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<document id="680948D5B3505E68A6DA47F315C1FAE6" ID-DOI="10.11646/phytotaxa.400.4.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13716305" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725649528446" checkinUser="felipe" docAuthor="Calvo, Joel &amp; Romoleroux, Katya" docDate="2019" docId="03C82F5EFFAAB570D2F4FF46FDD104AE" docLanguage="en" docName="phytotaxa.400.4.4.pdf" docOrigin="Phytotaxa 400 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.400.4.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Monticalia speciosa J. Calvo &amp; Romol. 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="247" masterDocId="FFF15726FFABB571D27CFFB2FFEF0305" masterDocTitle="Monticalia speciosa (Senecioneae, Compositae), a noteworthy new species from Ecuador" masterLastPageNumber="250" masterPageNumber="246" pageNumber="247" updateTime="1725655066763" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="E48A36B810E4DF022B6796870ED1AF7E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EDA66373FC08AF957A7F4D38EDB21D4B">
<mods:title id="1AAB3D757C6D7952C378E67FBD122C9A">Monticalia speciosa (Senecioneae, Compositae), a noteworthy new species from Ecuador</mods:title>
</mods:titleInfo>
<mods:name id="3639DABF8C8C4375827D8A9DF003E799" type="personal">
<mods:role id="3A823F71601FA279DD17527700B1F746">
<mods:roleTerm id="8CA72015961C7B51CAF18FC8D3EF9B1C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9F90825BBD0C5D467A9B2DFCE30CF3F">Calvo, Joel</mods:namePart>
<mods:affiliation id="500786C1E6DA34C6C4E1736AAAEB4427">Instituto de Geografía, Facultad de Ciencias del Mar y Geografía, Pontificia Universidad Católica de Valparaíso, Avenida Brasil 2241, 2362807 Valparaíso, Chile.</mods:affiliation>
</mods:name>
<mods:name id="86E2A49A542B4D2495E256FC93A16228" type="personal">
<mods:role id="DAF16F75EA4641A3187F8D55A7EDD252">
<mods:roleTerm id="8237D8E8356CDA84B1165FA12455ED7B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1EF67F14F473CD34C9CC1D2099C90173">Romoleroux, Katya</mods:namePart>
<mods:affiliation id="50013790955D2A50A7708504341093BC">Herbario QCA, Escuela de Ciencias Biológicas, Pontificia Universidad Católica del Ecuador, Av. 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="45355DCF110CF408109F2A65E181A842">text</mods:typeOfResource>
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<mods:titleInfo id="E517BB9DCF12132B851A2B90EDA541B7">
<mods:title id="BA3CE6B166FCD10FD002343D59DA01CD">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="A23396FBC26493CC8170404483B1C61C">
<mods:date id="721513B9EF97E896232E36E25BD6374B">2019</mods:date>
<mods:detail id="424D030CB409385EDA250C51607128CE" type="pubDate">
<mods:number id="B37A0FB093906F2C5C141113A7B9C3DD">2019-04-08</mods:number>
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<mods:number id="08A2E106F28A7584C97C78F8E0C23F0A">400</mods:number>
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<mods:number id="07AC06F3EA34D10DC1027FAAABB7EFB7">4</mods:number>
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<mods:start id="2F002997CF22A18B658E309192BAD3CA">246</mods:start>
<mods:end id="351FA0E5439F88728786335E41595FBD">250</mods:end>
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<mods:location id="DE5453ECCCEBA69F30A603D5C09F02F3">
<mods:url id="1FF9329D33BBE1944FBE5248F0A15E85">http://dx.doi.org/10.11646/phytotaxa.400.4.4</mods:url>
</mods:location>
<mods:classification id="DD56D493C6854A45BFAB10E57CC5A1AF">journal article</mods:classification>
<mods:identifier id="8CE70686657041F444691242E2AC4807" type="DOI">10.11646/phytotaxa.400.4.4</mods:identifier>
<mods:identifier id="20FE22704FEE8543A5FA2E129072D981" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="7738BCBE78596063A2B8820C91A7F3B9" type="Zenodo-Dep">13716305</mods:identifier>
</mods:mods>
<treatment id="03C82F5EFFAAB570D2F4FF46FDD104AE" LSID="urn:lsid:plazi:treatment:03C82F5EFFAAB570D2F4FF46FDD104AE" httpUri="http://treatment.plazi.org/id/03C82F5EFFAAB570D2F4FF46FDD104AE" lastPageNumber="247" pageId="1" pageNumber="247">
<subSubSection id="C37BCDC3FFAAB570D2F4FF46FCCB020A" box="[136,804,244,271]" pageId="1" pageNumber="247" type="nomenclature">
<paragraph id="8BDE9E48FFAAB570D2F4FF46FCCB020A" blockId="1.[136,804,244,271]" box="[136,804,244,271]" pageId="1" pageNumber="247">
<heading id="D0962924FFAAB570D2F4FF46FCCB020A" box="[136,804,244,271]" fontSize="11" level="2" pageId="1" pageNumber="247" reason="3">
<taxonomicName id="4C61E5CBFFAAB570D2F4FF46FDA4020A" ID-CoL="7XCS5" authority="J. Calvo &amp; Romol." authorityName="J. Calvo &amp; Romol." authorityYear="2019" box="[136,587,244,271]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa" status="sp. nov.">
<emphasis id="B915425AFFAAB570D2F4FF46FE82020B" bold="true" box="[136,365,244,271]" italics="true" pageId="1" pageNumber="247">Monticalia speciosa</emphasis>
J. Calvo &amp; Romol.
</taxonomicName>
,
<emphasis id="B915425AFFAAB570D029FF46FD43020B" bold="true" box="[597,684,244,270]" italics="true" pageId="1" pageNumber="247">
<taxonomicNameLabel id="A226FF21FFAAB570D029FF46FD43020B" box="[597,684,244,270]" pageId="1" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135A82CDFFAAB570D0C7FF44FD11020A" box="[699,766,246,271]" captionStart="FIGURE 1" captionStartId="2.[136,229,1954,1975]" captionTargetBox="[140,1446,187,1925]" captionTargetId="figure-14@2.[140,1446,187,1925]" captionTargetPageId="2" captionText="FIGURE 1. Monticalia speciosa. A. Habitat. B. Stems in bloom. C. Detail of the capitula. D. Detail of the leaves. Photographs by J. Calvo at the locus classicus." figureDoi="http://doi.org/10.5281/zenodo.13716307" httpUri="https://zenodo.org/record/13716307/files/figure.png" pageId="1" pageNumber="247">Fig. 1</figureCitation>
,
<figureCitation id="135A82CDFFAAB570D170FF44FCF4020A" box="[780,795,246,271]" captionStart="FIGURE 2" captionStartId="3.[136,229,1957,1978]" captionTargetBox="[151,1435,190,1932]" captionTargetId="figure-16@3.[151,1436,190,1932]" captionTargetPageId="3" captionText="FIGURE 2. Monticalia speciosa (plant details on the left) compared to M. peruviana (plant details on the right). A. Ray florets. B. Involucres. C. Leaves. D. Synflorescences. Photographs by J. Calvo at the locus classicus." figureDoi="http://doi.org/10.5281/zenodo.13716309" httpUri="https://zenodo.org/record/13716309/files/figure.png" pageId="1" pageNumber="247">2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37BCDC3FFAAB570D2F4FEAEFC4A00DA" pageId="1" pageNumber="247" type="materials_examined">
<paragraph id="8BDE9E48FFAAB570D2F4FEAEFC180250" blockId="1.[136,1452,284,413]" pageId="1" pageNumber="247">
<emphasis id="B915425AFFAAB570D2F4FEAEFF2D0234" bold="true" box="[136,194,284,305]" pageId="1" pageNumber="247">Type:</emphasis>
—
<materialsCitation id="3B099415FFAAB570D2A7FEAEFC1C0250" collectingDate="2018-10-18" collectionCode="QCA" collectorName="de Anteojos &amp; K. Romoleroux &amp; J. Calvo" country="Ecuador" elevation="3935" latitude="-0.31166667" location="Papallacta" longLatPrecision="21" longitude="-78.165" pageId="1" pageNumber="247" specimenCode="QCA 241524" specimenCount="2" stateProvince="Napo" typeStatus="holotype">
<collectingCountry id="F376DED8FFAAB570D2A7FEAEFEBD0234" box="[219,338,284,305]" name="Ecuador" pageId="1" pageNumber="247">ECUADOR</collectingCountry>
.
<collectingRegion id="49A550AAFFAAB570D325FEAEFE7E0234" box="[345,401,284,305]" country="Ecuador" name="Napo" pageId="1" pageNumber="247">Napo</collectingRegion>
:
<location id="8EBEC893FFAAB570D3E4FEAEFE120234" LSID="urn:lsid:plazi:treatment:03C82F5EFFAAB570D2F4FF46FDD104AE:8EBEC893FFAAB570D3E4FEAEFE120234" box="[408,509,284,305]" country="Ecuador" latitude="-0.31166667" longLatPrecision="21" longitude="-78.165" name="Papallacta" pageId="1" pageNumber="247" stateProvince="Napo">Papallacta</location>
, sendero “Agua y Vida”, lagunas
<collectorName id="2694FB9EFFAAB570D145FEAEFC430234" box="[825,940,284,305]" pageId="1" pageNumber="247">de Anteojos</collectorName>
,
<geoCoordinate id="EE55F88FFFAAB570D1C8FEAEFBF40234" box="[948,1051,284,305]" degrees="0" direction="south" minutes="18" orientation="latitude" pageId="1" pageNumber="247" precision="15" seconds="42" value="-0.31166667">0º18’42’’S</geoCoordinate>
<geoCoordinate id="EE55F88FFFAAB570D65CFEAEFB720234" box="[1056,1181,284,305]" degrees="78" direction="west" minutes="09" orientation="longitude" pageId="1" pageNumber="247" precision="15" seconds="54" value="-78.165">78º09’54’’W</geoCoordinate>
,
<quantity id="4C9933ADFFAAB570D6D9FEAEFB010234" box="[1189,1262,284,305]" metricMagnitude="3" metricUnit="m" metricValue="3.935" pageId="1" pageNumber="247" unit="m" value="3935.0">
<elevation id="004C797BFFAAB570D6D9FEAEFB010234" box="[1189,1262,284,305]" metricMagnitude="3" metricUnit="m" metricValue="3.935" pageId="1" pageNumber="247" unit="m" value="3935.0">3935 m</elevation>
</quantity>
, páramo de pajonal,
<date id="FFDFB888FFAAB570D2C1FEF2FED90250" box="[189,310,320,341]" pageId="1" pageNumber="247" value="2018-10-18">
<collectingDate id="EF9B4160FFAAB570D2C1FEF2FED90250" box="[189,310,320,341]" pageId="1" pageNumber="247" value="2018-10-18">18 Oct 2018</collectingDate>
</date>
,
<emphasis id="B915425AFFAAB570D33CFEF2FD9A0250" box="[320,629,320,341]" italics="true" pageId="1" pageNumber="247">
<collectorName id="2694FB9EFFAAB570D33CFEF2FE3F0250" box="[320,464,320,341]" pageId="1" pageNumber="247">K. Romoleroux</collectorName>
&amp;
<collectorName id="2694FB9EFFAAB570D393FEF2FDD00250" box="[495,575,320,341]" pageId="1" pageNumber="247">J. Calvo</collectorName>
6168
</emphasis>
(
<typeStatus id="54DA20EAFFAAB570D0FEFEF2FD340250" box="[642,731,320,341]" pageId="1" pageNumber="247" type="holotype">holotype</typeStatus>
:
<specimenCode id="DBC73633FFAAB570D098FEF2FC890250" box="[740,870,320,341]" collectionCode="QCA" country="Ecuador" lsid="urn:lsid:biocol.org:col:15650" name="Pontificia Universidad Católica del Ecuador" pageId="1" pageNumber="247" type="Herbarium">QCA 241524</specimenCode>
;
<typeStatus id="54DA20EAFFAAB570D10CFEF2FC560250" box="[880,953,320,341]" pageId="1" pageNumber="247" type="isotype">isotype</typeStatus>
: MO)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDE9E48FFAAB570D2F4FED6FCA70298" blockId="1.[136,1452,284,413]" pageId="1" pageNumber="247">
It differs from the other radiate species of
<taxonomicName id="4C61E5CBFFAAB570D05CFED6FD66027C" authorityName="Jeffrey" authorityYear="1992" box="[544,649,356,377]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="genus">
<emphasis id="B915425AFFAAB570D05CFED6FD66027C" box="[544,649,356,377]" italics="true" pageId="1" pageNumber="247">Monticalia</emphasis>
</taxonomicName>
by its minute leaves, which are sessile to subsessile, elliptic, ciliate on the margins and midrib, and concolorous, and by having eight involucral bracts.
</paragraph>
<paragraph id="8BDE9E48FFAAB570D2F4FE7CFC4A00DA" blockId="1.[136,1452,462,1963]" pageId="1" pageNumber="247">
Shrub
<quantity id="4C9933ADFFAAB570D2A9FE7CFEDC02E2" box="[213,307,462,487]" metricMagnitude="0" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="1" pageNumber="247" unit="m" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 m</quantity>
tall; stem to
<quantity id="4C9933ADFFAAB570D3ACFE7CFDF802E2" box="[464,535,462,487]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="247" unit="cm" value="15.0">15 cm</quantity>
in diam. at the base; young stems slightly corrugated, covered by multicellular trichomes
<quantity id="4C9933ADFFAAB570D37EFE40FE7F010E" box="[258,400,498,523]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.9" metricValueMin="0.9" pageId="1" pageNumber="247" unit="mm" value="1.4" valueMax="1.9" valueMin="0.9">0.9–1.9 mm</quantity>
. Leaves alternate, with short internodes
<quantity id="4C9933ADFFAAB570D117FE40FC17010E" box="[875,1016,498,523]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="1" pageNumber="247" unit="mm" value="1.15" valueMax="1.5" valueMin="0.8">0.8–1.5 mm</quantity>
, sessile to subsessile; lamina elliptic,
<quantity id="4C9933ADFFAAB570D2F4FDA4FEFE012A" box="[136,273,534,559]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="4.1" metricValueMin="3.1" pageId="1" pageNumber="247" unit="mm" value="3.5999999999999996" valueMax="4.1" valueMin="3.1">3.1–4.1 mm</quantity>
long,
<quantity id="4C9933ADFFAAB570D320FDA4FE0A012A" box="[348,485,534,559]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.9" metricValueMin="1.2" pageId="1" pageNumber="247" unit="mm" value="1.5499999999999998" valueMax="1.9" valueMin="1.2">1.2–1.9 mm</quantity>
wide, rather obtuse at the apex with a callous-like acumen, plane, somewhat fleshy, ciliate on the margins and midrib (cilia ca.
<quantity id="4C9933ADFFAAB570D013FD88FD290156" box="[623,710,570,595]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="247" unit="mm" value="0.4">0.4 mm</quantity>
long), concolorous, light green, outspreading, somewhat reflexed with age, with a gibbous-base ca.
<quantity id="4C9933ADFFAAB570D06FFDECFD830172" box="[531,620,606,631]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="247" unit="mm" value="0.3">0.3 mm</quantity>
long at attachment point with the stem. Synflorescence corymbiform or subumbelliform, with bracts similar to the upper cauline leaves that decrease in size upwards and become linear-oblong. Capitula heterogamous, radiate, with pedicels covered by multicellular trichomes; involucre obconical,
<quantity id="4C9933ADFFAAB570D711FD14FF5B01E6" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="1" pageNumber="247" unit="mm" value="8.25" valueMax="8.5" valueMin="8.0">8–8.5 mm</quantity>
long,
<quantity id="4C9933ADFFAAB570D281FD78FEB401E6" box="[253,347,714,739]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="247" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
wide at the apex. Involucral bracts 8, linear-oblong,
<quantity id="4C9933ADFFAAB570D1C8FD78FBD301E6" box="[948,1084,714,739]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.3" metricValueMin="7.2" pageId="1" pageNumber="247" unit="mm" value="7.25" valueMax="7.3" valueMin="7.2">7.2–7.3 mm</quantity>
long,
<quantity id="4C9933ADFFAAB570D6F8FD78FAE301E6" box="[1156,1292,714,739]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="1" pageNumber="247" unit="mm" value="1.2" valueMax="1.5" valueMin="0.9">0.9–1.5 mm</quantity>
wide, smooth or 1-keeled, glabrescent, usually with the apex dark-purplish. Supplementary bracts 4–6, linear-subulate,
<quantity id="4C9933ADFFAAB570D759FD5CFA430002" box="[1317,1452,750,775]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="5.4" metricValueMin="3.4" pageId="1" pageNumber="247" unit="mm" value="4.4" valueMax="5.4" valueMin="3.4">3.4–5.4 mm</quantity>
long,
<quantity id="4C9933ADFFAAB570D2B5FCA0FEBF002E" box="[201,336,786,811]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="1" pageNumber="247" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
wide, smooth, a half to two-thirds as long as the involucral bracts, with scattered trichomes on the margins and the apex usually dark-purplish. Ray florets 8, female,
<quantity id="4C9933ADFFAAB570D100FC84FC12004A" box="[892,1021,822,847]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="10.0" metricValueMin="9.5" pageId="1" pageNumber="247" unit="mm" value="9.75" valueMax="10.0" valueMin="9.5">9.5–10 mm</quantity>
long,
<quantity id="4C9933ADFFAAB570D639FC84FB22004A" box="[1093,1229,822,847]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="1" pageNumber="247" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
wide (limb
<quantity id="4C9933ADFFAAB570D724FC84FF5B0076" metricMagnitude="-3" metricUnit="m" metricValue="6.7" metricValueMax="6.9" metricValueMin="6.5" pageId="1" pageNumber="247" unit="mm" value="6.7" valueMax="6.9" valueMin="6.5">6.5–6.9 mm</quantity>
long, 4-nerved, 3-toothed at the apex), yellow. Disc florets 9–14,
<quantity id="4C9933ADFFAAB570D1C8FCE8FBD10076" box="[948,1086,858,883]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.7" metricValueMin="7.1" pageId="1" pageNumber="247" unit="mm" value="7.4" valueMax="7.7" valueMin="7.1">7.1–7.7 mm</quantity>
long,
<quantity id="4C9933ADFFAAB570D6F6FCE8FB110076" box="[1162,1278,858,883]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="1" pageNumber="247" unit="mm" value="0.95" valueMax="1.0" valueMin="0.9">0.9–1 mm</quantity>
wide, 5-lobed, yellow. Anthers ca.
<quantity id="4C9933ADFFAAB570D316FCCCFE2E0092" box="[362,449,894,919]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="247" unit="mm" value="2.2">2.2 mm</quantity>
long, auriculate to very shortly caudate (ca. 1/6 as long as the filament collar); filament collar balusterform. Style-branches truncate with a crown of sweeping hairs. Achenes (immature) glabrous; pappus
<quantity id="4C9933ADFFAAB570D2F4FC74FEE000DA" box="[136,271,966,991]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.6" metricValueMin="6.5" pageId="1" pageNumber="247" unit="mm" value="7.05" valueMax="7.6" valueMin="6.5">6.5–7.6 mm</quantity>
long, barbellate, whitish. Chromosome number: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37BCDC3FFAAB570D2C1FC58FD5507B2" pageId="1" pageNumber="247" type="distribution">
<paragraph id="8BDE9E48FFAAB570D2C1FC58FD630796" blockId="1.[136,1452,462,1963]" pageId="1" pageNumber="247">
<emphasis id="B915425AFFAAB570D2C1FC58FE1F0706" bold="true" box="[189,496,1002,1027]" pageId="1" pageNumber="247">Distribution and habitat:</emphasis>
—
<collectingCountry id="F376DED8FFAAB570D071FC58FD850706" box="[525,618,1002,1027]" name="Ecuador" pageId="1" pageNumber="247">Ecuador</collectingCountry>
(
<collectingRegion id="49A550AAFFAAB570D000FC58FD500706" box="[636,703,1002,1027]" country="Ecuador" name="Napo" pageId="1" pageNumber="247">Napo</collectingRegion>
). This species is known from the highlands of Papallacta, in the southwestern part of the Cayambe-Coca National Park.
<taxonomicName id="4C61E5CBFFAAB570D16CFBBCFC1A0722" box="[784,1013,1038,1063]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D16CFBBCFC1A0722" box="[784,1013,1038,1063]" italics="true" pageId="1" pageNumber="247">Monticalia speciosa</emphasis>
</taxonomicName>
thrives in humid grassy páramos and around the transition between high montane forest and páramo, at elevations of
<quantity id="4C9933ADFFAAB570D183FB80FB72074E" box="[1023,1181,1074,1099]" metricMagnitude="3" metricUnit="m" metricValue="3.9" metricValueMax="4.05" metricValueMin="3.75" pageId="1" pageNumber="247" unit="m" value="3900.0" valueMax="4050.0" valueMin="3750.0">3750–4050 m</quantity>
. Some species that were observed in the same habitat are
<taxonomicName id="4C61E5CBFFAAB570D38CFBE4FD22076A" box="[496,717,1110,1135]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="padifolia">
<emphasis id="B915425AFFAAB570D38CFBE4FD22076A" box="[496,717,1110,1135]" italics="true" pageId="1" pageNumber="247">Baccharis padifolia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C61E5CBFFAAB570D0A6FBE4FC00076A" box="[730,1007,1110,1135]" class="Magnoliopsida" family="Asteraceae" genus="Diplostephium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="hartwegii">
<emphasis id="B915425AFFAAB570D0A6FBE4FC00076A" box="[730,1007,1110,1135]" italics="true" pageId="1" pageNumber="247">Diplostephium hartwegii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C61E5CBFFAAB570D180FBE4FB7C076A" box="[1020,1171,1110,1135]" class="Magnoliopsida" family="Asteraceae" genus="Gynoxys" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B915425AFFAAB570D180FBE4FBB3076A" box="[1020,1116,1110,1135]" italics="true" pageId="1" pageNumber="247">Gynoxys</emphasis>
spp.
</taxonomicName>
,
<emphasis id="B915425AFFAAB570D6E1FBE4FEEF0796" italics="true" pageId="1" pageNumber="247">
<taxonomicName id="4C61E5CBFFAAB570D6E1FBE4FA6C076A" authorityName="Jeffrey" authorityYear="1992" baseAuthorityName="Turczaninow" baseAuthorityYear="1851" box="[1181,1411,1110,1135]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="andicola">Monticalia andicola</taxonomicName>
,
<taxonomicName id="4C61E5CBFFAAB570D7F1FBE4FEEF0796" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="arbutifolia">M. arbutifolia</taxonomicName>
</emphasis>
,
<emphasis id="B915425AFFAAB570D372FBC8FE570796" box="[270,440,1146,1171]" italics="true" pageId="1" pageNumber="247">
<taxonomicName id="4C61E5CBFFAAB570D372FBC8FE5B0796" authorityName="Nordenstam" authorityYear="1996" baseAuthorityName="Cuatrecasas" baseAuthorityYear="1953" box="[270,436,1146,1171]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="befarioides">M. befarioides</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C61E5CBFFAAB570D393FBC8FD690796" authorityName="Jeffrey" authorityYear="1992" baseAuthorityName="Persoon" baseAuthorityYear="1807" box="[495,646,1146,1171]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B915425AFFAAB570D393FBC8FD690796" box="[495,646,1146,1171]" italics="true" pageId="1" pageNumber="247">M. peruviana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDE9E48FFAAB570D2C1FB2CFD5507B2" blockId="1.[136,1452,462,1963]" box="[189,698,1182,1207]" pageId="1" pageNumber="247">
<emphasis id="B915425AFFAAB570D2C1FB2CFEAC07B2" bold="true" box="[189,323,1182,1207]" pageId="1" pageNumber="247">Phenology:</emphasis>
—Collected in bloom in October.
</paragraph>
</subSubSection>
<subSubSection id="C37BCDC3FFAAB570D2C1FB70FED307FA" pageId="1" pageNumber="247" type="etymology">
<paragraph id="8BDE9E48FFAAB570D2C1FB70FED307FA" blockId="1.[136,1452,462,1963]" pageId="1" pageNumber="247">
<emphasis id="B915425AFFAAB570D2C1FB70FEA707DE" bold="true" box="[189,328,1218,1243]" pageId="1" pageNumber="247">Etymology:</emphasis>
—The epithet
<taxonomicName id="4C61E5CBFFAAB570D393FB70FDA007DE" box="[495,591,1218,1243]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D393FB70FDA007DE" box="[495,591,1218,1243]" italics="true" pageId="1" pageNumber="247">speciosa</emphasis>
</taxonomicName>
means showy, splendid, which refers to the outstanding habit of this plant when it blooms.
</paragraph>
</subSubSection>
<subSubSection id="C37BCDC3FFAAB570D2C1FAB8FDD104AE" pageId="1" pageNumber="247" type="discussion">
<paragraph id="8BDE9E48FFAAB570D2C1FAB8FF3C068A" blockId="1.[136,1452,462,1963]" pageId="1" pageNumber="247">
<emphasis id="B915425AFFAAB570D2C1FAB8FEAA0626" bold="true" box="[189,325,1290,1315]" pageId="1" pageNumber="247">Discussion:</emphasis>
—The new species can be easily identified by its minute leaves, which are sessile to subsessile, elliptic, ciliate on the margins and midrib, and plainly concolorous. The leaves of
<taxonomicName id="4C61E5CBFFAAB570D638FA9CFB230642" box="[1092,1228,1326,1351]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D638FA9CFB230642" box="[1092,1228,1326,1351]" italics="true" pageId="1" pageNumber="247">M. speciosa</emphasis>
</taxonomicName>
are gibbous-based, character that has not been observed in other species. It is also characterized by its radiate capitula with eight involucral bracts.
</paragraph>
<paragraph id="8BDE9E48FFAAB570D2C1FA28FD05041E" blockId="1.[136,1452,462,1963]" pageId="1" pageNumber="247">
The distribution of
<taxonomicName id="4C61E5CBFFAAB570D3F1FA28FDFF06B6" box="[397,528,1434,1459]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D3F1FA28FDFF06B6" box="[397,528,1434,1459]" italics="true" pageId="1" pageNumber="247">M. speciosa</emphasis>
</taxonomicName>
overlaps with
<taxonomicName id="4C61E5CBFFAAB570D0D1FA28FAE906B6" authority="(Turczaninow 1851: 91) Jeffrey (1992: 69)" authorityName="Jeffrey" authorityPageNumber="69" authorityYear="1992" baseAuthorityName="Turczaninow" baseAuthorityPageNumber="91" baseAuthorityYear="1851" box="[685,1286,1434,1459]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="andicola">
<emphasis id="B915425AFFAAB570D0D1FA28FCDE06B6" box="[685,817,1434,1459]" italics="true" pageId="1" pageNumber="247">M.andicola</emphasis>
(
<bibRefCitation id="EFF0E3B9FFAAB570D141FA28FBD906B6" author="Turczaninow, N." box="[829,1078,1434,1459]" pageId="1" pageNumber="247" pagination="59 - 95" refId="ref2498" refString="Turczaninow, N. (1851) Synanthereae quaedam hucusque indescriptae. Bulletin de la Societe Imperiale des Naturalistes de Moscou 24 (3): 59 - 95." type="journal article" year="1851">Turczaninow1851: 91</bibRefCitation>
)
<bibRefCitation id="EFF0E3B9FFAAB570D63EFA28FAE906B6" author="Jeffrey, C." box="[1090,1286,1434,1459]" pageId="1" pageNumber="247" pagination="49 - 109" refId="ref2110" refString="Jeffrey, C. (1992) The tribe Senecioneae (Compositae) in the Mascarene Islands with an annotated world check-list of the genera of the tribe. Notes on Compositae: VI. Kew Bulletin 47 (1): 49 - 109. https: // doi. org / 10.2307 / 4110768" type="journal article" year="1992">Jeffrey (1992: 69)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C61E5CBFFAAB570D76DFA28FA4306B6" box="[1297,1452,1434,1459]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="arbutifolia">
<emphasis id="B915425AFFAAB570D76DFA28FA4306B6" box="[1297,1452,1434,1459]" italics="true" pageId="1" pageNumber="247">M. arbutifolia</emphasis>
</taxonomicName>
(Kunth in
<bibRefCitation id="EFF0E3B9FFAAB570D280FA0CFDC806D2" author="Humboldt, F. W. H. A. &amp; Bonpland, A. J. A. &amp; Kunth, K. S." box="[252,551,1470,1495]" pageId="1" pageNumber="247" pagination="121 - 168" refId="ref2051" refString="Humboldt, F. W. H. A., Bonpland, A. J. A. &amp; Kunth, K. S. (1818) Nova genera et species plantarum, vol. 4, part 16 (ed. fol.). Lutetiae Parisiorum, Paris, pp. 121 - 168." type="book chapter" year="1818">
Humboldt
<emphasis id="B915425AFFAAB570D309FA0CFE4206D2" box="[373,429,1470,1495]" italics="true" pageId="1" pageNumber="247">et al.</emphasis>
1818: 143
</bibRefCitation>
)
<bibRefCitation id="EFF0E3B9FFAAB570D04AFA0CFCEC06D2" author="Jeffrey, C." box="[566,771,1470,1495]" pageId="1" pageNumber="247" pagination="49 - 109" refId="ref2110" refString="Jeffrey, C. (1992) The tribe Senecioneae (Compositae) in the Mascarene Islands with an annotated world check-list of the genera of the tribe. Notes on Compositae: VI. Kew Bulletin 47 (1): 49 - 109. https: // doi. org / 10.2307 / 4110768" type="journal article" year="1992">Jeffrey (1992: 70)</bibRefCitation>
,
<taxonomicName id="4C61E5CBFFAAB570D173FA0CFF4206FE" authority="(Cuatrecasas 1953: 134) Nordenstam (1996: 49)" authorityName="Nordenstam" authorityPageNumber="49" authorityYear="1996" baseAuthorityName="Cuatrecasas" baseAuthorityPageNumber="134" baseAuthorityYear="1953" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="befarioides">
<emphasis id="B915425AFFAAB570D173FA0CFC5C06D2" box="[783,947,1470,1495]" italics="true" pageId="1" pageNumber="247">M. befarioides</emphasis>
(
<bibRefCitation id="EFF0E3B9FFAAB570D1BDFA0CFB2D06D2" author="Cuatrecasas, J." box="[961,1218,1470,1495]" pageId="1" pageNumber="247" pagination="120 - 153" refId="ref1902" refString="Cuatrecasas, J. (1953) Neue und bemerkenswerte andine Compositen. Feddes Repertorium specierum novarum regni vegetabilis 55: 120 - 153. https: // doi. org / 10.1002 / fedr. 19530550203" type="journal article" year="1953">Cuatrecasas 1953: 134</bibRefCitation>
)
<bibRefCitation id="EFF0E3B9FFAAB570D6AEFA0CFF4206FE" author="Nordenstam, B." pageId="1" pageNumber="247" pagination="47 - 50" refId="ref2166" refString="Nordenstam, B. (1996) New combinations in Ecuadorean Senecioneae. Compositae Newsletter 29: 47 - 50." type="journal article" year="1996">Nordenstam (1996: 49)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C61E5CBFFAAB570D296FA50FCD306FE" authority="(Persoon 1807: 436) Jeffrey (1992: 72)" authorityName="Jeffrey" authorityPageNumber="72" authorityYear="1992" baseAuthorityName="Persoon" baseAuthorityPageNumber="436" baseAuthorityYear="1807" box="[234,828,1506,1531]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B915425AFFAAB570D296FA50FE6D06FE" box="[234,386,1506,1531]" italics="true" pageId="1" pageNumber="247">M. peruviana</emphasis>
(
<bibRefCitation id="EFF0E3B9FFAAB570D3F3FA50FD8A06FE" author="Persoon, C. H." box="[399,613,1506,1531]" pageId="1" pageNumber="247" pagination="273 - 657" refId="ref2239" refString="Persoon, C. H. (1807) Synopsis plantarum, vol. 2 (2). Apud Bibliopolas Treuttel et Wurtz, Parisiis Lutetiorum; apud J. G. Cottam, Tubingae, pp. 273 - 657." type="book chapter" year="1807">Persoon 1807: 436</bibRefCitation>
)
<bibRefCitation id="EFF0E3B9FFAAB570D00FFA50FCD306FE" author="Jeffrey, C." box="[627,828,1506,1531]" pageId="1" pageNumber="247" pagination="49 - 109" refId="ref2110" refString="Jeffrey, C. (1992) The tribe Senecioneae (Compositae) in the Mascarene Islands with an annotated world check-list of the genera of the tribe. Notes on Compositae: VI. Kew Bulletin 47 (1): 49 - 109. https: // doi. org / 10.2307 / 4110768" type="journal article" year="1992">Jeffrey (1992: 72)</bibRefCitation>
</taxonomicName>
. Among these, it only might be confused with the latter species; however, several characters are useful to discriminate them from each other (
<figureCitation id="135A82CDFFAAB570D644F9B4FB93051A" box="[1080,1148,1542,1567]" captionStart="FIGURE 2" captionStartId="3.[136,229,1957,1978]" captionTargetBox="[151,1435,190,1932]" captionTargetId="figure-16@3.[151,1436,190,1932]" captionTargetPageId="3" captionText="FIGURE 2. Monticalia speciosa (plant details on the left) compared to M. peruviana (plant details on the right). A. Ray florets. B. Involucres. C. Leaves. D. Synflorescences. Photographs by J. Calvo at the locus classicus." figureDoi="http://doi.org/10.5281/zenodo.13716309" httpUri="https://zenodo.org/record/13716309/files/figure.png" pageId="1" pageNumber="247">Fig. 2</figureCitation>
).
<taxonomicName id="4C61E5CBFFAAB570D6F3F9B4FA6E051A" authorityName="Jeffrey" authorityYear="1992" baseAuthorityName="Persoon" baseAuthorityYear="1807" box="[1167,1409,1542,1567]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B915425AFFAAB570D6F3F9B4FA6E051A" box="[1167,1409,1542,1567]" italics="true" pageId="1" pageNumber="247">Monticalia peruviana</emphasis>
</taxonomicName>
is a smaller shrub with discolorous leaves due to the whitish lanate indumentum beneath. The margins are revolute and not ciliate as in
<taxonomicName id="4C61E5CBFFAAB570D36CF9FCFE790562" box="[272,406,1614,1639]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D36CF9FCFE790562" box="[272,406,1614,1639]" italics="true" pageId="1" pageNumber="247">M. speciosa</emphasis>
</taxonomicName>
. The color of the leaves is also very different in the living plants (dark green in
<taxonomicName id="4C61E5CBFFAAB570D762F9FCFA430562" authorityName="Jeffrey" authorityYear="1992" baseAuthorityName="Persoon" baseAuthorityYear="1807" box="[1310,1452,1614,1639]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B915425AFFAAB570D762F9FCFA430562" box="[1310,1452,1614,1639]" italics="true" pageId="1" pageNumber="247">P. peruviana</emphasis>
</taxonomicName>
vs. light green in
<taxonomicName id="4C61E5CBFFAAB570D32FF9C0FE34058E" box="[339,475,1650,1675]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D32FF9C0FE34058E" box="[339,475,1650,1675]" italics="true" pageId="1" pageNumber="247">M. speciosa</emphasis>
</taxonomicName>
). The involucre of
<taxonomicName id="4C61E5CBFFAAB570D0C7F9C0FCBA058E" authorityName="Jeffrey" authorityYear="1992" baseAuthorityName="Persoon" baseAuthorityYear="1807" box="[699,853,1650,1675]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B915425AFFAAB570D0C7F9C0FCBA058E" box="[699,853,1650,1675]" italics="true" pageId="1" pageNumber="247">M. peruviana</emphasis>
</taxonomicName>
has 12–13 involucral bracts and ray florets whereas
<taxonomicName id="4C61E5CBFFAAB570D2F4F924FEE105AA" box="[136,270,1686,1711]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D2F4F924FEE105AA" box="[136,270,1686,1711]" italics="true" pageId="1" pageNumber="247">M. speciosa</emphasis>
</taxonomicName>
has only eight. In
<collectingCountry id="F376DED8FFAAB570D39DF924FDD105AA" box="[481,574,1686,1711]" name="Ecuador" pageId="1" pageNumber="247">Ecuador</collectingCountry>
there is another species with minute leaves, i.e.,
<taxonomicName id="4C61E5CBFFAAB570D61BF924FE2705D6" authority="(Cuatrecasas 1954: 41) Jeffrey (1992: 70)" authorityName="Jeffrey" authorityPageNumber="70" authorityYear="1992" baseAuthorityName="Cuatrecasas" baseAuthorityPageNumber="41" baseAuthorityYear="1954" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="empetroides">
<emphasis id="B915425AFFAAB570D61BF924FAFA05AA" box="[1127,1301,1686,1711]" italics="true" pageId="1" pageNumber="247">M. empetroides</emphasis>
(
<bibRefCitation id="EFF0E3B9FFAAB570D758F924FF0305D6" author="Cuatrecasas, J." pageId="1" pageNumber="247" pagination="39 - 49" refId="ref1939" refString="Cuatrecasas, J. (1954) New species of Compositae from Ecuador, collected by W. H. Camp. Brittonia 8: 39 - 49. https: // doi. org / 10.2307 / 2804909" type="journal article" year="1954">Cuatrecasas 1954: 41</bibRefCitation>
)
<bibRefCitation id="EFF0E3B9FFAAB570D287F908FE2705D6" author="Jeffrey, C." box="[251,456,1722,1747]" pageId="1" pageNumber="247" pagination="49 - 109" refId="ref2110" refString="Jeffrey, C. (1992) The tribe Senecioneae (Compositae) in the Mascarene Islands with an annotated world check-list of the genera of the tribe. Notes on Compositae: VI. Kew Bulletin 47 (1): 49 - 109. https: // doi. org / 10.2307 / 4110768" type="journal article" year="1992">Jeffrey (1992: 70)</bibRefCitation>
</taxonomicName>
. Any confusion is unlikely because this latter species has strongly revolute leaves, which are whitish lanate beneath. The number of involucral bracts is also different (12 vs.
<quantity id="4C9933ADFFAAB570D652F96CFBB605F2" box="[1070,1113,1758,1783]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="1" pageNumber="247" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="4C61E5CBFFAAB570D623F96CFB0B05F2" box="[1119,1252,1758,1783]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D623F96CFB0B05F2" box="[1119,1252,1758,1783]" italics="true" pageId="1" pageNumber="247">M. speciosa</emphasis>
</taxonomicName>
). In
<collectingCountry id="F376DED8FFAAB570D766F96CFA6705F2" box="[1306,1416,1758,1783]" name="Colombia" pageId="1" pageNumber="247">Colombia</collectingCountry>
no radiate species with eight involucral bracts are known.
</paragraph>
<paragraph id="8BDE9E48FFAAB570D2C1F894FDD104AE" blockId="1.[136,1452,462,1963]" pageId="1" pageNumber="247">
It is important to note that any collection belonging to the new species was found among the
<taxonomicName id="4C61E5CBFFAAB570D6B2F894FAA6043A" authorityName="Jeffrey" authorityYear="1992" box="[1230,1353,1830,1855]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="genus">
<emphasis id="B915425AFFAAB570D6B2F894FAA6043A" box="[1230,1353,1830,1855]" italics="true" pageId="1" pageNumber="247">Monticalia</emphasis>
</taxonomicName>
material (and related genera) kept at the Ecuadorian herbaria HA,
<collectingRegion id="49A550AAFFAAB570D175F8F8FCBF0466" box="[777,848,1866,1891]" country="Ecuador" name="Loja" pageId="1" pageNumber="247">LOJA</collectingRegion>
, Q, QAP, QCA, QCNE, and QPLS. Taking in account that
<taxonomicName id="4C61E5CBFFAAB570D2C7F8DCFEAD0482" box="[187,322,1902,1927]" class="Magnoliopsida" family="Asteraceae" genus="Monticalia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="247" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B915425AFFAAB570D2C7F8DCFEAD0482" box="[187,322,1902,1927]" italics="true" pageId="1" pageNumber="247">M. speciosa</emphasis>
</taxonomicName>
is a large shrub reaching up to
<quantity id="4C9933ADFFAAB570D0D3F8DCFD1E0482" box="[687,753,1902,1927]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="247" unit="m" value="3.5">3.5 m</quantity>
tall and displaying showy yellow capitula, it is odd how this species has been overlooked until now.
</paragraph>
</subSubSection>
</treatment>
</document>
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