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<document id="9BEE238A0417D272CA6CD68B6B95EDCC" ID-CLB-Dataset="46319" ID-DOI="10.5281/zenodo.278726" ID-GBIF-Dataset="d1323970-6ef7-47dc-8b45-2ccd538acc7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="278726" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460390935971" checkinUser="plazi" docAuthor="Seniczak, Stanisław, Penttinen, Ritva &amp; Seniczak, Anna" docDate="2011" docId="03C8020FFFBAFFE282E9DD78FCEFFA35" docLanguage="en" docName="zt03034p031.pdf" docOrigin="Zootaxa 3034" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cosmochthonius reticulatus Grandjean 1947" docType="treatment" docVersion="10" lastPageNumber="12" masterDocId="FFF17A77FFBBFFE9827ED86DFFA9FFB9" masterDocTitle="The ontogeny of morphological traits in three European species of Cosmochthonius Berlese, 1910 (Acari: Oribatida: Cosmochthoniidae)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="2" updateTime="1698298215618" updateUser="ExternalLinkService">
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<mods:title id="873DFDBA2B1C667B216412C2E4367618">The ontogeny of morphological traits in three European species of Cosmochthonius Berlese, 1910 (Acari: Oribatida: Cosmochthoniidae)</mods:title>
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<treatment id="03C8020FFFBAFFE282E9DD78FCEFFA35" ID-DOI="http://doi.org/10.5281/zenodo.5696410" ID-GBIF-Taxon="119416789" ID-Zenodo-Dep="5696410" LSID="urn:lsid:plazi:treatment:03C8020FFFBAFFE282E9DD78FCEFFA35" httpUri="http://treatment.plazi.org/id/03C8020FFFBAFFE282E9DD78FCEFFA35" lastPageId="11" lastPageNumber="12" pageId="1" pageNumber="2">
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<emphasis id="B9156F0BFFBAFFE882E9DD78FC6FFA97" bold="true" box="[151,966,1301,1326]" pageId="1" pageNumber="2">
<taxonomicName id="4C61C89AFFBAFFE882E9DD78FD02FA97" ID-CoL="7SN9R" authority="Grandjean, 1947" authorityName="Grandjean" authorityYear="1947" box="[151,683,1301,1326]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFBAFFE882E9DD78FE72FA97" bold="true" box="[151,475,1301,1326]" italics="true" pageId="1" pageNumber="2">Cosmochthonius reticulatus</emphasis>
Grandjean, 1947
</taxonomicName>
(
<figureCitation id="135AAF9CFFBAFFE880C7DD78FCB6FA97" box="[697,799,1301,1326]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="4.[151,250,2002,2025]" captionStartId-1="5.[151,250,1753,1776]" captionStartId-2="6.[151,250,1832,1855]" captionStartId-3="7.[151,250,1957,1980]" captionStartId-4="8.[151,250,1806,1829]" captionStartId-5="9.[151,250,1943,1966]" captionStartId-6="10.[151,250,1996,2019]" captionTargetBox-0="[252,1260,198,1984]" captionTargetBox-1="[289,1303,182,1718]" captionTargetBox-2="[155,1426,405,1811]" captionTargetBox-3="[299,1294,216,1912]" captionTargetBox-4="[156,1432,213,1771]" captionTargetBox-5="[361,1226,228,1907]" captionTargetBox-6="[371,1238,331,1972]" captionTargetId-0="figure@4.[239,1268,189,1989]" captionTargetId-1="figure@5.[280,1309,155,1732]" captionTargetId-2="figure@6.[151,1436,401,1811]" captionTargetId-3="figure@7.[286,1301,199,1919]" captionTargetId-4="figure@8.[151,1436,193,1785]" captionTargetId-5="figure@9.[343,1243,213,1920]" captionTargetId-6="figure@10.[344,1244,310,1978]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionText-0="FIGURE 1. Cosmochthonius reticulatus, adult with cerotegument, dorsal aspect, legs partially drawn, scale bar 50 μm." captionText-1="FIGURE 2. Cosmochthonius reticulatus, adult. (A) Notogaster, cerotegument removed, scale bar 50 μm; (B) fragment of pygidium." captionText-2="FIGURE 3. Cosmochthonius reticulatus, adult, anogenital region, legs partially drawn, scale bar 50 μm." captionText-3="FIGURE 4. Cosmochthonius reticulatus, larva, dorsal aspect, legs partially drawn, scale bar 50 μm." captionText-4="FIGURE 5. Cosmochthonius reticulatus, lateral aspect, legs partially drawn, scale bars 50 μm. (A) Larva; (B) tritonymph." captionText-5="FIGURE 6. Cosmochthonius reticulatus, scale bars 50 μm. (A) Anal region of larva; (B) anogenital region of tritonymph." captionText-6="FIGURE 7. Cosmochthonius reticulatus, tritonymph, dorsal aspect, legs partially drawn, scale bar 50 μm." httpUri-0="https://zenodo.org/record/278727/files/figure.png" httpUri-1="https://zenodo.org/record/278728/files/figure.png" httpUri-2="https://zenodo.org/record/278729/files/figure.png" httpUri-3="https://zenodo.org/record/278730/files/figure.png" httpUri-4="https://zenodo.org/record/278731/files/figure.png" httpUri-5="https://zenodo.org/record/278732/files/figure.png" httpUri-6="https://zenodo.org/record/278733/files/figure.png" pageId="1" pageNumber="2">Figs 1–7</figureCitation>
,
<figureCitation id="135AAF9CFFBAFFE88152DD78FC93FA97" box="[812,826,1301,1326]" captionStart="FIGURE 8" captionStartId="12.[151,250,1693,1716]" captionTargetBox="[162,1423,187,1671]" captionTargetId="figure@12.[151,1436,169,1688]" captionTargetPageId="12" captionText="FIGURE 8. Hypertrophied setae of tritonymph, scale bars 50 μm. (A) Cosmochthonius reticulatus; (B) C. foliatus." httpUri="https://zenodo.org/record/278734/files/figure.png" pageId="1" pageNumber="2">8</figureCitation>
A, 9A, 10A)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37BE092FFBAFFE882E9DD30FD5DFA76" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="8BDEB319FFBAFFE882E9DD30FD48FA09" blockId="1.[151,1436,1373,1487]" pageId="1" pageNumber="2">
<taxonomicName id="4C61C89AFFBAFFE882E9DD30FDDDFACA" ID-CoL="7SN9R" authority="Grandjean 1947" authorityName="Grandjean" authorityYear="1947" box="[151,628,1373,1395]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFBAFFE882E9DD30FE1AFACA" box="[151,435,1373,1395]" italics="true" pageId="1" pageNumber="2">Cosmochthonius reticulatus</emphasis>
:
<bibRefCitation id="EFF0CEE8FFBAFFE883BDDD30FDDDFACA" author="Grandjean" box="[451,628,1373,1395]" pageId="1" pageNumber="2" refString="Grandjean, F. (1947 b) L'origine de la pince mandibulaire chez les Acariens actinochitineux. Archives des Sciences physiques et naturelles, Geneve, (5), 29, 305 - 355." type="journal article" year="1947" yearSuffix="b">Grandjean 1947b</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFF0CEE8FFBAFFE880FADD30FD55FACA" author="Trave" box="[644,764,1373,1395]" pageId="1" pageNumber="2" refString="Trave, J. (1956) Contributions a l' etude de la fauna de la Massane (Deuxieme note). Oribates (Acarien), 1 re partie. Vie et Milieu, 7 (1), 77 - 94." type="journal article" year="1956">Travé 1956</bibRefCitation>
,
<bibRefCitation id="EFF0CEE8FFBAFFE88176DD33FC97FACA" author="Trave" box="[776,830,1374,1395]" pageId="1" pageNumber="2" refString="Trave, J. (1984) Contribution a l'etude des Oribates (Acariens) de l'Ile de Port-Cros (Parc National). Travaux Scientifiques du Parc National de Port-Cros, France, 10, 119 - 150." type="journal article" year="1984">1984</bibRefCitation>
;
<bibRefCitation id="EFF0CEE8FFBAFFE88134DD30FC5BFACA" author="Grandjean" box="[842,1010,1373,1395]" pageId="1" pageNumber="2" refString="Grandjean, F. (1962) Nouvelles observations sur les Oribates (2 e serie). Acarologia, 4, 396 - 422." type="journal article" year="1962">Grandjean 1962</bibRefCitation>
;
<bibRefCitation id="EFF0CEE8FFBAFFE88183DD30FAB0FACA" author="Balogh" box="[1021,1305,1373,1395]" pageId="1" pageNumber="2" refString="Balogh, J. &amp; Mahunka, S. (1983) Primitive Oribatids of the Palaearctic Region. In: Balogh, J. &amp; Mahunka, S. (Eds.): The soil mites of the world. Akademiai Kiado, Budapest, vol. 1, 1 - 372." type="journal article" year="1983">Balogh and Mahunka 1983</bibRefCitation>
;
<bibRefCitation id="EFF0CEE8FFBAFFE8875BDD30FECBFA2B" author="Perez-Inigo" pageId="1" pageNumber="2" refString="Perez-Inigo, C. &amp; Pena, M. A. (1995) Oribatidos (Acari, Oribatei) de Fuerteventura (Islas Canarias). Graellsia, 51, 143 - 164." type="journal article" year="1995">Pérez-Iñigo and Peña 1995</bibRefCitation>
;
<bibRefCitation id="EFF0CEE8FFBAFFE88313DD11FD71FA28" author="Mahunka" box="[365,728,1404,1426]" pageId="1" pageNumber="2" refString="Mahunka, S. &amp; Mahunka-Papp, L. (2003) Contribution to the knowledge of the Hungarian Oribatida fauna (Acari). I. Acta Zoologica Academiae Scientiarum Hungaricae, 49 (4), 255 - 260." type="journal article" year="2003">Mahunka and Mahunka-Papp 2003</bibRefCitation>
,
<bibRefCitation id="EFF0CEE8FFBAFFE8809ADD11FCB3FA28" author="Mahunka-Papp" box="[740,794,1404,1425]" pageId="1" pageNumber="2" refString="Mahunka, S. &amp; Mahunka-Papp, L. (2010) New and little known oribatid mites from the Carpathian Basin and the Balkan Peninsula (Acari: Oribatida). Acta Zoologica Academiae Scientiarum Hungaricae, 56 (3), 211 - 234." type="journal article" year="2010">2010</bibRefCitation>
;
<bibRefCitation id="EFF0CEE8FFBAFFE8815BDD10FB9AFA28" author="Karasawa" box="[805,1075,1404,1426]" pageId="1" pageNumber="2" refString="Karasawa, S. &amp; Hijii, N. (2004 a) Effects of microhabitat diversity and geographical isolation on oribatid mite (Acari: Oribatida) communities in mangrove forests. Pedobiologia, 48 (3), 245 - 256." type="journal article" year="2004" yearSuffix="a">Karasawa and Hijii 2004a</bibRefCitation>
,
<bibRefCitation id="EFF0CEE8FFBAFFE88641DD11FB28FA2B" author="Karasawa" box="[1087,1153,1404,1426]" pageId="1" pageNumber="2" refString="Karasawa, S. &amp; Hijii, N. (2004 b) Morphological modifications among oribatid mites (Acari: Oribatida) in relation to habitat differentiation in mangrove forests. Pedobiologia, 48 (3), 383 - 394." type="journal article" year="2004" yearSuffix="b">2004b</bibRefCitation>
;
<bibRefCitation id="EFF0CEE8FFBAFFE886F2DD11FAA5FA28" author="Subias" box="[1164,1292,1404,1426]" pageId="1" pageNumber="2" refString="Subias, L. S. (2004) Systematic, synonymic and biogeographical check - list of the world's oribatid mites (Acariformes, Oribatida) (1758 - 2002). Graellsia, 60, 3 - 305. Online version accessed in February 2011. 557 pp. http: // www. ucm. es / info / zoo / Artropodos / Catalogo. pdf." type="book chapter" year="2004">Subías 2004</bibRefCitation>
, 2011;
<bibRefCitation id="EFF0CEE8FFBAFFE88726DD10FECDFA09" author="Karasawa" pageId="1" pageNumber="2" refString="Karasawa, S., Gotoh, K., Sasaki, T. &amp; Hijii, N. (2005) Wind-based dispersal of oribatid mites (Acari: Oribatida) in a subtropical forest in Japan. Journal of the Acarological Society of Japan, 14 (2), 117 - 122." type="journal article" year="2005">
Karasawa
<emphasis id="B9156F0BFFBAFFE88288DDF1FE80FA08" box="[246,297,1435,1457]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
2005
</bibRefCitation>
;
<bibRefCitation id="EFF0CEE8FFBAFFE88310DDF7FD34FA09" author="Penttinen" box="[366,669,1434,1456]" pageId="1" pageNumber="2" refString="Penttinen, R. &amp; Gordeeva, E. (2005) Redescription of Cosmochthonius foliatus Subias, 1982 and supplements to the descriptions of C. reticulatus Grandjean, 1947 and C. lanatus (Michael, 1995) (Acari: Oribatida: Cosmochthoniidae). Acarologia, 46 (4), 349 - 361." type="journal article" year="2005">Penttinen and Gordeeva 2005</bibRefCitation>
,
<bibRefCitation id="EFF0CEE8FFBAFFE880D9DDF6FD74FA09" author="Penttinen" box="[679,733,1435,1456]" pageId="1" pageNumber="2" refString="Penttinen, R. &amp; Gordeeva, E. (2009) Distribution of Cosmochthonius species (Oribatida: Cosmochthoniidae) in the eastern part of the Mediterranean, Ukraine and Tajikistan. In: Sabelis, M. W. &amp; Bruin, J. (Eds.): Trends in Acarology, pp. 171 - 174." type="book chapter" year="2009">2009</bibRefCitation>
.
</paragraph>
<paragraph id="8BDEB319FFBAFFE882E9DDD4FD5DFA76" blockId="1.[151,1436,1373,1487]" box="[151,756,1465,1487]" pageId="1" pageNumber="2">
<taxonomicName id="4C61C89AFFBAFFE882E9DDD4FDEDFA76" ID-CoL="7SNBF" authority="Sergienko, 1991" authorityName="Sergienko" authorityYear="1991" box="[151,580,1465,1487]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="trivialis">
<emphasis id="B9156F0BFFBAFFE882E9DDD4FE3CFA76" box="[151,405,1465,1487]" italics="true" pageId="1" pageNumber="2">Cosmochthonius trivialis</emphasis>
<bibRefCitation id="EFF0CEE8FFBAFFE883E5DDD4FDEDFA76" author="Sergienko" box="[411,580,1465,1487]" pageId="1" pageNumber="2" refString="Sergienko, G. D. (1991) New species of primitive oribatids (Oribatei, Archoribatida) from Ukraine. Vestnik Zoologii, 1991 (6), 26 - 33 (in Russian)." type="journal article" year="1991">Sergienko, 1991</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="EFF0CEE8FFBAFFE88030DDD4FD59FA76" author="Sergienko" box="[590,752,1465,1487]" pageId="1" pageNumber="2" refString="Sergienko, G. D. (1994) Lower Oribatids. In: Topacevskij, B. A. (Ed.): Fauna of the Ukraine 25. Naukova Dumka, Kiev, p. 5 - 204 (In Russian)." type="journal article" year="1994">Sergienko 1994</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37BE092FFBAFFE882E9DD96FAD6F81A" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BDEB319FFBAFFE882E9DD96FE14F8AA" blockId="1.[151,1436,1531,2027]" pageId="1" pageNumber="2">
<emphasis id="B9156F0BFFBAFFE882E9DD96FEB8F9AD" bold="true" box="[151,273,1531,1556]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
Adult small (266–340 µm), light brown, with typical characters of
<taxonomicName id="4C61C89AFFBAFFE88671DD91FB63F9AD" box="[1039,1226,1532,1556]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9156F0BFFBAFFE88671DD91FB63F9AD" box="[1039,1226,1532,1556]" italics="true" pageId="1" pageNumber="2">Cosmochthonius</emphasis>
</taxonomicName>
(for adult see also
<bibRefCitation id="EFF0CEE8FFBAFFE882E9DE4DFE7BF981" author="Penttinen" box="[151,466,1568,1592]" pageId="1" pageNumber="2" refString="Penttinen, R. &amp; Gordeeva, E. (2005) Redescription of Cosmochthonius foliatus Subias, 1982 and supplements to the descriptions of C. reticulatus Grandjean, 1947 and C. lanatus (Michael, 1995) (Acari: Oribatida: Cosmochthoniidae). Acarologia, 46 (4), 349 - 361." type="journal article" year="2005">Penttinen &amp; Gordeeva 2005</bibRefCitation>
). Seta
<emphasis id="B9156F0BFFBAFFE88061DE4CFD82F981" box="[543,555,1569,1592]" italics="true" pageId="1" pageNumber="2">c</emphasis>
3 inserted on lateral border of plate
<emphasis id="B9156F0BFFBAFFE881CBDE4CFC7FF981" box="[949,982,1569,1592]" italics="true" pageId="1" pageNumber="2">Na</emphasis>
(
<figureCitation id="135AAF9CFFBAFFE8819BDE4DFB85F981" box="[997,1068,1568,1592]" captionStart="FIGURE 1" captionStartId="4.[151,250,2002,2025]" captionTargetBox="[252,1260,198,1984]" captionTargetId="figure@4.[239,1268,189,1989]" captionTargetPageId="4" captionText="FIGURE 1. Cosmochthonius reticulatus, adult with cerotegument, dorsal aspect, legs partially drawn, scale bar 50 μm." httpUri="https://zenodo.org/record/278727/files/figure.png" pageId="1" pageNumber="2">Figs 1</figureCitation>
,
<figureCitation id="135AAF9CFFBAFFE88647DE4DFBE3F981" box="[1081,1098,1568,1592]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1776]" captionTargetBox="[289,1303,182,1718]" captionTargetId="figure@5.[280,1309,155,1732]" captionTargetPageId="5" captionText="FIGURE 2. Cosmochthonius reticulatus, adult. (A) Notogaster, cerotegument removed, scale bar 50 μm; (B) fragment of pygidium." httpUri="https://zenodo.org/record/278728/files/figure.png" pageId="1" pageNumber="2">2</figureCitation>
). Setae of
<emphasis id="B9156F0BFFBAFFE886BFDE4CFB64F981" box="[1217,1229,1569,1592]" italics="true" pageId="1" pageNumber="2">e</emphasis>
-series with 12–13 pairs of cilia, those of
<emphasis id="B9156F0BFFBAFFE883E1DE2AFE0EF9E6" box="[415,423,1607,1631]" italics="true" pageId="1" pageNumber="2">f</emphasis>
-series shorter (
<tableCitation id="C6E386A2FFBAFFE88027DE2AFD06F9E6" box="[601,687,1607,1631]" captionStart="TABLE 1" captionStartId="2.[151,239,295,319]" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages of Cosmochthonius reticulatus, C. foliatus, and C. ugamaensis (mean measurements of 4 – 10 specimens in μm); larva (L), protonymph (PN), deutonymph (DN), tritonymph (TN), adult (AD); Nd – not developed." pageId="1" pageNumber="2">Table 1</tableCitation>
), with 10–11 pairs of cilia. Whole body covered with reticulate, porous cerotegument, cerotegumental collar absent. Cuticle with polygonal or hexagonal foveae in SEM micrographs, reticulate in light microscopy (
<figureCitation id="135AAF9CFFBAFFE8802ADEE2FD32F91E" box="[596,667,1679,1703]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1776]" captionTargetBox="[289,1303,182,1718]" captionTargetId="figure@5.[280,1309,155,1732]" captionTargetPageId="5" captionText="FIGURE 2. Cosmochthonius reticulatus, adult. (A) Notogaster, cerotegument removed, scale bar 50 μm; (B) fragment of pygidium." httpUri="https://zenodo.org/record/278728/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
). Genital plate larger than anal plate, with 10 pairs of setae, six of which inserted on inner border of plate (
<figureCitation id="135AAF9CFFBAFFE88024DEDEFD37F972" box="[602,670,1715,1739]" captionStart="FIGURE 3" captionStartId="6.[151,250,1832,1855]" captionTargetBox="[155,1426,405,1811]" captionTargetId="figure@6.[151,1436,401,1811]" captionTargetPageId="6" captionText="FIGURE 3. Cosmochthonius reticulatus, adult, anogenital region, legs partially drawn, scale bar 50 μm." httpUri="https://zenodo.org/record/278729/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
). Formulae of setae (+solenidia) of legs (trochanter to tarsus) as follows: I – 0-5-5-(6+1)-(19+1),
<date id="FFDF95D9FFBAFFE88393DEB5FDEBF956" box="[493,578,1751,1775]" bridgedPair="-" pageId="1" pageNumber="2">II – 1-6</date>
-5-(6+1)-(17+1);
<date id="FFDF95D9FFBAFFE88177DEB5FCC3F956" box="[777,874,1751,1775]" bridgedPair="-" pageId="1" pageNumber="2">III – 2-3</date>
-4-(4+1)-15;
<date id="FFDF95D9FFBAFFE88180DEB5FBC9F956" box="[1022,1120,1751,1775]" bridgedPair="-" pageId="1" pageNumber="2">IV – 2-3</date>
-4-(4+1)-14. Tarsus I bidactyle, tarsi II–IV tridactyle.
</paragraph>
<paragraph id="8BDEB319FFBAFFE882B9DF4DFAD6F81A" blockId="1.[151,1436,1531,2027]" pageId="1" pageNumber="2">
Larva unpigmented, nymphs light brown. Most prodorsal setae bushy, sensillus with long, barbed head. Gastronotum with three transverse scissures. Larva with 14 pairs of gastronotal setae, nymphs with 16 pairs. Setae of
<emphasis id="B9156F0BFFBAFFE887F8DF29FA3BF8E2" box="[1414,1426,1860,1883]" italics="true" pageId="1" pageNumber="2">e</emphasis>
- and
<emphasis id="B9156F0BFFBAFFE882B8DF0AFF67F8C6" box="[198,206,1895,1919]" italics="true" pageId="1" pageNumber="2">f</emphasis>
-series hypertrophied, and pinnate; other setae distinctly shorter and barbed. Hypertrophied setae of larva with 6–7 pairs of cilia, those of nymphs with 8–9 pairs. Other morphological characters of juveniles given in
<tableCitation id="C6E386A2FFBAFFE88758DFE6FAD5F81A" box="[1318,1404,1931,1955]" captionStart="TABLE 2" captionStartId="3.[151,239,151,175]" captionTargetBox="[151,1436,238,590]" captionTargetPageId="3" captionText="TABLE 2. Selected morphological characters of Cosmochthonius reticulatus, C. foliatus, C. ugamaensis, and C. ponticus." httpUri="http://table.plazi.org/id/DF1EE391FFB8FFEA82E9D8FAFF68FF6B" pageId="1" pageNumber="2" tableUuid="DF1EE391FFB8FFEA82E9D8FAFF68FF6B">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37BE092FFBAFFEB82B9DFC3FA3BF872" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="8BDEB319FFBAFFE882B9DFC3FA35F852" blockId="1.[151,1436,1531,2027]" pageId="1" pageNumber="2">
<emphasis id="B9156F0BFFBAFFE882B9DFC3FD21F87E" bold="true" box="[199,648,1966,1991]" pageId="1" pageNumber="2">Description of larva and tritonymph:</emphasis>
Larva small (
<tableCitation id="C6E386A2FFBAFFE88159DFC2FCD2F87E" box="[807,891,1967,1991]" captionStart="TABLE 1" captionStartId="2.[151,239,295,319]" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages of Cosmochthonius reticulatus, C. foliatus, and C. ugamaensis (mean measurements of 4 – 10 specimens in μm); larva (L), protonymph (PN), deutonymph (DN), tritonymph (TN), adult (AD); Nd – not developed." pageId="1" pageNumber="2">Table 1</tableCitation>
), unpigmented, often distended in alcohol samples. Prodorsum relatively long, subtriangular, with wide rounded rostrum, with small, thin fenestrate areas (
<figureCitation id="135AAF9CFFBAFFE88710DFBEFA35F852" box="[1390,1436,2003,2027]" captionStart="FIGURE 4" captionStartId="7.[151,250,1957,1980]" captionTargetBox="[299,1294,216,1912]" captionTargetId="figure@7.[286,1301,199,1919]" captionTargetPageId="7" captionText="FIGURE 4. Cosmochthonius reticulatus, larva, dorsal aspect, legs partially drawn, scale bar 50 μm." httpUri="https://zenodo.org/record/278730/files/figure.png" pageId="1" pageNumber="2">Fig.</figureCitation>
</paragraph>
<paragraph id="8BDEB319FFB9FFEB82E9D8FAFF7AFF4E" blockId="2.[151,1436,151,247]" pageId="2" pageNumber="3">
4). Prodorsal setae
<emphasis id="B9156F0BFFB9FFEB8313D8F4FE2DFF09" box="[365,388,153,176]" italics="true" pageId="2" pageNumber="3">ro</emphasis>
,
<emphasis id="B9156F0BFFB9FFEB83EDD8F5FE0EFF09" box="[403,423,152,176]" italics="true" pageId="2" pageNumber="3">le</emphasis>
,
<emphasis id="B9156F0BFFB9FFEB83CBD8F4FE62FF09" box="[437,459,153,176]" italics="true" pageId="2" pageNumber="3">in</emphasis>
and
<emphasis id="B9156F0BFFB9FFEB807CD8F4FD8FFF09" box="[514,550,153,176]" italics="true" pageId="2" pageNumber="3">exs</emphasis>
bushy, curved, usually uniramous, spines single or bifurcate; seta
<emphasis id="B9156F0BFFB9FFEB8772D8F4FA84FF09" box="[1292,1325,153,176]" italics="true" pageId="2" pageNumber="3">exi</emphasis>
distinctly shorter and rather smooth (
<figureCitation id="135AAF9CFFB9FFEB83B6D8D1FDA2FF6D" box="[456,523,188,212]" captionStart="FIGURE 5" captionStartId="8.[151,250,1806,1829]" captionTargetBox="[156,1432,213,1771]" captionTargetId="figure@8.[151,1436,193,1785]" captionTargetPageId="8" captionText="FIGURE 5. Cosmochthonius reticulatus, lateral aspect, legs partially drawn, scale bars 50 μm. (A) Larva; (B) tritonymph." httpUri="https://zenodo.org/record/278731/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
A). Bothridium rounded, sensillus slightly curved, and with long, narrow, barbed head.
</paragraph>
<caption id="DF1EE391FFB9FFEB82E9D94AFD1BFEC6" pageId="2" pageNumber="3">
<paragraph id="8BDEB319FFB9FFEB82E9D94AFD1BFEC6" blockId="2.[151,1436,295,383]" pageId="2" pageNumber="3">
<emphasis id="B9156F0BFFB9FFEB82E9D94AFEA0FE86" bold="true" box="[151,265,295,319]" pageId="2" pageNumber="3">TABLE 1.</emphasis>
Measurements of some morphological characters of juvenile stages of
<taxonomicName id="4C61C89AFFB9FFEB8674D945FA9DFE86" box="[1034,1332,296,319]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB9FFEB8674D945FA9DFE86" box="[1034,1332,296,319]" italics="true" pageId="2" pageNumber="3">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C61C89AFFB9FFEB873CD945FF1FFEE6" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B9156F0BFFB9FFEB873CD945FAFDFE86" box="[1346,1364,296,319]" italics="true" pageId="2" pageNumber="3">C</emphasis>
.
<emphasis id="B9156F0BFFB9FFEB871CD945FF1FFEE6" italics="true" pageId="2" pageNumber="3">foliatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C61C89AFFB9FFEB828ED925FE25FEE6" box="[240,396,328,351]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ugamaensis">
<emphasis id="B9156F0BFFB9FFEB828ED925FEA8FEE6" box="[240,257,328,351]" italics="true" pageId="2" pageNumber="3">C</emphasis>
.
<emphasis id="B9156F0BFFB9FFEB8370D925FE25FEE6" box="[270,396,328,351]" italics="true" pageId="2" pageNumber="3">ugamaensis</emphasis>
</taxonomicName>
(mean measurements of 4–10 specimens in μm); larva (L), protonymph (PN), deutonymph (DN), tritonymph (TN), adult (AD); Nd – not developed.
</paragraph>
</caption>
<paragraph id="8BDEB319FFB9FFEB82DED9F2FA3BFE0D" blockId="2.[160,1427,415,1995]" box="[160,1426,415,437]" pageId="2" pageNumber="3">Species Morphological characters L PN DN TN AD</paragraph>
<paragraph id="8BDEB319FFB9FFEB82DED9A4FA3BFC05" blockId="2.[160,1427,415,1995]" pageId="2" pageNumber="3">
<taxonomicName id="4C61C89AFFB9FFEB82DED9A4FDAAFE67" authority="Body" authorityName="Body" box="[160,515,456,478]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB9FFEB82DED9A4FF18FE67" box="[160,177,457,478]" italics="true" pageId="2" pageNumber="3">C</emphasis>
.
<emphasis id="B9156F0BFFB9FFEB82C0D9A5FE83FE67" box="[190,298,456,478]" italics="true" pageId="2" pageNumber="3">reticulatus</emphasis>
Body
</taxonomicName>
length 160 243 264 330 330 Body width 96 120 136 182 173 Length of: sensillus 48 51 58 65 67 seta
<emphasis id="B9156F0BFFB9FFEB81E3DA28FC01FDE3" box="[925,936,581,602]" italics="true" pageId="2" pageNumber="3">c</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DA3FFBD8FDE3" box="[937,1137,581,606]" pageId="2" pageNumber="3" value="1924-01-22">1 22 24</date>
38 43 50 seta
<emphasis id="B9156F0BFFB9FFEB81E3DA1EFC01FD31" box="[925,936,627,648]" italics="true" pageId="2" pageNumber="3">c</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DA12FBD8FD3E" box="[937,1137,625,651]" pageId="2" pageNumber="3" value="1916-03-21">3 21 16</date>
32 42 45 seta
<emphasis id="B9156F0BFFB9FFEB81E6DACDFC19FD0C" box="[920,944,672,693]" italics="true" pageId="2" pageNumber="3">cp</emphasis>
25 32 45 52 53 seta
<emphasis id="B9156F0BFFB9FFEB81E2DAA5FC00FD67" box="[924,937,712,734]" italics="true" pageId="2" pageNumber="3">d</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DABBFBD8FD64" box="[937,1137,712,738]" pageId="2" pageNumber="3" value="1913-01-17">1 17 13</date>
15 17 31 seta
<emphasis id="B9156F0BFFB9FFEB81E2DA98FC00FCB2" box="[924,937,757,779]" italics="true" pageId="2" pageNumber="3">d</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DB6EFBD8FCB2" box="[937,1137,758,783]" pageId="2" pageNumber="3" value="1922-02-23">2 23 22</date>
22 23 35 seta
<geoCoordinate id="EE55D5DEFFB9FFEB81E3DB4EFBD8FC81" box="[925,1137,802,828]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="15" value="2.0702777">
<emphasis id="B9156F0BFFB9FFEB81E3DB4EFC01FC81" box="[925,936,803,824]" italics="true" pageId="2" pageNumber="3">e</emphasis>
1 63 73
</geoCoordinate>
103 144 176 seta
<emphasis id="B9156F0BFFB9FFEB81DCDB22FC00FCDC" box="[930,937,847,869]" italics="true" pageId="2" pageNumber="3">f</emphasis>
1 54 64 92 136 162 genital opening Nd 22 35 48 82 analopening 23 30 42 55 50
</paragraph>
<paragraph id="8BDEB319FFB9FFEB82DEDBBDFA3BFA7A" blockId="2.[160,1427,415,1995]" pageId="2" pageNumber="3">
<taxonomicName id="4C61C89AFFB9FFEB82DEDBBDFDAAFC5C" authority="Body" authorityName="Body" box="[160,515,975,997]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B9156F0BFFB9FFEB82DEDBBDFF18FC5C" box="[160,177,976,997]" italics="true" pageId="2" pageNumber="3">C</emphasis>
.
<emphasis id="B9156F0BFFB9FFEB82C0DBA2FEA3FC5C" box="[190,266,975,997]" italics="true" pageId="2" pageNumber="3">foliatus</emphasis>
Body
</taxonomicName>
length 149 182 198 231 297 Body width 88 102 112 120 152 Length of: sensillus 48 51 62 72 65 seta
<emphasis id="B9156F0BFFB9FFEB81E3DC20FC01FBDB" box="[925,936,1101,1122]" italics="true" pageId="2" pageNumber="3">c</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DC34FBD8FBD8" box="[937,1137,1100,1125]" pageId="2" pageNumber="3" value="1924-01-22">1 22 24</date>
37 42 39 seta
<emphasis id="B9156F0BFFB9FFEB81E3DC17FC01FB36" box="[925,936,1146,1167]" italics="true" pageId="2" pageNumber="3">c</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DCEAFBD8FB37" box="[937,1137,1145,1171]" pageId="2" pageNumber="3" value="1917-03-22">3 22 17</date>
30 33 39 seta
<emphasis id="B9156F0BFFB9FFEB81E6DCCAFC19FB05" box="[920,944,1191,1212]" italics="true" pageId="2" pageNumber="3">cp</emphasis>
25 17 41 51 57 seta
<emphasis id="B9156F0BFFB9FFEB81E2DCA2FC00FB5C" box="[924,937,1231,1253]" italics="true" pageId="2" pageNumber="3">d</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DCB0FBD8FB5C" box="[937,1137,1231,1257]" pageId="2" pageNumber="3" value="1915-01-17">1 17 15</date>
15 21 29 seta
<emphasis id="B9156F0BFFB9FFEB81E2DC90FC00FAAA" box="[924,937,1277,1299]" italics="true" pageId="2" pageNumber="3">d</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DD67FBD8FAAB" box="[937,1137,1276,1302]" pageId="2" pageNumber="3" value="1922-02-25">2 25 22</date>
25 27 39 seta
<geoCoordinate id="EE55D5DEFFB9FFEB81E3DD46FBD8FAF9" box="[925,1137,1322,1348]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="15" value="2.0208333">
<emphasis id="B9156F0BFFB9FFEB81E3DD46FC01FAF9" box="[925,936,1323,1344]" italics="true" pageId="2" pageNumber="3">e</emphasis>
1 60 75
</geoCoordinate>
103 126 168 seta
<emphasis id="B9156F0BFFB9FFEB81DCDD3AFC00FAD4" box="[930,937,1367,1389]" italics="true" pageId="2" pageNumber="3">f</emphasis>
1 51 65 83 112 146 genital opening Nd 21 32 40 70 analopening 20 32 39 45 54
</paragraph>
<paragraph id="8BDEB319FFB9FFEB82DEDDB5FA3BF872" blockId="2.[160,1427,415,1995]" pageId="2" pageNumber="3">
<taxonomicName id="4C61C89AFFB9FFEB82DEDDB5FDE0FA54" authority="Bodylength" authorityName="Bodylength" box="[160,585,1495,1517]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ugamaensis">
<emphasis id="B9156F0BFFB9FFEB82DEDDB5FF18FA54" box="[160,177,1496,1517]" italics="true" pageId="2" pageNumber="3">C</emphasis>
.
<emphasis id="B9156F0BFFB9FFEB82C0DDB5FE9CFA54" box="[190,309,1496,1517]" italics="true" pageId="2" pageNumber="3">ugamaensis</emphasis>
Bodylength
</taxonomicName>
159 211 244 310 304 Body width 96 125 144 188 153 Length of: sensillus 49 56 63 64 63 seta
<emphasis id="B9156F0BFFB9FFEB81E3DE39FC01F9D0" box="[925,936,1620,1641]" italics="true" pageId="2" pageNumber="3">c</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DE0CFBD8F9D0" box="[937,1137,1620,1645]" pageId="2" pageNumber="3" value="1927-01-17">1 17 27</date>
33 45 50 seta
<emphasis id="B9156F0BFFB9FFEB81E3DEECFC01F92F" box="[925,936,1665,1686]" italics="true" pageId="2" pageNumber="3">c</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DEE3FBD8F92F" box="[937,1137,1664,1690]" pageId="2" pageNumber="3" value="1915-03-20">3 20 15</date>
29 37 49 seta
<emphasis id="B9156F0BFFB9FFEB81E6DEC2FC19F97D" box="[920,944,1711,1732]" italics="true" pageId="2" pageNumber="3">cp</emphasis>
27 27 39 48 55 seta
<emphasis id="B9156F0BFFB9FFEB81E2DEBAFC00F954" box="[924,937,1751,1773]" italics="true" pageId="2" pageNumber="3">d</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DE88FBD8F955" box="[937,1137,1751,1777]" pageId="2" pageNumber="3" value="1918-01-17">1 17 18</date>
20 23 31 seta
<emphasis id="B9156F0BFFB9FFEB81E2DF69FC00F8A3" box="[924,937,1796,1818]" italics="true" pageId="2" pageNumber="3">d</emphasis>
<date id="FFDF95D9FFB9FFEB81D7DF7FFBD8F8A0" box="[937,1137,1796,1822]" pageId="2" pageNumber="3" value="1920-02-18">2 18 20</date>
25 32 46 seta
<geoCoordinate id="EE55D5DEFFB9FFEB81E3DF5EFBD9F8FE" box="[925,1136,1841,1867]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="15" value="2.0241666">
<emphasis id="B9156F0BFFB9FFEB81E3DF5EFC01F8F1" box="[925,936,1843,1864]" italics="true" pageId="2" pageNumber="3">e</emphasis>
1 60 87
</geoCoordinate>
95 133 155 seta
<emphasis id="B9156F0BFFB9FFEB81DCDF32FC00F8CC" box="[930,937,1887,1909]" italics="true" pageId="2" pageNumber="3">f</emphasis>
1 49 70 87 109 143 genital opening Nd 20 28 51 70 analopening 22 26 35 46 54
</paragraph>
</subSubSection>
<caption id="DF1EE391FFB8FFEA82E9D8FAFF68FF6B" ID-Table-UUID="DF1EE391FFB8FFEA82E9D8FAFF68FF6B" httpUri="http://table.plazi.org/id/DF1EE391FFB8FFEA82E9D8FAFF68FF6B" pageId="3" pageNumber="4" targetBox="[151,1436,238,590]" targetIsTable="true" targetPageId="3">
<paragraph id="8BDEB319FFB8FFEA82E9D8FAFF68FF6B" blockId="3.[151,1436,151,210]" pageId="3" pageNumber="4">
<emphasis id="B9156F0BFFB8FFEA82E9D8FAFEA0FF16" bold="true" box="[151,265,151,175]" pageId="3" pageNumber="4">TABLE 2.</emphasis>
Selected morphological characters of
<taxonomicName id="4C61C89AFFB8FFEA80D4D8F5FC7DFF16" box="[682,980,152,175]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB8FFEA80D4D8F5FC7DFF16" box="[682,980,152,175]" italics="true" pageId="3" pageNumber="4">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C61C89AFFB8FFEA819DD8F5FBFAFF16" box="[995,1107,152,175]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B9156F0BFFB8FFEA819DD8F5FC5CFF16" box="[995,1013,152,175]" italics="true" pageId="3" pageNumber="4">C</emphasis>
.
<emphasis id="B9156F0BFFB8FFEA867DD8F5FBFAFF16" box="[1027,1107,152,175]" italics="true" pageId="3" pageNumber="4">foliatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C61C89AFFB8FFEA861FD8F5FAABFF16" box="[1121,1282,152,175]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ugamaensis">
<emphasis id="B9156F0BFFB8FFEA861FD8F5FBDAFF16" box="[1121,1139,152,175]" italics="true" pageId="3" pageNumber="4">C</emphasis>
.
<emphasis id="B9156F0BFFB8FFEA86FFD8F5FAABFF16" box="[1153,1282,152,175]" italics="true" pageId="3" pageNumber="4">ugamaensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C61C89AFFB8FFEA8742D8F5FF12FF6B" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis id="B9156F0BFFB8FFEA8742D8F5FF12FF6B" italics="true" pageId="3" pageNumber="4">C. ponticus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BDEB319FFB8FFEA82D1D883FAB9FD96" pageId="3" pageNumber="4">
<table id="F96141B9FFB8001682E9D883FA35FDF7" box="[151,1436,238,590]" gridcols="4" gridrows="6" pageId="3" pageNumber="4">
<tr id="3551B15BFFB8001682E9D883FA35FE9A" box="[151,1436,238,291]" gridrow="0" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9D883FDE0FE9A" box="[151,585,238,291]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">
Morphological char-
<taxonomicName id="4C61C89AFFB8FFEA83BED882FDE0FEBC" box="[448,585,239,261]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB8FFEA83BED882FDE0FEBC" box="[448,585,239,261]" italics="true" pageId="3" pageNumber="4">C. reticulatus</emphasis>
</taxonomicName>
acters
</th>
<th id="7680D827FFB8001680B8D883FC99FE9A" box="[710,816,238,291]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="4C61C89AFFB8FFEA80B8D882FC99FEBC" box="[710,816,239,261]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B9156F0BFFB8FFEA80B8D882FD7EFEBD" box="[710,727,239,260]" italics="true" pageId="3" pageNumber="4">C</emphasis>
.
<emphasis id="B9156F0BFFB8FFEA809AD882FC99FEBC" box="[740,816,239,261]" italics="true" pageId="3" pageNumber="4">foliatus</emphasis>
</taxonomicName>
</th>
<th id="7680D827FFB8001681C7D883FBE7FE9A" box="[953,1102,238,291]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="4C61C89AFFB8FFEA81C7D882FBE7FEBC" box="[953,1102,239,261]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ugamaensis">
<emphasis id="B9156F0BFFB8FFEA81C7D882FC63FEBD" box="[953,970,239,260]" italics="true" pageId="3" pageNumber="4">C</emphasis>
.
<emphasis id="B9156F0BFFB8FFEA81A8D89DFBE7FEBC" box="[982,1102,240,261]" italics="true" pageId="3" pageNumber="4">ugamaensis</emphasis>
</taxonomicName>
</th>
<th id="7680D827FFB8001686B6D883FA35FE9A" box="[1224,1436,238,291]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="4C61C89AFFB8FFEA86B6D89FFA92FEBE" box="[1224,1339,242,263]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis id="B9156F0BFFB8FFEA86B6D89FFA92FEBE" box="[1224,1339,242,263]" italics="true" pageId="3" pageNumber="4">C. ponticus</emphasis>
</taxonomicName>
1
</th>
</tr>
<tr id="3551B15BFFB8001682E9D95BFA35FEF5" box="[151,1436,310,332]" gridrow="1" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="7680D827FFB8001682E9D95BFDE0FEF5" box="[151,585,310,332]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">Adult:</th>
</tr>
<tr id="3551B15BFFB8001682E9D90DFA35FE2D" box="[151,1436,352,404]" gridrow="2" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9D90DFDE0FE2D" box="[151,585,352,404]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">
Pairs of cilia on
<emphasis id="B9156F0BFFB8FFEA832BD90CFEC9FECF" box="[341,352,353,374]" italics="true" pageId="3" pageNumber="4">e</emphasis>
- 12–13 series setae
</th>
<td id="7680D827FFB8001680B8D90DFC99FE2D" box="[710,816,352,404]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">23–25</td>
<td id="7680D827FFB8001681C7D90DFBE7FE2D" box="[953,1102,352,404]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">16–18</td>
<td id="7680D827FFB8001686B6D90DFA35FE2D" box="[1224,1436,352,404]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">13–14</td>
</tr>
<tr id="3551B15BFFB8001682E9D9C5FA35FE65" box="[151,1436,424,476]" gridrow="3" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9D9C5FDE0FE65" box="[151,585,424,476]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">
Pairs of cilia on
<emphasis id="B9156F0BFFB8FFEA832BD9C5FEF5FE07" box="[341,348,424,446]" italics="true" pageId="3" pageNumber="4">f</emphasis>
- 10–11 series setae
</th>
<td id="7680D827FFB8001680B8D9C5FC99FE65" box="[710,816,424,476]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">22–24</td>
<td id="7680D827FFB8001681C7D9C5FBE7FE65" box="[953,1102,424,476]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">13–14</td>
<td id="7680D827FFB8001686B6D9C5FA35FE65" box="[1224,1436,424,476]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">10–11</td>
</tr>
<tr id="3551B15BFFB8001682E9D99DFA35FDBF" box="[151,1436,496,518]" gridrow="4" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9D99DFDE0FDBF" box="[151,585,496,518]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">Form of cerotegument Thick</th>
<td id="7680D827FFB8001680B8D99DFC99FDBF" box="[710,816,496,518]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">Thick</td>
<td id="7680D827FFB8001681C7D99DFBE7FDBF" box="[953,1102,496,518]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">Thin, granular</td>
<td id="7680D827FFB8001686B6D99DFA35FDBF" box="[1224,1436,496,518]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">Thin, granular</td>
</tr>
<tr id="3551B15BFFB8001682E9DA74FA35FDF7" box="[151,1436,537,590]" gridrow="5" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DA74FDE0FDF7" box="[151,585,537,590]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">Presence of cerotegu- Absent mental collar</th>
<td id="7680D827FFB8001680B8DA74FC99FDF7" box="[710,816,537,590]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">Present</td>
<td id="7680D827FFB8001681C7DA74FBE7FDF7" box="[953,1102,537,590]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">Absent</td>
<td id="7680D827FFB8001686B6DA74FA35FDF7" box="[1224,1436,537,590]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">Absent</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDEB319FFB8FFEA82D1DA0FFB53F954" pageId="3" pageNumber="4">
<table id="F96141B9FFB8001682E9DA0CFA35F948" box="[151,1436,609,1777]" gridcols="5" gridrows="19" pageId="3" pageNumber="4">
<tr id="3551B15BFFB8001682E9DA0CFA35FD2F" box="[151,1436,609,662]" gridrow="0" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DA0CFE3DFD2F" box="[151,404,609,662]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">Pattern of cuticle on pygidium</th>
<th id="7680D827FFB8001683C8DA0CFD3FFD2F" box="[438,662,609,662]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">Reticulate</th>
<th id="7680D827FFB8001680C2DA0CFC20FD2F" box="[700,905,609,662]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">With foveae</th>
<th id="7680D827FFB8001681D1DA0CFB3BFD2F" box="[943,1170,609,662]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">Small circles, semi- circles</th>
<th id="7680D827FFB8001686C1DA0CFA35FD2F" box="[1215,1436,609,662]" gridcol="4" gridrow="0" pageId="3" pageNumber="4">Foveae of various size</th>
</tr>
<tr id="3551B15BFFB8001682E9DAC4FA35FD59" box="[151,1436,681,736]" gridrow="1" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DAC4FE3DFD59" box="[151,404,681,736]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">
Shape of setae
<emphasis id="B9156F0BFFB8FFEA8339DAC7FEFBFD06" box="[327,338,682,703]" italics="true" pageId="3" pageNumber="4">c</emphasis>
1,
<emphasis id="B9156F0BFFB8FFEA8318DAC7FED8FD06" box="[358,369,682,703]" italics="true" pageId="3" pageNumber="4">c</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA82A4DAA7FF4EFD59" box="[218,231,714,736]" italics="true" pageId="3" pageNumber="4">d</emphasis>
-series
</th>
<td id="7680D827FFB8001683C8DAC4FD3FFD59" box="[438,662,681,736]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001680C2DAC4FC20FD59" box="[700,905,681,736]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001681D1DAC4FB3BFD59" box="[943,1170,681,736]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001686C1DAC4FA35FD59" box="[1215,1436,681,736]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">Bushy</td>
</tr>
<tr id="3551B15BFFB8001682E9DA9EFA35FC97" box="[151,1436,755,814]" gridrow="2" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DA9EFE3DFC97" box="[151,404,755,814]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">
Shape of setae
<emphasis id="B9156F0BFFB8FFEA8339DA99FEFAFCB3" box="[327,339,756,778]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1 and
<emphasis id="B9156F0BFFB8FFEA82D1DB7BFF15FC92" box="[175,188,790,811]" italics="true" pageId="3" pageNumber="4">p</emphasis>
1
</th>
<td id="7680D827FFB8001683C8DA9EFD3FFC97" box="[438,662,755,814]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">
Thicker than
<emphasis id="B9156F0BFFB8FFEA8039DA99FDFDFCB3" box="[583,596,756,778]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA83BEDB78FE65FC92" box="[448,460,789,811]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001680C2DA9EFC20FC97" box="[700,905,755,814]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">
Thicker than
<emphasis id="B9156F0BFFB8FFEA8135DA99FCFEFCB3" box="[843,855,756,778]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA80B8DB78FD7AFC92" box="[710,723,789,811]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001681D1DA9EFB3BFC97" box="[943,1170,755,814]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">
Thicker than
<emphasis id="B9156F0BFFB8FFEA8641DA99FBE5FCB3" box="[1087,1100,756,778]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA8601DA99FB22FCB3" box="[1151,1163,756,778]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001686C1DA9EFA35FC97" box="[1215,1436,755,814]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">
Similar as
<emphasis id="B9156F0BFFB8FFEA874ADA99FAE8FCB3" box="[1332,1345,756,778]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA86B6DB78FB7CFC92" box="[1224,1237,789,811]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
</tr>
<tr id="3551B15BFFB8001682E9DB2CFA35FCCC" box="[151,1436,833,885]" gridrow="3" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DB2CFE3DFCCC" box="[151,404,833,885]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">Setae on inner border of genital plate</th>
<td id="7680D827FFB8001683C8DB2CFD3FFCCC" box="[438,662,833,885]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">6 pairs</td>
<td id="7680D827FFB8001680C2DB2CFC20FCCC" box="[700,905,833,885]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">6 pairs</td>
<td id="7680D827FFB8001681D1DB2CFB3BFCCC" box="[943,1170,833,885]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">7 pairs</td>
<td id="7680D827FFB8001686C1DB2CFA35FCCC" box="[1215,1436,833,885]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">6 pairs</td>
</tr>
<tr id="3551B15BFFB8001682E9DBE4FA35FC26" box="[151,1436,905,927]" gridrow="4" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7680D827FFB8001682E9DBE4FE3DFC26" box="[151,404,905,927]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">Nymphs:</th>
</tr>
<tr id="3551B15BFFB8001682E9DBDFFA35FC5E" box="[151,1436,946,999]" gridrow="5" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DBDFFE3DFC5E" box="[151,404,946,999]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">
Pairs of cilia on
<emphasis id="B9156F0BFFB8FFEA832BDBDEFEC9FC71" box="[341,352,947,968]" italics="true" pageId="3" pageNumber="4">e</emphasis>
- series setae
</th>
<td id="7680D827FFB8001683C8DBDFFD3FFC5E" box="[438,662,946,999]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">8–9</td>
<td id="7680D827FFB8001680C2DBDFFC20FC5E" box="[700,905,946,999]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">19–23</td>
<td id="7680D827FFB8001681D1DBDFFB3BFC5E" box="[943,1170,946,999]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">14–15</td>
<td id="7680D827FFB8001686C1DBDFFA35FC5E" box="[1215,1436,946,999]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">18–20</td>
</tr>
<tr id="3551B15BFFB8001682E9DB97FA35FB96" box="[151,1436,1018,1071]" gridrow="6" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DB97FE3DFB96" box="[151,404,1018,1071]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">
Pairs of cilia on
<emphasis id="B9156F0BFFB8FFEA832BDB97FEF5FBA9" box="[341,348,1018,1040]" italics="true" pageId="3" pageNumber="4">f</emphasis>
- series setae
</th>
<td id="7680D827FFB8001683C8DB97FD3FFB96" box="[438,662,1018,1071]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">8–9</td>
<td id="7680D827FFB8001680C2DB97FC20FB96" box="[700,905,1018,1071]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">19–22</td>
<td id="7680D827FFB8001681D1DB97FB3BFB96" box="[943,1170,1018,1071]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">11–13</td>
<td id="7680D827FFB8001686C1DB97FA35FB96" box="[1215,1436,1018,1071]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">11–12</td>
</tr>
<tr id="3551B15BFFB8001682E9DC2FFA35FBE5" box="[151,1436,1090,1116]" gridrow="7" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DC2FFE3DFBE5" box="[151,404,1090,1116]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">
Basal cilia of seta
<emphasis id="B9156F0BFFB8FFEA8316DC2FFEC6FBE1" box="[360,367,1090,1112]" italics="true" pageId="3" pageNumber="4">f</emphasis>
1
</th>
<td id="7680D827FFB8001683C8DC2FFD3FFBE5" box="[438,662,1090,1116]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">
Longer than
<emphasis id="B9156F0BFFB8FFEA803FDC2EFDE5FBE1" box="[577,588,1091,1112]" italics="true" pageId="3" pageNumber="4">e</emphasis>
1
</td>
<td id="7680D827FFB8001680C2DC2FFC20FBE5" box="[700,905,1090,1116]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">
Similar to
<emphasis id="B9156F0BFFB8FFEA814FDC2EFC95FBE1" box="[817,828,1091,1112]" italics="true" pageId="3" pageNumber="4">e</emphasis>
1
</td>
<td id="7680D827FFB8001681D1DC2FFB3BFBE5" box="[943,1170,1090,1116]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">
Longer than
<emphasis id="B9156F0BFFB8FFEA8644DC2EFBECFBE1" box="[1082,1093,1091,1112]" italics="true" pageId="3" pageNumber="4">e</emphasis>
1
</td>
<td id="7680D827FFB8001686C1DC2FFA35FBE5" box="[1215,1436,1090,1116]" gridcol="4" gridrow="7" pageId="3" pageNumber="4">
Similar to
<emphasis id="B9156F0BFFB8FFEA874DDC2EFA97FBE1" box="[1331,1342,1091,1112]" italics="true" pageId="3" pageNumber="4">e</emphasis>
1
</td>
</tr>
<tr id="3551B15BFFB8001682E9DC03FA35FB1B" box="[151,1436,1134,1186]" gridrow="8" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DC03FE3DFB1B" box="[151,404,1134,1186]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">
Shape of setae of
<emphasis id="B9156F0BFFB8FFEA831DDC03FED9FB3D" box="[355,368,1134,1156]" italics="true" pageId="3" pageNumber="4">d</emphasis>
- series
</th>
<td id="7680D827FFB8001683C8DC03FD3FFB1B" box="[438,662,1134,1186]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001680C2DC03FC20FB1B" box="[700,905,1134,1186]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001681D1DC03FB3BFB1B" box="[943,1170,1134,1186]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001686C1DC03FA35FB1B" box="[1215,1436,1134,1186]" gridcol="4" gridrow="8" pageId="3" pageNumber="4">Bushy</td>
</tr>
<tr id="3551B15BFFB8001682E9DCDBFA35FB69" box="[151,1436,1206,1232]" gridrow="9" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DCDBFE3DFB69" box="[151,404,1206,1232]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">
Shape of seta
<emphasis id="B9156F0BFFB8FFEA8342DCDAFEE1FB75" box="[316,328,1207,1228]" italics="true" pageId="3" pageNumber="4">p</emphasis>
1
</th>
<td id="7680D827FFB8001683C8DCDBFD3FFB69" box="[438,662,1206,1232]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">Rather pinnate</td>
<td id="7680D827FFB8001680C2DCDBFC20FB69" box="[700,905,1206,1232]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">Bushy</td>
<td id="7680D827FFB8001681D1DCDBFB3BFB69" box="[943,1170,1206,1232]" gridcol="3" gridrow="9" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001686C1DCDBFA35FB69" box="[1215,1436,1206,1232]" gridcol="4" gridrow="9" pageId="3" pageNumber="4">Bushy</td>
</tr>
<tr id="3551B15BFFB8001682E9DC8FFA35FAAF" box="[151,1436,1250,1302]" gridrow="10" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DC8FFE3DFAAF" box="[151,404,1250,1302]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">
Shape of
<emphasis id="B9156F0BFFB8FFEA8370DC8FFEB3FB41" box="[270,282,1250,1272]" italics="true" pageId="3" pageNumber="4">h</emphasis>
-series setae
</th>
<td id="7680D827FFB8001683C8DC8FFD3FFAAF" box="[438,662,1250,1302]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001680C2DC8FFC20FAAF" box="[700,905,1250,1302]" gridcol="2" gridrow="10" pageId="3" pageNumber="4">Bushy</td>
<td id="7680D827FFB8001681D1DC8FFB3BFAAF" box="[943,1170,1250,1302]" gridcol="3" gridrow="10" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001686C1DC8FFA35FAAF" box="[1215,1436,1250,1302]" gridcol="4" gridrow="10" pageId="3" pageNumber="4">Bushy</td>
</tr>
<tr id="3551B15BFFB8001682E9DD47FA35FAFD" box="[151,1436,1322,1348]" gridrow="11" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DD47FE3DFAFD" box="[151,404,1322,1348]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">Length of basal cilia</th>
<td id="7680D827FFB8001683C8DD47FD3FFAFD" box="[438,662,1322,1348]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">
<emphasis id="B9156F0BFFB8FFEA83BEDD47FE65FAF9" box="[448,460,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1 longer than
<emphasis id="B9156F0BFFB8FFEA802DDD47FDC9FAF9" box="[595,608,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2,
<emphasis id="B9156F0BFFB8FFEA800EDD47FDD4FAF9" box="[624,637,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001680C2DD47FC20FAFD" box="[700,905,1322,1348]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">
<emphasis id="B9156F0BFFB8FFEA80B8DD47FD7AFAF9" box="[710,723,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1 longer than
<emphasis id="B9156F0BFFB8FFEA8126DD47FCCCFAF9" box="[856,869,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2,
<emphasis id="B9156F0BFFB8FFEA810BDD47FC2BFAF9" box="[885,898,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001681D1DD47FB3BFAFD" box="[943,1170,1322,1348]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">
<emphasis id="B9156F0BFFB8FFEA81C7DD47FC6CFAF9" box="[953,965,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1,
<emphasis id="B9156F0BFFB8FFEA81A7DD47FC4CFAF9" box="[985,997,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 longer than
<emphasis id="B9156F0BFFB8FFEA8612DD47FBD0FAF9" box="[1132,1145,1322,1344]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001686C1DD47FA35FAFD" box="[1215,1436,1322,1348]" gridcol="4" gridrow="11" pageId="3" pageNumber="4">Similar</td>
</tr>
<tr id="3551B15BFFB8001682E9DD36FA35FA23" box="[151,1436,1371,1434]" gridrow="12" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DD36FE3DFA23" box="[151,404,1371,1434]" gridcol="0" gridrow="12" pageId="3" pageNumber="4">
Shape of setae
<emphasis id="B9156F0BFFB8FFEA8334DD36FEFEFAC8" box="[330,343,1371,1393]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1 and
<emphasis id="B9156F0BFFB8FFEA82CCDDEFFF17FA2E" box="[178,190,1410,1431]" italics="true" pageId="3" pageNumber="4">p</emphasis>
1
</th>
<td id="7680D827FFB8001683C8DD36FD3FFA23" box="[438,662,1371,1434]" gridcol="1" gridrow="12" pageId="3" pageNumber="4">
Thicker than
<emphasis id="B9156F0BFFB8FFEA8043DD36FDE3FAC8" box="[573,586,1371,1393]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA80FCDD36FD26FAC8" box="[642,655,1371,1393]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001680C2DD36FC20FA23" box="[700,905,1371,1434]" gridcol="2" gridrow="12" pageId="3" pageNumber="4">
Thicker than
<emphasis id="B9156F0BFFB8FFEA813ADD36FCF8FAC8" box="[836,849,1371,1393]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA80C2DDEDFD60FA2F" box="[700,713,1408,1430]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001681D1DD36FB3BFA23" box="[943,1170,1371,1434]" gridcol="3" gridrow="12" pageId="3" pageNumber="4">
Thicker than
<emphasis id="B9156F0BFFB8FFEA8648DD36FBEAFAC8" box="[1078,1091,1371,1393]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA8605DD36FB21FAC8" box="[1147,1160,1371,1393]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
<td id="7680D827FFB8001686C1DD36FA35FA23" box="[1215,1436,1371,1434]" gridcol="4" gridrow="12" pageId="3" pageNumber="4">
Similar as
<emphasis id="B9156F0BFFB8FFEA8755DD36FA91FAC8" box="[1323,1336,1371,1393]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and
<emphasis id="B9156F0BFFB8FFEA870EDD36FAD4FAC8" box="[1392,1405,1371,1393]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3
</td>
</tr>
<tr id="3551B15BFFB8001682E9DDDCFA35FA7F" box="[151,1436,1457,1478]" gridrow="13" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7680D827FFB8001682E9DDDCFE3DFA7F" box="[151,404,1457,1478]" gridcol="0" gridrow="13" pageId="3" pageNumber="4">Larva:</th>
</tr>
<tr id="3551B15BFFB8001682E9DDB1FA35F9AD" box="[151,1436,1500,1556]" gridrow="14" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DDB1FE3DF9AD" box="[151,404,1500,1556]" gridcol="0" gridrow="14" pageId="3" pageNumber="4">
Pairs of cilia on setae of
<emphasis id="B9156F0BFFB8FFEA82B5DD92FF7FF9AD" box="[203,214,1535,1556]" italics="true" pageId="3" pageNumber="4">e</emphasis>
-series
</th>
<td id="7680D827FFB8001683C8DDB1FD3FF9AD" box="[438,662,1500,1556]" gridcol="1" gridrow="14" pageId="3" pageNumber="4">6–7</td>
<td id="7680D827FFB8001680C2DDB1FC20F9AD" box="[700,905,1500,1556]" gridcol="2" gridrow="14" pageId="3" pageNumber="4">10–11</td>
<td id="7680D827FFB8001681D1DDB1FB3BF9AD" box="[943,1170,1500,1556]" gridcol="3" gridrow="14" pageId="3" pageNumber="4">15–16</td>
<td id="7680D827FFB8001686C1DDB1FA35F9AD" box="[1215,1436,1500,1556]" gridcol="4" gridrow="14" pageId="3" pageNumber="4">17–18</td>
</tr>
<tr id="3551B15BFFB8001682E9DE47FA35F9D8" box="[151,1436,1578,1633]" gridrow="15" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DE47FE3DF9D8" box="[151,404,1578,1633]" gridcol="0" gridrow="15" pageId="3" pageNumber="4">
Pairs of cilia on setae of
<emphasis id="B9156F0BFFB8FFEA82B5DE26FF7BF9D8" box="[203,210,1611,1633]" italics="true" pageId="3" pageNumber="4">f</emphasis>
-series
</th>
<td id="7680D827FFB8001683C8DE47FD3FF9D8" box="[438,662,1578,1633]" gridcol="1" gridrow="15" pageId="3" pageNumber="4">6–7</td>
<td id="7680D827FFB8001680C2DE47FC20F9D8" box="[700,905,1578,1633]" gridcol="2" gridrow="15" pageId="3" pageNumber="4">10–11</td>
<td id="7680D827FFB8001681D1DE47FB3BF9D8" box="[943,1170,1578,1633]" gridcol="3" gridrow="15" pageId="3" pageNumber="4">11–12</td>
<td id="7680D827FFB8001686C1DE47FA35F9D8" box="[1215,1436,1578,1633]" gridcol="4" gridrow="15" pageId="3" pageNumber="4">12–13</td>
</tr>
<tr id="3551B15BFFB8001682E9DE1AFA35F928" box="[151,1436,1655,1681]" gridrow="16" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DE1AFE3DF928" box="[151,404,1655,1681]" gridcol="0" gridrow="16" pageId="3" pageNumber="4">
Basal cilia of seta
<emphasis id="B9156F0BFFB8FFEA8317DE1AFED9F934" box="[361,368,1655,1677]" italics="true" pageId="3" pageNumber="4">f</emphasis>
1
</th>
<td id="7680D827FFB8001683C8DE1AFD3FF928" box="[438,662,1655,1681]" gridcol="1" gridrow="16" pageId="3" pageNumber="4">
Longer than
<emphasis id="B9156F0BFFB8FFEA8046DE15FDEAF934" box="[568,579,1656,1677]" italics="true" pageId="3" pageNumber="4">e</emphasis>
1
</td>
<td id="7680D827FFB8001680C2DE1AFC20F928" box="[700,905,1655,1681]" gridcol="2" gridrow="16" pageId="3" pageNumber="4">
Similar to
<emphasis id="B9156F0BFFB8FFEA8159DE15FC9BF934" box="[807,818,1656,1677]" italics="true" pageId="3" pageNumber="4">e</emphasis>
1
</td>
<td id="7680D827FFB8001681D1DE1AFB3BF928" box="[943,1170,1655,1681]" gridcol="3" gridrow="16" pageId="3" pageNumber="4">
Longer than
<emphasis id="B9156F0BFFB8FFEA864EDE15FB92F934" box="[1072,1083,1656,1677]" italics="true" pageId="3" pageNumber="4">e</emphasis>
1
</td>
<td id="7680D827FFB8001686C1DE1AFA35F928" box="[1215,1436,1655,1681]" gridcol="4" gridrow="16" pageId="3" pageNumber="4">
Longer than
<emphasis id="B9156F0BFFB8FFEA873EDE15FAE2F934" box="[1344,1355,1656,1677]" italics="true" pageId="3" pageNumber="4">e</emphasis>
1
</td>
</tr>
<tr id="3551B15BFFB8001682E9DECAFA35F978" box="[151,1436,1703,1729]" gridrow="17" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DECAFE3DF978" box="[151,404,1703,1729]" gridcol="0" gridrow="17" pageId="3" pageNumber="4">
Shape of seta
<emphasis id="B9156F0BFFB8FFEA8345DECAFEEEF904" box="[315,327,1703,1725]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1
</th>
<td id="7680D827FFB8001683C8DECAFD3FF978" box="[438,662,1703,1729]" gridcol="1" gridrow="17" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001680C2DECAFC20F978" box="[700,905,1703,1729]" gridcol="2" gridrow="17" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001681D1DECAFB3BF978" box="[943,1170,1703,1729]" gridcol="3" gridrow="17" pageId="3" pageNumber="4">Pinnate</td>
<td id="7680D827FFB8001686C1DECAFA35F978" box="[1215,1436,1703,1729]" gridcol="4" gridrow="17" pageId="3" pageNumber="4">Bushy</td>
</tr>
<tr id="3551B15BFFB8001682E9DEBAFA35F948" box="[151,1436,1751,1777]" gridrow="18" pageId="3" pageNumber="4">
<th id="7680D827FFB8001682E9DEBAFE3DF948" box="[151,404,1751,1777]" gridcol="0" gridrow="18" pageId="3" pageNumber="4">
Pairs of cilia on seta
<emphasis id="B9156F0BFFB8FFEA83FFDEBAFE27F954" box="[385,398,1751,1773]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1
</th>
<td id="7680D827FFB8001683C8DEBAFD3FF948" box="[438,662,1751,1777]" gridcol="1" gridrow="18" pageId="3" pageNumber="4">4–5</td>
<td id="7680D827FFB8001680C2DEBAFC20F948" box="[700,905,1751,1777]" gridcol="2" gridrow="18" pageId="3" pageNumber="4">8</td>
<td id="7680D827FFB8001681D1DEBAFB3BF948" box="[943,1170,1751,1777]" gridcol="3" gridrow="18" pageId="3" pageNumber="4">6</td>
<td id="7680D827FFB8001686C1DEBAFA35F948" box="[1215,1436,1751,1777]" gridcol="4" gridrow="18" pageId="3" pageNumber="4">Many</td>
</tr>
</table>
</paragraph>
<subSubSection id="C37BE092FFB8FFE282E9DF7CFE38FCD7" lastPageId="11" lastPageNumber="12" pageId="3" pageNumber="4" type="key">
<paragraph id="8BDEB319FFB8FFEF82E9DF7CFB5AFE84" blockId="3.[151,593,1809,1832]" lastBlockId="6.[151,1436,151,323]" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
<keyLead id="3090102CFFB8FFEA82E9DF7CFDF8F891" box="[151,593,1809,1832]" pageId="3" pageNumber="4">
<key id="FFF0E428FFB8FFEA82E9DF7CFDF8F891" box="[151,593,1809,1832]" pageId="3" pageNumber="4">
<keyStep id="3095ABBCFFB8FFEA82E9DF7CFDF8F891" box="[151,593,1809,1832]" pageId="3" pageNumber="4">
1 According to
<bibRefCitation id="EFF0CEE8FFB8FFEA835ADF7EFDF8F891" author="Seniczak" box="[292,593,1811,1832]" pageId="3" pageNumber="4" refString="Seniczak, S. &amp; Seniczak, A. (2009 a) Differentiation of body form of Protoplophoroidea (Acari: Oribatida) in the light of ontogeny of three species. Journal of Natural History, 44 (7), 389 - 419." type="journal article" year="2009" yearSuffix="a">Seniczak and Seniczak (2009a)</bibRefCitation>
</keyStep>
</key>
</keyLead>
Gastronotum with 14 pairs of setae (
<figureCitation id="135AAF9CFFBEFFEC8016DF39FD1BF8D5" box="[616,690,1876,1900]" captionStart="FIGURE 4" captionStartId="7.[151,250,1957,1980]" captionTargetBox="[299,1294,216,1912]" captionTargetId="figure@7.[286,1301,199,1919]" captionTargetPageId="7" captionText="FIGURE 4. Cosmochthonius reticulatus, larva, dorsal aspect, legs partially drawn, scale bar 50 μm." httpUri="https://zenodo.org/record/278730/files/figure.png" pageId="5" pageNumber="6">Figs 4</figureCitation>
,
<figureCitation id="135AAF9CFFBEFFEC80C1DF39FD64F8D5" box="[703,717,1876,1900]" captionStart="FIGURE 5" captionStartId="8.[151,250,1806,1829]" captionTargetBox="[156,1432,213,1771]" captionTargetId="figure@8.[151,1436,193,1785]" captionTargetPageId="8" captionText="FIGURE 5. Cosmochthonius reticulatus, lateral aspect, legs partially drawn, scale bars 50 μm. (A) Larva; (B) tritonymph." httpUri="https://zenodo.org/record/278731/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
A), including inguinal seta
<emphasis id="B9156F0BFFBEFFEC867BDF38FBBAF8D4" box="[1029,1043,1877,1901]" italics="true" pageId="5" pageNumber="6">h</emphasis>
4 positioned anterior to paraproctal valves (segment PS). Gastronotum with three transverse scissures, which divide it into four parts. Plate
<emphasis id="B9156F0BFFBEFFEC873EDF17FAC9F828" box="[1344,1376,1914,1937]" italics="true" pageId="5" pageNumber="6">Na</emphasis>
with four pairs of setae:
<emphasis id="B9156F0BFFBEFFEC830CDFF6FED7F80B" box="[370,382,1947,1970]" italics="true" pageId="5" pageNumber="6">c</emphasis>
1–
<emphasis id="B9156F0BFFBEFFEC83EADFF6FE09F80B" box="[404,416,1947,1970]" italics="true" pageId="5" pageNumber="6">c</emphasis>
<quantity id="4C991EFCFFBEFFEC83DEDFC4FE6FF80B" box="[416,454,1946,1975]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="6" unit="in" value="3.0">3 in</quantity>
anterior row,
<emphasis id="B9156F0BFFBEFFEC801ADFF6FDD6F80B" box="[612,639,1947,1970]" italics="true" pageId="5" pageNumber="6">cp</emphasis>
in posterior row; all setae rather long, and barbed. Plate
<emphasis id="B9156F0BFFBEFFEC877CDFF6FA80F80B" box="[1282,1321,1947,1970]" italics="true" pageId="5" pageNumber="6">Nm</emphasis>
1 with two pairs of setae (
<emphasis id="B9156F0BFFBEFFEC8340DFD3FEE5F86F" box="[318,332,1982,2006]" italics="true" pageId="5" pageNumber="6">d</emphasis>
1,
<emphasis id="B9156F0BFFBEFFEC831ADFD3FEDBF86F" box="[356,370,1982,2006]" italics="true" pageId="5" pageNumber="6">d</emphasis>
2), shorter than
<emphasis id="B9156F0BFFBEFFEC805EDFD2FD85F86F" box="[544,556,1983,2006]" italics="true" pageId="5" pageNumber="6">c</emphasis>
-series. Setae of
<emphasis id="B9156F0BFFBEFFEC8097DFD2FD5CF86F" box="[745,757,1983,2006]" italics="true" pageId="5" pageNumber="6">e</emphasis>
- and
<emphasis id="B9156F0BFFBEFFEC8146DFD3FCE9F86F" box="[824,832,1982,2006]" italics="true" pageId="5" pageNumber="6">f</emphasis>
-series hypertrophied, pinnate, inserted on intercalary sclerites between plates
<emphasis id="B9156F0BFFBDFFEF83D7D8F4FE79FF09" box="[425,464,153,176]" italics="true" pageId="6" pageNumber="7">Nm</emphasis>
1 and
<emphasis id="B9156F0BFFBDFFEF806ED8F4FD9EFF09" box="[528,567,153,176]" italics="true" pageId="6" pageNumber="7">Nm</emphasis>
2, and
<emphasis id="B9156F0BFFBDFFEF8001D8F4FD0FFF09" box="[639,678,153,176]" italics="true" pageId="6" pageNumber="7">Nm</emphasis>
2 and pygidium (
<emphasis id="B9156F0BFFBDFFEF8121D8F4FCD4FF09" box="[863,893,153,176]" italics="true" pageId="6" pageNumber="7">Py</emphasis>
), respectively; each seta with 6–7 pairs of cilia. Pygidium weakly developed, with four pairs of slightly curved and barbed setae of
<emphasis id="B9156F0BFFBDFFEF8648D8D1FBEDFF6D" box="[1078,1092,188,212]" italics="true" pageId="6" pageNumber="7">h</emphasis>
-series, length decreasing from
<emphasis id="B9156F0BFFBDFFEF82E9D8B0FF0CFF4C" box="[151,165,221,245]" italics="true" pageId="6" pageNumber="7">h</emphasis>
1 to
<emphasis id="B9156F0BFFBDFFEF82B0D8B0FF75FF4C" box="[206,220,221,245]" italics="true" pageId="6" pageNumber="7">h</emphasis>
4 (
<figureCitation id="135AAF9CFFBDFFEF8291D8B0FE9FFF4C" box="[239,310,221,245]" captionStart="FIGURE 5" captionStartId="8.[151,250,1806,1829]" captionTargetBox="[156,1432,213,1771]" captionTargetId="figure@8.[151,1436,193,1785]" captionTargetPageId="8" captionText="FIGURE 5. Cosmochthonius reticulatus, lateral aspect, legs partially drawn, scale bars 50 μm. (A) Larva; (B) tritonymph." httpUri="https://zenodo.org/record/278731/files/figure.png" pageId="6" pageNumber="7">Figs 5</figureCitation>
A, 6A). Seta
<emphasis id="B9156F0BFFBDFFEF83B9D8B0FE7CFF4C" box="[455,469,221,245]" italics="true" pageId="6" pageNumber="7">h</emphasis>
1 with 4–5 cilia, other setae barbed. Paraproctal valves with four pairs of barbed setae, similar in shape to
<emphasis id="B9156F0BFFBDFFEF830DD96CFE28FEA0" box="[371,385,257,281]" italics="true" pageId="6" pageNumber="7">h</emphasis>
4. Cupule
<emphasis id="B9156F0BFFBDFFEF838FD96FFDAEFEA0" box="[497,519,258,281]" italics="true" pageId="6" pageNumber="7">ia</emphasis>
posteroventral to seta
<emphasis id="B9156F0BFFBDFFEF8172D96FFCB1FEA0" box="[780,792,258,281]" italics="true" pageId="6" pageNumber="7">c</emphasis>
3, cupule
<emphasis id="B9156F0BFFBDFFEF81FDD96FFC36FEA0" box="[899,927,258,281]" italics="true" pageId="6" pageNumber="7">im</emphasis>
posteroventral to seta
<emphasis id="B9156F0BFFBDFFEF86DDD96FFB14FEA0" box="[1187,1213,258,281]" italics="true" pageId="6" pageNumber="7">cp</emphasis>
, cupule
<emphasis id="B9156F0BFFBDFFEF875ED96FFA9FFEA0" box="[1312,1334,258,281]" italics="true" pageId="6" pageNumber="7">ip</emphasis>
between setae
<emphasis id="B9156F0BFFBDFFEF82A8D94BFF77FE87" box="[214,222,294,318]" italics="true" pageId="6" pageNumber="7">f</emphasis>
2 and
<emphasis id="B9156F0BFFBDFFEF8365D948FE80FE84" box="[283,297,293,317]" italics="true" pageId="6" pageNumber="7">h</emphasis>
3, cupule
<emphasis id="B9156F0BFFBDFFEF83EFD948FE0EFE84" box="[401,423,293,317]" italics="true" pageId="6" pageNumber="7">ih</emphasis>
posterior to seta
<emphasis id="B9156F0BFFBDFFEF8016D948FDDFFE84" box="[616,630,293,317]" italics="true" pageId="6" pageNumber="7">h</emphasis>
4 (
<figureCitation id="135AAF9CFFBDFFEF80F3D948FD79FE84" box="[653,720,293,317]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1966]" captionTargetBox="[361,1226,228,1907]" captionTargetId="figure@9.[343,1243,213,1920]" captionTargetPageId="9" captionText="FIGURE 6. Cosmochthonius reticulatus, scale bars 50 μm. (A) Anal region of larva; (B) anogenital region of tritonymph." httpUri="https://zenodo.org/record/278732/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
A). Ventral parts of gastronotum weakly striated.
</paragraph>
<caption id="DF1EE391FFBFFFED82E9DFBFFAF9F85E" box="[151,1360,2002,2025]" httpUri="https://zenodo.org/record/278727/files/figure.png" pageId="4" pageNumber="5" targetBox="[252,1260,198,1984]" targetPageId="4">
<paragraph id="8BDEB319FFBFFFED82E9DFBFFAF9F85E" blockId="4.[151,1360,2002,2025]" box="[151,1360,2002,2025]" pageId="4" pageNumber="5">
<emphasis id="B9156F0BFFBFFFED82E9DFBFFEA5F851" bold="true" box="[151,268,2002,2025]" pageId="4" pageNumber="5">FIGURE 1</emphasis>
.
<taxonomicName id="4C61C89AFFBFFFED8367DFBEFD9AF850" ID-CoL="7SN9R" box="[281,563,2003,2025]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFBFFFED8367DFBEFD9AF850" box="[281,563,2003,2025]" italics="true" pageId="4" pageNumber="5">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
, adult with cerotegument, dorsal aspect, legs partially drawn, scale bar 50 μm.
</paragraph>
</caption>
<caption id="DF1EE391FFBEFFEC82E9DEB4FF56F8B7" httpUri="https://zenodo.org/record/278728/files/figure.png" pageId="5" pageNumber="6" targetBox="[289,1303,182,1718]" targetPageId="5">
<paragraph id="8BDEB319FFBEFFEC82E9DEB4FF56F8B7" blockId="5.[151,1436,1753,1806]" pageId="5" pageNumber="6">
<emphasis id="B9156F0BFFBEFFEC82E9DEB4FEA6F956" bold="true" box="[151,271,1753,1776]" pageId="5" pageNumber="6">FIGURE 2</emphasis>
.
<taxonomicName id="4C61C89AFFBEFFEC835EDEB7FD97F949" ID-CoL="7SN9R" box="[288,574,1754,1776]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFBEFFEC835EDEB7FD97F949" box="[288,574,1754,1776]" italics="true" pageId="5" pageNumber="6">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
, adult. (A) Notogaster, cerotegument removed, scale bar 50 μm; (B) fragment of pygidium.
</paragraph>
</caption>
<caption id="DF1EE391FFBDFFEF82E9DF45FB13F884" box="[151,1210,1832,1855]" httpUri="https://zenodo.org/record/278729/files/figure.png" pageId="6" pageNumber="7" targetBox="[155,1426,405,1811]" targetPageId="6">
<paragraph id="8BDEB319FFBDFFEF82E9DF45FB13F884" blockId="6.[151,1210,1832,1855]" box="[151,1210,1832,1855]" pageId="6" pageNumber="7">
<emphasis id="B9156F0BFFBDFFEF82E9DF45FEBBF887" bold="true" box="[151,274,1832,1855]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
<taxonomicName id="4C61C89AFFBDFFEF8367DF44FD9AF886" ID-CoL="7SN9R" box="[281,563,1833,1855]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFBDFFEF8367DF44FD9AF886" box="[281,563,1833,1855]" italics="true" pageId="6" pageNumber="7">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
, adult, anogenital region, legs partially drawn, scale bar 50 μm.
</paragraph>
</caption>
<caption id="DF1EE391FFBCFFEE82E9DFC8FB24F803" box="[151,1165,1957,1980]" httpUri="https://zenodo.org/record/278730/files/figure.png" pageId="7" pageNumber="8" targetBox="[299,1294,216,1912]" targetPageId="7">
<paragraph id="8BDEB319FFBCFFEE82E9DFC8FB24F803" blockId="7.[151,1165,1957,1980]" box="[151,1165,1957,1980]" pageId="7" pageNumber="8">
<emphasis id="B9156F0BFFBCFFEE82E9DFC8FEBAF802" bold="true" box="[151,275,1957,1980]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
<taxonomicName id="4C61C89AFFBCFFEE8367DFCBFD9AF805" ID-CoL="7SN9R" box="[281,563,1958,1980]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFBCFFEE8367DFCBFD9AF805" box="[281,563,1958,1980]" italics="true" pageId="7" pageNumber="8">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
, larva, dorsal aspect, legs partially drawn, scale bar 50 μm.
</paragraph>
</caption>
<caption id="DF1EE391FFB3FFE182E9DF63FAD9F89D" box="[151,1392,1805,1829]" httpUri="https://zenodo.org/record/278731/files/figure.png" pageId="8" pageNumber="9" targetBox="[156,1432,213,1771]" targetPageId="8">
<paragraph id="8BDEB319FFB3FFE182E9DF63FAD9F89D" blockId="8.[151,1392,1805,1829]" box="[151,1392,1805,1829]" pageId="8" pageNumber="9">
<emphasis id="B9156F0BFFB3FFE182E9DF63FEBBF89D" bold="true" box="[151,274,1806,1829]" pageId="8" pageNumber="9">FIGURE 5.</emphasis>
<taxonomicName id="4C61C89AFFB3FFE18367DF63FD9AF89D" ID-CoL="7SN9R" box="[281,563,1806,1828]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB3FFE18367DF63FD9AF89D" box="[281,563,1806,1828]" italics="true" pageId="8" pageNumber="9">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
, lateral aspect, legs partially drawn, scale bars 50 μm. (A) Larva; (B) tritonymph.
</paragraph>
</caption>
<caption id="DF1EE391FFB2FFE082E9DFFAFAC5F814" box="[151,1388,1942,1966]" httpUri="https://zenodo.org/record/278732/files/figure.png" pageId="9" pageNumber="10" targetBox="[361,1226,228,1907]" targetPageId="9">
<paragraph id="8BDEB319FFB2FFE082E9DFFAFAC5F814" blockId="9.[151,1388,1942,1966]" box="[151,1388,1942,1966]" pageId="9" pageNumber="10">
<emphasis id="B9156F0BFFB2FFE082E9DFFAFEBBF814" bold="true" box="[151,274,1943,1966]" pageId="9" pageNumber="10">FIGURE 6.</emphasis>
<taxonomicName id="4C61C89AFFB2FFE08367DFFAFD9AF814" ID-CoL="7SN9R" box="[281,563,1943,1965]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB2FFE08367DFFAFD9AF814" box="[281,563,1943,1965]" italics="true" pageId="9" pageNumber="10">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
, scale bars 50 μm. (A) Anal region of larva; (B) anogenital region of tritonymph.
</paragraph>
</caption>
<paragraph id="8BDEB319FFB1FFE282B9D8FAFB89FE67" blockId="10.[151,1436,151,284]" lastBlockId="11.[151,1437,151,1420]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Prodorsum of tritonymph (
<figureCitation id="135AAF9CFFB1FFE3838DD8FAFD9EFF16" box="[499,567,151,175]" captionStart="FIGURE 7" captionStartId="10.[151,250,1996,2019]" captionTargetBox="[371,1238,331,1972]" captionTargetId="figure@10.[344,1244,310,1978]" captionTargetPageId="10" captionText="FIGURE 7. Cosmochthonius reticulatus, tritonymph, dorsal aspect, legs partially drawn, scale bar 50 μm." httpUri="https://zenodo.org/record/278733/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
) relatively smaller, and
<emphasis id="B9156F0BFFB1FFE38138D8F4FCCDFF09" box="[838,868,153,176]" italics="true" pageId="10" pageNumber="11">Py</emphasis>
longer than in larva. Rostrum with small dens and two transverse rows of small, thin fenestrate areas. Prodorsal setae
<emphasis id="B9156F0BFFB1FFE381F1D8D0FC0CFF6D" box="[911,933,189,212]" italics="true" pageId="10" pageNumber="11">ro</emphasis>
,
<emphasis id="B9156F0BFFB1FFE381C8D8D1FC63FF6D" box="[950,970,188,212]" italics="true" pageId="10" pageNumber="11">le</emphasis>
,
<emphasis id="B9156F0BFFB1FFE381A4D8D0FC59FF6D" box="[986,1008,189,212]" italics="true" pageId="10" pageNumber="11">in</emphasis>
and
<emphasis id="B9156F0BFFB1FFE38656D8D0FBE5FF6D" box="[1064,1100,189,212]" italics="true" pageId="10" pageNumber="11">exs</emphasis>
bushy, biramous, with longer single, and bifurcate spines than in larva; seta
<emphasis id="B9156F0BFFB1FFE380DBD88CFD6FFF41" box="[677,710,225,248]" italics="true" pageId="10" pageNumber="11">exi</emphasis>
barbed. Bothridium and sensillus as in larva. Gastronotum with 16 pairs of setae, including hypertrophied, and pinnate setae of
<emphasis id="B9156F0BFFB1FFE38123D968FCC0FEA5" box="[861,873,261,284]" italics="true" pageId="10" pageNumber="11">e</emphasis>
- and
<emphasis id="B9156F0BFFB1FFE381D4D969FC1BFEA5" box="[938,946,260,284]" italics="true" pageId="10" pageNumber="11">f</emphasis>
-series, inserted on intercalary sclerites, and curved posterior (
<figureCitation id="135AAF9CFFB0FFE2831CD8FAFE0EFF16" box="[354,423,151,175]" captionStart="FIGURE 5" captionStartId="8.[151,250,1806,1829]" captionTargetBox="[156,1432,213,1771]" captionTargetId="figure@8.[151,1436,193,1785]" captionTargetPageId="8" captionText="FIGURE 5. Cosmochthonius reticulatus, lateral aspect, legs partially drawn, scale bars 50 μm. (A) Larva; (B) tritonymph." httpUri="https://zenodo.org/record/278731/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
B); other setae distinctly shorter and barbed. Hypertrophied setae with 8–9 pairs of cilia (
<figureCitation id="135AAF9CFFB0FFE282E1D8D1FF48FF6D" box="[159,225,188,212]" captionStart="FIGURE 8" captionStartId="12.[151,250,1693,1716]" captionTargetBox="[162,1423,187,1671]" captionTargetId="figure@12.[151,1436,169,1688]" captionTargetPageId="12" captionText="FIGURE 8. Hypertrophied setae of tritonymph, scale bars 50 μm. (A) Cosmochthonius reticulatus; (B) C. foliatus." httpUri="https://zenodo.org/record/278734/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
A), basal cilia of
<emphasis id="B9156F0BFFB0FFE283D8D8D1FE07FF6D" box="[422,430,188,212]" italics="true" pageId="11" pageNumber="12">f</emphasis>
-series longer than those of
<emphasis id="B9156F0BFFB0FFE28096D8D0FD5DFF6D" box="[744,756,189,212]" italics="true" pageId="11" pageNumber="12">e</emphasis>
-series; length of cilia decreasing from basal to distal part of setae. Seta
<emphasis id="B9156F0BFFB0FFE28369D88CFE8AFF41" box="[279,291,225,248]" italics="true" pageId="11" pageNumber="12">c</emphasis>
3 bushy and shorter than other pinnate setae of
<emphasis id="B9156F0BFFB0FFE28141D88CFCE2FF41" box="[831,843,225,248]" italics="true" pageId="11" pageNumber="12">c</emphasis>
-series, setae of
<emphasis id="B9156F0BFFB0FFE2867BD88DFBBAFF41" box="[1029,1043,224,248]" italics="true" pageId="11" pageNumber="12">d</emphasis>
-series shorter than
<emphasis id="B9156F0BFFB0FFE2868AD88CFAA9FF41" box="[1268,1280,225,248]" italics="true" pageId="11" pageNumber="12">c</emphasis>
3, and pinnate (Fig. 9A). Pygidium with setae of
<emphasis id="B9156F0BFFB0FFE2806AD96AFD8BFEA6" box="[532,546,263,287]" italics="true" pageId="11" pageNumber="12">h</emphasis>
- and
<emphasis id="B9156F0BFFB0FFE2801FD965FDC6FEA6" box="[609,623,264,287]" italics="true" pageId="11" pageNumber="12">p</emphasis>
-series; all curved and heavily barbed, but
<emphasis id="B9156F0BFFB0FFE28636D965FBFFFEA6" box="[1096,1110,264,287]" italics="true" pageId="11" pageNumber="12">p</emphasis>
1 and
<emphasis id="B9156F0BFFB0FFE286EAD96AFB0BFEA6" box="[1172,1186,263,287]" italics="true" pageId="11" pageNumber="12">h</emphasis>
1 thicker than
<emphasis id="B9156F0BFFB0FFE28745D96AFAE0FEA6" box="[1339,1353,263,287]" italics="true" pageId="11" pageNumber="12">h</emphasis>
2 and
<emphasis id="B9156F0BFFB0FFE287F8D96AFA3DFEA6" box="[1414,1428,263,287]" italics="true" pageId="11" pageNumber="12">h</emphasis>
3 (
<figureCitation id="135AAF9CFFB0FFE282E1D940FF58FEFC" box="[159,241,301,325]" captionStart="FIGURE 10" captionStartId="14.[151,250,1964,1987]" captionTargetBox="[222,1348,281,1920]" captionTargetId="figure@14.[215,1372,268,1922]" captionTargetPageId="14" captionText="FIGURE 10. Posterior margin of gastronotum, scale bar 20 μm. (A) Cosmochthonius reticulatus; (B) C. foliatus; (C) C. ugamaensis; (D) C. ponticus." httpUri="https://zenodo.org/record/278735/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
A). Setae of
<emphasis id="B9156F0BFFB0FFE28300D940FE33FEFC" box="[382,410,301,325]" italics="true" pageId="11" pageNumber="12">ad</emphasis>
-series barbed,
<emphasis id="B9156F0BFFB0FFE2803BD940FDC8FEFC" box="[581,609,301,325]" italics="true" pageId="11" pageNumber="12">ad</emphasis>
1 approximately as long as
<emphasis id="B9156F0BFFB0FFE281EED942FC37FEFF" box="[912,926,303,326]" italics="true" pageId="11" pageNumber="12">p</emphasis>
2, length decreasing from
<emphasis id="B9156F0BFFB0FFE286C0D940FB73FEFC" box="[1214,1242,301,325]" italics="true" pageId="11" pageNumber="12">ad</emphasis>
1 to
<emphasis id="B9156F0BFFB0FFE2877DD940FAB6FEFC" box="[1283,1311,301,325]" italics="true" pageId="11" pageNumber="12">ad</emphasis>
4 (
<figureCitation id="135AAF9CFFB0FFE28748D940FAD3FEFC" box="[1334,1402,301,325]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1966]" captionTargetBox="[361,1226,228,1907]" captionTargetId="figure@9.[343,1243,213,1920]" captionTargetPageId="9" captionText="FIGURE 6. Cosmochthonius reticulatus, scale bars 50 μm. (A) Anal region of larva; (B) anogenital region of tritonymph." httpUri="https://zenodo.org/record/278732/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
B). Paraproctal valves with four pairs of barbed setae, slightly shorter than
<emphasis id="B9156F0BFFB0FFE281CDD939FC66FED5" box="[947,975,340,364]" italics="true" pageId="11" pageNumber="12">ad</emphasis>
4. Cupules
<emphasis id="B9156F0BFFB0FFE28638D938FBF5FED5" box="[1094,1116,341,364]" italics="true" pageId="11" pageNumber="12">ia</emphasis>
and
<emphasis id="B9156F0BFFB0FFE286ECD938FB07FED5" box="[1170,1198,341,364]" italics="true" pageId="11" pageNumber="12">im</emphasis>
as in larva, cupule
<emphasis id="B9156F0BFFB0FFE287F8D938FA35FED5" box="[1414,1436,341,364]" italics="true" pageId="11" pageNumber="12">ip</emphasis>
posterolaterally to seta
<emphasis id="B9156F0BFFB0FFE283DFD916FE06FE2A" box="[417,431,379,403]" italics="true" pageId="11" pageNumber="12">h</emphasis>
2, cupule
<emphasis id="B9156F0BFFB0FFE28067D916FD94FE2A" box="[537,573,379,403]" italics="true" pageId="11" pageNumber="12">iad</emphasis>
lateral to anterior part of anal valves, cupules
<emphasis id="B9156F0BFFB0FFE2862AD911FBDCFE2A" box="[1108,1141,380,403]" italics="true" pageId="11" pageNumber="12">ips</emphasis>
and
<emphasis id="B9156F0BFFB0FFE286D1D916FB6CFE2A" box="[1199,1221,379,403]" italics="true" pageId="11" pageNumber="12">ih</emphasis>
displaced laterally from cupule
<emphasis id="B9156F0BFFB0FFE2835BD9CFFEE0FE03" box="[293,329,418,442]" italics="true" pageId="11" pageNumber="12">iad</emphasis>
(
<figureCitation id="135AAF9CFFB0FFE28326D9CFFE32FE03" box="[344,411,418,442]" captionStart="FIGURE 5" captionStartId="8.[151,250,1806,1829]" captionTargetBox="[156,1432,213,1771]" captionTargetId="figure@8.[151,1436,193,1785]" captionTargetPageId="8" captionText="FIGURE 5. Cosmochthonius reticulatus, lateral aspect, legs partially drawn, scale bars 50 μm. (A) Larva; (B) tritonymph." httpUri="https://zenodo.org/record/278731/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
B). Ventral parts of gastronotum weakly striated. Shape of tibia and tarsus I, solenidia φ and ω, and famulus ε similar to that of
<taxonomicName id="4C61C89AFFB0FFE28062D9AAFD35FE67" ID-CoL="7SN9Q" box="[540,668,455,478]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis id="B9156F0BFFB0FFE28062D9AAFD86FE67" box="[540,559,455,478]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B9156F0BFFB0FFE28043D9AAFD35FE67" box="[573,668,455,478]" italics="true" pageId="11" pageNumber="12">ponticus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFF0CEE8FFB0FFE280A9D9ABFBB8FE67" author="Seniczak" box="[727,1041,454,478]" pageId="11" pageNumber="12" refString="Seniczak, S. &amp; Seniczak, A. (2009 a) Differentiation of body form of Protoplophoroidea (Acari: Oribatida) in the light of ontogeny of three species. Journal of Natural History, 44 (7), 389 - 419." type="journal article" year="2009" yearSuffix="a">Seniczak &amp; Seniczak 2009a</bibRefCitation>
).
</paragraph>
<caption id="DF1EE391FFB1FFE382E9DFA1FB63F858" box="[151,1226,1996,2019]" httpUri="https://zenodo.org/record/278733/files/figure.png" pageId="10" pageNumber="11" targetBox="[371,1238,331,1972]" targetPageId="10">
<paragraph id="8BDEB319FFB1FFE382E9DFA1FB63F858" blockId="10.[151,1226,1996,2019]" box="[151,1226,1996,2019]" pageId="10" pageNumber="11">
<emphasis id="B9156F0BFFB1FFE382E9DFA1FEBAF85B" bold="true" box="[151,275,1996,2019]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
<taxonomicName id="4C61C89AFFB1FFE38367DFA0FD9AF85A" ID-CoL="7SN9R" box="[281,563,1997,2019]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB1FFE38367DFA0FD9AF85A" box="[281,563,1997,2019]" italics="true" pageId="10" pageNumber="11">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
, tritonymph, dorsal aspect, legs partially drawn, scale bar 50 μm.
</paragraph>
</caption>
<paragraph id="8BDEB319FFB0FFE282B9D984FE38FCD7" blockId="11.[151,1437,151,1420]" pageId="11" pageNumber="12">
<emphasis id="B9156F0BFFB0FFE282B9D984FD68FDBB" bold="true" box="[199,705,489,514]" pageId="11" pageNumber="12">Summary of ontogenetic transformations:</emphasis>
The ontogeny of
<taxonomicName id="4C61C89AFFB0FFE281F1D986FB81FDBB" ID-CoL="7SN9R" box="[911,1064,490,514]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB0FFE281F1D986FC0BFDBB" box="[911,930,491,514]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B9156F0BFFB0FFE281CFD987FB81FDBB" box="[945,1064,490,514]" italics="true" pageId="11" pageNumber="12">reticulatus</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C61C89AFFB0FFE2876AD986FA3CFDBB" ID-CoL="7SN9Q" box="[1300,1429,491,514]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis id="B9156F0BFFB0FFE2876AD986FA8EFDBB" box="[1300,1319,491,514]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B9156F0BFFB0FFE28748D986FA3CFDBB" box="[1334,1429,491,514]" italics="true" pageId="11" pageNumber="12">ponticus</emphasis>
</taxonomicName>
, which was described earlier (
<bibRefCitation id="EFF0CEE8FFB0FFE283A5DA63FCB3FD9F" author="Seniczak" box="[475,794,526,550]" pageId="11" pageNumber="12" refString="Seniczak, S. &amp; Seniczak, A. (2009 a) Differentiation of body form of Protoplophoroidea (Acari: Oribatida) in the light of ontogeny of three species. Journal of Natural History, 44 (7), 389 - 419." type="journal article" year="2009" yearSuffix="a">Seniczak &amp; Seniczak 2009a</bibRefCitation>
). The number of prodorsal setae is constant during ontogeny (six pairs, including the sensillus), and the shape of setae remains similar in all instars. In contrast, the number of gastronotal setae increases in ontogeny, from 14 pairs in larva to 16 pairs in protonymph (
<emphasis id="B9156F0BFFB0FFE28697DA3BFB5EFDD7" box="[1257,1271,598,622]" italics="true" pageId="11" pageNumber="12">h</emphasis>
4 lost,
<emphasis id="B9156F0BFFB0FFE2873BDA3AFAFAFDD7" box="[1349,1363,599,622]" italics="true" pageId="11" pageNumber="12">p</emphasis>
-series gained), and these setae remain in the nymphs and adult. All instars have three transverse scissures on the dorsal part of hysterosoma, and hypertrophied and pinnate setae of the
<emphasis id="B9156F0BFFB0FFE28118DACFFCDBFD00" box="[870,882,674,697]" italics="true" pageId="11" pageNumber="12">e</emphasis>
- and
<emphasis id="B9156F0BFFB0FFE281CCDACCFC13FD00" box="[946,954,673,697]" italics="true" pageId="11" pageNumber="12">f</emphasis>
-series, which are positioned on intercalary sclerites. Aggenital setae are absent throughout. The formulae of setae are as follows: gastronotal setae – 14-16-16- 16-16 (larva to adult); segments PS–AN – 43333-4444-444; genital setae –
<date id="FFDF95D9FFB0FFE28182DA85FBF7FCB9" box="[1020,1118,744,768]" pageId="11" pageNumber="12" value="2010-02-01" valueMax="2010-07-01" valueMin="2010-02-01">1-2-7-10</date>
(protonymph to adult); and coxisternal setae – 3-2-2 (larva), 3-2-2-1 (protonymph), 3-2-3-3 (deutonymph) and 3-2-3-4 (tritonymph and adult). All formulae are consistent with patterns presented by
<bibRefCitation id="EFF0CEE8FFB0FFE28080DB5DFC61FCF1" author="Grandjean" box="[766,968,816,840]" pageId="11" pageNumber="12" refString="Grandjean, F. (1949) Formules anales, gastronotiques, genitales et aggenitales du developpement numerique des poils chez les Oribates. Bulletin Societe Zoologique France, 74, 201 - 225." type="journal article" year="1949">Grandjean (1949)</bibRefCitation>
. In the larva seta
<emphasis id="B9156F0BFFB0FFE286F3DB5CFB01FCF0" box="[1165,1192,817,841]" italics="true" pageId="11" pageNumber="12">1c</emphasis>
is scaliform and covers Claparéde’s organ.
</paragraph>
</subSubSection>
<subSubSection id="C37BE092FFB0FFE282B9DB14FCEFFA35" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BDEB319FFB0FFE282B9DB14FCEFFA35" blockId="11.[151,1437,151,1420]" pageId="11" pageNumber="12">
<emphasis id="B9156F0BFFB0FFE282B9DB14FE52FC2B" bold="true" box="[199,507,889,914]" pageId="11" pageNumber="12">Distribution and ecology:</emphasis>
<taxonomicName id="4C61C89AFFB0FFE2807BDB14FCE9FC28" box="[517,832,889,913]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB0FFE2807BDB14FCE9FC28" box="[517,832,889,913]" italics="true" pageId="11" pageNumber="12">Cosmochthonius reticulatus</emphasis>
</taxonomicName>
is a Mediterranean occidental species (
<bibRefCitation id="EFF0CEE8FFB0FFE28779DB14FA31FC28" author="Subias" box="[1287,1432,889,913]" pageId="11" pageNumber="12" refString="Subias, L. S. (2004) Systematic, synonymic and biogeographical check - list of the world's oribatid mites (Acariformes, Oribatida) (1758 - 2002). Graellsia, 60, 3 - 305. Online version accessed in February 2011. 557 pp. http: // www. ucm. es / info / zoo / Artropodos / Catalogo. pdf." type="book chapter" year="2004">Subías 2004</bibRefCitation>
, 2011), with type-locality in Périgueux city, north-west of Bordeaux,
<collectingCountry id="F376F389FFB0FFE281EEDBF3FC75FC0C" box="[912,988,926,949]" name="France" pageId="11" pageNumber="12">France</collectingCountry>
(
<bibRefCitation id="EFF0CEE8FFB0FFE28192DBF0FB04FC0C" author="Grandjean" box="[1004,1197,925,949]" pageId="11" pageNumber="12" refString="Grandjean, F. (1947 b) L'origine de la pince mandibulaire chez les Acariens actinochitineux. Archives des Sciences physiques et naturelles, Geneve, (5), 29, 305 - 355." type="journal article" year="1947" yearSuffix="b">Grandjean 1947b</bibRefCitation>
). According to
<bibRefCitation id="EFF0CEE8FFB0FFE28722DBF0FF72FC60" author="Trave" pageId="11" pageNumber="12" refString="Trave, J. (1956) Contributions a l' etude de la fauna de la Massane (Deuxieme note). Oribates (Acarien), 1 re partie. Vie et Milieu, 7 (1), 77 - 94." type="journal article" year="1956">Travé (1956</bibRefCitation>
,
<bibRefCitation id="EFF0CEE8FFB0FFE28296DBACFE88FC60" author="Trave" box="[232,289,961,985]" pageId="11" pageNumber="12" refString="Trave, J. (1984) Contribution a l'etude des Oribates (Acariens) de l'Ile de Port-Cros (Parc National). Travaux Scientifiques du Parc National de Port-Cros, France, 10, 119 - 150." type="journal article" year="1984">1984</bibRefCitation>
) this species prefers warm, south-slope biotopes in pine forests (
<taxonomicName id="4C61C89AFFB0FFE28672DBAEFAA8FC60" authority="Mill." authorityName="Mill." box="[1036,1281,961,986]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="alepensis">
<emphasis id="B9156F0BFFB0FFE28672DBAEFB15FC63" box="[1036,1212,962,986]" italics="true" pageId="11" pageNumber="12">Pinus alepensis</emphasis>
Mill.
</taxonomicName>
), but it is not very common.
<bibRefCitation id="EFF0CEE8FFB0FFE28336DB88FD3FFC44" author="Perez-Inigo" box="[328,662,997,1021]" pageId="11" pageNumber="12" refString="Perez-Inigo, C. &amp; Pena, M. A. (1995) Oribatidos (Acari, Oribatei) de Fuerteventura (Islas Canarias). Graellsia, 51, 143 - 164." type="journal article" year="1995">Pérez-Iñigo and Peña (1995)</bibRefCitation>
found it in soil with mosses, while
<bibRefCitation id="EFF0CEE8FFB0FFE28636DB88FF72FB98" author="Mahunka" pageId="11" pageNumber="12" refString="Mahunka, S. &amp; Mahunka-Papp, L. (2003) Contribution to the knowledge of the Hungarian Oribatida fauna (Acari). I. Acta Zoologica Academiae Scientiarum Hungaricae, 49 (4), 255 - 260." type="journal article" year="2003">Mahunka and Mahunka-Papp (2003</bibRefCitation>
,
<bibRefCitation id="EFF0CEE8FFB0FFE2829BDC64FEB6FB98" author="Mahunka-Papp" box="[229,287,1033,1057]" pageId="11" pageNumber="12" refString="Mahunka, S. &amp; Mahunka-Papp, L. (2010) New and little known oribatid mites from the Carpathian Basin and the Balkan Peninsula (Acari: Oribatida). Acta Zoologica Academiae Scientiarum Hungaricae, 56 (3), 211 - 234." type="journal article" year="2010">2010</bibRefCitation>
) recorded
<taxonomicName id="4C61C89AFFB0FFE283E8DC67FD84FB9B" box="[406,557,1034,1058]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB0FFE283E8DC67FE00FB98" box="[406,425,1034,1057]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B9156F0BFFB0FFE283C8DC67FD84FB9B" box="[438,557,1034,1058]" italics="true" pageId="11" pageNumber="12">reticulatus</emphasis>
</taxonomicName>
from dead tree trunk and oak forest.
<bibRefCitation id="EFF0CEE8FFB0FFE281B6DC64FA8CFB98" author="Penttinen" box="[968,1317,1033,1057]" pageId="11" pageNumber="12" refString="Penttinen, R. &amp; Gordeeva, E. (2009) Distribution of Cosmochthonius species (Oribatida: Cosmochthoniidae) in the eastern part of the Mediterranean, Ukraine and Tajikistan. In: Sabelis, M. W. &amp; Bruin, J. (Eds.): Trends in Acarology, pp. 171 - 174." type="book chapter" year="2009">Penttinen and Gordeeva (2009)</bibRefCitation>
found this species more frequently in litter under juniper [
<taxonomicName id="4C61C89AFFB0FFE280C0DC42FBDFFBFC" authority="Sibth. &amp; Sm." authorityName="Sibth. &amp; Sm." box="[702,1142,1069,1094]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="macrocarpa">
<emphasis id="B9156F0BFFB0FFE280C0DC42FC16FBFF" box="[702,959,1071,1094]" italics="true" pageId="11" pageNumber="12">Juniperus macrocarpa</emphasis>
(Sibth. &amp; Sm.)
</taxonomicName>
] in Lesvos (
<collectingCountry id="F376F389FFB0FFE28776DC40FAF4FBFC" box="[1288,1373,1069,1093]" name="Greece" pageId="11" pageNumber="12">Greece</collectingCountry>
), and under cypress (
<taxonomicName id="4C61C89AFFB0FFE28337DC3FFDF2FBD3" box="[329,603,1106,1130]" class="Pinopsida" family="Cupressaceae" genus="Cupressus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="sempervirens">
<emphasis id="B9156F0BFFB0FFE28337DC3FFDF2FBD3" box="[329,603,1106,1130]" italics="true" pageId="11" pageNumber="12">Cupressus sempervirens</emphasis>
</taxonomicName>
L.) and oak trees (
<taxonomicName id="4C61C89AFFB0FFE2813CDC3FFBBBFBD3" box="[834,1042,1106,1130]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="coccifera">
<emphasis id="B9156F0BFFB0FFE2813CDC3FFBBBFBD3" box="[834,1042,1106,1130]" italics="true" pageId="11" pageNumber="12">Quercus coccifera</emphasis>
</taxonomicName>
L.) in Rhodes (
<collectingCountry id="F376F389FFB0FFE286AEDC3CFA8CFBD0" box="[1232,1317,1105,1129]" name="Greece" pageId="11" pageNumber="12">Greece</collectingCountry>
), and also under juniper (
<taxonomicName id="4C61C89AFFB0FFE2833FDC1AFDE3FB34" authority="Bieb" authorityName="Bieb" box="[321,586,1141,1166]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="B9156F0BFFB0FFE2833FDC1AFDA3FB37" box="[321,522,1142,1166]" italics="true" pageId="11" pageNumber="12">Juniperus excelsa</emphasis>
Bieb
</taxonomicName>
) in
<collectingCountry id="F376F389FFB0FFE28007DC18FD7FFB34" box="[633,726,1141,1165]" name="Ukraine" pageId="11" pageNumber="12">Ukraine</collectingCountry>
, and considered it common in the eastern Mediterranean area.
<bibRefCitation id="EFF0CEE8FFB0FFE282E9DCF7FE7BFB08" author="Karasawa" box="[151,466,1177,1201]" pageId="11" pageNumber="12" refString="Karasawa, S. &amp; Hijii, N. (2004 a) Effects of microhabitat diversity and geographical isolation on oribatid mite (Acari: Oribatida) communities in mangrove forests. Pedobiologia, 48 (3), 245 - 256." type="journal article" year="2004" yearSuffix="a">Karasawa and Hijii (2004a)</bibRefCitation>
found
<taxonomicName id="4C61C89AFFB0FFE2805ADCF7FD17FB0B" box="[548,702,1178,1202]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB0FFE2805ADCF7FD9EFB08" box="[548,567,1178,1201]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B9156F0BFFB0FFE28039DCF7FD17FB0B" box="[583,702,1178,1202]" italics="true" pageId="11" pageNumber="12">reticulatus</emphasis>
</taxonomicName>
on tree branches in a typical mangrove forest about 4.5 m high, with dominant
<taxonomicName id="4C61C89AFFB0FFE2833FDCD2FDE2FB6F" box="[321,587,1214,1238]" class="Magnoliopsida" family="Rhizophoraceae" genus="Bruguiera" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="gymnorrhiza">
<emphasis id="B9156F0BFFB0FFE2833FDCD2FDE2FB6F" box="[321,587,1214,1238]" italics="true" pageId="11" pageNumber="12">Bruguiera gymnorrhiza</emphasis>
</taxonomicName>
(L.) Lam. and rare
<taxonomicName id="4C61C89AFFB0FFE28156DCD3FB97FB6C" authority="Griff" authorityName="Griff" box="[808,1086,1213,1238]" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="stylosa">
<emphasis id="B9156F0BFFB0FFE28156DCD3FBA9FB6F" box="[808,1024,1214,1238]" italics="true" pageId="11" pageNumber="12">Rhizophora stylosa</emphasis>
Griff
</taxonomicName>
in the Ryukyu Islands (
<collectingCountry id="F376F389FFB0FFE28734DCD0FA39FB6C" box="[1354,1424,1213,1237]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
). Interestingly, these adults were slightly smaller (
<bibRefCitation id="EFF0CEE8FFB0FFE280C5DC8CFC7DFB40" author="Karasawa" box="[699,980,1249,1273]" pageId="11" pageNumber="12" refString="Karasawa, S. &amp; Hijii, N. (2004 b) Morphological modifications among oribatid mites (Acari: Oribatida) in relation to habitat differentiation in mangrove forests. Pedobiologia, 48 (3), 383 - 394." type="journal article" year="2004" yearSuffix="b">Karasawa &amp; Hijii 2004b</bibRefCitation>
) than those investigated here.
<bibRefCitation id="EFF0CEE8FFB0FFE28750DC8FFE8DFAA4" pageId="11" pageNumber="12" refString="Karasawa, S., Gotoh, K., Sasaki, T. &amp; Hijii, N. (2005) Wind-based dispersal of oribatid mites (Acari: Oribatida) in a subtropical forest in Japan. Journal of the Acarological Society of Japan, 14 (2), 117 - 122." type="journal article">
Karasawa
<emphasis id="B9156F0BFFB0FFE282E9DD6AFF78FAA7" box="[151,209,1286,1310]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2005)
</bibRefCitation>
also found
<taxonomicName id="4C61C89AFFB0FFE283D3DD6BFDEFFAA7" box="[429,582,1286,1310]" class="Arachnida" family="Cosmochthoniidae" genus="Cosmochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9156F0BFFB0FFE283D3DD6BFE69FAA4" box="[429,448,1286,1309]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B9156F0BFFB0FFE283B1DD6BFDEFFAA7" box="[463,582,1286,1310]" italics="true" pageId="11" pageNumber="12">reticulatus</emphasis>
</taxonomicName>
in window traps mounted above the soil and at canopy level. This species was abundant in cypress tree litter near Villa del Casale (Central Sicilia,
<collectingCountry id="F376F389FFB0FFE281B2DD44FBA9FAF8" box="[972,1024,1321,1345]" name="Italy" pageId="11" pageNumber="12">Italy</collectingCountry>
,
<geoCoordinate id="EE55D5DEFFB0FFE28672DD44FBC4FAF8" box="[1036,1133,1321,1345]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="37.35">N37°21’</geoCoordinate>
52,
<geoCoordinate id="EE55D5DEFFB0FFE286E6DD44FB5CFAF8" box="[1176,1269,1321,1345]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="14.333333">E14°20’</geoCoordinate>
04,
<quantity id="4C991EFCFFB0FFE2875EDD44FAC1FAF8" box="[1312,1384,1321,1345]" metricMagnitude="2" metricUnit="m" metricValue="5.38" pageId="11" pageNumber="12" unit="m" value="538.0">538 m</quantity>
a. s. l.), achieving on
<date id="FFDF95D9FFB0FFE28322DD3CFE1FFAD1" box="[348,438,1360,1385]" pageId="11" pageNumber="12" value="1927-05">May 27</date>
of 2009 a density of 1,903 individuals per
<quantity id="4C991EFCFFB0FFE281CCDD3DFBA1FAD1" box="[946,1032,1360,1384]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="cm" value="500.0">500 cm</quantity>
3, with a large fraction of juveniles (62.7% of population), including larvae (27.5% of juveniles).
</paragraph>
</subSubSection>
</treatment>
</document>
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