treatments-xml/data/03/C8/6C/03C86C16EB375749FF6EDBDAFD7CFF7D.xml

<document id="573370C868602B198CD0FD75DF66CCA2" ID-CLB-Dataset="21673" ID-DOI="10.11646/zootaxa.4789.1.1" ID-GBIF-Dataset="c12aaed8-264c-459c-b1aa-09eaeb686670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884610" ID-ZooBank="311A27A2-A053-4D20-A43E-EF57DBE55166" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591601966137" checkinUser="plazi" docAuthor="Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit &amp; Katoh, Toru" docDate="2020" docId="03C86C16EB375749FF6EDBDAFD7CFF7D" docLanguage="en" docName="zootaxa.4789.1.1.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phortica (Ashima) andreagigoni Toda &amp; Banziger 2020, sp. nov." docType="treatment" docVersion="10" lastPageNumber="20" masterDocId="FFF1146EEB26575AFFF9DD08FFEBFFAA" masterDocTitle="Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="18" updateTime="1698842174822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C2F86C2F5CCEF7D151D613F8F2DB5DCC">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="605B90FCE1B0E1E95240ACE230628E31">Toda, Masanori J.</mods:namePart>
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<mods:namePart id="EFBE6E38BE24E44B834FE8CBC0201EE6">Bänziger, Hans</mods:namePart>
<mods:affiliation id="F2367B7E8D989730C00A94E6782A8A83">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="88943000FCA2598FA4ADAC1BB409ECA1">Sati, Pradeep C.</mods:namePart>
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<mods:affiliation id="ABD9473B4836FE281C93714E417F5AF3">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:namePart id="1B6ABE0E3A4053CD30C612C394A08FEF">Suwito, Awit</mods:namePart>
<mods:affiliation id="AD69F2447F7CC8E9563D305FD0F9846B">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<treatment id="03C86C16EB375749FF6EDBDAFD7CFF7D" ID-DOI="http://doi.org/10.5281/zenodo.5221290" ID-GBIF-Taxon="164612784" ID-Zenodo-Dep="5221290" LSID="urn:lsid:plazi:treatment:03C86C16EB375749FF6EDBDAFD7CFF7D" httpUri="http://treatment.plazi.org/id/03C86C16EB375749FF6EDBDAFD7CFF7D" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<heading id="D0966A6CEB37574BFF6EDBDAFC7EF947" bold="true" box="[151,917,1746,1773]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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13) 
<taxonomicName id="4C61A683EB37574BFF3FDBDAFCC0F947" authority="Toda &amp; Banziger" authorityName="Toda &amp; Banziger" authorityYear="2020" box="[198,811,1746,1773]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="andreagigoni" status="sp. nov." subGenus="Ashima">
<emphasis id="B9150112EB37574BFF3FDBDAFEC5F947" bold="true" box="[198,302,1746,1773]" italics="true" pageId="17" pageNumber="18">Phortica</emphasis>
(
<emphasis id="B9150112EB37574BFEC6DBDAFE77F947" bold="true" box="[319,412,1746,1773]" italics="true" pageId="17" pageNumber="18">Ashima</emphasis>
) 
<emphasis id="B9150112EB37574BFE54DBDAFDA5F947" bold="true" box="[429,590,1746,1773]" italics="true" pageId="17" pageNumber="18">andreagigoni</emphasis>
Toda &amp; Bänziger
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, 
<taxonomicNameLabel id="A226BC69EB37574BFCCEDBDBFC7EF947" box="[823,917,1747,1773]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C37B8E8BEB37574BFF6EDBFFFF00F8BB" box="[151,235,1783,1809]" pageId="17" pageNumber="18" type="description">
<paragraph id="8BDEDD00EB37574BFF6EDBFFFF00F8BB" blockId="17.[151,917,1746,1809]" box="[151,235,1783,1809]" pageId="17" pageNumber="18">
(
<figureCitation id="135AC185EB37574BFF59DBFFFF08F8BB" box="[160,227,1783,1809]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB37574BFF6EDA36FB21F86F" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BDEDD00EB37574BFF6EDA36FB21F86F" blockId="17.[151,1437,1854,2025]" pageId="17" pageNumber="18">
<emphasis id="B9150112EB37574BFF6EDA36FEF9F8F3" bold="true" box="[151,274,1854,1881]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
Some interfrontal setulae longer than lowest setulae on ocellar triangle (ch.3-2); arista without apical expansion (ch.7-0; 
<figureCitation id="135AC185EB37574BFE8BDA6BFE26F8D7" box="[370,461,1891,1917]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="17" pageNumber="18">Fig. 4A</figureCitation>
); pregonite strongly curved ventrad distally (
<figureCitation id="135AC185EB37574BFC34DA6BFBC3F8D7" box="[973,1064,1891,1917]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="17" pageNumber="18">Fig. 4C</figureCitation>
); posterior postgonites on lateral lobes of aedeagal sheath medially fused to each other (ch.66-1; 
<figureCitation id="135AC185EB37574BFC8FDA8FFC39F80B" box="[886,978,1927,1953]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="17" pageNumber="18">Fig. 4D</figureCitation>
); left and right lateral lobes of aedeagal sheath with 1 and 2 anterior, somewhat triangular, plate-like postgonites, respectively (
<figureCitation id="135AC185EB37574BFB9ADAA3FB55F86F" box="[1123,1214,1963,1989]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="17" pageNumber="18">Fig. 4D</figureCitation>
).
</paragraph>
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<subSubSection id="C37B8E8BEB375748FF3EDAC6FDB4FE21" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8BDEDD00EB375748FF3EDAC6FC80FF51" blockId="17.[151,1437,1854,2025]" lastBlockId="18.[151,1437,153,503]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B9150112EB37574BFF3EDAC6FEBAF843" bold="true" box="[199,337,1998,2025]" pageId="17" pageNumber="18">Description</emphasis>
(not repeating characters common to 
<taxonomicName id="4C61A683EB37574BFCFDDAD8FC94F843" authorityName=": Chen &amp; Maca" authorityYear="2012" box="[772,895,1999,2025]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="foliiseta">
<emphasis id="B9150112EB37574BFCFDDAD8FCFEF843" box="[772,789,2000,2025]" italics="true" pageId="17" pageNumber="18">P</emphasis>
. 
<emphasis id="B9150112EB37574BFCDCDAC7FC94F843" box="[805,895,1999,2025]" italics="true" pageId="17" pageNumber="18">foliiseta</emphasis>
</taxonomicName>
). Supracervical setae 14–15. Dorsomedial, tentorial apodeme 2/3 as long as basal, parallel portion of dorsolateral, tentorial apodeme. Longest, dorsal branch of arista shorter than longest seta on pedicel (
<figureCitation id="135AC185EB345748FD85DDB5FD33FF7D" box="[636,728,189,215]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="18" pageNumber="19">Fig. 4A</figureCitation>
). Fore and hind tarsomere Is slightly longer than 3 succeeding tarsomeres together; mid one as long as rest tarsomeres together.
</paragraph>
<paragraph id="8BDEDD00EB345748FF3EDC0DFB8CFEB5" blockId="18.[151,1437,153,503]" box="[199,1127,260,287]" pageId="18" pageNumber="19">
Measurements (
<typeStatus id="54DA63A2EB345748FE80DC0DFE34FEB5" box="[377,479,261,287]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
, in mm): BL = 3.69, ThL = 1.51, WL = 2.46, WW = 1.10.
</paragraph>
<paragraph id="8BDEDD00EB345748FF3EDC21FDB4FE21" blockId="18.[151,1437,153,503]" pageId="18" pageNumber="19">
Indices (
<typeStatus id="54DA63A2EB345748FED0DC21FE64FEE9" box="[297,399,297,323]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
, in ratio): FW/HW = 0.46, ch/o = 0.07, prorb = 1.26, rcorb = 0.53, vb = 0.29, dcl = 0.55, presctl = 0.46, sctl = (n/a), sterno = 1.00, orbito = 2.37, dcp = 0.33, sctlp = 1.32, C = 2.01, 4c = 1.69, 4v = 2.99, 5x = 0.97, ac = 3.45, M = 0.64, C3F = 0.58.
</paragraph>
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<paragraph id="8BDEDD00EB345748FF3EDC9DFED9FE79" blockId="18.[151,1437,153,503]" pageId="18" pageNumber="19">
<materialsCitation id="3B09D75DEB345748FF3EDC9DFED9FE79" ID-GBIF-Occurrence="2636268378" collectingDate="2009-07-05" collectionCode="SEHU" collectorName="H. Banziger" country="Thailand" elevation="800" location="Chang Khian Stream" municipality="Doi Suthep" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="holotype">
<emphasis id="B9150112EB345748FF3EDC9DFED0FE05" bold="true" box="[199,315,405,431]" pageId="18" pageNumber="19">
<typeStatus id="54DA63A2EB345748FF3EDC9DFEDDFE05" box="[199,310,405,431]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D671689EB345748FEB8DC9DFEBDFE05" box="[321,342,405,431]" pageId="18" pageNumber="19" type="male">♂</specimenCount>
(
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), “ 
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gully, 
<collectingMunicipality id="6BBA477AEB345748FCF1DC9DFC62FE05" box="[776,905,405,431]" pageId="18" pageNumber="19">Doi Suthep</collectingMunicipality>
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, 
<collectingCountry id="F3769D90EB345748FBDCDC9DFB60FE05" box="[1061,1163,405,431]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
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a.s.l.
</elevation>
, 
<date id="FFDFFBC0EB345748FADDDC9CFA73FE05" box="[1316,1432,404,431]" pageId="18" pageNumber="19" value="2009-07-05">
<collectingDate id="EF9B0228EB345748FADDDC9CFA73FE05" box="[1316,1432,404,431]" pageId="18" pageNumber="19" value="2009-07-05">5.vii.2009</collectingDate>
</date>
, 
<collectorName id="2694B8D6EB345748FF6EDCB1FECFFE79" box="[151,292,441,467]" pageId="18" pageNumber="19">H. Bänziger</collectorName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB345748FF3EDCD4FE39FE5D" box="[199,466,476,503]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BDEDD00EB345748FF3EDCD4FE39FE5D" blockId="18.[151,1437,153,503]" box="[199,466,476,503]" pageId="18" pageNumber="19">
<emphasis id="B9150112EB345748FF3EDCD4FE8AFE5D" bold="true" box="[199,353,476,503]" pageId="18" pageNumber="19">Distribution.</emphasis>
<collectingCountry id="F3769D90EB345748FE9EDCD5FE25FE5D" box="[359,462,477,503]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF1E8D88EB345748FF6EDA04FD31F8C6" ID-DOI="http://doi.org/10.5281/zenodo.3884622" ID-Zenodo-Dep="3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1804,1829]" targetBox="[151,1436,525,1774]" targetPageId="18">
<paragraph id="8BDEDD00EB345748FF6EDA04FD31F8C6" blockId="18.[151,1436,1804,1901]" pageId="18" pageNumber="19">
<emphasis id="B9150112EB345748FF6EDA04FEF8F88F" bold="true" box="[151,275,1804,1829]" pageId="18" pageNumber="19">FIGURE 4.</emphasis>
<taxonomicName id="4C61A683EB345748FEE3DA05FCF0F88F" authority="Toda &amp; Banziger" authorityName="Toda &amp; Banziger" authorityYear="2020" box="[282,795,1804,1829]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="andreagigoni" status="sp. nov." subGenus="Ashima">
<emphasis id="B9150112EB345748FEE3DA05FE98F88F" box="[282,371,1805,1829]" italics="true" pageId="18" pageNumber="19">Phortica</emphasis>
(
<emphasis id="B9150112EB345748FE7BDA05FE25F88F" box="[386,462,1805,1829]" italics="true" pageId="18" pageNumber="19">Ashima</emphasis>
) 
<emphasis id="B9150112EB345748FE24DA05FD8EF88F" box="[477,613,1805,1829]" italics="true" pageId="18" pageNumber="19">andreagigoni</emphasis>
Toda &amp; Bänziger
</taxonomicName>
, 
<emphasis id="B9150112EB345748FCDCDA05FC9DF88F" bold="true" box="[805,886,1805,1829]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A226BC69EB345748FCDCDA05FC9DF88F" box="[805,886,1805,1829]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C37B8E8BEB345748FF3EDA93FDC0F857" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BDEDD00EB345748FF3EDA93FDC0F857" blockId="18.[151,1436,1947,2045]" pageId="18" pageNumber="19">
<emphasis id="B9150112EB345748FF3EDA93FED0F81F" bold="true" box="[199,315,1947,1973]" pageId="18" pageNumber="19">Remarks.</emphasis>
This species resembles 
<taxonomicName id="4C61A683EB345748FDB3DA94FD28F81F" authorityName=": Chen &amp; Maca" authorityYear="2012" box="[586,707,1947,1974]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="foliiseta">
<emphasis id="B9150112EB345748FDB3DA94FDB0F81F" box="[586,603,1948,1973]" italics="true" pageId="18" pageNumber="19">P</emphasis>
. 
<emphasis id="B9150112EB345748FD90DA93FD28F81F" box="[617,707,1947,1973]" italics="true" pageId="18" pageNumber="19">foliiseta</emphasis>
</taxonomicName>
within the cluster of the four species, in having the dorsomedial, tentorial apodeme ≥ 2/3 basal, parallel portion of dorsolateral, tentorial apodeme (ch.1-0), but can be distinguished from it by the diagnostic characters.
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB355749FF3EDD91FD7CFF7D" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8BDEDD00EB355749FF3EDD91FD7CFF7D" blockId="19.[151,1436,153,215]" pageId="19" pageNumber="20">
<emphasis id="B9150112EB355749FF3EDD91FEA5FF19" bold="true" box="[199,334,153,179]" pageId="19" pageNumber="20">Etymology.</emphasis>
Composite patronym with which H.B. wants to honor Prof. Dr.Andrea Gigon for his friendship and creation of the Blue Lists (
<bibRefCitation id="EFF0A0F1EB355749FE38DDB5FD67FF7D" author="Gigon" box="[449,652,189,215]" etAl="et al." firstAuthor="Gigon" pageId="19" pageNumber="20" pagination="402 - 413" refId="ref34778" refString="Gigon, A., Langenauer, R., Meier, C. &amp; Nievergelt, B. (2000) Blue Lists of threatened species with stabilized or increasing abundance: A new instrument for conservation. Conservation Biology, 14, 402 - 413. https: // doi. org / 10.1046 / j. 1523 - 1739.2000.98443. x" type="journal article" year="2000">
Gigon 
<emphasis id="B9150112EB355749FDE9DDB6FDAAFF7D" box="[528,577,189,215]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 2000
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>