gulf of Aqaba (J. E. Randall), (c)
<emphasis id="449673278208FFADFDD9FBE4FD70FB8F" box="[532,706,1094,1115]" italics="true" pageId="3" pageNumber="64">
<taxonomicName id="B1E2D4B68208FFADFDD9FBE4FD0CFB8F" box="[532,702,1094,1115]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="mascareinsis">U. mascareinsis</taxonomicName>
,
</emphasis>
SAIAB 81951, SL 16.2 cm, Mozambique (P. C. Heemstra), (d)
<emphasis id="449673278208FFADFA4EFBE4FEB9FBAD" italics="true" pageId="3" pageNumber="64">
<taxonomicName id="B1E2D4B68208FFADFA4EFBE4FEB5FBAD" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="subvittatus">U. subvittatus</taxonomicName>
,
</emphasis>
BPBM 32266, SL 20.5 cm, Moluccas, Indonesia (J. E. Randall); (e)
<emphasis id="449673278208FFADFC0CFBC6FBE6FBAD" box="[961,1108,1123,1145]" italics="true" pageId="3" pageNumber="64">
<taxonomicName id="B1E2D4B68208FFADFC0CFBC6FBE2FBAD" box="[961,1104,1123,1145]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="subvittatus">U. subvittatus</taxonomicName>
,
</emphasis>
MZB PB389, SL 16.6 cm, Java, Indonesia (W. White); (f)
<emphasis id="449673278208FFADFE50FB21FDBFFB4C" box="[413,525,1155,1176]" italics="true" pageId="3" pageNumber="64">
<taxonomicName id="B1E2D4B68208FFADFE50FB21FDBBFB4C" box="[413,521,1155,1176]" class="Actinopterygii" family="Mullidae" genus="Mullus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="vittatus">U. vittatus</taxonomicName>
,
</emphasis>
CSIRO KD395, SL14.1 cm, Bali, Indonesia (W. White). The photographs of
<taxonomicName id="B1E2D4B68208FFADFAEDFB21FA2EFB4C" box="[1312,1436,1155,1176]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="stenopsis">
<emphasis id="449673278208FFADFAEDFB21FA2EFB4C" box="[1312,1436,1155,1176]" italics="true" pageId="3" pageNumber="64">U. stenopsis</emphasis>
</taxonomicName>
and
<taxonomicName id="B1E2D4B68208FFADFF0FFB00FE9EFB63" box="[194,300,1186,1207]" class="Actinopterygii" family="Mullidae" genus="Mullus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="449673278208FFADFF0FFB00FE9EFB63" box="[194,300,1186,1207]" italics="true" pageId="3" pageNumber="64">U. vittatus</emphasis>
</taxonomicName>
are shown mirror-inverted in order to facilitate comparison.
</paragraph>
</caption>
<subSubSection id="3EF8FCBE8208FFADFF0AFB7FFE14FACE" pageId="3" pageNumber="64" type="distribution">
<paragraph id="765DAF358208FFADFF0AFB7FFE14FACE" blockId="3.[151,1436,1245,2027]" pageId="3" pageNumber="64">
<emphasis id="449673278208FFADFF0AFB7FFEEBFB22" bold="true" box="[199,345,1245,1270]" pageId="3" pageNumber="64">Distribution</emphasis>
. N to NE
<collectingCountry id="0EF5EFA58208FFADFE13FB7CFDFAFB22" box="[478,584,1246,1270]" name="Australia" pageId="3" pageNumber="64">Australia</collectingCountry>
; Quezon,
<collectingCountry id="0EF5EFA58208FFADFD0FFB7CFCF6FB22" box="[706,836,1246,1270]" name="Philippines" pageId="3" pageNumber="64">Philippines</collectingCountry>
; first record of a deep-water goatfish of the genus
<taxonomicName id="B1E2D4B68208FFADFF5AFAA1FF4BFACE" box="[151,249,1283,1306]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="449673278208FFADFF5AFAA1FF4BFACE" box="[151,249,1283,1306]" italics="true" pageId="3" pageNumber="64">Upeneus</emphasis>
</taxonomicName>
for the Pacific.
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE8208FFADFF0AFA84FC8AF8AA" pageId="3" pageNumber="64" type="etymology">
<paragraph id="765DAF358208FFADFF0AFA84FBFBFAB6" blockId="3.[151,1436,1245,2027]" pageId="3" pageNumber="64">
<emphasis id="449673278208FFADFF0AFA84FEF4FAEA" bold="true" box="[199,326,1318,1342]" pageId="3" pageNumber="64">Etymology</emphasis>
. The name “
<taxonomicName id="B1E2D4B68208FFADFE13FA85FDF7FAEA" box="[478,581,1319,1342]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="stenopsis">
<emphasis id="449673278208FFADFE13FA85FDF7FAEA" box="[478,581,1319,1342]" italics="true" pageId="3" pageNumber="64">stenopsis</emphasis>
</taxonomicName>
” derives from the greek word “stenos”, meaning “narrow”, to refer to the rather narrow (both in depth and width) caudal peduncle of
<taxonomicName id="B1E2D4B68208FFADFCF9FAE9FBB1FAB6" box="[820,1027,1355,1378]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="stenopsis" status="sp. nov.">
<emphasis id="449673278208FFADFCF9FAE9FBB1FAB6" box="[820,1027,1355,1378]" italics="true" pageId="3" pageNumber="64">Upeneus stenopsis</emphasis>
</taxonomicName>
<emphasis id="449673278208FFADFBC7FAE8FBFBFAB6" bold="true" box="[1034,1097,1354,1378]" pageId="3" pageNumber="64">
<taxonomicNameLabel id="5FA5CE5C8208FFADFBC7FAE8FBFBFAB6" box="[1034,1097,1354,1378]" pageId="3" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="765DAF358208FFADFF0AFACFFC8AF8AA" blockId="3.[151,1436,1245,2027]" pageId="3" pageNumber="64">
<emphasis id="449673278208FFADFF0AFACFFED4FA52" bold="true" box="[199,358,1389,1414]" pageId="3" pageNumber="64">Comparisons</emphasis>
(see also Table 1):
<taxonomicName id="B1E2D4B68208FFADFD9CFACDFC96FA52" box="[593,804,1391,1414]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="stenopsis" status="sp. nov.">
<emphasis id="449673278208FFADFD9CFACDFC96FA52" box="[593,804,1391,1414]" italics="true" pageId="3" pageNumber="64">Upeneus stenopsis</emphasis>
</taxonomicName>
<emphasis id="449673278208FFADFCE2FACCFCC3FA52" bold="true" box="[815,881,1390,1414]" pageId="3" pageNumber="64">
<taxonomicNameLabel id="5FA5CE5C8208FFADFCE2FACCFCC3FA52" box="[815,881,1390,1414]" pageId="3" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from all other congeneric species by its thinner caudal peduncle (peduncle depth and width in combination) and from the most closely related and/or cooccurring species of the
<taxonomicName id="B1E2D4B68208FFADFE67FA15FE4FFA1A" box="[426,509,1463,1486]" class="Actinopterygii" family="Mullidae" genus="Mullus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="449673278208FFADFE67FA15FE4FFA1A" box="[426,509,1463,1486]" italics="true" pageId="3" pageNumber="64">vittatus</emphasis>
</taxonomicName>
group as follows: from
<taxonomicName id="B1E2D4B68208FFADFCC1FA15FC1CFA1A" box="[780,942,1462,1486]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="davidaromi">
<emphasis id="449673278208FFADFCC1FA15FC1CFA1A" box="[780,942,1462,1486]" italics="true" pageId="3" pageNumber="64">U. davidaromi</emphasis>
</taxonomicName>
by lesser body depth (body depth at dorsalfin origin 2225 vs 2428 %SL; body depth at anal-fin origin 1821 vs 2023 %SL), shallower head (maximum head depth 2223 vs 2427 %SL; head depth through eye 1819 vs 2024 %SL), and shorter caudal fin (caudal-fin length 2628 vs 2830 %SL); from
<taxonomicName id="B1E2D4B68208FFADFDE2F981FD51F9EE" box="[559,739,1571,1594]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="mascareinsis">
<emphasis id="449673278208FFADFDE2F981FD51F9EE" box="[559,739,1571,1594]" italics="true" pageId="3" pageNumber="64">U. mascareinsis</emphasis>
</taxonomicName>
in shorter interdorsal distance (1215 vs 1518 %SL), longer pectoral fins (pectoral-fin length 2325 vs 2124 %SL), and lower number of gill rakers (2528 vs 2730 total number of gill rakers); from
<taxonomicName id="B1E2D4B68208FFADFE2CF9C9FDCCF956" box="[481,638,1642,1666]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="subvittatus">
<emphasis id="449673278208FFADFE2CF9C9FDCCF956" box="[481,638,1642,1666]" italics="true" pageId="3" pageNumber="64">U. subvittatus</emphasis>
</taxonomicName>
in lesser body depth (body depth at dorsal-fin origin 2225 vs 2528 %SL; body depth at anal-fin origin 1821 vs 2223 %SL), larger eyes (orbit length 9.29.9 vs 7.88.8 %SL), and shorter interdorsal distance (1215 vs 1418 %SL); and from
<taxonomicName id="B1E2D4B68208FFADFC85F911FC09F91E" box="[840,955,1715,1738]" class="Actinopterygii" family="Mullidae" genus="Mullus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="449673278208FFADFC85F911FC09F91E" box="[840,955,1715,1738]" italics="true" pageId="3" pageNumber="64">U. vittatus</emphasis>
</taxonomicName>
in lesser body depth (body depth at dorsalfin origin 2225 vs 2430 %SL; body depth at anal-fin origin 1821 vs 2124 %SL), longer head (head length 3334 vs 2932 %SL, larger eyes (orbit length 9.29.9 vs 6.68.8 %SL), longer jaws (upper-jaw length 1415 vs 1113 %SL, lower-jaw length (1314 vs 1112 %SL), longer barbels (barbel length 2022 vs 1621 %SL), lesser first dorsal-fin height (2023 vs 2228 %SL), lower caudal-fin bars and white interspaces not varying in width, first dorsal black tip smaller, and lateral body stripes absent.
</paragraph>
</subSubSection>
<subSubSection id="3EF8FCBE8208FFABFF0AF828FC72FD50" lastPageId="5" lastPageNumber="66" pageId="3" pageNumber="64" type="discussion">
<paragraph id="765DAF358208FFAAFF0AF828FE7BFEC8" blockId="3.[151,1436,1245,2027]" lastBlockId="4.[151,1436,151,284]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">
<emphasis id="449673278208FFADFF0AF828FE89F876" bold="true" box="[199,315,1930,1954]" pageId="3" pageNumber="64">Remarks.</emphasis>
<taxonomicName id="B1E2D4B68208FFADFE8BF829FDA8F876" box="[326,538,1931,1954]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="stenopsis" status="sp. nov.">
<emphasis id="449673278208FFADFE8BF829FDA8F876" box="[326,538,1931,1954]" italics="true" pageId="3" pageNumber="64">Upeneus stenopsis</emphasis>
</taxonomicName>
<emphasis id="449673278208FFADFDEBF828FDD8F876" bold="true" box="[550,618,1930,1954]" pageId="3" pageNumber="64">
<taxonomicNameLabel id="5FA5CE5C8208FFADFDEBF828FDD8F876" box="[550,618,1930,1954]" pageId="3" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
shares with the three most closely related species
<taxonomicName id="B1E2D4B68208FFADFB0AF829FADCF876" box="[1223,1390,1930,1954]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="davidaromi">
<emphasis id="449673278208FFADFB0AF829FADCF876" box="[1223,1390,1930,1954]" italics="true" pageId="3" pageNumber="64">U. davidaromi</emphasis>
</taxonomicName>
,
<taxonomicName id="B1E2D4B68208FFADFA4CF829FE9AF812" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="mascareinsis">
<emphasis id="449673278208FFADFA4CF829FE9AF812" italics="true" pageId="3" pageNumber="64">U. mascareinsis</emphasis>
</taxonomicName>
, and
<taxonomicName id="B1E2D4B68208FFADFEA5F80DFDB6F812" box="[360,516,1966,1990]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="subvittatus">
<emphasis id="449673278208FFADFEA5F80DFDB6F812" box="[360,516,1966,1990]" italics="true" pageId="3" pageNumber="64">U. subvittatus</emphasis>
</taxonomicName>
the following features: no lateral body stripes in life, a shallow caudal peduncle, and a long head with long jaws and large eyes. Sufficient distinction among those species is reached when comparing the following characters in combination (
<figureCitation id="EED9B3B0820FFFAAFD22FF35FC85FF7B" box="[751,823,151,175]" captionStart="FIGURE 2" captionStartId="4.[151,250,1916,1939]" captionTargetBox="[162,1424,342,1883]" captionTargetId="figure@4.[151,1436,337,1895]" captionTargetPageId="4" captionText="FIGURE 2. Relationships among morphometric and meristic characters in four closely related species of the vittatu s group, Upeneus davidaromi, U. mascareinsis, U. stenopsis n. sp., and U. subvittatus. Specimens &lt;80 mm SL are distinguished as juveniles by crossed symbols." httpUri="https://zenodo.org/record/208423/files/figure.png" pageId="4" pageNumber="65">Fig. 2</figureCitation>
): Number of gill rakers on lower limb, body depth at anal-fin origin, caudal-peduncle depth and width, and maximum head depth. While
<taxonomicName id="B1E2D4B6820FFFAAFB92FF1FFB4EFF00" box="[1119,1276,188,212]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="subvittatus">
<emphasis id="44967327820FFFAAFB92FF1FFB4EFF00" box="[1119,1276,188,212]" italics="true" pageId="4" pageNumber="65">U. subvittatus</emphasis>
</taxonomicName>
occurs above
<quantity id="B11A02D0820FFFAAFF5AFF7DFF52FF23" box="[151,224,223,247]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="65" unit="m" value="120.0">120 m</quantity>
depth, the other three species can all be characterized as lower-shelf and upper-slope dwellers, occurring mostly below
<quantity id="B11A02D0820FFFAAFEF8FEA6FECEFEC8" box="[309,380,260,284]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="65" unit="m" value="100.0">100 m</quantity>
depth.
</paragraph>
<caption id="229DFFBD820FFFAAFF5AF8DEFE74F804" httpUri="https://zenodo.org/record/208423/files/figure.png" pageId="4" pageNumber="65" targetBox="[162,1424,342,1883]" targetPageId="4">
<paragraph id="765DAF35820FFFAAFF5AF8DEFE74F804" blockId="4.[151,1436,1916,2000]" pageId="4" pageNumber="65">
<emphasis id="44967327820FFFAAFF5AF8DEFEA6F846" bold="true" box="[151,276,1916,1939]" pageId="4" pageNumber="65">FIGURE 2.</emphasis>
Relationships among morphometric and meristic characters in four closely related species of the
<emphasis id="44967327820FFFAAFAC5F8DCFAFBF847" box="[1288,1353,1918,1939]" italics="true" pageId="4" pageNumber="65">vittatu</emphasis>
s group,
<taxonomicName id="B1E2D4B6820FFFAAFF5AF83FFEDEF866" box="[151,364,1948,1970]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="davidaromi">
<emphasis id="44967327820FFFAAFF5AF83FFEDEF866" box="[151,364,1948,1970]" italics="true" pageId="4" pageNumber="65">Upeneus davidaromi</emphasis>
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820FFFAAFEB6F83FFDADF866" box="[379,543,1949,1970]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="mascareinsis">
<emphasis id="44967327820FFFAAFEB6F83FFDADF866" box="[379,543,1949,1970]" italics="true" pageId="4" pageNumber="65">U. mascareinsis</emphasis>
</taxonomicName>
,
<taxonomicName id="B1E2D4B6820FFFAAFDE3F83FFD1EF866" box="[558,684,1949,1970]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="stenopsis" status="sp. nov.">
<emphasis id="44967327820FFFAAFDE3F83FFD1EF866" box="[558,684,1949,1970]" italics="true" pageId="4" pageNumber="65">U. stenopsis</emphasis>
</taxonomicName>
<emphasis id="44967327820FFFAAFD78F83EFD42F865" bold="true" box="[693,752,1948,1969]" pageId="4" pageNumber="65">
<taxonomicNameLabel id="5FA5CE5C820FFFAAFD78F83EFD42F865" box="[693,752,1948,1969]" pageId="4" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="B1E2D4B6820FFFAAFCE1F83FFC09F866" box="[812,955,1948,1970]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="subvittatus">
<emphasis id="44967327820FFFAAFCE1F83FFC09F866" box="[812,955,1948,1970]" italics="true" pageId="4" pageNumber="65">U. subvittatus</emphasis>
</taxonomicName>
. Specimens &lt;80 mm SL are distinguished as juveniles by crossed symbols.
</paragraph>
</caption>
<paragraph id="765DAF35820EFFABFF0AFF35FBEFFEEB" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="66">
The single juvenile specimen from Quezon largely overlaps in most morphometric and meristic characters with the four adult
<taxonomicName id="B1E2D4B6820EFFABFEF1FF1FFE7AFF00" box="[316,456,189,212]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="66" phylum="Chordata" rank="species" species="stenopsis" status="sp. nov.">
<emphasis id="44967327820EFFABFEF1FF1FFE7AFF00" box="[316,456,189,212]" italics="true" pageId="5" pageNumber="66">U. stenopsis</emphasis>
</taxonomicName>
<emphasis id="44967327820EFFABFE1CFF1EFDA0FF00" bold="true" box="[465,530,188,212]" pageId="5" pageNumber="66">
<taxonomicNameLabel id="5FA5CE5C820EFFABFE1CFF1EFDA0FF00" box="[465,530,188,212]" pageId="5" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Table 1,
<figureCitation id="EED9B3B0820EFFABFD46FF1EFD62FF00" box="[651,720,188,212]" captionStart="FIGURE 2" captionStartId="4.[151,250,1916,1939]" captionTargetBox="[162,1424,342,1883]" captionTargetId="figure@4.[151,1436,337,1895]" captionTargetPageId="4" captionText="FIGURE 2. Relationships among morphometric and meristic characters in four closely related species of the vittatu s group, Upeneus davidaromi, U. mascareinsis, U. stenopsis n. sp., and U. subvittatus. Specimens &lt;80 mm SL are distinguished as juveniles by crossed symbols." httpUri="https://zenodo.org/record/208423/files/figure.png" pageId="5" pageNumber="66">Fig. 2</figureCitation>
) and hence has been assigned to this species. Among the few minor deviations (Table 1,
<figureCitation id="EED9B3B0820EFFABFE03FF7DFDA1FF23" box="[462,531,223,247]" captionStart="FIGURE 2" captionStartId="4.[151,250,1916,1939]" captionTargetBox="[162,1424,342,1883]" captionTargetId="figure@4.[151,1436,337,1895]" captionTargetPageId="4" captionText="FIGURE 2. Relationships among morphometric and meristic characters in four closely related species of the vittatu s group, Upeneus davidaromi, U. mascareinsis, U. stenopsis n. sp., and U. subvittatus. Specimens &lt;80 mm SL are distinguished as juveniles by crossed symbols." httpUri="https://zenodo.org/record/208423/files/figure.png" pageId="5" pageNumber="66">Fig. 2</figureCitation>
) the shallower head through eye, the smaller orbit, and the shorter jaws in the juvenile may reflect developmental changes. Other differences like the wider caudal peduncle (Table 1,
<figureCitation id="EED9B3B0820EFFABFAD2FEA6FAD3FEC8" box="[1311,1377,260,284]" captionStart="FIGURE 2" captionStartId="4.[151,250,1916,1939]" captionTargetBox="[162,1424,342,1883]" captionTargetId="figure@4.[151,1436,337,1895]" captionTargetPageId="4" captionText="FIGURE 2. Relationships among morphometric and meristic characters in four closely related species of the vittatu s group, Upeneus davidaromi, U. mascareinsis, U. stenopsis n. sp., and U. subvittatus. Specimens &lt;80 mm SL are distinguished as juveniles by crossed symbols." httpUri="https://zenodo.org/record/208423/files/figure.png" pageId="5" pageNumber="66">Fig. 2</figureCitation>
) will need further studies to understand the overall intraspecific variability of this character.
</paragraph>
<paragraph id="765DAF35820EFFABFF0AFEEEFD30FD8B" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="66">
Among the four juveniles of
<taxonomicName id="B1E2D4B6820EFFABFDEDFEEFFD65FEB0" box="[544,727,333,356]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="66" phylum="Chordata" rank="species" species="mascareinsis">
<emphasis id="44967327820EFFABFDEDFEEFFD65FEB0" box="[544,727,333,356]" italics="true" pageId="5" pageNumber="66">U. mascareinsis</emphasis>
</taxonomicName>
one specimen (SAIAB 82328) was earlier referred to as
<taxonomicName id="B1E2D4B6820EFFABFAB2FEEEFE9AFE5C" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="66" phylum="Chordata" rank="species" species="mascareinsis">
cf.
<emphasis id="44967327820EFFABFF5AFED3FE9AFE5C" box="[151,296,369,392]" italics="true" pageId="5" pageNumber="66">mascareinsis</emphasis>
</taxonomicName>
in
<bibRefCitation id="1273D2C4820EFFABFE81FECDFD25FE53" author="Uiblein" box="[332,663,367,391]" pageId="5" pageNumber="66" refString="Uiblein F, &amp; Heemstra, P. C (2010) A taxonomic review of the Western Indian Ocean goatfishes of the genus Upeneus (Family Mullidae) with descriptions of four new species. Smithiana Bulletin, 11, 35 - 71." type="journal article" year="2010">Uiblein and Heemstra (2010)</bibRefCitation>
. Here, the full assignment to this species is based on the documented co-occurrence of
<specimenCount id="60E464BC820EFFABFE91FE36FE06FE78" box="[348,436,404,428]" pageId="5" pageNumber="66" type="adult">6 adults</specimenCount>
(&gt;
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SL) with
<specimenCount id="60E464BC820EFFABFD53FE36FCABFE78" box="[670,793,404,428]" pageId="5" pageNumber="66" type="juvenile">3 juveniles</specimenCount>
at two collecting stations off
<collectingCountry id="0EF5EFA5820EFFABFBA4FE37FB5FFE78" box="[1129,1261,404,428]" name="Réunion" pageId="5" pageNumber="66">La Reunion</collectingCountry>
(MNHN
<date id="025C89F5820EFFABFA96FE36FF04FE1B" pageId="5" pageNumber="66" value="1965" valueMax="1979">1965- 79</date>
, RMNH 24949), the close overlap in geographical and depth distribution, the general similarity among the four juveniles, and the consistency of deviations in morphometric but not meristic characters from adult conspecifics. The major deviations are a shallower body, a smaller head with shorter jaws, shorter barbels, and shorter anal and paired fins in the four juvenile
<taxonomicName id="B1E2D4B6820EFFABFE39FD87FD15FDE8" box="[500,679,549,572]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="66" phylum="Chordata" rank="species" species="mascareinsis">
<emphasis id="44967327820EFFABFE39FD87FD15FDE8" box="[500,679,549,572]" italics="true" pageId="5" pageNumber="66">U. mascareinsis</emphasis>
</taxonomicName>
compared to the adults (Table 1,
<figureCitation id="EED9B3B0820EFFABFBECFD86FBD4FDE8" box="[1057,1126,548,572]" captionStart="FIGURE 2" captionStartId="4.[151,250,1916,1939]" captionTargetBox="[162,1424,342,1883]" captionTargetId="figure@4.[151,1436,337,1895]" captionTargetPageId="4" captionText="FIGURE 2. Relationships among morphometric and meristic characters in four closely related species of the vittatu s group, Upeneus davidaromi, U. mascareinsis, U. stenopsis n. sp., and U. subvittatus. Specimens &lt;80 mm SL are distinguished as juveniles by crossed symbols." httpUri="https://zenodo.org/record/208423/files/figure.png" pageId="5" pageNumber="66">Fig. 2</figureCitation>
). The eyes are only smaller in the initial stages of the ontogeny (
<figureCitation id="EED9B3B0820EFFABFDFDFDE5FDC7FD8B" box="[560,629,583,607]" captionStart="FIGURE 2" captionStartId="4.[151,250,1916,1939]" captionTargetBox="[162,1424,342,1883]" captionTargetId="figure@4.[151,1436,337,1895]" captionTargetPageId="4" captionText="FIGURE 2. Relationships among morphometric and meristic characters in four closely related species of the vittatu s group, Upeneus davidaromi, U. mascareinsis, U. stenopsis n. sp., and U. subvittatus. Specimens &lt;80 mm SL are distinguished as juveniles by crossed symbols." httpUri="https://zenodo.org/record/208423/files/figure.png" pageId="5" pageNumber="66">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="765DAF35820EFFABFF0AFDCFFC72FD50" blockId="5.[151,1437,151,645]" box="[199,960,620,645]" pageId="5" pageNumber="66">
<taxonomicName id="B1E2D4B6820EFFABFF0AFDCFFE25FD50" box="[199,407,621,644]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="66" phylum="Chordata" rank="species" species="stenopsis" status="sp. nov.">
<emphasis id="44967327820EFFABFF0AFDCFFE25FD50" box="[199,407,621,644]" italics="true" pageId="5" pageNumber="66">Upeneus stenopsis</emphasis>
</taxonomicName>
<emphasis id="44967327820EFFABFE50FDCEFE6EFD50" bold="true" box="[413,476,620,644]" pageId="5" pageNumber="66">
<taxonomicNameLabel id="5FA5CE5C820EFFABFE50FDCEFE6EFD50" box="[413,476,620,644]" pageId="5" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
attains
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SL; depth range
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127
<specimenCount id="60E464BC820EFFABFCBCFDCEFC09FD50" box="[881,955,620,644]" pageId="5" pageNumber="66" type="generic">275 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>