treatments-xml/data/03/F5/87/03F587F6FFDDCD7FFCFBFE10FD4FFA20.xml

<document id="21F29B33CE74D436DF423992AC0E1571" ID-DOI="10.5252/z2011n4a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="5166897" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628228810046" checkinUser="felipe" docAuthor="Cigliano, María Marta, Pocco, Martina E. &amp; Lange, Carlos E." docDate="2011" docId="03F587F6FFDDCD7FFCFBFE10FD4FFA20" docLanguage="en" docName="Zoosystema.33.4.523-544.pdf" docOrigin="Zoosystema 33 (4)" docSource="http://dx.doi.org/10.5252/z2011n4a5" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.6:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="6" docTitle="Tiyantiyana Cigliano &amp; Pocco &amp; Lange 2011, n. gen." docType="treatment" docVersion="3" lastPageNumber="530" masterDocId="FFCCFF8EFFDFCD78FFDAFF86FFCCFFA7" masterDocTitle="Grasshoppers of the Andes: new Melanoplinae and Gomphocerinae taxa (Insecta, Orthoptera, Acrididae) from Huascarán National Park and Callejón de Huaylas, Ancash, Peru" masterLastPageNumber="544" masterPageNumber="523" pageNumber="525" updateTime="1699036276121" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BB0252AF069C7313DF5878994DD7355F">Grasshoppers of the Andes: new Melanoplinae and Gomphocerinae taxa (Insecta, Orthoptera, Acrididae) from Huascarán National Park and Callejón de Huaylas, Ancash, Peru</mods:title>
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<mods:namePart id="9B493C2B707CB4A7A472100B35F089B3">Cigliano, María Marta</mods:namePart>
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<mods:namePart id="07A82E7AF5387D75E87861437A421BD9">Pocco, Martina E.</mods:namePart>
<mods:affiliation id="E995C08F3A075118ED5EB98791542851">Museo de La Plata, División Entomología, Paseo del Bosque 1900, La Plata (Argentina) cigliano @ fcnym. unlp. edu. ar martinapocco @ fcnym. unlp. edu. ar</mods:affiliation>
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<mods:namePart id="11A0F1A561CE90FA5195B0C792A408A1">Lange, Carlos E.</mods:namePart>
<mods:affiliation id="2B1CFAF75626E7D0A8C504741B304383">Comisión de Investigaciones Científicas (CIC) de la provincia de Buenos Aires, CCT La Plata, CEPAVE, CONICET-UNLP, calle 2 # 584,1900, La Plata (Argentina) carlosl @ cepave. edu. ar</mods:affiliation>
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<subSubSection id="C346656BFFDDCD7AFCFBFE10FBF2FE17" box="[801,1086,406,432]" pageId="2" pageNumber="525" type="nomenclature">
<paragraph id="8BE336E0FFDDCD7AFCFBFE10FBF2FE17" blockId="2.[801,1086,406,432]" box="[801,1086,406,432]" pageId="2" pageNumber="525">
<heading id="D0AB818CFFDDCD7AFCFBFE10FBF2FE17" box="[801,1086,406,432]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="525" reason="2">
Genus 
<taxonomicName id="4C5C4D63FFDDCD7AFCB4FE11FC20FE17" ID-CoL="8P27Y" authority="Cigliano &amp; Pocco &amp; Lange, 2011" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[878,1004,407,432]" class="Insecta" family="Acrididae" genus="Tiyantiyana" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="525" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF2FFDDCD7AFCB4FE11FC20FE17" box="[878,1004,407,432]" italics="true" pageId="2" pageNumber="525">Tiyantiyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5789FFDDCD7AFC29FE10FBF2FE17" box="[1011,1086,406,432]" pageId="2" pageNumber="525" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C346656BFFDDCD7AFD7FFE49FB64FE41" box="[677,1192,463,487]" pageId="2" pageNumber="525" type="materials_examined">
<paragraph id="8BE336E0FFDDCD7AFD7FFE49FB64FE41" blockId="2.[677,1192,463,487]" box="[677,1192,463,487]" pageId="2" pageNumber="525">
<collectionCode id="ED4DAE25FFDDCD7AFD7FFE49FCEFFE42" box="[677,803,463,486]" pageId="2" pageNumber="525">
<typeStatus id="54E78842FFDDCD7AFD7FFE49FD19FE42" box="[677,725,463,486]" pageId="2" pageNumber="525">TYPE</typeStatus>
SPECIES
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. 
<emphasis id="B928EAF2FFDDCD7AFCF5FE57FBD0FE40" box="[815,1052,464,487]" italics="true" pageId="2" pageNumber="525">
<emphasis id="B928EAF2FFDDCD7AFCF5FE57FC84FE41" bold="true" box="[815,840,465,486]" italics="true" pageId="2" pageNumber="525">—</emphasis>
<taxonomicName id="4C5C4D63FFDDCD7AFC94FE56FBD0FE40" authority="Cigliano &amp; Pocco &amp; Lange, 2011" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[846,1052,464,487]" class="Insecta" family="Acrididae" genus="Tiyantiyana" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="525" phylum="Arthropoda" rank="species" species="sunipenis" status="n. gen., n. sp.">Tiyantiyana sunipenis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A21B5789FFDDCD7AFBF8FE49FB64FE41" box="[1058,1192,463,487]" pageId="2" pageNumber="525" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C346656BFFDDCD7AFD7FFE79FC9AFDEC" pageId="2" pageNumber="525" type="etymology">
<paragraph id="8BE336E0FFDDCD7AFD7FFE79FC9AFDEC" blockId="2.[675,1210,510,587]" pageId="2" pageNumber="525">ETYMOLOGY. — “Tiyantiyan” means cricket in Quechua native language. In the Huascarán region, grasshoppers are called crickets.</paragraph>
</subSubSection>
<subSubSection id="C346656BFFDDCD7AFD7FFDE2FD2DFD31" pageId="2" pageNumber="525" type="distribution">
<paragraph id="8BE336E0FFDDCD7AFD7FFDE2FD2DFD31" blockId="2.[677,1209,611,662]" pageId="2" pageNumber="525">
DISTRIBUTION. — 
<collectingCountry id="F34B7670FFDDCD7AFCB2FDE2FC55FDDC" box="[872,921,612,635]" name="Peru" pageId="2" pageNumber="525">Peru</collectingCountry>
, 
<collectingRegion id="4998F802FFDDCD7AFC7FFDE5FC22FDDC" box="[933,1006,611,635]" country="Peru" name="Ancash" pageId="2" pageNumber="525">Ancash</collectingRegion>
(see distribution in OSF).
</paragraph>
</subSubSection>
<subSubSection id="C346656BFFDDCD7AFD7FFD28FC57FCED" pageId="2" pageNumber="525" type="diagnosis">
<paragraph id="8BE336E0FFDDCD7AFD7FFD28FC57FCED" blockId="2.[675,1211,685,842]" pageId="2" pageNumber="525">
DIAGNOSIS. — Among the South American 
<taxonomicName id="4C5C4D63FFDDCD7AFB97FD2BFD05FD78" class="Insecta" family="Acrididae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="525" phylum="Arthropoda" rank="subFamily" subFamily="Melanoplinae">Melanoplinae</taxonomicName>
, 
<taxonomicName id="4C5C4D63FFDDCD7AFD09FD4EFC89FD78" authority="Cigliano &amp; Pocco &amp; Lange, 2011" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[723,837,712,735]" class="Insecta" family="Acrididae" genus="Tiyantiyana" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="525" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF2FFDDCD7AFD09FD4EFC89FD78" box="[723,837,712,735]" italics="true" pageId="2" pageNumber="525">Tiyantiyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5789FFDDCD7AFC96FD4EFC5CFD47" box="[844,912,712,736]" pageId="2" pageNumber="525" rank="genus">n. gen.</taxonomicNameLabel>
is characterized by the unique shape of the aedeagus with the ventral valves very long, protruding beneath the pallium, highly surpassing the level of the epiproct, sheath of aedeagus with numerous spines (
<figureCitation id="13672A65FFDDCD7AFD31FCB7FCEEFCED" box="[747,802,817,842]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Figs 1</figureCitation>
I-L; 4C-D).
</paragraph>
</subSubSection>
<subSubSection id="C346656BFFDDCD7FFD7FFCF4FD4FFA20" lastPageId="7" lastPageNumber="530" pageId="2" pageNumber="525" type="description">
<paragraph id="8BE336E0FFDDCD7AFD7FFCF4FCF9FC2D" blockId="2.[677,1209,882,971]" box="[677,821,882,908]" pageId="2" pageNumber="525">DESCRIPTION</paragraph>
<paragraph id="8BE336E0FFDDCD7AFD7FFC17FB65FC6C" blockId="2.[677,1209,882,971]" pageId="2" pageNumber="525">Small sized apterous insects.Body colour highly variable with multiple short carinulae on tegument.</paragraph>
<paragraph id="8BE336E0FFDDCD7FFD7FFC76FEAFFE48" blockId="2.[674,1212,1008,1700]" lastBlockId="7.[111,648,216,495]" lastPageId="7" lastPageNumber="530" pageId="2" pageNumber="525">
<emphasis id="B928EAF2FFDDCD7AFD7FFC76FD2EFBAD" box="[677,738,1008,1034]" italics="true" pageId="2" pageNumber="525">Males</emphasis>
Head globed-shape, fastigium declivent, with lateral carinae weakly indicated, rounded at the joint of frontal costa; eyes subcircular (
<figureCitation id="13672A65FFDDCD7AFC08FBC9FBC6FBCE" box="[978,1034,1103,1129]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Figs 1</figureCitation>
A-C; 4A). Lateral foveolae of fastigium visible from above (
<figureCitation id="13672A65FFDDCD7AFB8CFBE8FB66FB2E" box="[1110,1194,1134,1161]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Fig. 1C</figureCitation>
). Antennae short, shorter than head and pronotum, with flattened segments (
<figureCitation id="13672A65FFDDCD7AFC7CFB28FC38FB6F" box="[934,1012,1198,1224]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Fig. 1A</figureCitation>
; 
<figureCitation id="13672A65FFDDCD7AFC27FB28FBEDFB6F" box="[1021,1057,1198,1224]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[127,1196,224,1576]" captionTargetId="figure-20@6.[127,1196,215,1576]" captionTargetPageId="6" captionText="FIG. 4. — Tiyantiyana sunipenis n. gen., n. sp.: A, male habitus, lateral view; B, female habitus, lateral view; C, D, endophallic plates, cingulum and aedeagal valves with sheath, lateral and dorsal views; E, epiphallus, dorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5166917" httpUri="https://zenodo.org/record/5166917/files/figure.png" pageId="2" pageNumber="525">4A</figureCitation>
). Pronotal disk flat, trapezoidal, weakly constricted at the middle; lateral and mid longitudinal carinae weakly indicated; transverse sulci not indicated on pronotal disk, only evident on lateral lobes of pronotum; hind margin emarginated (
<figureCitation id="13672A65FFDDCD7AFCF5FACAFCB0FAC1" box="[815,892,1356,1382]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Fig.1C</figureCitation>
). Meso, metanoto and abdominal tergites with lateral carinulae at the sides of mid longitudinal carina. Prosternal process collar-shaped. Cerci conical,width tapering towards the apex,barely reaching the tip of epiproct (
<figureCitation id="13672A65FFDDCD7AFC09FA4DFB8BFA41" box="[979,1095,1483,1510]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Fig. 1D, E</figureCitation>
). Epiproct triangular;furculae not developed(
<figureCitation id="13672A65FFDDCD7AFC24FA6DFB80F9A2" box="[1022,1100,1515,1541]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Fig.1D</figureCitation>
). Abdominal terminalia swollen and upcurved.Pallium thick, highly elevated (
<figureCitation id="13672A65FFDDCD7AFC9FF9ACFC5DF9E3" box="[837,913,1578,1604]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Fig.1E</figureCitation>
). Subgenital plate short, with rounded distal margin, with the apex surpassing the tip of epiproct. Legs with femora robust, mostly the hind ones (
<figureCitation id="13672A65FFDDCD7AFCC8F90FFCAEF904" box="[786,866,1673,1699]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Figs 1A</figureCitation>
; 
<figureCitation id="13672A65FFDDCD7AFCB1F90FFC41F904" box="[875,909,1673,1699]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[127,1196,224,1576]" captionTargetId="figure-20@6.[127,1196,215,1576]" captionTargetPageId="6" captionText="FIG. 4. — Tiyantiyana sunipenis n. gen., n. sp.: A, male habitus, lateral view; B, female habitus, lateral view; C, D, endophallic plates, cingulum and aedeagal valves with sheath, lateral and dorsal views; E, epiphallus, dorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5166917" httpUri="https://zenodo.org/record/5166917/files/figure.png" pageId="2" pageNumber="525">4A</figureCitation>
). Phallic complex (
<figureCitation id="13672A65FFDDCD7AFB90F90FFB4EF904" box="[1098,1154,1673,1699]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="2" pageNumber="525">Figs 1</figureCitation>
I-M; 4C-E): cingulum with reduced rami, zygoma and apodemes well developed. Basal endophallic plates barely developed comparatively to the apical plates. Valves of aedeagus very long, protruding beneath the pallium, highly surpassing the level of the epiproct, apex bent downwards; sheath of aedeagus covering ⅔ of the valves of aedeagus with numerous spines; arch of cingulum large.Epiphallus with lophi widely developed horizontally.
</paragraph>
<caption id="DF236668FFDCCD7BFFABF999FEBAF906" ID-DOI="http://doi.org/10.5281/zenodo.5166902" ID-Zenodo-Dep="5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="3" pageNumber="526" startId="3.[113,124,1567,1584]" targetBox="[156,1167,216,1500]" targetPageId="3">
<paragraph id="8BE336E0FFDCCD7BFFABF999FEBAF906" blockId="3.[113,1211,1567,1697]" pageId="3" pageNumber="526">
FIG. 1. — 
<taxonomicName id="4C5C4D63FFDCCD7BFF18F999FEBEF997" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[194,370,1567,1584]" class="Insecta" family="Acrididae" genus="Tiyantiyana" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="sunipenis">
<emphasis id="B928EAF2FFDCCD7BFF18F999FEBEF997" box="[194,370,1567,1584]" italics="true" pageId="3" pageNumber="526">Tiyantiyana sunipenis</emphasis>
</taxonomicName>
n. gen, n. sp.: 
<emphasis id="B928EAF2FFDCCD7BFE35F999FE30F997" bold="true" box="[495,508,1567,1584]" pageId="3" pageNumber="526">A</emphasis>
, male habitus, lateral view; 
<emphasis id="B928EAF2FFDCCD7BFD39F999FD3CF997" bold="true" box="[739,752,1567,1584]" pageId="3" pageNumber="526">B</emphasis>
, 
<emphasis id="B928EAF2FFDCCD7BFD21F999FCC5F997" bold="true" box="[763,777,1567,1584]" pageId="3" pageNumber="526">C</emphasis>
, male head and pronotum, lateral and dorsal views; 
<emphasis id="B928EAF2FFDCCD7BFFABF9B3FFB3F9E1" bold="true" box="[113,127,1589,1606]" pageId="3" pageNumber="526">D</emphasis>
, 
<emphasis id="B928EAF2FFDCCD7BFF53F9B3FF59F9E1" bold="true" box="[137,149,1589,1606]" pageId="3" pageNumber="526">E</emphasis>
, male distal abdominal segments, dorsal and lateral views; 
<emphasis id="B928EAF2FFDCCD7BFD5EF9B3FD43F9E1" bold="true" box="[644,655,1589,1606]" pageId="3" pageNumber="526">F</emphasis>
, 
<emphasis id="B928EAF2FFDCCD7BFD43F9B3FD6BF9E1" bold="true" box="[665,679,1589,1606]" pageId="3" pageNumber="526">G</emphasis>
, female head and pronotum, lateral and dorsal views; 
<emphasis id="B928EAF2FFDCCD7BFBB0F9B3FBB4F9E1" bold="true" box="[1130,1144,1589,1606]" pageId="3" pageNumber="526">H</emphasis>
, female ovipositor valves; 
<emphasis id="B928EAF2FFDCCD7BFEDFF9CAFEC7F9FA" bold="true" box="[261,267,1612,1629]" pageId="3" pageNumber="526">I</emphasis>
, phallic complex,lateral view; 
<emphasis id="B928EAF2FFDCCD7BFDD9F9CAFDC1F9FA" bold="true" box="[515,525,1612,1629]" pageId="3" pageNumber="526">J</emphasis>
, 
<emphasis id="B928EAF2FFDCCD7BFDCCF9CAFDEDF9FA" bold="true" box="[534,545,1612,1629]" pageId="3" pageNumber="526">L</emphasis>
, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; 
<emphasis id="B928EAF2FFDCCD7BFF71F9E5FF74F9D3" bold="true" box="[171,184,1635,1652]" pageId="3" pageNumber="526">K</emphasis>
, distal portion of aedeagus and aedeagal valves, ventral view; 
<emphasis id="B928EAF2FFDCCD7BFD12F9E5FD15F9D3" bold="true" box="[712,729,1635,1652]" pageId="3" pageNumber="526">M</emphasis>
, epiphallus, dorsal view. Abbreviations: 
<emphasis id="B928EAF2FFDCCD7BFBF2F9E5FB8CF9D3" bold="true" box="[1064,1088,1635,1652]" pageId="3" pageNumber="526">Ap</emphasis>
, apodemes of cingulum; 
<emphasis id="B928EAF2FFDCCD7BFF1DF9FFFF17F92D" bold="true" box="[199,219,1657,1674]" pageId="3" pageNumber="526">Ar</emphasis>
, arch of aedeagus; 
<emphasis id="B928EAF2FFDCCD7BFEA6F9FFFE5EF92D" bold="true" box="[380,402,1657,1674]" pageId="3" pageNumber="526">Av</emphasis>
, aedeagal valves; 
<emphasis id="B928EAF2FFDCCD7BFDF3F9FFFD8DF92D" bold="true" box="[553,577,1657,1674]" pageId="3" pageNumber="526">Ep</emphasis>
, endophallic plates; 
<emphasis id="B928EAF2FFDCCD7BFD33F9FFFD38F92D" bold="true" box="[745,756,1657,1674]" pageId="3" pageNumber="526">L</emphasis>
, lophi of epiphallus; 
<emphasis id="B928EAF2FFDCCD7BFC47F9FFFC77F92D" bold="true" box="[925,955,1657,1674]" pageId="3" pageNumber="526">Rm</emphasis>
, rami; 
<emphasis id="B928EAF2FFDCCD7BFC28F9FFFBC5F92D" bold="true" box="[1010,1033,1657,1674]" pageId="3" pageNumber="526">Sh</emphasis>
, sheath of aedeagus, 
<emphasis id="B928EAF2FFDCCD7BFFABF916FF4BF906" bold="true" box="[113,135,1680,1697]" pageId="3" pageNumber="526">Zy</emphasis>
, zygoma. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF236668FFDBCD7CFFABF9B3FD74F906" ID-DOI="http://doi.org/10.5281/zenodo.5166909" ID-Zenodo-Dep="5166909" httpUri="https://zenodo.org/record/5166909/files/figure.png" pageId="4" pageNumber="527" startId="4.[113,124,1589,1606]" targetBox="[192,1131,216,1499]" targetPageId="4">
<paragraph id="8BE336E0FFDBCD7CFFABF9B3FD74F906" blockId="4.[113,1211,1589,1697]" pageId="4" pageNumber="527">
FIG. 2. — 
<emphasis id="B928EAF2FFDBCD7CFF1CF9B3FF24F9E1" bold="true" box="[198,232,1589,1606]" pageId="4" pageNumber="527">A -K</emphasis>
, 
<taxonomicName id="4C5C4D63FFDBCD7CFF29F9B3FE48F9E1" authority="Cigliano &amp; Pocco &amp; Lange, 2011" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[243,388,1589,1606]" class="Insecta" family="Acrididae" genus="Maeacris" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="chilikuti" status="sp. nov.">
<emphasis id="B928EAF2FFDBCD7CFF29F9B3FE48F9E1" box="[243,388,1589,1606]" italics="true" pageId="4" pageNumber="527">Maeacris chilikuti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5789FFDBCD7CFE56F9B3FE72F9E1" box="[396,446,1589,1606]" pageId="4" pageNumber="527" rank="species">n. sp.</taxonomicNameLabel>
; 
<emphasis id="B928EAF2FFDBCD7CFE1DF9B3FE18F9E1" bold="true" box="[455,468,1589,1606]" pageId="4" pageNumber="527">A</emphasis>
, male habitus, lateral view; 
<emphasis id="B928EAF2FFDBCD7CFD19F9B3FD1CF9E1" bold="true" box="[707,720,1589,1606]" pageId="4" pageNumber="527">B</emphasis>
, male head and pronotum, dorsal view; 
<emphasis id="B928EAF2FFDBCD7CFBF0F9B3FBF4F9E1" bold="true" box="[1066,1080,1589,1606]" pageId="4" pageNumber="527">C</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFB9EF9B3FB9EF9E1" bold="true" box="[1092,1106,1589,1606]" pageId="4" pageNumber="527">D</emphasis>
, male distal abdominal segments, dorsal and lateral views; 
<emphasis id="B928EAF2FFDBCD7CFE25F9CAFDC7F9FA" bold="true" box="[511,523,1612,1629]" pageId="4" pageNumber="527">E</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFDCDF9CAFDEEF9FA" bold="true" box="[535,546,1612,1629]" pageId="4" pageNumber="527">F</emphasis>
, female head and pronotum, dorsal and lateral views; 
<emphasis id="B928EAF2FFDBCD7CFC2AF9CAFBD8F9FA" bold="true" box="[1008,1044,1612,1629]" pageId="4" pageNumber="527">G -U</emphasis>
, phallic complex of 
<taxonomicName id="4C5C4D63FFDBCD7CFFABF9E5FF77F9D3" authorityName="Ronderos" authorityYear="1983" box="[113,187,1635,1652]" class="Insecta" family="Acrididae" genus="Maeacris" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="B928EAF2FFDBCD7CFFABF9E5FF77F9D3" box="[113,187,1635,1652]" italics="true" pageId="4" pageNumber="527">Maeacris</emphasis>
</taxonomicName>
, species as indicated; 
<emphasis id="B928EAF2FFDBCD7CFEA9F9E5FE4DF9D3" bold="true" box="[371,385,1635,1652]" pageId="4" pageNumber="527">G</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFE50F9E5FE59F9D3" bold="true" box="[394,405,1635,1652]" pageId="4" pageNumber="527">L</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFE44F9E5FE60F9D3" bold="true" box="[414,428,1635,1652]" pageId="4" pageNumber="527">Q</emphasis>
, endophallic plates,cingulum and aedeagal valves with sheath,lateral view; 
<emphasis id="B928EAF2FFDBCD7CFBCEF9E5FBEEF9D3" bold="true" box="[1044,1058,1635,1652]" pageId="4" pageNumber="527">H</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFBF1F9E5FBF0F9D3" bold="true" box="[1067,1084,1635,1652]" pageId="4" pageNumber="527">M</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFB9EF9E5FB9DF9D3" bold="true" box="[1092,1105,1635,1652]" pageId="4" pageNumber="527">R</emphasis>
, endophallic plates, cingulum and aedeagal valves with sheath, dorsal view; 
<emphasis id="B928EAF2FFDBCD7CFD5BF9FFFD4BF92D" bold="true" box="[641,647,1657,1674]" pageId="4" pageNumber="527">I</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFD4BF9FFFD53F92D" bold="true" box="[657,671,1657,1674]" pageId="4" pageNumber="527">N</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFD73F9FFFD79F92D" bold="true" box="[681,693,1657,1674]" pageId="4" pageNumber="527">S</emphasis>
, distal portion of aedeagus and aedeagal valves, ventral view; 
<emphasis id="B928EAF2FFDBCD7CFFABF916FFB7F906" bold="true" box="[113,123,1680,1697]" pageId="4" pageNumber="527">J</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFF5CF916FF5FF906" bold="true" box="[134,147,1680,1697]" pageId="4" pageNumber="527">K</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFF44F916FF61F906" bold="true" box="[158,173,1680,1697]" pageId="4" pageNumber="527">O</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFF6DF916FF0FF906" bold="true" box="[183,195,1680,1697]" pageId="4" pageNumber="527">P</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFF14F916FF15F906" bold="true" box="[206,217,1680,1697]" pageId="4" pageNumber="527">T</emphasis>
, 
<emphasis id="B928EAF2FFDBCD7CFF3EF916FF3EF906" bold="true" box="[228,242,1680,1697]" pageId="4" pageNumber="527">U</emphasis>
, epiphallus, dorsal and lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF236668FFDACD7DFFABF9CAFC5CF906" ID-DOI="http://doi.org/10.5281/zenodo.5166911" ID-Zenodo-Dep="5166911" httpUri="https://zenodo.org/record/5166911/files/figure.png" pageId="5" pageNumber="528" startId="5.[113,124,1612,1629]" targetBox="[172,1152,216,1500]" targetPageId="5">
<paragraph id="8BE336E0FFDACD7DFFABF9CAFC5CF906" blockId="5.[113,1211,1612,1697]" pageId="5" pageNumber="528">
FIG. 3. — 
<taxonomicName id="4C5C4D63FFDACD7DFF1EF9CAFE4DF9FA" authority="Cigliano &amp; Pocco &amp; Lange, 2011" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[196,385,1612,1629]" class="Insecta" family="Acrididae" genus="Huaylasacris" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="528" phylum="Arthropoda" rank="species" species="maxicerci" status="n. gen., n. sp.">
<emphasis id="B928EAF2FFDACD7DFF1EF9CAFE4DF9FA" box="[196,385,1612,1629]" italics="true" pageId="5" pageNumber="528">Huaylasacris maxicerci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5789FFDACD7DFE5DF9CAFE33F9FA" box="[391,511,1612,1629]" pageId="5" pageNumber="528" rank="species">n. gen., n. sp.</taxonomicNameLabel>
: 
<emphasis id="B928EAF2FFDACD7DFDDCF9CAFDDFF9FA" bold="true" box="[518,531,1612,1629]" pageId="5" pageNumber="528">A</emphasis>
, male habitus, lateral view; 
<emphasis id="B928EAF2FFDACD7DFD24F9CAFCC7F9FA" bold="true" box="[766,779,1612,1629]" pageId="5" pageNumber="528">B</emphasis>
, male head and pronotum, dorsal view; 
<emphasis id="B928EAF2FFDACD7DFB85F9CAFBA1F9FA" bold="true" box="[1119,1133,1612,1629]" pageId="5" pageNumber="528">C</emphasis>
, 
<emphasis id="B928EAF2FFDACD7DFBA2F9CAFB4AF9FA" bold="true" box="[1144,1158,1612,1629]" pageId="5" pageNumber="528">D</emphasis>
, male distal abdominal segments, dorsal and lateral views; 
<emphasis id="B928EAF2FFDACD7DFDF4F9E5FDF6F9D3" bold="true" box="[558,570,1635,1652]" pageId="5" pageNumber="528">E</emphasis>
, 
<emphasis id="B928EAF2FFDACD7DFD9EF9E5FD83F9D3" bold="true" box="[580,591,1635,1652]" pageId="5" pageNumber="528">F</emphasis>
, female head and pronotum, dorsal and lateral views; 
<emphasis id="B928EAF2FFDACD7DFBC2F9E5FBEAF9D3" bold="true" box="[1048,1062,1635,1652]" pageId="5" pageNumber="528">G</emphasis>
, phallic complex, lateral view; 
<emphasis id="B928EAF2FFDACD7DFF02F9FFFF2AF92D" bold="true" box="[216,230,1657,1674]" pageId="5" pageNumber="528">H</emphasis>
, 
<emphasis id="B928EAF2FFDACD7DFF2AF9FFFF31F92D" bold="true" box="[240,253,1657,1674]" pageId="5" pageNumber="528">K</emphasis>
, endophallic plates, cingulum and aedeagal valves with sheath, dorsal and lateral views; 
<emphasis id="B928EAF2FFDACD7DFC05F9FFFC29F92D" bold="true" box="[991,997,1657,1674]" pageId="5" pageNumber="528">I</emphasis>
, 
<emphasis id="B928EAF2FFDACD7DFC34F9FFFC34F92D" bold="true" box="[1006,1016,1657,1674]" pageId="5" pageNumber="528">J</emphasis>
, epiphallus, dorsal and lateral views; 
<emphasis id="B928EAF2FFDACD7DFF38F916FF21F906" bold="true" box="[226,237,1680,1697]" pageId="5" pageNumber="528">L</emphasis>
, distal portion of aedeagus and aedeagal valves, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF236668FFD9CD7EFFABF9FFFBECF906" ID-DOI="http://doi.org/10.5281/zenodo.5166917" ID-Zenodo-Dep="5166917" httpUri="https://zenodo.org/record/5166917/files/figure.png" pageId="6" pageNumber="529" startId="6.[113,124,1657,1674]" targetBox="[127,1196,224,1576]" targetPageId="6">
<paragraph id="8BE336E0FFD9CD7EFFABF9FFFBECF906" blockId="6.[113,1211,1657,1697]" pageId="6" pageNumber="529">
FIG. 4. — 
<taxonomicName id="4C5C4D63FFD9CD7EFF18F9FFFEBDF92D" authority="Cigliano &amp; Pocco &amp; Lange, 2011" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[194,369,1657,1674]" class="Insecta" family="Acrididae" genus="Tiyantiyana" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="sunipenis" status="n. gen., n. sp.">
<emphasis id="B928EAF2FFD9CD7EFF18F9FFFEBDF92D" box="[194,369,1657,1674]" italics="true" pageId="6" pageNumber="529">Tiyantiyana sunipenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5789FFD9CD7EFEADF9FFFE21F92D" box="[375,493,1657,1674]" pageId="6" pageNumber="529" rank="species">n. gen., n. sp.</taxonomicNameLabel>
: 
<emphasis id="B928EAF2FFD9CD7EFE2EF9FFFDCDF92D" bold="true" box="[500,513,1657,1674]" pageId="6" pageNumber="529">A</emphasis>
, male habitus, lateral view; 
<emphasis id="B928EAF2FFD9CD7EFD32F9FFFD39F92D" bold="true" box="[744,757,1657,1674]" pageId="6" pageNumber="529">B</emphasis>
, female habitus, lateral view; 
<emphasis id="B928EAF2FFD9CD7EFC36F9FFFC36F92D" bold="true" box="[1004,1018,1657,1674]" pageId="6" pageNumber="529">C</emphasis>
, 
<emphasis id="B928EAF2FFD9CD7EFBDEF9FFFBDEF92D" bold="true" box="[1028,1042,1657,1674]" pageId="6" pageNumber="529">D</emphasis>
, endophallic plates, cingulum and aedeagal valves with sheath, lateral and dorsal views; 
<emphasis id="B928EAF2FFD9CD7EFD70F916FD7AF906" bold="true" box="[682,694,1680,1697]" pageId="6" pageNumber="529">E</emphasis>
, epiphallus, dorsal view. Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="8BE336E0FFD8CD7FFFABFD93FF0DFD88" blockId="7.[113,648,533,749]" box="[113,193,533,559]" pageId="7" pageNumber="530">
<emphasis id="B928EAF2FFD8CD7FFFABFD93FF0DFD88" box="[113,193,533,559]" italics="true" pageId="7" pageNumber="530">Females</emphasis>
</paragraph>
<paragraph id="8BE336E0FFD8CD7FFFABFDB2FE0DFD4A" blockId="7.[113,648,533,749]" pageId="7" pageNumber="530">
Similar to males, but more robust and with the pronotum constricted at the middle. Tegument with clearly indicated carinulae (
<figureCitation id="13672A65FFD8CD7FFE3AFDF2FDF4FD29" box="[480,568,628,654]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[127,1196,224,1576]" captionTargetId="figure-20@6.[127,1196,215,1576]" captionTargetPageId="6" captionText="FIG. 4. — Tiyantiyana sunipenis n. gen., n. sp.: A, male habitus, lateral view; B, female habitus, lateral view; C, D, endophallic plates, cingulum and aedeagal valves with sheath, lateral and dorsal views; E, epiphallus, dorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5166917" httpUri="https://zenodo.org/record/5166917/files/figure.png" pageId="7" pageNumber="530">Fig. 4B</figureCitation>
). Head with fastigium more prominent; eyes subtrigonal (
<figureCitation id="13672A65FFD8CD7FFFA0FD35FF01FD6A" box="[122,205,691,717]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="7" pageNumber="530">Figs 1F</figureCitation>
; 
<figureCitation id="13672A65FFD8CD7FFF01FD35FF31FD6A" box="[219,253,691,717]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[127,1196,224,1576]" captionTargetId="figure-20@6.[127,1196,215,1576]" captionTargetPageId="6" captionText="FIG. 4. — Tiyantiyana sunipenis n. gen., n. sp.: A, male habitus, lateral view; B, female habitus, lateral view; C, D, endophallic plates, cingulum and aedeagal valves with sheath, lateral and dorsal views; E, epiphallus, dorsal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5166917" httpUri="https://zenodo.org/record/5166917/files/figure.png" pageId="7" pageNumber="530">4B</figureCitation>
). Ovipositor valves strong and long, with serrate margins (
<figureCitation id="13672A65FFD8CD7FFE86FD55FE7FFD4A" box="[348,435,723,749]" captionStart="FIG" captionStartId="3.[113,124,1567,1584]" captionTargetBox="[156,1167,216,1500]" captionTargetId="figure-17@3.[155,1168,215,1500]" captionTargetPageId="3" captionText="FIG. 1. — Tiyantiyana sunipenis n. gen, n. sp.: A, male habitus, lateral view; B, C, male head and pronotum, lateral and dorsal views; D, E, male distal abdominal segments, dorsal and lateral views; F, G, female head and pronotum, lateral and dorsal views; H, female ovipositor valves;I, phallic complex,lateral view;J, L, endophallic plates,cingulum and aedeagal valves with sheath, dorsal and lateral views; K, distal portion of aedeagus and aedeagal valves, ventral view; M, epiphallus, dorsal view. Abbreviations: Ap, apodemes of cingulum; Ar, arch of aedeagus; Av, aedeagal valves; Ep, endophallic plates;L, lophi of epiphallus; Rm, rami; Sh, sheath of aedeagus, Zy, zygoma. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5166902" httpUri="https://zenodo.org/record/5166902/files/figure.png" pageId="7" pageNumber="530">Fig. 1H</figureCitation>
).
</paragraph>
<paragraph id="8BE336E0FFD8CD7FFFABFC95FED9FC8C" blockId="7.[111,649,787,1415]" box="[113,277,787,813]" pageId="7" pageNumber="530">RELATIONSHIPS</paragraph>
<paragraph id="8BE336E0FFD8CD7FFFABFCB4FD4FFA20" blockId="7.[111,649,787,1415]" pageId="7" pageNumber="530">
Despite the lack of the spur on the prosternum (that may be related to the apterism condition), 
<taxonomicName id="4C5C4D63FFD8CD7FFFA9FCF4FF23FC2C" authority="Cigliano &amp; Pocco &amp; Lange, 2011" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[115,239,882,907]" class="Insecta" family="Acrididae" genus="Tiyantiyana" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF2FFD8CD7FFFA9FCF4FF23FC2C" box="[115,239,882,907]" italics="true" pageId="7" pageNumber="530">Tiyantiyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5789FFD8CD7FFF28FCF4FEF6FC2B" box="[242,314,882,908]" pageId="7" pageNumber="530" rank="genus">n. gen.</taxonomicNameLabel>
shares the remaining diagnostic characters of the subfamily 
<taxonomicName id="4C5C4D63FFD8CD7FFE7FFC17FD8CFC0C" box="[421,576,913,939]" class="Insecta" family="Acrididae" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="530" phylum="Arthropoda" rank="subFamily" subFamily="Melanoplinae">Melanoplinae</taxonomicName>
: frons convex, profile of face and fastigium united in the same curve; male abdominal terminalia swollen and upcurved, palium coriaceous; male phallic complex with the middle part of the endophallic sclerites constituted by one dorso-lateral piece; cingulum with arch clearly developed. Based on the morphological evidence it was not possible to assign the genus to any of the three tribes described for the South American 
<taxonomicName id="4C5C4D63FFD8CD7FFEF9FB28FE7BFB6F" box="[291,439,1198,1224]" class="Insecta" family="Acrididae" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="530" phylum="Arthropoda" rank="subFamily" subFamily="Melanoplinae">Melanoplinae</taxonomicName>
(
<bibRefCitation id="EFCD4B11FFD8CD7FFE19FB28FF66FB4F" author="AMEDEGNATO C. &amp; CHAPCO W. &amp; LITZENBERGER G." pageId="7" pageNumber="530" pagination="115 - 119" refId="ref10910" refString="AMEDEGNATO C., CHAPCO W. &amp; LITZENBERGER G. 2003. - Out of South America? Additional evidence for a shouthern origin of melanopline grasshoppers. Molecular Phylogenetics and Evolution 29: 115 - 119." type="journal article" year="2003">
Amédégnato 
<emphasis id="B928EAF2FFD8CD7FFD89FB29FDB1FB6F" box="[595,637,1198,1224]" italics="true" pageId="7" pageNumber="530">et al</emphasis>
. 2003
</bibRefCitation>
; 
<bibRefCitation id="EFCD4B11FFD8CD7FFF69FB4BFE43FB4F" author="CARBONELL C. S. &amp; CIGLIANO M. M. &amp; LANGE C. E." box="[179,399,1229,1256]" pageId="7" pageNumber="530" refId="ref11155" refString="CARBONELL C. S., CIGLIANO M. M. &amp; LANGE C. E. 2006. - Acridomorph (Orthoptera) species of Argentina and Uruguay / Especies de Acridomorfos (Orthoptera) de Argentina y Uruguay. CD-ROM. Publications on Orthopteran Diversity. The Orthopterists' Society at the Museo de La Plata." type="book" year="2006">
Carbonell 
<emphasis id="B928EAF2FFD8CD7FFEC4FB48FE82FB40" box="[286,334,1229,1255]" italics="true" pageId="7" pageNumber="530">et al.</emphasis>
2006
</bibRefCitation>
): 
<taxonomicName id="4C5C4D63FFD8CD7FFE44FB4BFD4EFB4F" authority="Hebard, 1924" authorityName="Hebard" authorityYear="1924" box="[414,642,1229,1256]" class="Insecta" family="Acrididae" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="530" phylum="Arthropoda" rank="tribe" tribe="Jivarini">Jivarini Hebard, 1924</taxonomicName>
, 
<taxonomicName id="4C5C4D63FFD8CD7FFFB5FB6BFE10FAA0" authority="Rehn &amp; Randell, 1963" authorityName="Rehn &amp; Randell" authorityYear="1963" box="[111,476,1261,1287]" class="Insecta" family="Acrididae" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="530" phylum="Arthropoda" rank="tribe" tribe="Dichroplini">Dichroplini Rehn &amp; Randell, 1963</taxonomicName>
and Parascopini 
<bibRefCitation id="EFCD4B11FFD8CD7FFFB5FA8BFEE7FA80" author="RONDEROS R. A." box="[111,299,1293,1319]" pageId="7" pageNumber="530" pagination="1 - 4" refId="ref12073" refString="RONDEROS R. A. 1983. - Notas sobre Melanoplinae neotropicales (Orthoptera, Acrididae). Revista de la Sociedad Entomologica Argentina 42 (1 - 4): 131." type="journal article" year="1983">Ronderos, 1983</bibRefCitation>
. However, molecular analyses currently underway have shown that 
<taxonomicName id="4C5C4D63FFD8CD7FFDD0FAABFD44FAE1" authority="Cigliano &amp; Pocco &amp; Lange, 2011" authorityName="Cigliano &amp; Pocco &amp; Lange" authorityYear="2011" box="[522,648,1325,1350]" class="Insecta" family="Acrididae" genus="Tiyantiyana" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EAF2FFD8CD7FFDD0FAABFD44FAE1" box="[522,648,1325,1350]" italics="true" pageId="7" pageNumber="530">Tiyantiyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5789FFD8CD7FFFABFACBFF0DFAC0" box="[113,193,1357,1383]" pageId="7" pageNumber="530" rank="genus">n. gen.</taxonomicNameLabel>
is grouped with the remaining Andean melanoplines genera (unpublished observations).
</paragraph>
</subSubSection>
</treatment>
</document>