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<mods:title id="663F573C3A58C4B401EFCEC1FFD0AD9D">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:affiliation id="3B28CAF638B7F8D0414685584CB25DDC">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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12)
<taxonomicName id="4C61A683EB37574BFF3FDFFAFD48FCA7" ID-CoL="4GS4N" authority="Duda, 1923" authorityName="Duda (Duda" authorityYear="1923" box="[198,675,754,781]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="foliiseta" subGenus="Ashima">
<emphasis id="B9150112EB37574BFF3FDFFAFEC5FCA7" bold="true" box="[198,302,754,781]" italics="true" pageId="17" pageNumber="18">Phortica</emphasis>
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<emphasis id="B9150112EB37574BFE54DFFAFDE6FCA7" bold="true" box="[429,525,754,781]" italics="true" pageId="17" pageNumber="18">foliiseta</emphasis>
<bibRefCitation id="EFF0A0F1EB37574BFDECDFFBFD48FCA7" author="Duda" box="[533,675,755,781]" firstAuthor="Duda" pageId="17" pageNumber="18" pagination="24 - 59" refId="ref34577" refString="Duda, O. (1923) Die orientalischen und australischen Drosophiliden-Arten (Dipteren) des Ungarischen National-Museums zu Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 20, 24 - 59." type="journal article" year="1923">Duda, 1923</bibRefCitation>
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<treatmentCitation id="0AC0FB11EB37574BFF6EDE3FFE05FCE5" author="Duda, O." box="[151,494,822,847]" page="35" pageId="17" pageNumber="18" year="1923">
<taxonomicName id="4C61A683EB37574BFF6EDE3FFE05FCE5" ID-CoL="4GS4N" authority="Duda, 1923: 35" authorityName="Duda (Duda" authorityPageNumber="35" authorityYear="1923" box="[151,494,822,847]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="foliiseta">
<emphasis id="B9150112EB37574BFF6EDE3FFEA3FCE5" box="[151,328,823,847]" italics="true" pageId="17" pageNumber="18">Phortica foliiseta</emphasis>
<bibRefCitation id="EFF0A0F1EB37574BFEB7DE3EFE05FCE5" author="Duda" box="[334,494,822,847]" firstAuthor="Duda" pageId="17" pageNumber="18" pagination="24 - 59" refId="ref34577" refString="Duda, O. (1923) Die orientalischen und australischen Drosophiliden-Arten (Dipteren) des Ungarischen National-Museums zu Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 20, 24 - 59." type="journal article" year="1923">Duda, 1923: 35</bibRefCitation>
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;
<treatmentCitation id="0AC0FB11EB37574BFE01DE3EFD02FCE5" author="Chen, H. W. &amp; Toda, M. J. &amp; Gao, J. J." box="[504,745,822,847]" page="420" pageId="17" pageNumber="18" year="2005">
<bibRefCitation id="EFF0A0F1EB37574BFE01DE3EFD02FCE5" author="Chen" box="[504,745,822,847]" etAl="et al." firstAuthor="Chen" pageId="17" pageNumber="18" pagination="419 - 429" refId="ref34109" refString="Chen, H. W., Toda, M. J. &amp; Gao, J. J. (2005 b) The Phortica (s. str.) foliiseta species-complex (Diptera, Drosophilidae) from China and its adjacent countries. Acta Zootaxonomica Sinica, 30, 419 - 429." type="journal article" year="2005" yearSuffix="b">
Chen
<emphasis id="B9150112EB37574BFDCDDE30FD8AFCE5" box="[564,609,823,847]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
., 2005b: 420
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<treatmentCitation id="0AC0FB11EB37574BFF6EDE5EFD3BFCC4" author="Tsacas, L. &amp; Okada, T." box="[151,720,853,878]" page="230" pageId="17" pageNumber="18" year="1983">
<taxonomicName id="4C61A683EB37574BFF6EDE5EFD3BFCC4" authority=": Tsacas &amp; Okada, 1983: 230" authorityName=": Tsacas &amp; Okada" authorityPageNumber="230" authorityYear="1983" box="[151,720,853,878]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="foliiseta" subGenus="Phortica">
<emphasis id="B9150112EB37574BFF6EDE5EFF0BFCC7" box="[151,224,854,877]" italics="true" pageId="17" pageNumber="18">Amiota</emphasis>
(
<emphasis id="B9150112EB37574BFF16DE5EFEA3FCC4" box="[239,328,854,878]" italics="true" pageId="17" pageNumber="18">Phortica</emphasis>
)
<emphasis id="B9150112EB37574BFEAEDE5EFE42FCC4" box="[343,425,854,878]" italics="true" pageId="17" pageNumber="18">foliiseta</emphasis>
:
<bibRefCitation id="EFF0A0F1EB37574BFE4CDE5EFD3BFCC4" author="Tsacas &amp; Okada" box="[437,720,853,878]" firstAuthor="Tsacas" pageId="17" pageNumber="18" pagination="228 - 237" refId="ref37303" refString="Tsacas, L. &amp; Okada, T. (1983) On the Oriental and New Guinean species of the genus Amiota Loew originally described by Duda as Phortica foliiseta, with descriptions of three new species (Diptera, Drosophilidae). Kontyu, 51, 228 - 237." type="journal article" year="1983">Tsacas &amp; Okada, 1983: 230</bibRefCitation>
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<treatmentCitation id="0AC0FB11EB37574BFF6EDE7CFD57FC27" author="Chen, H. W. &amp; Maca, J." box="[151,700,884,909]" page="507" pageId="17" pageNumber="18" year="2012">
<taxonomicName id="4C61A683EB37574BFF6EDE7CFD57FC27" ID-CoL="4GS4N" authority=": Chen &amp; Maca, 2012: 507" authorityName=": Chen &amp; Maca" authorityPageNumber="507" authorityYear="2012" box="[151,700,884,909]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="foliiseta" subGenus="Ashima">
<emphasis id="B9150112EB37574BFF6EDE7CFF1BFC26" box="[151,240,884,908]" italics="true" pageId="17" pageNumber="18">Phortica</emphasis>
(
<emphasis id="B9150112EB37574BFF06DE7CFEA0FC26" box="[255,331,884,908]" italics="true" pageId="17" pageNumber="18">Ashima</emphasis>
)
<emphasis id="B9150112EB37574BFEA0DE7CFE40FC26" box="[345,427,884,908]" italics="true" pageId="17" pageNumber="18">foliiseta</emphasis>
:
<bibRefCitation id="EFF0A0F1EB37574BFE41DE7CFD57FC27" author="Chen &amp; Maca" box="[440,700,884,909]" firstAuthor="Chen" pageId="17" pageNumber="18" pagination="493 - 509" refId="ref33941" refString="Chen, H. W. &amp; Maca, J. (2012) Ten new species of the genus Phortica from the Afrotropical and Oriental regions (Diptera: Drosophilidae). Zootaxa, 3478 (1), 493 - 509. https: // doi. org / 10.11646 / zootaxa. 3478.1.42" type="journal article" year="2012">Chen &amp; Máca, 2012: 507</bibRefCitation>
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<subSubSection id="C37B8E8BEB37574BFF6EDEB2FC57FA5B" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BDEDD00EB37574BFF6EDEB2FAF7FBB7" blockId="17.[151,1437,954,1666]" pageId="17" pageNumber="18">
<emphasis id="B9150112EB37574BFF6EDEB2FEF9FC7F" bold="true" box="[151,274,954,981]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
Dorsomedial, tentorial apodeme 3/4 as long as basal, parallel portion of dorsolateral, tentorial apodeme; longest, dorsal branch of arista as long as longest seta on pedicel; each lateral lobe of aedeagal sheath with apically more or less pointed, sclerotized, anterior and posterior postgonites (“
<figureCitation id="135AC185EB37574BFC5BD90BFC03FBB7" box="[930,1000,1027,1053]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
” in
<bibRefCitation id="EFF0A0F1EB37574BFBEFD90BFAFBFBB7" author="Tsacas &amp; Okada" box="[1046,1296,1027,1053]" firstAuthor="Tsacas" pageId="17" pageNumber="18" pagination="228 - 237" refId="ref37303" refString="Tsacas, L. &amp; Okada, T. (1983) On the Oriental and New Guinean species of the genus Amiota Loew originally described by Duda as Phortica foliiseta, with descriptions of three new species (Diptera, Drosophilidae). Kontyu, 51, 228 - 237." type="journal article" year="1983">Tsacas &amp; Okada 1983</bibRefCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB37574BFF3ED92EFEADFAB0" blockId="17.[151,1437,954,1666]" pageId="17" pageNumber="18">
<emphasis id="B9150112EB37574BFF3ED92EFDFAFBEB" bold="true" box="[199,529,1062,1089]" pageId="17" pageNumber="18">Supplementary description.</emphasis>
Supracervical setae approximately 12. Some interfrontal setulae as long as lowest setulae on ocellar triangle (ch.3-0). Arista with arb = 12/0; apical, leaf-like expansion narrower than long, apically tapering and pointed (ch.7-1). Cibarial, medial sensilla approximately 7 per side; posterior sensilla approximately 6 per side. Fore and mid tibiae entirely yellow (ch.17,18-0). All tarsi entirely yellow. Abdominal sternite III longer than wide (ch.22-0), not wider than IV (ch.23-0), without dense setulae on medial surface (ch.26-0). Pregonite gently curved ventrad distally. Posterior postgonites on lateral lobes of aedeagal sheath medially separated from each other (ch.66-0).
</paragraph>
<paragraph id="8BDEDD00EB37574BFF3ED82BFDEAFA2F" blockId="17.[151,1437,954,1666]" pageId="17" pageNumber="18">
The antisymmetry is observed in the postgonites: in A-type, the right, posterior postgonite is slightly curved but the left one nearly straight, and the right, anterior postgonite is thicker than the left one (“
<figureCitation id="135AC185EB37574BFB8CD84FFB51FACB" box="[1141,1210,1351,1377]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
” in
<bibRefCitation id="EFF0A0F1EB37574BFB1FD84FFF38FA2F" author="Tsacas &amp; Okada" firstAuthor="Tsacas" pageId="17" pageNumber="18" pagination="228 - 237" refId="ref37303" refString="Tsacas, L. &amp; Okada, T. (1983) On the Oriental and New Guinean species of the genus Amiota Loew originally described by Duda as Phortica foliiseta, with descriptions of three new species (Diptera, Drosophilidae). Kontyu, 51, 228 - 237." type="journal article" year="1983">Tsacas &amp; Okada 1983</bibRefCitation>
); but in B-type,
<emphasis id="B9150112EB37574BFE70D864FE11FA2F" box="[393,506,1388,1413]" italics="true" pageId="17" pageNumber="18">vice versa</emphasis>
.
</paragraph>
<paragraph id="8BDEDD00EB37574BFF3ED886FC57FA5B" blockId="17.[151,1437,954,1666]" pageId="17" pageNumber="18">
<emphasis id="B9150112EB37574BFF3ED886FE2DFA03" bold="true" box="[199,454,1422,1449]" pageId="17" pageNumber="18">Specimens examined.</emphasis>
<materialsCitation id="3B09D75DEB37574BFE32D887FB50FA03" ID-GBIF-Occurrence="2636268400" box="[459,1211,1423,1449]" collectingDate="1992-10-16" collectionCode="SEHU" collectorName="M. J. Toda" country="Taiwan" location="Shanlinxi" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2">
<collectingCountry id="F3769D90EB37574BFE32D887FDCAFA03" box="[459,545,1423,1449]" name="Taiwan" pageId="17" pageNumber="18">Taiwan</collectingCountry>
:
<specimenCount id="9D671689EB37574BFDD2D887FDA5FA03" box="[555,590,1423,1449]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
(A-type),
<location id="8EBE8BDBEB37574BFD3BD887FCDBFA03" LSID="urn:lsid:plazi:treatment:03C86C16EB37574BFF6EDFFAFAB1F92B:8EBE8BDBEB37574BFD3BD887FCDBFA03" box="[706,816,1423,1449]" country="Taiwan" name="Shanlinxi" pageId="17" pageNumber="18">Shanlinxi</location>
,
<date id="FFDFFBC0EB37574BFCC3D887FC46FA03" box="[826,941,1423,1449]" pageId="17" pageNumber="18" value="1992-10-16">
<collectingDate id="EF9B0228EB37574BFCC3D887FC46FA03" box="[826,941,1423,1449]" pageId="17" pageNumber="18" value="1992-10-16">16.x.1992</collectingDate>
</date>
,
<collectorName id="2694B8D6EB37574BFC4ED887FBCCFA03" box="[951,1063,1423,1449]" pageId="17" pageNumber="18">M.J. Toda</collectorName>
leg. (
<collectionCode id="ED7045C5EB37574BFB9DD887FB58FA03" box="[1124,1203,1423,1449]" pageId="17" pageNumber="18">SEHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B09D75DEB37574BFB3FD887FD6DFA67" ID-GBIF-Occurrence="2636268352" collectingDate="1992-10-15" collectingDateMax="1992-10-16" collectingDateMin="1992-10-15" collectionCode="SEHU" collectorName="M. J. Toda" country="Taiwan" pageId="17" pageNumber="18" specimenCount="3" specimenCount-male="3">
<specimenCount id="9D671689EB37574BFB3FD887FB02FA03" box="[1222,1257,1423,1449]" pageId="17" pageNumber="18" type="male">3♂</specimenCount>
(B-type), Shan- linxi,
<date id="FFDFFBC0EB37574BFF21D8BAFE9EFA67" box="[216,373,1458,1485]" pageId="17" pageNumber="18" value="1992-10-15" valueMax="1992-10-16" valueMin="1992-10-15">
<collectingDate id="EF9B0228EB37574BFF21D8BAFE9EFA67" box="[216,373,1458,1485]" pageId="17" pageNumber="18" value="1992-10-15" valueMax="1992-10-16" valueMin="1992-10-15">1516.x.1992</collectingDate>
</date>
,
<collectorName id="2694B8D6EB37574BFE79D8BBFE1AFA67" box="[384,497,1459,1485]" pageId="17" pageNumber="18">M.J. Toda</collectorName>
leg. (
<collectionCode id="ED7045C5EB37574BFDD6D8BBFD95FA67" box="[559,638,1459,1485]" pageId="17" pageNumber="18">SEHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B09D75DEB37574BFD6BD8BBFB1AFA67" ID-GBIF-Occurrence="2636268343" box="[658,1265,1459,1485]" collectingDate="1992-10-08" collectionCode="SEHU" collectorName="M. J. Toda" country="Taiwan" location="Wulai" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D671689EB37574BFD6BD8BBFD5EFA67" box="[658,693,1459,1485]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
(A-type),
<location id="8EBE8BDBEB37574BFCD3D8BBFC9BFA67" LSID="urn:lsid:plazi:treatment:03C86C16EB37574BFF6EDFFAFAB1F92B:8EBE8BDBEB37574BFCD3D8BBFC9BFA67" box="[810,880,1459,1485]" country="Taiwan" name="Wulai" pageId="17" pageNumber="18">Wulai</location>
,
<date id="FFDFFBC0EB37574BFC82D8BBFC34FA67" box="[891,991,1459,1485]" pageId="17" pageNumber="18" value="1992-10-08">
<collectingDate id="EF9B0228EB37574BFC82D8BBFC34FA67" box="[891,991,1459,1485]" pageId="17" pageNumber="18" value="1992-10-08">8.x.1992</collectingDate>
</date>
,
<collectorName id="2694B8D6EB37574BFC12D8BBFBB0FA67" box="[1003,1115,1459,1485]" pageId="17" pageNumber="18">M.J. Toda</collectorName>
leg. (
<collectionCode id="ED7045C5EB37574BFB63D8BBFB02FA67" box="[1178,1257,1459,1485]" pageId="17" pageNumber="18">SEHU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B09D75DEB37574BFB02D8BBFC53FA5B" ID-GBIF-Occurrence="2636268358" collectingDate="1987-03-04" collectingDateMax="1987-03-10" collectingDateMin="1987-03-04" collectionCode="T, SEHU" collectorName="X. Peng" country="China" location="Dinghushan" municipality="Btype" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong">
<collectingCountry id="F3769D90EB37574BFB02D8BBFAA9FA67" box="[1275,1346,1459,1485]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
:
<specimenCount id="9D671689EB37574BFAB4D8BBFA9BFA67" box="[1357,1392,1459,1485]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
(
<collectingMunicipality id="6BBA477AEB37574BFA86D8BBFF20FA5B" pageId="17" pageNumber="18">Btype</collectingMunicipality>
),
<location id="8EBE8BDBEB37574BFF26D8DFFE82FA5B" LSID="urn:lsid:plazi:treatment:03C86C16EB37574BFF6EDFFAFAB1F92B:8EBE8BDBEB37574BFF26D8DFFE82FA5B" box="[223,361,1495,1521]" country="China" municipality="Btype" name="Dinghushan" pageId="17" pageNumber="18" stateProvince="Guangdong">Dinghushan</location>
,
<collectingRegion id="49A513E2EB37574BFE8DD8DFFE11FA5B" box="[372,506,1495,1521]" country="China" name="Guangdong" pageId="17" pageNumber="18">Guangdong</collectingRegion>
,
<date id="FFDFFBC0EB37574BFDFCD8DFFD76FA5B" box="[517,669,1495,1521]" pageId="17" pageNumber="18" value="1987-03-04" valueMax="1987-03-10" valueMin="1987-03-04">
<collectingDate id="EF9B0228EB37574BFDFCD8DFFD76FA5B" box="[517,669,1495,1521]" pageId="17" pageNumber="18" value="1987-03-04" valueMax="1987-03-10" valueMin="1987-03-04">410.iii.1987</collectingDate>
</date>
,
<collectionCode id="ED7045C5EB37574BFD51D8DFFD50FA5B" box="[680,699,1495,1521]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="18">T</collectionCode>
.-
<collectorName id="2694B8D6EB37574BFD3ED8DFFCCAFA5B" box="[711,801,1495,1521]" pageId="17" pageNumber="18">X. Peng</collectorName>
leg. (
<collectionCode id="ED7045C5EB37574BFC99D8DFFC44FA5B" box="[864,943,1495,1521]" pageId="17" pageNumber="18">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB37574BFF3ED8F2FCF4F9BF" box="[199,799,1530,1558]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BDEDD00EB37574BFF3ED8F2FCF4F9BF" blockId="17.[151,1437,954,1666]" box="[199,799,1530,1558]" pageId="17" pageNumber="18">
<emphasis id="B9150112EB37574BFF3ED8F2FE8AF9BF" bold="true" box="[199,353,1530,1557]" pageId="17" pageNumber="18">Distribution.</emphasis>
<collectingCountry id="F3769D90EB37574BFE9ED8F3FE57F9BF" box="[359,444,1531,1557]" name="Taiwan" pageId="17" pageNumber="18">Taiwan</collectingCountry>
,
<collectingCountry id="F3769D90EB37574BFE3ED8F3FDE1F9BF" box="[455,522,1531,1557]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
(
<collectingRegion id="49A513E2EB37574BFDE0D8F3FD49F9BC" box="[537,674,1531,1558]" country="China" name="Guangdong" pageId="17" pageNumber="18">Guangdong</collectingRegion>
),
<collectingCountry id="F3769D90EB37574BFD4DD8F3FCF0F9BF" box="[692,795,1531,1557]" name="Thailand" pageId="17" pageNumber="18">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB37574BFF3EDB17FAB1F92B" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BDEDD00EB37574BFF3EDB17FAB1F92B" blockId="17.[151,1437,954,1666]" pageId="17" pageNumber="18">
<emphasis id="B9150112EB37574BFF3EDB17FED0F993" bold="true" box="[199,315,1567,1593]" pageId="17" pageNumber="18">Remarks.</emphasis>
This species and the subsequent three species form a cluster (BP = 88%) with three synapomorphies (
<figureCitation id="135AC185EB37574BFF59DB4BFF08F9F7" box="[160,227,1603,1629]" captionStart="FIGURE 1" captionStartId="7.[151,250,1683,1708]" captionTargetBox="[151,1434,779,1658]" captionTargetId="figure@7.[151,1436,777,1658]" captionTargetPageId="7" captionText="FIGURE 1. The strict consensus tree resulting from the parsimony analysis I (PAUP* v4.0a166) of the data matrix of 40 spp. × 66 morphological characters (Appendix 1) for the genus Phortica (especially focusing on the subgenus Ashima). Synapomorphies (solid circle: nonhomoplastic; open circle: homoplastic) inferred from both ACCTRAN and DELTRAN character optimization are indicated on each internal branch along with support values (bootstrap frequency %)." figureDoi="http://doi.org/10.5281/zenodo.3884614" httpUri="https://zenodo.org/record/3884614/files/figure.png" pageId="17" pageNumber="18">Fig. 1</figureCitation>
): abdominal sternite III longer than wide (ch.22-0), not wider than IV (ch.23-0), without dense setulae on medial surface (ch.26-0). However, this species can be distinguished from them by the diagnostic characters.
</paragraph>
</subSubSection>
</treatment>
</document>