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<document id="AFF56B67BC38EC428D6535EFE8AB4024" ID-CLB-Dataset="7979" ID-DOI="10.11646/zootaxa.4933.3.4" ID-GBIF-Dataset="a3bf6440-2b88-402b-9bbe-107a257286e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4554966" ID-ZooBank="92A9D8F4-88CF-4971-9759-276BB230E208" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613986682380" checkinUser="plazi" docAuthor="Heffern, Daniel, Santos-Silva, Antonio &amp; Nascimento, Francisco E. L." docDate="2021" docId="03C89A5DBD14CE6B52CAFE8F58FBDB40" docLanguage="en" docName="zootaxa.4933.3.4.pdf" docOrigin="Zootaxa 4933 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nyssicus Pascoe 1859" docType="key" docVersion="8" lastPageNumber="364" masterDocId="FFF1E225BD17CE68525DFFA85D67DF18" masterDocTitle="American Cerambycidae (Coleoptera): Four new species, new records and taxonomic notes" masterLastPageNumber="378" masterPageNumber="361" pageNumber="364" updateTime="1698931602904" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0F5B8181E97DD2A0CD941AC402202933">American Cerambycidae (Coleoptera): Four new species, new records and taxonomic notes</mods:title>
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<mods:namePart id="C4E56A5DC7463B6C35DCBBA1E5CAA913">Heffern, Daniel</mods:namePart>
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<mods:affiliation id="5D0D712B476826A76C16CE904155CC71">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil &amp; toncriss @ uol. com. br; https: // orcid. org / 0000 - 0001 - 7128 - 1418</mods:affiliation>
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<mods:namePart id="EEBDD424B0C4DAC0CD190D756D801B10">Nascimento, Francisco E. L.</mods:namePart>
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<mods:affiliation id="9F995A72779A8207A8CE0827A2051641">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil &amp; eribnascimentofl @ gmail. com; https: // orcid. org / 0000 - 0002 - 5047 - 8921</mods:affiliation>
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<paragraph id="8BDE2B4BBD14CE6B52CAFE8F5CBFDE59" blockId="3.[151,472,294,321]" box="[151,472,294,321]" pageId="3" pageNumber="364">
<heading id="D0969C27BD14CE6B52CAFE8F5CBFDE59" bold="true" box="[151,472,294,321]" fontSize="11" level="1" pageId="3" pageNumber="364" reason="1">
<emphasis id="B915F759BD14CE6B52CAFE8F5CBFDE59" bold="true" box="[151,472,294,321]" pageId="3" pageNumber="364">
Key to species of
<taxonomicName id="4C6150C8BD14CE6B5333FE8F5CBFDE59" ID-CoL="665M" authorityName="Pascoe" authorityYear="1859" box="[366,472,295,321]" class="Insecta" family="Cerambycidae" genus="Nyssicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="B915F759BD14CE6B5333FE8F5CBFDE59" bold="true" box="[366,472,295,321]" italics="true" pageId="3" pageNumber="364">Nyssicus</emphasis>
</taxonomicName>
</emphasis>
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</paragraph>
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<subSubSection id="C37B78C0BD14CE6B52CAFECD58FBDB40" pageId="3" pageNumber="364" type="key">
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<paragraph id="8BDE2B4BBD14CE6B52CAFECD58FBDE62" blockId="3.[151,1436,357,408]" box="[151,1436,357,379]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFECD58FBDE62" box="[151,1436,357,379]" pageId="3" pageNumber="364">1) Outer spine of the elytral apex black....................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BDE2B4BBD14CE6B52CAFE2B58FBDE80" blockId="3.[151,1436,357,408]" box="[151,1436,386,408]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFE2B58FBDE80" box="[151,1436,386,408]" pageId="3" pageNumber="364"> Outer spine of the elytral apex not black, concolor with adjacent area............................................ 5</keyLead>
</paragraph>
</keyStep>
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<keyLead id="3090887EBD14CE6B52CAFE1558FBDEE8" pageId="3" pageNumber="364">
2(1). Pronotum with blackish macula close to center of anterior margin.
<collectingCountry id="F3766BDBBD14CE6B5101FE155E8BDECB" box="[860,1004,445,467]" name="French Guiana" pageId="3" pageNumber="364">French Guiana</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B51A8FE155957DECB" box="[1013,1072,445,467]" name="Brazil" pageId="3" pageNumber="364">Brazil</collectingCountry>
(
<collectingRegion id="49A5E5A9BD14CE6B5666FE15590EDECB" box="[1083,1129,445,467]" country="Brazil" name="Para" pageId="3" pageNumber="364">Pará</collectingRegion>
,
<collectingRegion id="49A5E5A9BD14CE6B562CFE1559A5DECB" box="[1137,1218,445,467]" country="Brazil" name="Alagoas" pageId="3" pageNumber="364">Alagoas</collectingRegion>
,
<collectingRegion id="49A5E5A9BD14CE6B5696FE155862DECB" box="[1227,1285,445,467]" country="Brazil" name="Bahia" pageId="3" pageNumber="364">Bahia</collectingRegion>
,
<collectingRegion id="49A5E5A9BD14CE6B5753FE1558FEDECA" box="[1294,1433,445,467]" country="Brazil" name="Espirito Santo" pageId="3" pageNumber="364">Espírito Santo</collectingRegion>
,
<collectingRegion id="49A5E5A9BD14CE6B52BFFE725C08DEE8" box="[226,367,474,496]" country="Brazil" name="Rio de Janeiro" pageId="3" pageNumber="364">Rio de Janeiro</collectingRegion>
, S„o Paulo).........................................................
<taxonomicName id="4C6150C8BD14CE6B56C8FE7358FBDEE8" ID-CoL="487Q2" authority="Martins, 2005" authorityName="Martins" authorityYear="2005" box="[1173,1436,474,496]" class="Insecta" family="Cerambycidae" genus="Nyssicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="mendosus">
<emphasis id="B915F759BD14CE6B56C8FE735877DEF7" box="[1173,1296,474,496]" italics="true" pageId="3" pageNumber="364">N. mendosus</emphasis>
Martins, 2005
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE2B4BBD14CE6B52CAFE5058FBDD15" blockId="3.[151,1437,445,526]" box="[151,1436,504,526]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFE5058FBDD15" box="[151,1436,504,526]" pageId="3" pageNumber="364"> Pronotum lacking blackish macula close to anterior margin.................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="309533EEBD14CE6B52CAFD9A58FBDD9A" pageId="3" pageNumber="364">
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<keyLead id="3090887EBD14CE6B52CAFD9A58FBDD7D" pageId="3" pageNumber="364">
3(2). Internal tubercle of the scape in male placed about middle, wide lobe-shaped.
<collectingCountry id="F3766BDBBD14CE6B5192FD9A593ADD50" box="[975,1117,562,584]" name="French Guiana" pageId="3" pageNumber="364">French Guiana</collectingCountry>
................................................................................................
<taxonomicName id="4C6150C8BD14CE6B566FFDF858FBDD7D" ID-CoL="487PZ" authority="Tavakilian &amp; Dalens, 2014" authorityName="Tavakilian &amp; Dalens" authorityYear="2014" box="[1074,1436,592,614]" class="Insecta" family="Cerambycidae" genus="Nyssicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="lobatus">
<emphasis id="B915F759BD14CE6B566FFDF859F2DD7D" box="[1074,1173,592,613]" italics="true" pageId="3" pageNumber="364">N. lobatus</emphasis>
<bibRefCitation id="EFF056BABD14CE6B56C7FDF858FBDD7D" author="Tavakilian, G. L. &amp; Dalens, P. - H." box="[1178,1436,592,614]" pageId="3" pageNumber="364" pagination="23 - 47" refId="ref14435" refString="Tavakilian, G. L. &amp; Dalens, P. - H. (2014) Revision du genre Nyssicus Pascoe, 1859 et creation du genre Heteronyssicus n. g. (Coleoptera, Cerambycidae, Cerambycinae). Les Cahiers Magellanes, New Series, 15, 23 - 47." type="journal article" year="2014">Tavakilian &amp; Dalens, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE2B4BBD14CE6B52CAFDC558FBDD9A" blockId="3.[151,1436,562,643]" box="[151,1436,621,643]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFDC558FBDD9A" box="[151,1436,621,643]" pageId="3" pageNumber="364"> Internal tubercle of the scape in male placed before middle, slender............................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="309533EEBD14CE6B52CAFD0058FBDC0E" pageId="3" pageNumber="364">
<paragraph id="8BDE2B4BBD14CE6B52CAFD0058FBDDC2" blockId="3.[151,1436,680,790]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFD0058FBDDC2" pageId="3" pageNumber="364">
4(3). Ventral surface of the metafemoral club with abundant setigerous granules.
<collectingCountry id="F3766BDBBD14CE6B51E5FD00592FDDA6" box="[952,1096,680,702]" name="French Guiana" pageId="3" pageNumber="364">French Guiana</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B560FFD0059EADDA6" box="[1106,1165,680,702]" name="Brazil" pageId="3" pageNumber="364">Brazil</collectingCountry>
(
<collectingRegion id="49A5E5A9BD14CE6B56C4FD005985DDA6" box="[1177,1250,680,702]" country="Brazil" name="Amapa" pageId="3" pageNumber="364">Amapá</collectingRegion>
, Amazonas)........................................................................
<taxonomicName id="4C6150C8BD14CE6B564DFD6E58FBDDC2" ID-CoL="487PY" authority="Tavakilian &amp; Dalens, 2014" authorityName="Tavakilian &amp; Dalens" authorityYear="2014" box="[1040,1436,709,731]" class="Insecta" family="Cerambycidae" genus="Nyssicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B915F759BD14CE6B564DFD6E59F3DDC2" box="[1040,1172,709,731]" italics="true" pageId="3" pageNumber="364">N. granulatus</emphasis>
<bibRefCitation id="EFF056BABD14CE6B56C7FD6D58FBDDC2" author="Tavakilian, G. L. &amp; Dalens, P. - H." box="[1178,1436,709,731]" pageId="3" pageNumber="364" pagination="23 - 47" refId="ref14435" refString="Tavakilian, G. L. &amp; Dalens, P. - H. (2014) Revision du genre Nyssicus Pascoe, 1859 et creation du genre Heteronyssicus n. g. (Coleoptera, Cerambycidae, Cerambycinae). Les Cahiers Magellanes, New Series, 15, 23 - 47." type="journal article" year="2014">Tavakilian &amp; Dalens, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE2B4BBD14CE6B52CAFD4B58FBDC0E" blockId="3.[151,1436,680,790]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFD4B58FBDC0E" pageId="3" pageNumber="364">
Ventral surface of the metafemoral club with a few setigerous granules.
<collectingCountry id="F3766BDBBD14CE6B51C7FD4A5E9DDDE0" box="[922,1018,738,760]" name="Honduras" pageId="3" pageNumber="364">Honduras</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B5659FD4A5931DDE0" box="[1028,1110,738,760]" name="Ecuador" pageId="3" pageNumber="364">Ecuador</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B5602FD4A59A3DDE0" box="[1119,1220,738,760]" name="Venezuela" pageId="3" pageNumber="364">Venezuela</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B5693FD4B599BDDE0" box="[1230,1276,739,760]" name="Peru" pageId="3" pageNumber="364">Peru</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B575BFD4A5828DDE0" box="[1286,1359,738,760]" name="Bolivia" pageId="3" pageNumber="364">Bolivia</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B5704FD4A5C4DDC0E" name="French Guiana" pageId="3" pageNumber="364">French Guiana</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B536EFCA85C09DC0E" box="[307,366,768,790]" name="Brazil" pageId="3" pageNumber="364">Brazil</collectingCountry>
(Amazonas,
<collectingRegion id="49A5E5A9BD14CE6B53B0FCA85F7DDC0E" box="[493,538,768,790]" country="Brazil" name="Para" pageId="3" pageNumber="364">Pará</collectingRegion>
,
<collectingRegion id="49A5E5A9BD14CE6B5079FCA85FC5DC0D" box="[548,674,768,789]" country="Brazil" name="Mato Grosso" pageId="3" pageNumber="364">Mato Grosso</collectingRegion>
,
<collectingRegion id="49A5E5A9BD14CE6B50F1FCA85E6BDC0E" box="[684,780,768,790]" country="Brazil" name="Rondonia" pageId="3" pageNumber="364">Rondônia</collectingRegion>
, Maranh„o)................
<taxonomicName id="4C6150C8BD14CE6B5619FCA858FBDC0E" ID-CoL="487QF" authority="(Swederus, 1787)" baseAuthorityName="Swederus" baseAuthorityYear="1787" box="[1092,1436,768,790]" class="Insecta" family="Cerambycidae" genus="Nyssicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="quadriguttatus">
<emphasis id="B915F759BD14CE6B5619FCA8598ADC0D" box="[1092,1261,768,789]" italics="true" pageId="3" pageNumber="364">N. quadriguttatus</emphasis>
(Swederus, 1787)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309533EEBD14CE6B52CAFC9258FBDC93" pageId="3" pageNumber="364">
<paragraph id="8BDE2B4BBD14CE6B52CAFC9258FBDC75" blockId="3.[151,1437,826,907]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFC9258FBDC75" pageId="3" pageNumber="364">
5(1). Scape in male lacking internal tubercle; apex of the antennomeres IIIV blackish; sutural spine slightly shorter than outer spine.
<collectingCountry id="F3766BDBBD14CE6B52BFFCF05C4BDC76" box="[226,300,856,878]" name="Bolivia" pageId="3" pageNumber="364">Bolivia</collectingCountry>
......................................................................
<taxonomicName id="4C6150C8BD14CE6B562FFCF058FBDC75" ID-CoL="487PW" authority="Martins, 2005" authorityName="Martins" authorityYear="2005" box="[1138,1436,855,878]" class="Insecta" family="Cerambycidae" genus="Nyssicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="contaminatus">
<emphasis id="B915F759BD14CE6B562FFCF05868DC75" box="[1138,1295,856,877]" italics="true" pageId="3" pageNumber="364">N. contaminatus</emphasis>
Martins, 2005
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE2B4BBD14CE6B52CAFCDD58FBDC93" blockId="3.[151,1437,826,907]" box="[151,1436,885,907]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFCDD58FBDC93" box="[151,1436,885,907]" pageId="3" pageNumber="364"> Scape in male with internal tubercle; apex of the antennomeres not blackish; sutural spine much shorter than outer spine... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="309533EEBD14CE6B52CAFC0758FBDB40" pageId="3" pageNumber="364">
<paragraph id="8BDE2B4BBD14CE6B52CAFC0758FBDCE7" blockId="3.[151,1437,943,1112]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFC0758FBDCE7" pageId="3" pageNumber="364">
6(5). Anterior margin of the internal tubercle of the scape in male as an extension of the anterior margin of the scape; anterior yellowish elytral macula subtriangular; posterior yellowish elytral macula distinctly longer than wide.
<collectingCountry id="F3766BDBBD14CE6B56C6FC655999DCFB" box="[1179,1278,973,995]" name="Venezuela" pageId="3" pageNumber="364">Venezuela</collectingCountry>
...................................................................................
<taxonomicName id="4C6150C8BD14CE6B5661FC4358FBDCE7" ID-CoL="487PX" authority="Joly &amp; Martinez, 1981" authorityName="Joly &amp; Martinez" authorityYear="1981" box="[1084,1436,1002,1024]" class="Insecta" family="Cerambycidae" genus="Nyssicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="B915F759BD14CE6B5661FC4359DADCE7" box="[1084,1213,1002,1024]" italics="true" pageId="3" pageNumber="364">N. fernandezi</emphasis>
Joly &amp; Martínez, 1981
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE2B4BBD14CE6B52CAFBA058FBDB40" blockId="3.[151,1437,943,1112]" pageId="3" pageNumber="364">
<keyLead id="3090887EBD14CE6B52CAFBA058FBDB40" pageId="3" pageNumber="364">
Anterior margin of the internal tubercle of the scape in male not as extension of the anterior margin of the scape (internal margin of the scape distinctly concave base and tubercle); anterior yellowish elytral macula subrounded; posterior yellowish elytral macula slightly longer than wide.
<collectingCountry id="F3766BDBBD14CE6B5075FBEA5FEFDB40" box="[552,648,1090,1112]" name="Honduras" pageId="3" pageNumber="364">Honduras</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B50CFFBEA5F90DB40" box="[658,759,1090,1112]" name="Nicaragua" pageId="3" pageNumber="364">Nicaragua</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B515DFBEA5E0CDB40" box="[768,875,1090,1112]" name="Costa Rica" pageId="3" pageNumber="364">Costa Rica</collectingCountry>
,
<collectingCountry id="F3766BDBBD14CE6B5128FBEB5EA6DB40" box="[885,961,1091,1112]" name="Panama" pageId="3" pageNumber="364">Panama</collectingCountry>
............
<taxonomicName id="4C6150C8BD14CE6B5600FBEB58FBDB40" ID-CoL="487RQ" authority="(Linsley, 1935)" baseAuthorityName="Linsley" baseAuthorityYear="1935" box="[1117,1436,1090,1112]" class="Insecta" family="Cerambycidae" genus="Nyssicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="topographicus">
<emphasis id="B915F759BD14CE6B5600FBEB5863DB4F" box="[1117,1284,1090,1112]" italics="true" pageId="3" pageNumber="364">N. topographicus</emphasis>
(
<bibRefCitation id="EFF056BABD14CE6B574DFBEA58F2DB40" author="Linsley, E. G." box="[1296,1429,1090,1112]" pageId="3" pageNumber="364" pagination="67 - 102" refId="ref13892" refString="Linsley, E. G. (1935) Studies in the Longicornia of Mexico (Coleoptera: Cerambycidae). Transactions of the American Entomological Society, 61, 67 - 102." type="journal article" year="1935">Linsley, 1935</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>