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<mods:title id="7D91473F981DC4D0259848EE2B2F4F80">Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)</mods:title>
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<mods:namePart id="7920DE279FAF4B4B423EB387E4429349">Gruendel, Joachim</mods:namePart>
<mods:affiliation id="C62628C5C45923E06B225227309BCFEF">Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:namePart id="5075CD95AF279945A1CB9A464AD09450">Keupp, Helmut</mods:namePart>
<mods:affiliation id="58A7A446C4D486C47AE3552A032B4483">Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:namePart id="744604B27463F185A72159347E0DF6C3">Lang, Fritz</mods:namePart>
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<mods:namePart id="19DA16F20183EE589112A4AAA616BA70">Nuetzel, Alexander</mods:namePart>
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<mods:affiliation id="E2D0BCFA4E346DDE9AE106482B96906E">SNSB-Bayerische Staatssammlung fuer Palaeontologie und Geologie, Richard-Wagner-Str. 10, 80333 Muenchen, Germany &amp; Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany</mods:affiliation>
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<treatment id="49FC536A2D495A479354C1047C51E141" ID-GBIF-Taxon="206232336" LSID="urn:lsid:plazi:treatment:49FC536A2D495A479354C1047C51E141" httpUri="http://treatment.plazi.org/id/49FC536A2D495A479354C1047C51E141" lastPageNumber="179" pageId="0" pageNumber="179" scope_class="Gastropoda" scope_family="Ptygmatididae" scope_subClass="Heterobranchia" scope_subFamily="Ptygmatidinae" scope_superFamily="Nerineoidea">
<subSubSection id="76B3D45B2BCB84BA0826900385DAF7A6" pageId="0" pageNumber="179" type="nomenclature">
<paragraph id="D026EAFC9BECD94FD739DF0D81090461" pageId="0" pageNumber="179">
<taxonomicName id="D7278FB1CC4452D5B60143BACAB130A1" ID-CoL="C88N6" LSID="49FC536A-2D49-5A47-9354-C1047C51E141" authority="(Quenstedt, 1884)" baseAuthorityName="Quenstedt" baseAuthorityYear="1884" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">Ptygmatis? polyspira (Quenstedt, 1884)</taxonomicName>
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<paragraph id="2AC4DC593A7BCB934E4A780331D41548" pageId="0" pageNumber="179">
<figureCitation id="FEE728EE517A551396177F1629164F81" captionStart="Plate 12" captionStartId="F12" captionText="Plate 12. (1 - 3) Ptygmatis clio (d'Orbigny, 1852). (1 - 2) SNSB-BSPG 2021 XV 59, Saal (collection Lang); (1) section of fragment, height 21 mm; (2) detail, height 3.7 mm. (3) SNSB-BSPG 2021 XV 60, Saal (collection Lang), lateral view, height 108 mm. (4) Ptygmatis? polyspira (Quenstedt, 1881 - 1884), holotype, collection University Tuebingen (collection Quenstedt); Nattheim, lateral view, height 32 mm (original to Quenstedt 1881 - 1884, pl. 207, fig. 3. (11 - 12) Bactroptyxis teres (Muenster in Goldfuss, 1844), collection Sauerborn, Nattheim area. (5 - 7) Ptygmatis? ursicina (Thurmann in Thurmann &amp; Etallon, 1861). (5 - 6) SNSB-BSPG 2021 XV 30, Saal (collection Lang); (5) last whorl with aperture, height 4 mm; (6) lateral view, height 9.5 mm. (7) SNSB-BSPG 2021 XV 31, Saal (collection Lang), aperture and ornament of base, width 2.5 mm." figureDoi="10.3897/zitteliana.96.e84187.plate12" httpUri="https://binary.pensoft.net/fig/780390" pageId="0" pageNumber="179">Plate 12: fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="69D5B892FEEEED19ACB1A48AC93D2924" pageId="0" pageNumber="179" type="reference_group">
<paragraph id="BEA3D551FBE5D8E98D32DF2F2AC7A39E" pageId="0" pageNumber="179">
<taxonomicName id="E509ECB7E36A78965875521DD979FB87" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">v*1881-1884 - Nerinea polyspira</taxonomicName>
- Quenstedt: 554, pl. 207, fig. 3.
</paragraph>
<paragraph id="B944718FA3F37911B21246CEF49904E6" pageId="0" pageNumber="179">
<taxonomicName id="AD9DB762E3D752FED9A7EC49268158B2" class="Gastropoda" family="Ptygmatididae" genus="Aphanoptyxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphanoptyxis polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">1901 - Aphanoptyxis polyspira</taxonomicName>
Quenstedt - Geiger: 301.
</paragraph>
<paragraph id="06993BEA724780F07ED05B5D0B5960E2" pageId="0" pageNumber="179">
<taxonomicName id="DDBB33A03FF5427B34999315F4551CBD" class="Gastropoda" family="Ptygmatididae" genus="Aphanoptyxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphanoptyxis polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">1997 - Aphanoptyxis polyspira</taxonomicName>
(Quenstedt, 1884) -
<normalizedToken id="4401275DE0190EC607D8F7B07C3609C5" originalValue="Hägele">Haegele</normalizedToken>
: 133, fig. p.133, lower left.
</paragraph>
</subSubSection>
<subSubSection id="6CA7C98E857E3B3BC38D7A55B38C75FA" pageId="0" pageNumber="179" type="materials_examined">
<paragraph id="68A19BD9B68459D241098D3D803DD78F" pageId="0" pageNumber="179">Material.</paragraph>
<paragraph id="0680377441186E99C62744D5D1394E2A" pageId="0" pageNumber="179">
<bibRefCitation id="0AFA261C8F6DDAC7D26C5FBA112A5BED" author="Quenstedt, FA" journalOrPublisher="Fues, Leipzig" pageId="0" pageNumber="179" refId="B82" refString="Quenstedt, FA, 1881-1884. Petrefaktenkunde Deutschlands. Erste Abteilung, Band 7: Gasteropoden. Fues, Leipzig" title="Petrefaktenkunde Deutschlands. Erste Abteilung, Band 7: Gasteropoden." year="1881 - 1884">
<normalizedToken id="10D8864CBAF47849FD749047EB6B3EAB" originalValue="Quenstedts">Quenstedt's</normalizedToken>
(1881-1884)
</bibRefCitation>
figured specimen (
<typeStatus id="15DFAA873D37C008346C30F239D408E9">holotype</typeStatus>
by monotypy) from Nattheim (
<normalizedToken id="4521629DC488446432FCE9A976ED05A6" originalValue="Tübingen">Tuebingen</normalizedToken>
, Quenstedt collection).
</paragraph>
</subSubSection>
<subSubSection id="CC14AB26DAB85F6A9D76A30746C7E437" pageId="0" pageNumber="179" type="description">
<paragraph id="538FB8E0FEC1CA832C6DBDAF322109C4" pageId="0" pageNumber="179">Description.</paragraph>
<paragraph id="FF1B1EAEB8BC3CB7BE4D4ABBAAA9A637" pageId="0" pageNumber="179">The specimen consists of 7 whorls and is 32 mm high (apex missing). The shell is moderately slender and the whorls are increasing regularly in width. The sutures are somewhat pronounced by a subsutural bulge. The whorl face is straight and entirely covered by spiral cords (7-8 spiral cords on last whorl). The transition from whorl face to base is angular. The aperture is not preserved, plaits are not visible.</paragraph>
</subSubSection>
<subSubSection id="965655C7BE8CA5369FE1946A04B8E1C4" pageId="0" pageNumber="179" type="remarks">
<paragraph id="EA146CB1C8FF55E0075F388086CABA2F" pageId="0" pageNumber="179">Remarks.</paragraph>
<paragraph id="5391B3DAC0B31DC2CF0DD4371FA95850" pageId="0" pageNumber="179">
The studied holotype of
<taxonomicName id="3425492156591CF7D0895E99B4C864BE" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">
<emphasis id="D583F7FCBA972C75C5676CA8A03DEBCC" italics="true" pageId="0" pageNumber="179">Nerinea polyspira</emphasis>
</taxonomicName>
Quenstedt, 1884 is a poorly preserved specimen. Its systematic and taxonomic position remain unclear because aperture and plaits are unknown.
</paragraph>
</subSubSection>
<subSubSection id="EECC3751F9A08A34625A3A0D74D7D76B" pageId="0" pageNumber="179" type="relationships">
<paragraph id="9B0AF14FF9D44F299D1A27AFAD57185A" pageId="0" pageNumber="179">Relationships.</paragraph>
<paragraph id="6993A0FF9D062C335A32AF251A61C8A2" pageId="0" pageNumber="179">
<taxonomicName id="3A619A0CC42F7916391573E7891F5B43" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">
<emphasis id="B7055C373DD125A0EBC0C99D600A0D11" italics="true" pageId="0" pageNumber="179">Nerinea ursicina</emphasis>
</taxonomicName>
Thurmann, 1861 (in
<bibRefCitation id="662C7249B04FE465877C1105AE2A60D9" author="Thurmann, J" journalOrPublisher="Memoires de la Societe du museum d'histoire naturelle de Strasbourg" pageId="0" pageNumber="179" refId="B100" refString="Thurmann, J, Etallon, A, 1861-1864. Lethea bruntrutana ou etudes paleontologiques et stratigraphiques sur le Jura bernois et en particulier les environs de Porrentruy. Neue Denkschrift der allgemeinen Schweizerischen Gesellschaft fuer die Naturwissenschaften 18-20: 1-500." title="Lethea bruntrutana ou etudes paleontologiques et stratigraphiques sur le Jura bernois et en particulier les environs de Porrentruy. Neue Denkschrift der allgemeinen Schweizerischen Gesellschaft fuer die Naturwissenschaften 18 - 20: 1 - 500." year="1861 - 1864">
Thurmann and
<normalizedToken id="14D262A7448EE8097B12C4393F03C7A6" originalValue="Étallon">Etallon</normalizedToken>
1861-1864
</bibRefCitation>
) differs in having a strong adapical bulge, making the whorl face distinctly concave, fewer spiral cords, and four apertural plaits.
<taxonomicName id="9620C666862C2E7DA78FDA3AA64C122D" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea punctata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="punctata">
<emphasis id="3D2CA3D93C3BFFB35E5535BD1D5788FD" italics="true" pageId="0" pageNumber="179">Nerinea punctata</emphasis>
</taxonomicName>
Voltz sensu
<bibRefCitation id="F9C43495349255CA586FC7A7883FC893" author="Bronn, HG" journalOrPublisher="Neues Jahrbuch fuer Mineralogie, Geognosie, Geologie und Petrefaktenkunde" pageId="0" pageNumber="179" pagination="544 - 566" refId="B5" refString="Bronn, HG, 1836. Uebersicht und Abbildungen der bis jetzt bekannten Nerinea -Arten. Neues Jahrbuch fuer Mineralogie, Geognosie, Geologie und Petrefaktenkunde 4: 544 - 566" title="Uebersicht und Abbildungen der bis jetzt bekannten Nerinea - Arten." volume="4" year="1836">Bronn (1836)</bibRefCitation>
has a narrow but distinct ramp, and only three spiral cords on its whorl face.
<taxonomicName id="0DF58F0E1A4E9E9829613FF80689927B" class="Gastropoda" family="Eunerineidae" genus="Nerinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinella calliope" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="calliope">
<emphasis id="9F4C15A360D000D827C3734AFB638D51" italics="true" pageId="0" pageNumber="179">Nerinella calliope</emphasis>
</taxonomicName>
<normalizedToken id="E9ED776DD8B59BA7B5C206D17DDAEB2D" originalValue="dOrbigny">d'Orbigny</normalizedToken>
sensu
<bibRefCitation id="5F5C77A004A5DCF8B59453FB478E9215" author="Cossmann, M" journalOrPublisher="The author and Comptoir Geologique, Paris" pageId="0" pageNumber="179" refId="B12" refString="Cossmann, M, 1898. Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1-179." title="Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1 - 179." year="1898">Cossmann (1898)</bibRefCitation>
has 5-6 spiral cords on the whorl face, some of them having fine knobs. Its aperture has three plaits.
<taxonomicName id="7C8E3FFD4AC3BD49CF1BDDBF25778475" class="Gastropoda" family="Eunerineidae" genus="Nerinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinella turritella" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="turritella">
<emphasis id="0974CD81C62313781947E04F7C58C42A" italics="true" pageId="0" pageNumber="179">Nerinella turritella</emphasis>
</taxonomicName>
Voltz sensu
<bibRefCitation id="203EF8908F1E401DA2999085A0348F66" author="Cossmann, M" journalOrPublisher="The author and Comptoir Geologique, Paris" pageId="0" pageNumber="179" refId="B12" refString="Cossmann, M, 1898. Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1-179." title="Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1 - 179." year="1898">Cossmann (1898)</bibRefCitation>
lacks a bulge and has four strong, knobby spiral cords on the whorl face and additional weaker cords between them.
<taxonomicName id="38953D68E9BEBF3827CB8140D16A6C17" class="Gastropoda" family="Eunerineidae" genus="Nerinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinella cyane" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="cyane">
<emphasis id="6A4235204F508FEB3367DE9614287979" italics="true" pageId="0" pageNumber="179">Nerinella cyane</emphasis>
</taxonomicName>
Loriol in Loriol &amp; Pellat, 1874 has higher whorls with a smooth portion above the suture.
</paragraph>
</subSubSection>
</treatment>
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