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<mods:title id="8FD1738C72EA12B3B2DEC914FF5DC118">Taxonomy of the katydids (Orthoptera: Tettigoniidae) from East Asia and adjacent islands. Communication 13</mods:title>
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<subSubSection id="5096652AFF88FFF72FEA3C07FCD3B57B" box="[490,863,1374,1398]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="183336A1FF88FFF72FEA3C07FCD3B57B" blockId="6.[490,863,1374,1398]" box="[490,863,1374,1398]" pageId="6" pageNumber="7">
<heading id="437B81CDFF88FFF72FEA3C07FCD3B57B" box="[490,863,1374,1398]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="7" reason="2">
<emphasis id="2AF8EAB3FF88FFF72FEA3C07FCD3B57B" bold="true" box="[490,863,1374,1398]" pageId="6" pageNumber="7">
Genus
<taxonomicName id="DF8C4D22FF88FFF72C393C07FCD3B57B" ID-CoL="8NTJW" authority="Redtenbacher, 1891" authorityName="Redtenbacher" authorityYear="1891" box="[569,863,1374,1398]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF88FFF72C393C07FDFAB57B" bold="true" box="[569,630,1374,1398]" italics="true" pageId="6" pageNumber="7">Liara</emphasis>
Redtenbacher, 1891
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="5096652AFF88FFF62F093CC1FC33B271" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="183336A1FF88FFF72F093CC1FBEBB5A0" blockId="6.[227,1127,1431,1481]" box="[265,1127,1431,1453]" pageId="6" pageNumber="7">
<typeStatus id="C7378803FF88FFF72F093CC1FEB7B5A0" box="[265,315,1432,1453]" pageId="6" pageNumber="7">Type</typeStatus>
species:
<taxonomicName id="DF8C4D22FF88FFF72F9A3CC1FC8FB5A1" authority="Redtenbacher, 1891" authorityName="Redtenbacher" authorityYear="1891" box="[410,771,1431,1453]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="2AF8EAB3FF88FFF72F9A3CC1FDB8B5A1" box="[410,564,1431,1453]" italics="true" pageId="6" pageNumber="7">Liara rufescens</emphasis>
Redtenbacher, 1891
</taxonomicName>
, by subsequent designation (Kirby,
</paragraph>
<paragraph id="183336A1FF88FFF72EE33CEAFEAEB5C4" blockId="6.[227,1127,1431,1481]" box="[227,290,1459,1481]" pageId="6" pageNumber="7">1906).</paragraph>
<paragraph id="183336A1FF88FFF72F093CB3FBE4B63A" blockId="6.[227,1128,1514,1646]" pageId="6" pageNumber="7">
<collectionCode id="7E9DAE64FF88FFF72F093CB3FEC0B5F2" box="[265,332,1514,1535]" pageId="6" pageNumber="7">NOTE</collectionCode>
. Ingrisch (1998) included in this genus several species previously described in the genera
<taxonomicName id="DF8C4D22FF88FFF72F2B3F5FFEECB616" authorityName="Redtenbacher" authorityYear="1891" box="[299,352,1542,1563]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF88FFF72F2B3F5FFEECB616" box="[299,352,1542,1563]" italics="true" pageId="6" pageNumber="7">Liara</emphasis>
</taxonomicName>
,
<taxonomicName id="DF8C4D22FF88FFF72F6F3F5FFD15B616" authority="Karny, 1907" authorityName="Karny" authorityYear="1907" box="[367,665,1542,1563]" class="Insecta" family="Tettigoniidae" genus="Acanthocoryphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF88FFF72F6F3F5FFD9BB616" box="[367,535,1542,1563]" italics="true" pageId="6" pageNumber="7">Acanthocoryphus</emphasis>
Karny, 1907
</taxonomicName>
and
<taxonomicName id="DF8C4D22FF88FFF72CCC3F5FFB8CB616" authority="Redtenbacher, 1891" authorityName="Redtenbacher" authorityYear="1891" box="[716,1024,1542,1564]" class="Insecta" family="Tettigoniidae" genus="Oxystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF88FFF72CCC3F5FFCB9B616" box="[716,821,1542,1563]" italics="true" pageId="6" pageNumber="7">Oxystethus</emphasis>
Redtenbacher, 1891
</taxonomicName>
(the latter name was changed into
<emphasis id="2AF8EAB3FF88FFF72FD53F78FDB5B63B" box="[469,569,1569,1590]" italics="true" pageId="6" pageNumber="7">Unalianus</emphasis>
Koçak et Kemal,
<quantity id="DF749B44FF88FFF72CF23F78FCB1B63A" box="[754,829,1569,1591]" metricMagnitude="1" metricUnit="m" metricValue="5.10286" pageId="6" pageNumber="7" unit="in" value="2009.0">2009 in</quantity>
connection with homonymy).
</paragraph>
<paragraph id="183336A1FF88FFF62EE33F64FBEAB116" blockId="6.[227,1128,1514,1646]" lastBlockId="7.[227,1127,262,636]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Status of the second taxon was reduced by him up to subgeneric one. But for some species from the third taxon, the genus
<taxonomicName id="DF8C4D22FF88FFF72C113F01FDF6B660" authorityName="Redtenbacher" authorityYear="1891" box="[529,634,1624,1645]" class="Insecta" family="Tettigoniidae" genus="Oxystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF88FFF72C113F01FDF6B660" box="[529,634,1624,1645]" italics="true" pageId="6" pageNumber="7">Oxystethus</emphasis>
</taxonomicName>
was preserved, although these species have almost all their morphological characters very similar to those of
<taxonomicName id="DF8C4D22FF89FFF62D1C385EFCDDB111" authorityName="Redtenbacher" authorityYear="1891" box="[796,849,263,284]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="2AF8EAB3FF89FFF62D1C385EFCDDB111" box="[796,849,263,284]" italics="true" pageId="7" pageNumber="8">Liara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31CB57C8FF89FFF62D58385EFC01B111" box="[856,909,263,284]" pageId="7" pageNumber="8" sensu="stricto">s. str.</taxonomicNameLabel>
and
<taxonomicName id="DF8C4D22FF89FFF62DBE385FFBEAB116" authorityName="Karny" authorityYear="1907" box="[958,1126,262,283]" class="Insecta" family="Tettigoniidae" genus="Acanthocoryphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF89FFF62DBE385FFBEAB116" box="[958,1126,262,283]" italics="true" pageId="7" pageNumber="8">Acanthocoryphus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="183336A1FF89FFF62EE33878FBEBB13A" blockId="7.[227,1127,262,636]" box="[227,1127,289,311]" pageId="7" pageNumber="8">
(including very characteristic structure of the distal part of their male cerci; see
<figureCitation id="80B72A24FF89FFF62DEB3878FBD0B13A" box="[1003,1116,289,311]" captionStart="Figs 3150" captionStartId="9.[265,306,1481,1503]" captionTargetBox="[231,1118,456,1445]" captionTargetId="figure-82@9.[227,1123,451,1448]" captionTargetPageId="9" captionText="Figs 3150. Liara s. l.: 3136 L. (Liara) bifurcata sp. n.; 3743 L. (Liara) inaculeata" figureDoi="http://doi.org/10.5281/zenodo.7165410" httpUri="https://zenodo.org/record/7165410/files/figure.png" pageId="7" pageNumber="8">Figs 42, 44</figureCitation>
).
</paragraph>
<paragraph id="183336A1FF89FFF62EE33865FBEAB15F" blockId="7.[227,1127,262,636]" box="[227,1126,316,338]" pageId="7" pageNumber="8">
Probably this author understood that
<emphasis id="2AF8EAB3FF89FFF62C4B3865FD23B15C" box="[587,687,316,337]" italics="true" pageId="7" pageNumber="8">Unalianus</emphasis>
(=
<taxonomicName id="DF8C4D22FF89FFF62CD23865FCB7B15C" authorityName="Redtenbacher" authorityYear="1891" box="[722,827,316,337]" class="Insecta" family="Tettigoniidae" genus="Oxystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF89FFF62CD23865FCB7B15C" box="[722,827,316,337]" italics="true" pageId="7" pageNumber="8">Oxystethus</emphasis>
</taxonomicName>
) is also a subgenus of
<taxonomicName id="DF8C4D22FF89FFF62A1B3864FBDCB15F" authority="Gorochov, 2020" authorityName="Gorochov" authorityYear="2020" box="[1051,1104,317,338]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="s.">
<emphasis id="2AF8EAB3FF89FFF62A1B3864FBDCB15F" box="[1051,1104,317,338]" italics="true" pageId="7" pageNumber="8">Liara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31CB57C8FF89FFF62A573864FBEAB15F" box="[1111,1126,317,338]" pageId="7" pageNumber="8">s.</taxonomicNameLabel>
</paragraph>
<paragraph id="183336A1FF89FFF62EE33801FBEBB1B2" blockId="7.[227,1127,262,636]" pageId="7" pageNumber="8">
str., because he considered all species of these “genera” in the same key (but for the other genera, he prepared separate keys in the same monograph). Possibly, his opinion was based on small differences in shape of the male genitalia: in
<taxonomicName id="DF8C4D22FF89FFF62CEF38D7FCA8B1AE" authorityName="Gorochov" authorityYear="2020" box="[751,804,398,419]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="2AF8EAB3FF89FFF62CEF38D7FCA8B1AE" box="[751,804,398,419]" italics="true" pageId="7" pageNumber="8">Liara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31CB57C8FF89FFF62D2B38D7FCEDB1AE" box="[811,865,398,419]" pageId="7" pageNumber="8" sensu="stricto">s. str.</taxonomicNameLabel>
, these genitalia have a pair of rather large and semisclerotized lobes near middle or subapical parts of the genital sclerites,
</paragraph>
<paragraph id="183336A1FF89FFF62EE3389AFBEAB248" blockId="7.[227,1127,262,636]" pageId="7" pageNumber="8">
and apical parts of these sclerites are situated near each other; but in
<emphasis id="2AF8EAB3FF89FFF62D9E389AFB8EB1D5" box="[926,1026,451,472]" italics="true" pageId="7" pageNumber="8">Unalianus</emphasis>
, the male genitalia lack such lobes, and apical parts of the genital sclerites are widely separated from each other. However, these sclerites in
<taxonomicName id="DF8C4D22FF89FFF62C6738A3FC83B202" authorityName="Karny" authorityYear="1907" box="[615,783,506,527]" class="Insecta" family="Tettigoniidae" genus="Acanthocoryphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF89FFF62C6738A3FC83B202" box="[615,783,506,527]" italics="true" pageId="7" pageNumber="8">Acanthocoryphus</emphasis>
</taxonomicName>
are intermediate: with apical parts situated as in
<taxonomicName id="DF8C4D22FF89FFF62F6C3B4CFE2DB227" authorityName="Gorochov" authorityYear="2020" box="[364,417,533,554]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="2AF8EAB3FF89FFF62F6C3B4CFE2DB227" box="[364,417,533,554]" italics="true" pageId="7" pageNumber="8">Liara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31CB57C8FF89FFF62FAA3B4CFE6EB227" box="[426,482,533,554]" pageId="7" pageNumber="8" sensu="stricto">s. str.</taxonomicNameLabel>
, but without additional lobes (i.e. as in
<emphasis id="2AF8EAB3FF89FFF62D6C3B4CFC5CB227" box="[876,976,533,554]" italics="true" pageId="7" pageNumber="8">Unalianus</emphasis>
). It is a reason that the genus
<taxonomicName id="DF8C4D22FF89FFF62F773B69FE20B248" authorityName="Gorochov" authorityYear="2020" box="[375,428,560,581]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="2AF8EAB3FF89FFF62F773B69FE20B248" box="[375,428,560,581]" italics="true" pageId="7" pageNumber="8">Liara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31CB57C8FF89FFF62FB53B69FE56B24B" box="[437,474,560,582]" pageId="7" pageNumber="8" sensu="lato">s. l.</taxonomicNameLabel>
is here considered as consisting of three subgenera:
<taxonomicName id="DF8C4D22FF89FFF62DEC3B69FBADB248" authorityName="Gorochov" authorityYear="2020" box="[1004,1057,560,581]" class="Insecta" family="Tettigoniidae" genus="Liara" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="2AF8EAB3FF89FFF62DEC3B69FBADB248" box="[1004,1057,560,581]" italics="true" pageId="7" pageNumber="8">Liara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31CB57C8FF89FFF62A2A3B69FBEFB248" box="[1066,1123,560,581]" pageId="7" pageNumber="8" sensu="stricto">s. str.</taxonomicNameLabel>
,
</paragraph>
<paragraph id="183336A1FF89FFF62EE33B12FC33B271" blockId="7.[227,1127,262,636]" pageId="7" pageNumber="8">
<taxonomicName id="DF8C4D22FF89FFF62EE33B12FE07B26D" authorityName="Karny" authorityYear="1907" box="[227,395,587,608]" class="Insecta" family="Tettigoniidae" genus="Acanthocoryphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAB3FF89FFF62EE33B12FE07B26D" box="[227,395,587,608]" italics="true" pageId="7" pageNumber="8">Acanthocoryphus</emphasis>
</taxonomicName>
and
<emphasis id="2AF8EAB3FF89FFF62FBA3B12FD92B26D" box="[442,542,587,608]" italics="true" pageId="7" pageNumber="8">Unalianus</emphasis>
stat. n. These subgenera differ from each other in the abovementioned characters of male genitalia (for comparison see
<figureCitation id="80B72A24FF89FFF62D213B3FFC38B271" box="[801,948,614,636]" captionStart="Figs 3150" captionStartId="9.[265,306,1481,1503]" captionTargetBox="[231,1118,456,1445]" captionTargetId="figure-82@9.[227,1123,451,1448]" captionTargetPageId="9" captionText="Figs 3150. Liara s. l.: 3136 L. (Liara) bifurcata sp. n.; 3743 L. (Liara) inaculeata" figureDoi="http://doi.org/10.5281/zenodo.7165410" httpUri="https://zenodo.org/record/7165410/files/figure.png" pageId="7" pageNumber="8">Figs 34, 41, 50</figureCitation>
).
</paragraph>
</subSubSection>
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