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<document id="A3ABF4C5816365C843814A186B59375C" ID-DOI="10.5281/zenodo.195468" ID-GBIF-Dataset="e2290a51-8f8f-4a97-95a9-5a8710c4ece4" ID-ISSN="1175-5326" ID-Zenodo-Dep="195468" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460227508737" checkinUser="plazi" docAuthor="Siler, Cameron D., Diesmos, Arvin C., Linkem, Charles W., Diesmos, Mae L. &amp; Brown, Rafe M." docDate="2010" docId="03F587EFFFCAFFEC5BB593F6FDC2FD0D" docLanguage="en" docName="zt02482p063.pdf" docOrigin="Zootaxa 2482" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Platymantis biak Siler, Diesmos, Linkem, Diesmos &amp; Brown, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="59" masterDocId="FFCCFF97FFC8FFE65B22932BFFA2FFEF" masterDocTitle="A new species of limestone-forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae) from central Luzon Island, Philippines" masterLastPageNumber="63" masterPageNumber="49" pageNumber="51" updateTime="1698238669337" updateUser="plazi">
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<mods:title id="78DC3F942AD20E2924ABFFA2320B3658">A new species of limestone-forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae) from central Luzon Island, Philippines</mods:title>
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<mods:namePart id="3A0AE1E0F078A445ABA8CB974FC8FB78">Siler, Cameron D.</mods:namePart>
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<mods:namePart id="57324F7C32A248D292F7281B0952C9C2">Diesmos, Arvin C.</mods:namePart>
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<mods:namePart id="AE8DFA00961EF3E21F0A66F0A822BF74">Linkem, Charles W.</mods:namePart>
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<mods:namePart id="6C0A2F72222CD2CCD130009E62E44A6C">Diesmos, Mae L.</mods:namePart>
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<mods:namePart id="EEA49D98D66558D50889BAEC280A60E8">Brown, Rafe M.</mods:namePart>
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<mods:date id="D385089265DD947B5F49B469817CC4F4">2010</mods:date>
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<treatment id="03F587EFFFCAFFEC5BB593F6FDC2FD0D" ID-DOI="http://doi.org/10.5281/zenodo.5628517" ID-GBIF-Taxon="119386769" ID-Zenodo-Dep="5628517" LSID="urn:lsid:plazi:treatment:03F587EFFFCAFFEC5BB593F6FDC2FD0D" httpUri="http://treatment.plazi.org/id/03F587EFFFCAFFEC5BB593F6FDC2FD0D" lastPageId="10" lastPageNumber="59" pageId="2" pageNumber="51">
<subSubSection id="C3466572FFCAFFE45BB593F6FEA1FEF5" pageId="2" pageNumber="51" type="nomenclature">
<paragraph id="8BE336F9FFCAFFE45BB593F6FE64FF18" blockId="2.[151,454,221,282]" box="[151,454,221,247]" pageId="2" pageNumber="51">
<heading id="D0AB8195FFCAFFE45BB593F6FE64FF18" bold="true" box="[151,454,221,247]" fontSize="11" level="1" pageId="2" pageNumber="51" reason="1">
<emphasis id="B928EAEBFFCAFFE45BB593F6FE64FF18" bold="true" box="[151,454,221,247]" pageId="2" pageNumber="51">
<taxonomicName id="4C5C4D7AFFCAFFE45BB593F6FEC3FF18" ID-CoL="4JSPC" box="[151,353,221,247]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="biak" status="sp. nov.">
<emphasis id="B928EAEBFFCAFFE45BB593F6FEC3FF18" bold="true" box="[151,353,221,247]" italics="true" pageId="2" pageNumber="51">Platymantis biak</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B5790FFCAFFE45A4A93F6FE64FF18" box="[360,454,221,247]" pageId="2" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE336F9FFCAFFE45BB5922BFEA1FEF5" blockId="2.[151,454,221,282]" box="[151,259,256,282]" pageId="2" pageNumber="51">Figs. 24</paragraph>
</subSubSection>
<subSubSection id="C3466572FFCAFFE45BB59261FEBBFD5B" pageId="2" pageNumber="51" type="materials_examined">
<paragraph id="8BE336F9FFCAFFE45BB59261FE6DFE3B" blockId="2.[151,1437,330,1999]" pageId="2" pageNumber="51">
<emphasis id="B928EAEBFFCAFFE45BB59261FEB6FE8B" bold="true" box="[151,276,330,356]" pageId="2" pageNumber="51">
<typeStatus id="54E7885BFFCAFFE45BB59261FEB2FE8B" box="[151,272,330,356]" pageId="2" pageNumber="51" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F34B7669FFCAFFE45A3F9261FEFCFE8B" box="[285,350,330,356]" name="Panama" pageId="2" pageNumber="51">
<collectionCode id="ED4DAE3CFFCAFFE45A3F9261FEFCFE8B" box="[285,350,330,356]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="2" pageNumber="51">PNM</collectionCode>
</collectingCountry>
9679 (
<collectionCode id="ED4DAE3CFFCAFFE45A969261FE54FE8B" box="[436,502,330,356]" pageId="2" pageNumber="51">ACD</collectionCode>
Field No. 4543; KU 319973), adult male, collected on limestone outcrop in secondary-growth forest (20:00 hr) on
<date id="FFE21039FFCAFFE45959925BFCEAFE65" box="[635,840,368,394]" pageId="2" pageNumber="51" value="2009-01-17">
17 January
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</date>
in Barangay
<taxonomicName id="4C5C4D7AFFCAFFE458D2925BFB88FE65" box="[1008,1066,368,394]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="biak">Biak</taxonomicName>
na Bato, Municipality of San Miguel and Doña Remedios
<collectingCountry id="F34B7669FFCAFFE45AC992BEFDF0FE40" box="[491,594,405,431]" name="Trinidad and Tobago" pageId="2" pageNumber="51">Trinidad</collectingCountry>
, Bulacan Province, Luzon Island,
<collectingCountry id="F34B7669FFCAFFE458CB92BEFBCFFE40" box="[1001,1133,405,431]" name="Philippines" pageId="2" pageNumber="51">Philippines</collectingCountry>
(N 15.1084, E 121.0724;
<quantity id="4CA49B1CFFCAFFE45BB59291FF43FE3B" box="[151,225,442,468]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="51" unit="m" value="190.0">190 m</quantity>
elevation), by
<collectionCode id="ED4DAE3CFFCAFFE45AB39291FE6DFE3B" box="[401,463,442,468]" pageId="2" pageNumber="51">ACD</collectionCode>
</paragraph>
<paragraph id="8BE336F9FFCAFFE45BE692F4FEBBFD5B" blockId="2.[151,1437,330,1999]" pageId="2" pageNumber="51">
<emphasis id="B928EAEBFFCAFFE45BE692F4FE20FE16" bold="true" box="[196,386,479,505]" pageId="2" pageNumber="51">Paratopotypes.</emphasis>
Twenty-three males (KU 31997778, 31998182, 319987,
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<collectionCode id="ED4DAE3CFFCAFFE45F6C92CBFB2CFE15" box="[1102,1166,480,506]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="2" pageNumber="51">PNM</collectionCode>
</collectingCountry>
9680, 9681, 968283, 96849697), eight females (KU 31997475, 319979,
<collectingCountry id="F34B7669FFCAFFE4583D912EFCC2FDF0" box="[799,864,517,543]" name="Panama" pageId="2" pageNumber="51">
<collectionCode id="ED4DAE3CFFCAFFE4583D912EFCC2FDF0" box="[799,864,517,543]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="2" pageNumber="51">PNM</collectionCode>
</collectingCountry>
967475, 9676, 967778), and seven juveniles (KU 319976, 319980, 319983,
<collectingCountry id="F34B7669FFCAFFE459329101FDF3FDAB" box="[528,593,554,580]" name="Panama" pageId="2" pageNumber="51">
<collectionCode id="ED4DAE3CFFCAFFE459329101FDF3FDAB" box="[528,593,554,580]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="2" pageNumber="51">PNM</collectionCode>
</collectingCountry>
96989701) collected on karst formations in secondary-growth forest (18:0023:00),
<date id="FFE21039FFCAFFE45A6D917BFDA7FD85" box="[335,517,592,618]" pageId="2" pageNumber="51" value="2009-01" valueMax="2009-03-31" valueMin="2009-01-00">JanuaryMarch</date>
and November 2009, in Barangay
<taxonomicName id="4C5C4D7AFFCAFFE45885917BFC7DFD85" box="[935,991,592,618]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="biak">Biak</taxonomicName>
na Bato, Municipality of San Miguel and Doña Remedios
<collectingCountry id="F34B7669FFCAFFE45AAD915EFE55FD60" box="[399,503,629,655]" name="Trinidad and Tobago" pageId="2" pageNumber="51">Trinidad</collectingCountry>
, Bulacan Province, Luzon Island,
<collectingCountry id="F34B7669FFCAFFE458AD915EFBBAFD60" box="[911,1048,629,655]" name="Philippines" pageId="2" pageNumber="51">Philippines</collectingCountry>
; collected by
<collectionCode id="ED4DAE3CFFCAFFE45F9E915EFB5FFD60" box="[1212,1277,629,655]" pageId="2" pageNumber="51">ACD</collectionCode>
,
<collectionCode id="ED4DAE3CFFCAFFE45E28915EFAE4FD60" LSID="urn:lsid:biocol.org:col:15840" box="[1290,1350,629,655]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15840" name="Charles Darwin Research Station" pageId="2" pageNumber="51">CDS</collectionCode>
,
<collectionCode id="ED4DAE3CFFCAFFE45E71915EFA35FD60" box="[1363,1431,629,655]" pageId="2" pageNumber="51">CWL</collectionCode>
, and. MLD.
</paragraph>
</subSubSection>
<subSubSection id="C3466572FFCAFFE45BE69194FC98FBC5" pageId="2" pageNumber="51" type="diagnosis">
<paragraph id="8BE336F9FFCAFFE45BE69194FC98FBC5" blockId="2.[151,1437,330,1999]" pageId="2" pageNumber="51">
<emphasis id="B928EAEBFFCAFFE45BE69194FEE1FD36" bold="true" box="[196,323,703,729]" pageId="2" pageNumber="51">Diagnosis.</emphasis>
We assign the new species to the genus
<taxonomicName id="4C5C4D7AFFCAFFE4583C91EBFC08FD36" box="[798,938,704,729]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="B928EAEBFFCAFFE4583C91EBFC08FD36" box="[798,938,704,729]" italics="true" pageId="2" pageNumber="51">Platymantis</emphasis>
</taxonomicName>
on the basis of (1) its moderate body size, (2) absence of interdigital webbing, (3) the presence of a forked omosternum, (4) directly developing reproductive mode, and (5) terrestrial microhabitat preference (Brown and Alcala 1980;
<bibRefCitation id="EFCD4B08FFCAFFE45F979021FA35FCCB" author="Brown" box="[1205,1431,778,804]" pageId="2" pageNumber="61" refString="Brown, W. C., Brown, R. M. &amp; Alcala, A. C. (1997 a) Species of the hazelae group of Platymantis from the Philippines, with descriptions of two new species. Proceedings of the California Academy of Sciences, 49, 405 - 421." type="journal article" year="1997">
Brown
<emphasis id="B928EAEBFFCAFFE45E2F9021FAEBFCCC" box="[1293,1353,778,803]" italics="true" pageId="2" pageNumber="51">et al.</emphasis>
, 1997
</bibRefCitation>
; Brown, 2004).
<taxonomicName id="4C5C4D7AFFCAFFE45A71901BFD86FCA6" box="[339,548,816,841]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFCAFFE45A71901BFD86FCA6" box="[339,548,816,841]" italics="true" pageId="2" pageNumber="51">Platymantis biak</emphasis>
</taxonomicName>
can be distinguished from other
<collectingCountry id="F34B7669FFCAFFE458E3901BFBE2FCA5" box="[961,1088,816,842]" name="Philippines" pageId="2" pageNumber="51">Philippine</collectingCountry>
congeners by the following combination of characters: body size
<quantity id="4CA49B1CFFCAFFE45971907EFCA2FC80" box="[595,768,853,879]" metricMagnitude="-2" metricUnit="m" metricValue="3.6099999999999994" metricValueMax="3.9899999999999998" metricValueMin="3.2299999999999995" pageId="2" pageNumber="51" unit="mm" value="36.099999999999994" valueMax="39.9" valueMin="32.3">32.339.9 mm</quantity>
SVL for males,
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SVL for females; digital disc of Finger III 1.92.2
<emphasis id="B928EAEBFFCAFFE45AEF9051FE7FFC7C" box="[461,477,890,915]" italics="true" pageId="2" pageNumber="51">×</emphasis>
width of penultimate phalanx in males and
<quantity id="4CA49B1CFFCAFFE458CF9051FBC7FC7B" box="[1005,1125,890,916]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" metricValueMax="6.096" metricValueMin="3.5559999999999996" pageId="2" pageNumber="51" unit="in" value="1.9" valueMax="2.4" valueMin="1.4">1.42.4 in</quantity>
females (Fig. 3); Finger I slightly longer than Finger II; finger discs larger than toe discs; smooth skin; spotted flanks absent; supratympanic fold smooth, less protuberant; tympanum completely visible; dorsolateral stripes absent; dorsal folds, ridges, dermal tubercles absent; and a preference for terrestrial microhabitat in karst forest. Mensural and diagnostic differences are provided in
<tableCitation id="C6DE0342FFCAFFE459A8973BFD4EFBC5" box="[650,748,1040,1066]" captionStart="TABLE 1" captionStartId="6.[151,240,151,175]" captionTargetBox="[165,1412,367,1936]" captionTargetPageId="6" captionText="TABLE 1. Summary of qualitative diagnostic characters (present, absent) in Platymantis biak and specimens of other members of the P. guentheri Species Group. Sample size for each sex, body size, eye diameter, digit disk proportions, and general geographical distribution (PAIC = Pleistocene Aggregate Island Complexes, sensu Brown and Diesmos, 2002) are included for reference (SVL and ED given as range over mean ± standard deviation; digit disk proportions given as percentage over mean ± standard deviation). Only adult specimens in good preservation condition were used for morphometric analyses." httpUri="http://table.plazi.org/id/DF236671FFCEFFE05BB593BCFE34FEA0" pageId="2" pageNumber="51" tableUuid="DF236671FFCEFFE05BB593BCFE34FEA0">Tables 1</tableCitation>
and
<tableCitation id="C6DE0342FFCAFFE45807973BFC95FBC5" box="[805,823,1040,1066]" captionStart="TABLE 2" captionStartId="8.[151,240,151,175]" captionText="TABLE 2. Summary of mensural characters (in mm) and selected qualitative diagnostic characters (present, absent) in Platymantis biak and other Philippine limestone forest frog species. Sample size, body size, species group, and general geographical distribution are included for reference (SVL given as range)." pageId="2" pageNumber="51">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3466572FFCAFFEE5BE6971EFE29FB5C" lastPageId="8" lastPageNumber="57" pageId="2" pageNumber="51" type="description">
<paragraph id="8BE336F9FFCAFFE45BE6971EFEE8FA70" blockId="2.[151,1437,330,1999]" pageId="2" pageNumber="51">
<emphasis id="B928EAEBFFCAFFE45BE6971EFE59FBA0" bold="true" box="[196,507,1077,1103]" pageId="2" pageNumber="51">
Description of
<typeStatus id="54E7885BFFCAFFE45AA6971EFE57FBA0" box="[388,501,1077,1103]" pageId="2" pageNumber="51" type="holotype">holotype</typeStatus>
.
</emphasis>
A mature male; habitus robust; head broader than body, head length 35.5% snoutvent length, 90.4% head width; snout protruding moderately beyond lower jaw, snout tip comes to moderately sharp point in dorsal and lateral aspects; eyes moderate and protruding anterolaterally, eye diameter 12.7% snoutvent length, 35.9% head length; lips slightly flared and swollen (Fig. 2); interorbital region slightly convex; eye diameter 80.9% snout length, 134.2% eyenarial distance; pupil horizontally elliptical; canthus rostralis anteriorly concave in dorsal aspect; loreal region moderately concave, sloping ventrolaterally to labium; nostrils oriented posterolaterally; eyenarial distance 60.3% snout length; internarial region slightly concave; tympanic annulus distinct 56.8% eye diameter; supratympanic fold slightly protuberant, extending from posterior corner of eye across dorsal margin of tympanum to supraaxillary region.
</paragraph>
<paragraph id="8BE336F9FFCAFFE45BE69681FE6FF990" blockId="2.[151,1437,330,1999]" pageId="2" pageNumber="51">
Tongue elongate, wider posteriorly, with deep posterior notch and narrow anterior attachment; choanae situated at anterolateral edge of palate, oval, separated by distance two times greater than diameter of single choana; dentigerous process of vomer distinct, with no teeth visible on process; dentigerous process angled slightly anterolaterally, approximately at 2030º incline with closest (posterior) points separated by distance equal to 2.0
<emphasis id="B928EAEBFFCAFFE45A04956BFE94F9B6" box="[294,310,1600,1625]" italics="true" pageId="2" pageNumber="51">×</emphasis>
diameter of single choana, their most distant (anterior) ends separated by distance equal to 3.0
<emphasis id="B928EAEBFFCAFFE45EAE956BFA3EF9B6" box="[1420,1436,1600,1625]" italics="true" pageId="2" pageNumber="51">×</emphasis>
diameter of single choana.
</paragraph>
<paragraph id="8BE336F9FFCAFFE45BE695A1FC3DF820" blockId="2.[151,1437,330,1999]" pageId="2" pageNumber="51">Hand length 54.6% foot length; tibia length 113.0% foot length, 54.3% snoutvent length; fingers slender, long; terminal discs moderately expanded (Fig. 3A) with pronounced distal, circum-marginal groove; dorsal surfaces of terminal phalanges with small, cutaneous supra-articular flaps between ultimate and penultimate phalanges; relative lengths of fingers II &lt;I &lt;IV &lt;III; subarticular tubercles prominent, convex, smooth; one subarticular tubercle below Digits I and II, two tubercles under Digits III and IV; supernumerary tubercles present at bases of Digits IIV, distinct, round, prominent for Digits IIIV; inner metacarpal tubercle enlarged, elongate, oval shaped; outer metacarpal tubercle fully divided into round, slightly raised postaxial tubercle equal in size to supernumerary tubercle, and oval preaxial tubercle equal in size to preaxial metacarpal tubercle; nuptial pads absent; forearms slender, not hypertrophied.</paragraph>
<paragraph id="8BE336F9FFCBFFE55BB596BCFC22FA01" blockId="3.[151,1436,1431,1518]" pageId="3" pageNumber="52">
<emphasis id="B928EAEBFFCBFFE55BB596BCFE82FA40" bold="true" box="[151,288,1431,1455]" pageId="3" pageNumber="52">FIGURE. 2.</emphasis>
Photographs in life of
<taxonomicName id="4C5C4D7AFFCBFFE5593B96B3FD6CFA40" box="[537,718,1432,1455]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFCBFFE5593B96B3FD6CFA40" box="[537,718,1432,1455]" italics="true" pageId="3" pageNumber="52">Platymantis biak</emphasis>
</taxonomicName>
paratopotypes exhibiting (A) typical limestone-forest frog mottled color pattern,
<collectingCountry id="F34B7669FFCBFFE55A0C969CFECAFA21" box="[302,360,1463,1486]" name="Panama" pageId="3" pageNumber="52">PNM</collectingCountry>
9678, female, SVL =
<quantity id="4CA49B1CFFCBFFE5597B969CFD1EFA21" box="[601,700,1463,1486]" metricMagnitude="-2" metricUnit="m" metricValue="4.07" pageId="3" pageNumber="52" unit="mm" value="40.7">40.7 mm</quantity>
, and (B, C) perching behavior observed at
<typeStatus id="54E7885BFFCBFFE55FAF969CFB19FA21" box="[1165,1211,1463,1486]" pageId="3" pageNumber="52">type</typeStatus>
locality,
<collectingCountry id="F34B7669FFCBFFE55E3D969CFAFBFA21" box="[1311,1369,1463,1486]" name="Panama" pageId="3" pageNumber="52">PNM</collectingCountry>
9674, female, SVL =
<quantity id="4CA49B1CFFCBFFE55A1996FCFE3CFA01" box="[315,414,1495,1518]" metricMagnitude="-2" metricUnit="m" metricValue="4.08" pageId="3" pageNumber="52" unit="mm" value="40.8">40.8 mm</quantity>
. Photographs by A. Diesmos and C. Linkem.
</paragraph>
<paragraph id="8BE336F9FFCBFFE55BE69533FD52F903" blockId="3.[151,1437,1560,2034]" pageId="3" pageNumber="52">Skin of dorsal and ventral surfaces of head, trunk, and limbs smooth. Terminal discs of toes slightly expanded, with circum-marginal grooves; supra-articular cutaneous flaps above ultimate-penultimate phalangeal articulations; plantar surface of foot smooth, with well-developed, prominently rounded subarticular tubercles (Fig. 3B); supernumerary tubercles absent; relative lengths of toes I &lt;II &lt;V &lt;III &lt;IV; outer metatarsal tubercle small, round; inner metatarsal tubercle moderately prominent, elongate, length 1.7 x length of round, pointed, outer, metatarsal tubercle.</paragraph>
<paragraph id="8BE336F9FFCBFFE55BE695DCFEAFF81D" blockId="3.[151,1437,1560,2034]" pageId="3" pageNumber="52">
<emphasis id="B928EAEBFFCBFFE55BE695DCFD25F8FE" bold="true" box="[196,647,1783,1809]" pageId="3" pageNumber="52">
Measurements of
<typeStatus id="54E7885BFFCBFFE55A8495DCFDB1F8FE" box="[422,531,1783,1809]" pageId="3" pageNumber="52" type="holotype">holotype</typeStatus>
(in mm).
</emphasis>
Snoutvent length 39.9; head length 14.2; head width 15.7; snout length 6.3; upper arm length 9.0; forearm length 9.3; tibia length 21.7; femur length 20.4; tarsus length 9.4; foot length 19.2; hand length 10.5; Finger I length 7.6; Finger III length 7.6; Toe IV length 10.4; eyenarial distance 3.8; snoutnarial distance 2.0; interorbital distance 3.3; internarial distance 3.2; eyetympanum distance 0.8; eye diameter 5.1; horizontal tympanic annulus diameter 2.9; Finger I disc width 1.2; Finger III disc width 1.8; Toe IV disc width 1.2; Finger III penultimate phalange width 0.6; Toe IV penultimate phalange width 0.5.
</paragraph>
<paragraph id="8BE336F9FFCCFFE25BB59655FE63FA38" blockId="4.[151,1436,1406,1495]" pageId="4" pageNumber="53">
<emphasis id="B928EAEBFFCCFFE25BB59655FE82FA79" bold="true" box="[151,288,1406,1430]" pageId="4" pageNumber="53">FIGURE. 3.</emphasis>
Ventral views of hand (A) and foot (B) of female paratopotype of
<taxonomicName id="4C5C4D7AFFCCFFE258C59655FB39FA7A" box="[999,1179,1406,1429]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFCCFFE258C59655FB39FA7A" box="[999,1179,1406,1429]" italics="true" pageId="4" pageNumber="53">Platymantis biak</emphasis>
</taxonomicName>
(KU 319974), hand (C) and foot (D) of male
<taxonomicName id="4C5C4D7AFFCCFFE25A5F96B5FDF9FA5A" box="[381,603,1438,1461]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="B928EAEBFFCCFFE25A5F96B5FDF9FA5A" box="[381,603,1438,1461]" italics="true" pageId="4" pageNumber="53">Platymantis dorsalis</emphasis>
</taxonomicName>
(KU
<date id="FFE21039FFCCFFE259BB96B5FD4CFA5A" box="[665,750,1438,1461]" pageId="4" pageNumber="53" value="56-03-30">300356</date>
), and hand (E) and foot (F) of male
<taxonomicName id="4C5C4D7AFFCCFFE25F5596B5FAC7FA5A" box="[1143,1381,1438,1461]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B928EAEBFFCCFFE25F5596B5FAC7FA5A" box="[1143,1381,1438,1461]" italics="true" pageId="4" pageNumber="53">Platymantis guentheri</emphasis>
</taxonomicName>
(KU 306324). Scale bar =
<quantity id="4CA49B1CFFCCFFE25A5E96EBFE1FFA38" box="[380,445,1472,1495]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="53" unit="mm" value="2.0">2 mm</quantity>
.
</paragraph>
<paragraph id="8BE336F9FFCCFFE35BE69528FE81FECD" blockId="4.[151,1437,1539,2014]" lastBlockId="5.[151,1437,152,626]" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53">
<emphasis id="B928EAEBFFCCFFE25BE69528FD06F9F2" bold="true" box="[196,676,1539,1565]" pageId="4" pageNumber="53">
Coloration of
<typeStatus id="54E7885BFFCCFFE25A539528FE7EF9F2" box="[369,476,1539,1565]" pageId="4" pageNumber="53" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
The dominant dorsal color pattern on the head and body consists of dark mottled blotches of black interspersed with blotches of dark shades of brownish green. The dorsal surfaces of the limbs are banded with alternating black and dark green bars, separated by thin tan bands. An interorbital bar is present consisting of a thin, irregular, light-brown line. The head and snout region are darker than the dorsum, and nearly uniform black. A black bar extends across the dorsal half of the canthus rostralis to the anterior edge of eye. The tympanum is medium brownish-green with a dark blotch extending posterodorsally from the center. The temporal region is predominantly black with several small blotches of dark green. Lip coloration consists of alternating dark green and black vertical labial bars, bordered anteriorly by black lower edge of canthal and subocular regions. The transverse bars on the limbs have irregular widths, with an overall lighter coloration when compared to the dorsum due to the greater proportion of skin covered by tan and dark green bands. Arm banding patterns coalesce into mottled patterns on the elbows. Four black bars are present on the forearms, thighs, and tibia, with three bars present on the tarsus. Anterior flank coloration is predominantly black, progressively fading to lighter grayish-tan with dark brown blotches and spots, blending into a black, dark brown, and tan mottled pattern posteriorly. The dorsal surfaces of the hands are light brown, with alternating bands of light brown and grayish-tan on Fingers IIII, and dark brown and grayish-tan on Finger IV. The dorsal surfaces of the feet are colored with alternating bands of light brown and grayish-tan.
</paragraph>
<paragraph id="8BE336F9FFCDFFE35BE69206FE74FD9D" blockId="5.[151,1437,152,626]" pageId="5" pageNumber="54">The throat and chest are marbled light cream with several small tan speckles, while the periphery of the throat and lower lips are darker, with the anterior edge of the lower lip possessing an irregular outline of dark green blotches. The belly is homogeneous light tan bordered by light cream spots laterally. Ventral coloration on the arms consists of light tan coloration present on the upper arm and interior portions of the forearm, and dark green coloration present on the outer portions of the forearm and hand. Ventral surfaces of the thighs uniformly mottled cream anteriorly and light green-brown posteriorly. This coloration blends gradually down the legs into a homogenous dark brown color pattern on the tarsus and foot, similar to the dark coloration observed on the forearms. The subarticular tubercles of the hands and feet are light cream, and the iris is dark blue-gray around the pupil.</paragraph>
<paragraph id="8BE336F9FFCDFFE35BB594BBFEF8F827" blockId="5.[151,1437,1936,1992]" pageId="5" pageNumber="54">
<emphasis id="B928EAEBFFCDFFE35BB594BBFE82F847" bold="true" box="[151,288,1936,1960]" pageId="5" pageNumber="54">FIGURE. 4.</emphasis>
Photographs in preservative of
<taxonomicName id="4C5C4D7AFFCDFFE3595494BAFC89F847" box="[630,811,1937,1960]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="54" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFCDFFE3595494BAFC89F847" box="[630,811,1937,1960]" italics="true" pageId="5" pageNumber="54">Platymantis biak</emphasis>
</taxonomicName>
<typeStatus id="54E7885BFFCDFFE3581094BAFC2DF847" box="[818,911,1937,1960]" pageId="5" pageNumber="54" type="holotype">holotype</typeStatus>
(
<collectingCountry id="F34B7669FFCDFFE358BF94BAFC7AF847" box="[925,984,1937,1960]" name="Panama" pageId="5" pageNumber="54">PNM</collectingCountry>
9679) in dorsal and lateral views of head. Scale bar =
<quantity id="4CA49B1CFFCDFFE35A37949AFEF5F827" box="[277,343,1969,1992]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="54" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<caption id="DF236671FFCEFFE05BB593BCFE34FEA0" ID-Table-UUID="DF236671FFCEFFE05BB593BCFE34FEA0" httpUri="http://table.plazi.org/id/DF236671FFCEFFE05BB593BCFE34FEA0" pageId="6" pageNumber="55" targetBox="[165,1412,367,1936]" targetIsTable="true" targetPageId="6">
<paragraph id="8BE336F9FFCEFFE05BB593BCFE34FEA0" blockId="6.[151,1437,151,335]" pageId="6" pageNumber="55">
<emphasis id="B928EAEBFFCEFFE05BB593BCFEAEFF40" bold="true" box="[151,268,151,175]" pageId="6" pageNumber="55">TABLE 1.</emphasis>
Summary of qualitative diagnostic characters (present, absent) in
<taxonomicName id="4C5C4D7AFFCEFFE058C293B3FB3AFF40" box="[992,1176,152,175]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFCEFFE058C293B3FB3AFF40" box="[992,1176,152,175]" italics="true" pageId="6" pageNumber="55">Platymantis biak</emphasis>
</taxonomicName>
and specimens of other members of the
<taxonomicName id="4C5C4D7AFFCEFFE05A6A9393FE69FF20" box="[328,459,184,207]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B928EAEBFFCEFFE05A6A9393FE69FF20" box="[328,459,184,207]" italics="true" pageId="6" pageNumber="55">P. guentheri</emphasis>
</taxonomicName>
Species Group. Sample size for each sex, body size, eye diameter, digit disk proportions, and general geographical distribution (PAIC = Pleistocene Aggregate Island Complexes, sensu Brown and Diesmos, 2002) are included for reference (SVL and ED given as range over mean ± standard deviation; digit disk proportions given as percentage over mean ± standard deviation). Only adult specimens in good preservation condition were used for morphometric analyses.
</paragraph>
</caption>
<paragraph id="8BE336F9FFCEFFE05A88925BFA9BF89F" pageId="6" pageNumber="55">
<table id="F95CC459FFCE00195B879244FA26F87F" box="[165,1412,367,1936]" gridcols="6" gridrows="18" pageId="6" pageNumber="55" rowsContinueIn="7.[165,1409,215,1705]">
<tr id="356C34BBFFCE00195B879244FA26FE45" box="[165,1412,367,426]" gridrow="0" pageId="6" pageNumber="55" rowspan-0="1">
<th id="76BD5DC7FFCE00195A8B9244FD99FE45" box="[425,571,367,426]" gridcol="1" gridrow="0" pageId="6" pageNumber="55">
<taxonomicName id="4C5C4D7AFFCEFFE05A88925BFE71FE6A" box="[426,467,368,389]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="biak" status="sp. nov.">
<emphasis id="B928EAEBFFCEFFE05A88925BFE71FE6A" box="[426,467,368,389]" italics="true" pageId="6" pageNumber="55">biak</emphasis>
</taxonomicName>
<emphasis id="B928EAEBFFCEFFE05AFB9244FD85FE6B" bold="true" box="[473,551,367,388]" pageId="6" pageNumber="55">
<taxonomicNameLabel id="A21B5790FFCEFFE05AFB9244FD85FE6B" box="[473,551,367,388]" pageId="6" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(24m, 8f)
</th>
<th id="76BD5DC7FFCE001959AF9244FCBCFE45" box="[653,798,367,426]" gridcol="2" gridrow="0" pageId="6" pageNumber="55">
<taxonomicName id="4C5C4D7AFFCEFFE059AF925BFD6CFE6A" box="[653,718,368,389]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="bayani">
<emphasis id="B928EAEBFFCEFFE059AF925BFD6CFE6A" box="[653,718,368,389]" italics="true" pageId="6" pageNumber="55">bayani</emphasis>
</taxonomicName>
(9m, 9f)
</th>
<th id="76BD5DC7FFCE0019586F9244FC4FFE45" box="[845,1005,367,426]" gridcol="3" gridrow="0" pageId="6" pageNumber="55">
<taxonomicName id="4C5C4D7AFFCEFFE0586F925BFC06FE6A" box="[845,932,368,389]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="insulatus">
<emphasis id="B928EAEBFFCEFFE0586F925BFC06FE6A" box="[845,932,368,389]" italics="true" pageId="6" pageNumber="55">insulatus</emphasis>
</taxonomicName>
(8m, 6f)
</th>
<th id="76BD5DC7FFCE00195F349244FB60FE45" box="[1046,1218,367,426]" gridcol="4" gridrow="0" pageId="6" pageNumber="55">
<taxonomicName id="4C5C4D7AFFCEFFE05F34925BFB2AFE6A" box="[1046,1160,368,389]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="negrosensis">
<emphasis id="B928EAEBFFCEFFE05F34925BFB2AFE6A" box="[1046,1160,368,389]" italics="true" pageId="6" pageNumber="55">negrosensis</emphasis>
</taxonomicName>
(3m, 2f)
</th>
<th id="76BD5DC7FFCE00195FC19244FA26FE45" box="[1251,1412,367,426]" gridcol="5" gridrow="0" pageId="6" pageNumber="55">
<taxonomicName id="4C5C4D7AFFCEFFE05FC1925BFAE5FE6A" box="[1251,1351,368,389]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="luzonensis">
<emphasis id="B928EAEBFFCEFFE05FC1925BFAE5FE6A" box="[1251,1351,368,389]" italics="true" pageId="6" pageNumber="55">luzonensis</emphasis>
</taxonomicName>
(9m, 2f)
</th>
</tr>
<tr id="356C34BBFFCE00195B879294FA26FE1B" box="[165,1412,447,500]" gridrow="1" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B879294FE2EFE1B" box="[165,396,447,500]" gridcol="0" gridrow="1" pageId="6" pageNumber="55">Range</th>
<td id="76BD5DC7FFCE00195A8B9294FD99FE1B" box="[425,571,447,500]" gridcol="1" gridrow="1" pageId="6" pageNumber="55">Central Luzon Island</td>
<td id="76BD5DC7FFCE001959AF9294FCBCFE1B" box="[653,798,447,500]" gridcol="2" gridrow="1" pageId="6" pageNumber="55">Samar Island</td>
<td id="76BD5DC7FFCE0019586F9294FC4FFE1B" box="[845,1005,447,500]" gridcol="3" gridrow="1" pageId="6" pageNumber="55">Gigantes Islands</td>
<td id="76BD5DC7FFCE00195F349294FB60FE1B" box="[1046,1218,447,500]" gridcol="4" gridrow="1" pageId="6" pageNumber="55">Negros and Panay Islands</td>
<td id="76BD5DC7FFCE00195FC19294FA26FE1B" box="[1251,1412,447,500]" gridcol="5" gridrow="1" pageId="6" pageNumber="55">Luzon PAIC</td>
</tr>
<tr id="356C34BBFFCE00195B879133FA26FD7A" box="[165,1412,536,661]" gridrow="2" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B879133FE2EFD7A" box="[165,396,536,661]" gridcol="0" gridrow="2" pageId="6" pageNumber="55">SVL (m) SVL (f)</th>
<td id="76BD5DC7FFCE00195A8B9133FD99FD7A" box="[425,571,536,661]" gridcol="1" gridrow="2" pageId="6" pageNumber="55">32.339.9 (35.9 ± 1.8) 37.442.4 (40.1 ± 1.5)</td>
<td id="76BD5DC7FFCE001959AF9133FCBCFD7A" box="[653,798,536,661]" gridcol="2" gridrow="2" pageId="6" pageNumber="55">34.239.1 (37.1 ± 1.8) 44.349.8 (46.4 ± 2.1)</td>
<td id="76BD5DC7FFCE0019586F9133FC4FFD7A" box="[845,1005,536,661]" gridcol="3" gridrow="2" pageId="6" pageNumber="55">34.338.0 (36.7 ± 1.3) 40.242.9 (41.8 ± 1.0)</td>
<td id="76BD5DC7FFCE00195F349133FB60FD7A" box="[1046,1218,536,661]" gridcol="4" gridrow="2" pageId="6" pageNumber="55">28.335.7 (31.1 ± 4.0) 41.742.2 (42.0 ± 0.3)</td>
<td id="76BD5DC7FFCE00195FC19133FA26FD7A" box="[1251,1412,536,661]" gridcol="5" gridrow="2" pageId="6" pageNumber="55">24.132.5 (29.7 ± 2.4) 32.633.8 (33.2 ± 0.8)</td>
</tr>
<tr id="356C34BBFFCE00195B879181FA26FCC8" box="[165,1412,682,807]" gridrow="3" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B879181FE2EFCC8" box="[165,396,682,807]" gridcol="0" gridrow="3" pageId="6" pageNumber="55">ED (m) ED (f)</th>
<td id="76BD5DC7FFCE00195A8B9181FD99FCC8" box="[425,571,682,807]" gridcol="1" gridrow="3" pageId="6" pageNumber="55">3.35.2 (4.3 ± 0.5) 4.35.2 (4.9 ± 0.3)</td>
<td id="76BD5DC7FFCE001959AF9181FCBCFCC8" box="[653,798,682,807]" gridcol="2" gridrow="3" pageId="6" pageNumber="55">4.86.1 (5.5 ± 0.5) 5.46.8 (6.1 ± 0.4)</td>
<td id="76BD5DC7FFCE0019586F9181FC4FFCC8" box="[845,1005,682,807]" gridcol="3" gridrow="3" pageId="6" pageNumber="55">3.74.9 (4.2 ± 0.4) 5.15.7 (5.3 ± 0.2)</td>
<td id="76BD5DC7FFCE00195F349181FB60FCC8" box="[1046,1218,682,807]" gridcol="4" gridrow="3" pageId="6" pageNumber="55">3.14.1 (3.7 ± 0.6) 4.44.7 (4.6 ± 0.2)</td>
<td id="76BD5DC7FFCE00195FC19181FA26FCC8" box="[1251,1412,682,807]" gridcol="5" gridrow="3" pageId="6" pageNumber="55">2.73.9 (3.4 ± 0.4) 3.63.9 (3.7 ± 0.2)</td>
</tr>
<tr id="356C34BBFFCE00195B879017FA26FC56" box="[165,1412,828,953]" gridrow="4" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B879017FE2EFC56" box="[165,396,828,953]" gridcol="0" gridrow="4" pageId="6" pageNumber="55">Fin3DW/PpFin3 (m) Fin3DW/PpFin3 (f)</th>
<td id="76BD5DC7FFCE00195A8B9017FD99FC56" box="[425,571,828,953]" gridcol="1" gridrow="4" pageId="6" pageNumber="55">1.43.4 (2.1 ± 0.5) 1.42.7 (2.2 ± 0.4)</td>
<td id="76BD5DC7FFCE001959AF9017FCBCFC56" box="[653,798,828,953]" gridcol="2" gridrow="4" pageId="6" pageNumber="55">2.74.0 (3.3 ± 0.4) 2.84.5 (3.9 ± 0.5)</td>
<td id="76BD5DC7FFCE0019586F9017FC4FFC56" box="[845,1005,828,953]" gridcol="3" gridrow="4" pageId="6" pageNumber="55">2.73.1 (2.9 ± 0.1) 2.32.8 (2.7 ± 0.2)</td>
<td id="76BD5DC7FFCE00195F349017FB60FC56" box="[1046,1218,828,953]" gridcol="4" gridrow="4" pageId="6" pageNumber="55">2.62.8 (2.7 ± 0.1) 3.54.7 (4.1 ± 0.8)</td>
<td id="76BD5DC7FFCE00195FC19017FA26FC56" box="[1251,1412,828,953]" gridcol="5" gridrow="4" pageId="6" pageNumber="55">2.23.3 (2.8 ± 0.4) 3.03.2 (3.1 ± 0.2)</td>
</tr>
<tr id="356C34BBFFCE00195B8790E4FA26FBA4" box="[165,1412,975,1099]" gridrow="5" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B8790E4FE2EFBA4" box="[165,396,975,1099]" gridcol="0" gridrow="5" pageId="6" pageNumber="55">Toe4DW/PpToe4 (m) Toe4DW/PpToe4 (f)</th>
<td id="76BD5DC7FFCE00195A8B90E4FD99FBA4" box="[425,571,975,1099]" gridcol="1" gridrow="5" pageId="6" pageNumber="55">1.32.4 (1.7 ± 0.3) 1.63.1 (2.1 ± 0.5)</td>
<td id="76BD5DC7FFCE001959AF90E4FCBCFBA4" box="[653,798,975,1099]" gridcol="2" gridrow="5" pageId="6" pageNumber="55">2.13.3 (2.8 ± 0.4) 2.93.7 (3.1 ± 0.3)</td>
<td id="76BD5DC7FFCE0019586F90E4FC4FFBA4" box="[845,1005,975,1099]" gridcol="3" gridrow="5" pageId="6" pageNumber="55">1.93.4 (2.6 ± 0.4) 2.02.6 (2.3 ± 0.3)</td>
<td id="76BD5DC7FFCE00195F3490E4FB60FBA4" box="[1046,1218,975,1099]" gridcol="4" gridrow="5" pageId="6" pageNumber="55">1.81.9 (1.8 ± 0.1) 1.81.8 (1.8 ± 0.0)</td>
<td id="76BD5DC7FFCE00195FC190E4FA26FBA4" box="[1251,1412,975,1099]" gridcol="5" gridrow="5" pageId="6" pageNumber="55">1.62.2 (1.9 ± 0.2) 1.81.9 (1.8 ± 0.1)</td>
</tr>
<tr id="356C34BBFFCE00195B87974AFA26FB32" box="[165,1412,1121,1245]" gridrow="6" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B87974AFE2EFB32" box="[165,396,1121,1245]" gridcol="0" gridrow="6" pageId="6" pageNumber="55">Fin3DW/ Toe4DW (m) Fin3DW/Toe4DW (f)</th>
<td id="76BD5DC7FFCE00195A8B974AFD99FB32" box="[425,571,1121,1245]" gridcol="1" gridrow="6" pageId="6" pageNumber="55">0.91.7 (1.2 ± 0.2) 0.91.4 (1.2 ± 0.2)</td>
<td id="76BD5DC7FFCE001959AF974AFCBCFB32" box="[653,798,1121,1245]" gridcol="2" gridrow="6" pageId="6" pageNumber="55">1.11.5 (1.3 ± 0.1) 1.21.5 (1.3 ± 0.1)</td>
<td id="76BD5DC7FFCE0019586F974AFC4FFB32" box="[845,1005,1121,1245]" gridcol="3" gridrow="6" pageId="6" pageNumber="55">1.11.5 (1.3 ± 0.1) 1.21.6 (1.4 ± 0.1)</td>
<td id="76BD5DC7FFCE00195F34974AFB60FB32" box="[1046,1218,1121,1245]" gridcol="4" gridrow="6" pageId="6" pageNumber="55">1.82.0 (1.9 ± 0.1) 1.72.0 (1.9 ± 0.3)</td>
<td id="76BD5DC7FFCE00195FC1974AFA26FB32" box="[1251,1412,1121,1245]" gridcol="5" gridrow="6" pageId="6" pageNumber="55">1.62.0 (1.7 ± 0.1) 2.02.1 (2.1 ± 0.1)</td>
</tr>
<tr id="356C34BBFFCE00195B8797D8FA26FAC7" box="[165,1412,1267,1320]" gridrow="7" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B8797D8FE2EFAC7" box="[165,396,1267,1320]" gridcol="0" gridrow="7" pageId="6" pageNumber="55">Greatly expanded digital disks</th>
<td id="76BD5DC7FFCE00195A8B97D8FD99FAC7" box="[425,571,1267,1320]" gridcol="1" gridrow="7" pageId="6" pageNumber="55">+, moderate</td>
<td id="76BD5DC7FFCE001959AF97D8FCBCFAC7" box="[653,798,1267,1320]" gridcol="2" gridrow="7" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE0019586F97D8FC4FFAC7" box="[845,1005,1267,1320]" gridcol="3" gridrow="7" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE00195F3497D8FB60FAC7" box="[1046,1218,1267,1320]" gridcol="4" gridrow="7" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE00195FC197D8FA26FAC7" box="[1251,1412,1267,1320]" gridcol="5" gridrow="7" pageId="6" pageNumber="55">+</td>
</tr>
<tr id="356C34BBFFCE00195B879616FA26FABD" box="[165,1412,1341,1362]" gridrow="8" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B879616FE2EFABD" box="[165,396,1341,1362]" gridcol="0" gridrow="8" pageId="6" pageNumber="55">Dorsal skin rugosity</th>
<td id="76BD5DC7FFCE00195A8B9616FD99FABD" box="[425,571,1341,1362]" gridcol="1" gridrow="8" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE001959AF9616FCBCFABD" box="[653,798,1341,1362]" gridcol="2" gridrow="8" pageId="6" pageNumber="55"> to granular</td>
<td id="76BD5DC7FFCE0019586F9616FC4FFABD" box="[845,1005,1341,1362]" gridcol="3" gridrow="8" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE00195F349616FB60FABD" box="[1046,1218,1341,1362]" gridcol="4" gridrow="8" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE00195FC19616FA26FABD" box="[1251,1412,1341,1362]" gridcol="5" gridrow="8" pageId="6" pageNumber="55">+</td>
</tr>
<tr id="356C34BBFFCE00195B879643FA26FA92" box="[165,1412,1384,1405]" gridrow="9" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B879643FE2EFA92" box="[165,396,1384,1405]" gridcol="0" gridrow="9" pageId="6" pageNumber="55">Dorsal dermal tubercles</th>
<td id="76BD5DC7FFCE00195A8B9643FD99FA92" box="[425,571,1384,1405]" gridcol="1" gridrow="9" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE001959AF9643FCBCFA92" box="[653,798,1384,1405]" gridcol="2" gridrow="9" pageId="6" pageNumber="55">+, sparse</td>
<td id="76BD5DC7FFCE0019586F9643FC4FFA92" box="[845,1005,1384,1405]" gridcol="3" gridrow="9" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE00195F349643FB60FA92" box="[1046,1218,1384,1405]" gridcol="4" gridrow="9" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE00195FC19643FA26FA92" box="[1251,1412,1384,1405]" gridcol="5" gridrow="9" pageId="6" pageNumber="55">+</td>
</tr>
<tr id="356C34BBFFCE00195B8796B9FA26FA28" box="[165,1412,1426,1479]" gridrow="10" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B8796B9FE2EFA28" box="[165,396,1426,1479]" gridcol="0" gridrow="10" pageId="6" pageNumber="55">Tubercle color</th>
<td id="76BD5DC7FFCE00195A8B96B9FD99FA28" box="[425,571,1426,1479]" gridcol="1" gridrow="10" pageId="6" pageNumber="55">N/A</td>
<td id="76BD5DC7FFCE001959AF96B9FCBCFA28" box="[653,798,1426,1479]" gridcol="2" gridrow="10" pageId="6" pageNumber="55">Salmon</td>
<td id="76BD5DC7FFCE0019586F96B9FC4FFA28" box="[845,1005,1426,1479]" gridcol="3" gridrow="10" pageId="6" pageNumber="55">N/A</td>
<td id="76BD5DC7FFCE00195F3496B9FB60FA28" box="[1046,1218,1426,1479]" gridcol="4" gridrow="10" pageId="6" pageNumber="55">Cream</td>
<td id="76BD5DC7FFCE00195FC196B9FA26FA28" box="[1251,1412,1426,1479]" gridcol="5" gridrow="10" pageId="6" pageNumber="55">Dominant dorsal color</td>
</tr>
<tr id="356C34BBFFCE00195B8796F6FA26FA1D" box="[165,1412,1501,1522]" gridrow="11" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B8796F6FE2EFA1D" box="[165,396,1501,1522]" gridcol="0" gridrow="11" pageId="6" pageNumber="55">Dorsal folds/ridges</th>
<td id="76BD5DC7FFCE00195A8B96F6FD99FA1D" box="[425,571,1501,1522]" gridcol="1" gridrow="11" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE001959AF96F6FCBCFA1D" box="[653,798,1501,1522]" gridcol="2" gridrow="11" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE0019586F96F6FC4FFA1D" box="[845,1005,1501,1522]" gridcol="3" gridrow="11" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE00195F3496F6FB60FA1D" box="[1046,1218,1501,1522]" gridcol="4" gridrow="11" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE00195FC196F6FA26FA1D" box="[1251,1412,1501,1522]" gridcol="5" gridrow="11" pageId="6" pageNumber="55">+</td>
</tr>
<tr id="356C34BBFFCE00195B87952CFA26F9D3" box="[165,1412,1543,1596]" gridrow="12" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B87952CFE2EF9D3" box="[165,396,1543,1596]" gridcol="0" gridrow="12" pageId="6" pageNumber="55">Tympanum hidden</th>
<td id="76BD5DC7FFCE00195A8B952CFD99F9D3" box="[425,571,1543,1596]" gridcol="1" gridrow="12" pageId="6" pageNumber="55">No</td>
<td id="76BD5DC7FFCE001959AF952CFCBCF9D3" box="[653,798,1543,1596]" gridcol="2" gridrow="12" pageId="6" pageNumber="55">No</td>
<td id="76BD5DC7FFCE0019586F952CFC4FF9D3" box="[845,1005,1543,1596]" gridcol="3" gridrow="12" pageId="6" pageNumber="55">No</td>
<td id="76BD5DC7FFCE00195F34952CFB60F9D3" box="[1046,1218,1543,1596]" gridcol="4" gridrow="12" pageId="6" pageNumber="55">Posterodorsal edge</td>
<td id="76BD5DC7FFCE00195FC1952CFA26F9D3" box="[1251,1412,1543,1596]" gridcol="5" gridrow="12" pageId="6" pageNumber="55">Posteroventral edge</td>
</tr>
<tr id="356C34BBFFCE00195B879579FA26F969" box="[165,1412,1618,1670]" gridrow="13" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B879579FE2EF969" box="[165,396,1618,1670]" gridcol="0" gridrow="13" pageId="6" pageNumber="55">Supratympanic fold</th>
<td id="76BD5DC7FFCE00195A8B9579FD99F969" box="[425,571,1618,1670]" gridcol="1" gridrow="13" pageId="6" pageNumber="55">Smooth, less prominent</td>
<td id="76BD5DC7FFCE001959AF9579FCBCF969" box="[653,798,1618,1670]" gridcol="2" gridrow="13" pageId="6" pageNumber="55">Smooth, less prominent</td>
<td id="76BD5DC7FFCE0019586F9579FC4FF969" box="[845,1005,1618,1670]" gridcol="3" gridrow="13" pageId="6" pageNumber="55">Smooth, not prominent</td>
<td id="76BD5DC7FFCE00195F349579FB60F969" box="[1046,1218,1618,1670]" gridcol="4" gridrow="13" pageId="6" pageNumber="55">Rugose, prominent</td>
<td id="76BD5DC7FFCE00195FC19579FA26F969" box="[1251,1412,1618,1670]" gridcol="5" gridrow="13" pageId="6" pageNumber="55">Rugose, prominent</td>
</tr>
<tr id="356C34BBFFCE00195B8795B7FA26F95E" box="[165,1412,1692,1713]" gridrow="14" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B8795B7FE2EF95E" box="[165,396,1692,1713]" gridcol="0" gridrow="14" pageId="6" pageNumber="55">Dorsolateral stripes</th>
<td id="76BD5DC7FFCE00195A8B95B7FD99F95E" box="[425,571,1692,1713]" gridcol="1" gridrow="14" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE001959AF95B7FCBCF95E" box="[653,798,1692,1713]" gridcol="2" gridrow="14" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE0019586F95B7FC4FF95E" box="[845,1005,1692,1713]" gridcol="3" gridrow="14" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE00195F3495B7FB60F95E" box="[1046,1218,1692,1713]" gridcol="4" gridrow="14" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE00195FC195B7FA26F95E" box="[1251,1412,1692,1713]" gridcol="5" gridrow="14" pageId="6" pageNumber="55">+ or </td>
</tr>
<tr id="356C34BBFFCE00195B8795ECFA26F913" box="[165,1412,1735,1788]" gridrow="15" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B8795ECFE2EF913" box="[165,396,1735,1788]" gridcol="0" gridrow="15" pageId="6" pageNumber="55">Spotted flanks</th>
<td id="76BD5DC7FFCE00195A8B95ECFD99F913" box="[425,571,1735,1788]" gridcol="1" gridrow="15" pageId="6" pageNumber="55">, blotches anteriorly</td>
<td id="76BD5DC7FFCE001959AF95ECFCBCF913" box="[653,798,1735,1788]" gridcol="2" gridrow="15" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE0019586F95ECFC4FF913" box="[845,1005,1735,1788]" gridcol="3" gridrow="15" pageId="6" pageNumber="55">+</td>
<td id="76BD5DC7FFCE00195F3495ECFB60F913" box="[1046,1218,1735,1788]" gridcol="4" gridrow="15" pageId="6" pageNumber="55"></td>
<td id="76BD5DC7FFCE00195FC195ECFA26F913" box="[1251,1412,1735,1788]" gridcol="5" gridrow="15" pageId="6" pageNumber="55"></td>
</tr>
<tr id="356C34BBFFCE00195B87943AFA26F8A9" box="[165,1412,1809,1862]" gridrow="16" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B87943AFE2EF8A9" box="[165,396,1809,1862]" gridcol="0" gridrow="16" pageId="6" pageNumber="55">Limb bands</th>
<td id="76BD5DC7FFCE00195A8B943AFD99F8A9" box="[425,571,1809,1862]" gridcol="1" gridrow="16" pageId="6" pageNumber="55">+, well defined</td>
<td id="76BD5DC7FFCE001959AF943AFCBCF8A9" box="[653,798,1809,1862]" gridcol="2" gridrow="16" pageId="6" pageNumber="55">+, well defined</td>
<td id="76BD5DC7FFCE0019586F943AFC4FF8A9" box="[845,1005,1809,1862]" gridcol="3" gridrow="16" pageId="6" pageNumber="55">+ or , poorly defined</td>
<td id="76BD5DC7FFCE00195F34943AFB60F8A9" box="[1046,1218,1809,1862]" gridcol="4" gridrow="16" pageId="6" pageNumber="55">+, well defined</td>
<td id="76BD5DC7FFCE00195FC1943AFA26F8A9" box="[1251,1412,1809,1862]" gridcol="5" gridrow="16" pageId="6" pageNumber="55">+, well defined</td>
</tr>
<tr id="356C34BBFFCE00195B879470FA26F87F" box="[165,1412,1883,1936]" gridrow="17" pageId="6" pageNumber="55">
<th id="76BD5DC7FFCE00195B879470FE2EF87F" box="[165,396,1883,1936]" gridcol="0" gridrow="17" pageId="6" pageNumber="55">Microhabitat preference</th>
<td id="76BD5DC7FFCE00195A8B9470FD99F87F" box="[425,571,1883,1936]" gridcol="1" gridrow="17" pageId="6" pageNumber="55">Ground, karst habitat</td>
<td id="76BD5DC7FFCE001959AF9470FCBCF87F" box="[653,798,1883,1936]" gridcol="2" gridrow="17" pageId="6" pageNumber="55">Ground; karst habitat</td>
<td id="76BD5DC7FFCE0019586F9470FC4FF87F" box="[845,1005,1883,1936]" gridcol="3" gridrow="17" pageId="6" pageNumber="55">Ground; karst habitat</td>
<td id="76BD5DC7FFCE00195F349470FB60F87F" box="[1046,1218,1883,1936]" gridcol="4" gridrow="17" pageId="6" pageNumber="55">Arboreal</td>
<td id="76BD5DC7FFCE00195FC19470FA26F87F" box="[1251,1412,1883,1936]" gridcol="5" gridrow="17" pageId="6" pageNumber="55">Arboreal</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE336F9FFCFFFE15BB5938AFEF1FF59" blockId="7.[151,339,161,182]" box="[151,339,161,182]" pageId="7" pageNumber="56">
<emphasis id="B928EAEBFFCFFFE15BB5938AFF47FF59" bold="true" box="[151,229,161,182]" pageId="7" pageNumber="56">
<tableCitation id="C6DE0342FFCFFFE15BB5938AFF7DFF59" box="[151,223,161,182]" captionStart="TABLE 1" captionStartId="6.[151,240,151,175]" captionTargetBox="[165,1412,367,1936]" captionTargetPageId="6" captionText="TABLE 1. Summary of qualitative diagnostic characters (present, absent) in Platymantis biak and specimens of other members of the P. guentheri Species Group. Sample size for each sex, body size, eye diameter, digit disk proportions, and general geographical distribution (PAIC = Pleistocene Aggregate Island Complexes, sensu Brown and Diesmos, 2002) are included for reference (SVL and ED given as range over mean ± standard deviation; digit disk proportions given as percentage over mean ± standard deviation). Only adult specimens in good preservation condition were used for morphometric analyses." httpUri="http://table.plazi.org/id/DF236671FFCEFFE05BB593BCFE34FEA0" pageId="7" pageNumber="56" tableUuid="DF236671FFCEFFE05BB593BCFE34FEA0">Table 1</tableCitation>
.
</emphasis>
continued.
</paragraph>
<paragraph id="8BE336F9FFCFFFE15AB993FCFABBF946" pageId="7" pageNumber="56">
<table id="F95CC459FFCF00195B8793FCFA23F946" box="[165,1409,215,1705]" gridcols="6" gridrows="18" pageId="7" pageNumber="56" rowsContinueFrom="6.[165,1412,367,1936]">
<tr id="356C34BBFFCF00195B8793FCFA23FEFE" box="[165,1409,215,273]" gridrow="0" pageId="7" pageNumber="56" rowspan-0="1">
<th id="76BD5DC7FFCF00195AB893FCFDF3FEFE" box="[410,593,215,273]" gridcol="1" gridrow="0" pageId="7" pageNumber="56">
<taxonomicName id="4C5C4D7AFFCFFFE15AB993FCFE7AFF03" box="[411,472,215,236]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="56" phylum="Chordata" rank="species" species="rabori">
<emphasis id="B928EAEBFFCFFFE15AB993FCFE7AFF03" box="[411,472,215,236]" italics="true" pageId="7" pageNumber="56">rabori</emphasis>
</taxonomicName>
(2m, 6f)
</th>
<th id="76BD5DC7FFCF001959A093FCFC86FEFE" box="[642,804,215,273]" gridcol="2" gridrow="0" pageId="7" pageNumber="56">
<taxonomicName id="4C5C4D7AFFCFFFE159A193FCFD7DFF03" box="[643,735,215,236]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="56" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B928EAEBFFCFFFE159A193FCFD7DFF03" box="[643,735,215,236]" italics="true" pageId="7" pageNumber="56">guentheri</emphasis>
</taxonomicName>
(11m, 8f)
</th>
<th id="76BD5DC7FFCF0019587793FCFC56FEFE" box="[853,1012,215,273]" gridcol="3" gridrow="0" pageId="7" pageNumber="56">
<taxonomicName id="4C5C4D7AFFCFFFE1587793FCFC0BFF03" box="[853,937,215,236]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="56" phylum="Chordata" rank="species" species="banahao">
<emphasis id="B928EAEBFFCFFFE1587793FCFC0BFF03" box="[853,937,215,236]" italics="true" pageId="7" pageNumber="56">banahao</emphasis>
</taxonomicName>
(4m)
</th>
<th id="76BD5DC7FFCF00195F3293FCFB15FEFE" box="[1040,1207,215,273]" gridcol="4" gridrow="0" pageId="7" pageNumber="56">
<taxonomicName id="4C5C4D7AFFCFFFE15F3393FCFBC7FF03" box="[1041,1125,215,236]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="56" phylum="Chordata" rank="species" species="cornutus">
<emphasis id="B928EAEBFFCFFFE15F3393FCFBC7FF03" box="[1041,1125,215,236]" italics="true" pageId="7" pageNumber="56">cornutus</emphasis>
</taxonomicName>
(3m)
</th>
<th id="76BD5DC7FFCF00195FC093FCFA23FEFE" box="[1250,1409,215,273]" gridcol="5" gridrow="0" pageId="7" pageNumber="56">
<taxonomicName id="4C5C4D7AFFCFFFE15FC093FCFA94FF03" box="[1250,1334,215,236]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="56" phylum="Chordata" rank="species" species="diesmosi">
<emphasis id="B928EAEBFFCFFFE15FC093FCFA94FF03" box="[1250,1334,215,236]" italics="true" pageId="7" pageNumber="56">diesmosi</emphasis>
</taxonomicName>
(8m, 2f)
</th>
</tr>
<tr id="356C34BBFFCF00195B87920DFA23FED4" box="[165,1409,294,315]" gridrow="1" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B87920DFED9FED4" box="[165,379,294,315]" gridcol="0" gridrow="1" pageId="7" pageNumber="56">Range</th>
<td id="76BD5DC7FFCF00195AB8920DFDF3FED4" box="[410,593,294,315]" gridcol="1" gridrow="1" pageId="7" pageNumber="56">Mindanao PAIC</td>
<td id="76BD5DC7FFCF001959A0920DFC86FED4" box="[642,804,294,315]" gridcol="2" gridrow="1" pageId="7" pageNumber="56">Mindanao PAIC</td>
<td id="76BD5DC7FFCF00195877920DFC56FED4" box="[853,1012,294,315]" gridcol="3" gridrow="1" pageId="7" pageNumber="56">SE Luzon Island</td>
<td id="76BD5DC7FFCF00195F32920DFB15FED4" box="[1040,1207,294,315]" gridcol="4" gridrow="1" pageId="7" pageNumber="56">N. Luzon Island</td>
<td id="76BD5DC7FFCF00195FC0920DFA23FED4" box="[1250,1409,294,315]" gridcol="5" gridrow="1" pageId="7" pageNumber="56">SE Luzon Island</td>
</tr>
<tr id="356C34BBFFCF00195B87927AFA23FE22" box="[165,1409,337,461]" gridrow="2" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B87927AFED9FE22" box="[165,379,337,461]" gridcol="0" gridrow="2" pageId="7" pageNumber="56">SVL (m) SVL (f)</th>
<td id="76BD5DC7FFCF00195AB8927AFDF3FE22" box="[410,593,337,461]" gridcol="1" gridrow="2" pageId="7" pageNumber="56">30.433.5 (32.0 ± 2.2) 36.444.4 (41.5 ± 3.6)</td>
<td id="76BD5DC7FFCF001959A0927AFC86FE22" box="[642,804,337,461]" gridcol="2" gridrow="2" pageId="7" pageNumber="56">26.529.8 (28.6 ± 1.1) 32.337.3 (34.6 ± 2.0)</td>
<td id="76BD5DC7FFCF00195877927AFC56FE22" box="[853,1012,337,461]" gridcol="3" gridrow="2" pageId="7" pageNumber="56">32.436.3 (33.9 ± 1.7) N/A</td>
<td id="76BD5DC7FFCF00195F32927AFB15FE22" box="[1040,1207,337,461]" gridcol="4" gridrow="2" pageId="7" pageNumber="56">26.330.4 (27.8 ± 2.2) N/A</td>
<td id="76BD5DC7FFCF00195FC0927AFA23FE22" box="[1250,1409,337,461]" gridcol="5" gridrow="2" pageId="7" pageNumber="56">28.539.2 (35.3 ± 3.6) 49.052.6 (50.8 ± 2.6)</td>
</tr>
<tr id="356C34BBFFCF00195B8792C8FA23FDB0" box="[165,1409,483,607]" gridrow="3" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B8792C8FED9FDB0" box="[165,379,483,607]" gridcol="0" gridrow="3" pageId="7" pageNumber="56">ED (m) ED (f)</th>
<td id="76BD5DC7FFCF00195AB892C8FDF3FDB0" box="[410,593,483,607]" gridcol="1" gridrow="3" pageId="7" pageNumber="56">3.83.9 (3.9 ± 0.1) 4.05.7 (4.9 ± 0.7)</td>
<td id="76BD5DC7FFCF001959A092C8FC86FDB0" box="[642,804,483,607]" gridcol="2" gridrow="3" pageId="7" pageNumber="56">2.83.6 (3.3 ± 0.3) 3.74.5 (3.9 ± 0.3)</td>
<td id="76BD5DC7FFCF0019587792C8FC56FDB0" box="[853,1012,483,607]" gridcol="3" gridrow="3" pageId="7" pageNumber="56">3.64.1 (3.8 ± 0.2) N/A</td>
<td id="76BD5DC7FFCF00195F3292C8FB15FDB0" box="[1040,1207,483,607]" gridcol="4" gridrow="3" pageId="7" pageNumber="56">3.04.2 (3.5 ± 0.6) N/A</td>
<td id="76BD5DC7FFCF00195FC092C8FA23FDB0" box="[1250,1409,483,607]" gridcol="5" gridrow="3" pageId="7" pageNumber="56">3.75.4 (4.7 ± 0.5) 5.96.1 (6.0 ± 0.1)</td>
</tr>
<tr id="356C34BBFFCF00195B87915EFA23FD1D" box="[165,1409,629,754]" gridrow="4" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B87915EFED9FD1D" box="[165,379,629,754]" gridcol="0" gridrow="4" pageId="7" pageNumber="56">Fin3DW/ PpFin3 (m) Fin3DW/PpFin3 (f)</th>
<td id="76BD5DC7FFCF00195AB8915EFDF3FD1D" box="[410,593,629,754]" gridcol="1" gridrow="4" pageId="7" pageNumber="56">2.83.5 (3.2 ± 0.5) 3.34.2 (3.6 ± 0.3)</td>
<td id="76BD5DC7FFCF001959A0915EFC86FD1D" box="[642,804,629,754]" gridcol="2" gridrow="4" pageId="7" pageNumber="56">2.45.2 (3.2 ± 0.8) 2.65.7 (4.0 ± 1.2)</td>
<td id="76BD5DC7FFCF00195877915EFC56FD1D" box="[853,1012,629,754]" gridcol="3" gridrow="4" pageId="7" pageNumber="56">2.83.2 (3.0 ± 0.2) N/A</td>
<td id="76BD5DC7FFCF00195F32915EFB15FD1D" box="[1040,1207,629,754]" gridcol="4" gridrow="4" pageId="7" pageNumber="56">2.52.9 (2.7 ± 0.2) N/A</td>
<td id="76BD5DC7FFCF00195FC0915EFA23FD1D" box="[1250,1409,629,754]" gridcol="5" gridrow="4" pageId="7" pageNumber="56">1.52.1 (1.7 ± 0.2) 1.61.7 (1.6 ± 0.1)</td>
</tr>
<tr id="356C34BBFFCF00195B87902CFA23FC6B" box="[165,1409,775,900]" gridrow="5" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B87902CFED9FC6B" box="[165,379,775,900]" gridcol="0" gridrow="5" pageId="7" pageNumber="56">Toe4DW/PpToe4 (m) Toe4DW/PpToe4 (f)</th>
<td id="76BD5DC7FFCF00195AB8902CFDF3FC6B" box="[410,593,775,900]" gridcol="1" gridrow="5" pageId="7" pageNumber="56">2.02.7 (2.3 ± 0.5) 1.62.5 (2.3 ± 0.3)</td>
<td id="76BD5DC7FFCF001959A0902CFC86FC6B" box="[642,804,775,900]" gridcol="2" gridrow="5" pageId="7" pageNumber="56">1.53.5 (2.4 ± 0.6) 2.34.3 (3.2 ± 0.7)</td>
<td id="76BD5DC7FFCF00195877902CFC56FC6B" box="[853,1012,775,900]" gridcol="3" gridrow="5" pageId="7" pageNumber="56">1.92.3 (2.0 ± 0.2) N/A</td>
<td id="76BD5DC7FFCF00195F32902CFB15FC6B" box="[1040,1207,775,900]" gridcol="4" gridrow="5" pageId="7" pageNumber="56">1.62.0 (1.7 ± 0.2) N/A</td>
<td id="76BD5DC7FFCF00195FC0902CFA23FC6B" box="[1250,1409,775,900]" gridcol="5" gridrow="5" pageId="7" pageNumber="56">1.22.2 (1.7 ± 0.4) 1.72.1 (1.9 ± 0.3)</td>
</tr>
<tr id="356C34BBFFCF00195B8790B2FA23FBF9" box="[165,1409,921,1046]" gridrow="6" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B8790B2FED9FBF9" box="[165,379,921,1046]" gridcol="0" gridrow="6" pageId="7" pageNumber="56">Fin3DW/Toe4DW(m) Fin3DW/Toe4DW (f)</th>
<td id="76BD5DC7FFCF00195AB890B2FDF3FBF9" box="[410,593,921,1046]" gridcol="1" gridrow="6" pageId="7" pageNumber="56">1.31.6 (1.4 ± 0.2) 1.41.9 (1.6 ± 0.2)</td>
<td id="76BD5DC7FFCF001959A090B2FC86FBF9" box="[642,804,921,1046]" gridcol="2" gridrow="6" pageId="7" pageNumber="56">1.21.7 (1.4 ± 0.1) 1.11.7 (1.4 ± 0.2)</td>
<td id="76BD5DC7FFCF0019587790B2FC56FBF9" box="[853,1012,921,1046]" gridcol="3" gridrow="6" pageId="7" pageNumber="56">1.81.9 (1.8 ± 0.0) N/A</td>
<td id="76BD5DC7FFCF00195F3290B2FB15FBF9" box="[1040,1207,921,1046]" gridcol="4" gridrow="6" pageId="7" pageNumber="56">1.92.1 (2.0 ± 0.1) N/A</td>
<td id="76BD5DC7FFCF00195FC090B2FA23FBF9" box="[1250,1409,921,1046]" gridcol="5" gridrow="6" pageId="7" pageNumber="56">0.91.4 (1.1 ± 0.2) 0.80.8 (0.8 ± 0.0)</td>
</tr>
<tr id="356C34BBFFCF00195B879700FA23FB8F" box="[165,1409,1067,1120]" gridrow="7" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B879700FED9FB8F" box="[165,379,1067,1120]" gridcol="0" gridrow="7" pageId="7" pageNumber="56">Greatly expanded digital disks</th>
<td id="76BD5DC7FFCF00195AB89700FDF3FB8F" box="[410,593,1067,1120]" gridcol="1" gridrow="7" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF001959A09700FC86FB8F" box="[642,804,1067,1120]" gridcol="2" gridrow="7" pageId="7" pageNumber="56">+, moderate</td>
<td id="76BD5DC7FFCF001958779700FC56FB8F" box="[853,1012,1067,1120]" gridcol="3" gridrow="7" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF00195F329700FB15FB8F" box="[1040,1207,1067,1120]" gridcol="4" gridrow="7" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF00195FC09700FA23FB8F" box="[1250,1409,1067,1120]" gridcol="5" gridrow="7" pageId="7" pageNumber="56"></td>
</tr>
<tr id="356C34BBFFCF00195B87975DFA23FB64" box="[165,1409,1142,1163]" gridrow="8" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B87975DFED9FB64" box="[165,379,1142,1163]" gridcol="0" gridrow="8" pageId="7" pageNumber="56">Dorsal skin rugosity</th>
<td id="76BD5DC7FFCF00195AB8975DFDF3FB64" box="[410,593,1142,1163]" gridcol="1" gridrow="8" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF001959A0975DFC86FB64" box="[642,804,1142,1163]" gridcol="2" gridrow="8" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF00195877975DFC56FB64" box="[853,1012,1142,1163]" gridcol="3" gridrow="8" pageId="7" pageNumber="56"> to granular</td>
<td id="76BD5DC7FFCF00195F32975DFB15FB64" box="[1040,1207,1142,1163]" gridcol="4" gridrow="8" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF00195FC0975DFA23FB64" box="[1250,1409,1142,1163]" gridcol="5" gridrow="8" pageId="7" pageNumber="56"> to granular</td>
</tr>
<tr id="356C34BBFFCF00195B87978BFA23FB3A" box="[165,1409,1184,1237]" gridrow="9" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B87978BFED9FB3A" box="[165,379,1184,1237]" gridcol="0" gridrow="9" pageId="7" pageNumber="56">Dorsal dermal tubercles</th>
<td id="76BD5DC7FFCF00195AB8978BFDF3FB3A" box="[410,593,1184,1237]" gridcol="1" gridrow="9" pageId="7" pageNumber="56">+, dense</td>
<td id="76BD5DC7FFCF001959A0978BFC86FB3A" box="[642,804,1184,1237]" gridcol="2" gridrow="9" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF00195877978BFC56FB3A" box="[853,1012,1184,1237]" gridcol="3" gridrow="9" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF00195F32978BFB15FB3A" box="[1040,1207,1184,1237]" gridcol="4" gridrow="9" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF00195FC0978BFA23FB3A" box="[1250,1409,1184,1237]" gridcol="5" gridrow="9" pageId="7" pageNumber="56"></td>
</tr>
<tr id="356C34BBFFCF00195B8797C0FA23FAF0" box="[165,1409,1259,1311]" gridrow="10" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B8797C0FED9FAF0" box="[165,379,1259,1311]" gridcol="0" gridrow="10" pageId="7" pageNumber="56">Tubercle color</th>
<td id="76BD5DC7FFCF00195AB897C0FDF3FAF0" box="[410,593,1259,1311]" gridcol="1" gridrow="10" pageId="7" pageNumber="56">Dominant dorsal color</td>
<td id="76BD5DC7FFCF001959A097C0FC86FAF0" box="[642,804,1259,1311]" gridcol="2" gridrow="10" pageId="7" pageNumber="56">Dominant dorsal color</td>
<td id="76BD5DC7FFCF0019587797C0FC56FAF0" box="[853,1012,1259,1311]" gridcol="3" gridrow="10" pageId="7" pageNumber="56">Dominant dorsal color</td>
<td id="76BD5DC7FFCF00195F3297C0FB15FAF0" box="[1040,1207,1259,1311]" gridcol="4" gridrow="10" pageId="7" pageNumber="56">N/A</td>
<td id="76BD5DC7FFCF00195FC097C0FA23FAF0" box="[1250,1409,1259,1311]" gridcol="5" gridrow="10" pageId="7" pageNumber="56">N/A</td>
</tr>
<tr id="356C34BBFFCF00195B87961EFA23FAA5" box="[165,1409,1333,1354]" gridrow="11" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B87961EFED9FAA5" box="[165,379,1333,1354]" gridcol="0" gridrow="11" pageId="7" pageNumber="56">Dorsal folds/ridges</th>
<td id="76BD5DC7FFCF00195AB8961EFDF3FAA5" box="[410,593,1333,1354]" gridcol="1" gridrow="11" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF001959A0961EFC86FAA5" box="[642,804,1333,1354]" gridcol="2" gridrow="11" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF00195877961EFC56FAA5" box="[853,1012,1333,1354]" gridcol="3" gridrow="11" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF00195F32961EFB15FAA5" box="[1040,1207,1333,1354]" gridcol="4" gridrow="11" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF00195FC0961EFA23FAA5" box="[1250,1409,1333,1354]" gridcol="5" gridrow="11" pageId="7" pageNumber="56"></td>
</tr>
<tr id="356C34BBFFCF00195B879674FA23FA7B" box="[165,1409,1375,1428]" gridrow="12" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B879674FED9FA7B" box="[165,379,1375,1428]" gridcol="0" gridrow="12" pageId="7" pageNumber="56">Tympanum hidden</th>
<td id="76BD5DC7FFCF00195AB89674FDF3FA7B" box="[410,593,1375,1428]" gridcol="1" gridrow="12" pageId="7" pageNumber="56">Posterior edge</td>
<td id="76BD5DC7FFCF001959A09674FC86FA7B" box="[642,804,1375,1428]" gridcol="2" gridrow="12" pageId="7" pageNumber="56">Posteroventral edge</td>
<td id="76BD5DC7FFCF001958779674FC56FA7B" box="[853,1012,1375,1428]" gridcol="3" gridrow="12" pageId="7" pageNumber="56">Posterodorsal edge</td>
<td id="76BD5DC7FFCF00195F329674FB15FA7B" box="[1040,1207,1375,1428]" gridcol="4" gridrow="12" pageId="7" pageNumber="56">No</td>
<td id="76BD5DC7FFCF00195FC09674FA23FA7B" box="[1250,1409,1375,1428]" gridcol="5" gridrow="12" pageId="7" pageNumber="56">Posterodorsal edge</td>
</tr>
<tr id="356C34BBFFCF00195B879681FA23FA30" box="[165,1409,1450,1503]" gridrow="13" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B879681FED9FA30" box="[165,379,1450,1503]" gridcol="0" gridrow="13" pageId="7" pageNumber="56">Supratympanic fold</th>
<td id="76BD5DC7FFCF00195AB89681FDF3FA30" box="[410,593,1450,1503]" gridcol="1" gridrow="13" pageId="7" pageNumber="56">Rugose, prominent</td>
<td id="76BD5DC7FFCF001959A09681FC86FA30" box="[642,804,1450,1503]" gridcol="2" gridrow="13" pageId="7" pageNumber="56">Smooth, less prominent</td>
<td id="76BD5DC7FFCF001958779681FC56FA30" box="[853,1012,1450,1503]" gridcol="3" gridrow="13" pageId="7" pageNumber="56">Smooth, less prominent</td>
<td id="76BD5DC7FFCF00195F329681FB15FA30" box="[1040,1207,1450,1503]" gridcol="4" gridrow="13" pageId="7" pageNumber="56">Smooth, not prominent</td>
<td id="76BD5DC7FFCF00195FC09681FA23FA30" box="[1250,1409,1450,1503]" gridcol="5" gridrow="13" pageId="7" pageNumber="56">Smooth, less prominent</td>
</tr>
<tr id="356C34BBFFCF00195B8796DFFA23F9E6" box="[165,1409,1524,1545]" gridrow="14" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B8796DFFED9F9E6" box="[165,379,1524,1545]" gridcol="0" gridrow="14" pageId="7" pageNumber="56">Dorsolateral stripes</th>
<td id="76BD5DC7FFCF00195AB896DFFDF3F9E6" box="[410,593,1524,1545]" gridcol="1" gridrow="14" pageId="7" pageNumber="56">+ or </td>
<td id="76BD5DC7FFCF001959A096DFFC86F9E6" box="[642,804,1524,1545]" gridcol="2" gridrow="14" pageId="7" pageNumber="56">+ or </td>
<td id="76BD5DC7FFCF0019587796DFFC56F9E6" box="[853,1012,1524,1545]" gridcol="3" gridrow="14" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF00195F3296DFFB15F9E6" box="[1040,1207,1524,1545]" gridcol="4" gridrow="14" pageId="7" pageNumber="56">+</td>
<td id="76BD5DC7FFCF00195FC096DFFA23F9E6" box="[1250,1409,1524,1545]" gridcol="5" gridrow="14" pageId="7" pageNumber="56"></td>
</tr>
<tr id="356C34BBFFCF00195B879534FA23F9DB" box="[165,1409,1567,1588]" gridrow="15" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B879534FED9F9DB" box="[165,379,1567,1588]" gridcol="0" gridrow="15" pageId="7" pageNumber="56">Spotted flanks</th>
<td id="76BD5DC7FFCF00195AB89534FDF3F9DB" box="[410,593,1567,1588]" gridcol="1" gridrow="15" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF001959A09534FC86F9DB" box="[642,804,1567,1588]" gridcol="2" gridrow="15" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF001958779534FC56F9DB" box="[853,1012,1567,1588]" gridcol="3" gridrow="15" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF00195F329534FB15F9DB" box="[1040,1207,1567,1588]" gridcol="4" gridrow="15" pageId="7" pageNumber="56"></td>
<td id="76BD5DC7FFCF00195FC09534FA23F9DB" box="[1250,1409,1567,1588]" gridcol="5" gridrow="15" pageId="7" pageNumber="56"></td>
</tr>
<tr id="356C34BBFFCF00195B879562FA23F9B1" box="[165,1409,1609,1630]" gridrow="16" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B879562FED9F9B1" box="[165,379,1609,1630]" gridcol="0" gridrow="16" pageId="7" pageNumber="56">Limb bands</th>
<td id="76BD5DC7FFCF00195AB89562FDF3F9B1" box="[410,593,1609,1630]" gridcol="1" gridrow="16" pageId="7" pageNumber="56">+, well defined</td>
<td id="76BD5DC7FFCF001959A09562FC86F9B1" box="[642,804,1609,1630]" gridcol="2" gridrow="16" pageId="7" pageNumber="56">+, well defined</td>
<td id="76BD5DC7FFCF001958779562FC56F9B1" box="[853,1012,1609,1630]" gridcol="3" gridrow="16" pageId="7" pageNumber="56">+ or , faint</td>
<td id="76BD5DC7FFCF00195F329562FB15F9B1" box="[1040,1207,1609,1630]" gridcol="4" gridrow="16" pageId="7" pageNumber="56">+, poorly defined</td>
<td id="76BD5DC7FFCF00195FC09562FA23F9B1" box="[1250,1409,1609,1630]" gridcol="5" gridrow="16" pageId="7" pageNumber="56">+, well defined</td>
</tr>
<tr id="356C34BBFFCF00195B87955FFA23F946" box="[165,1409,1652,1705]" gridrow="17" pageId="7" pageNumber="56">
<th id="76BD5DC7FFCF00195B87955FFED9F946" box="[165,379,1652,1705]" gridcol="0" gridrow="17" pageId="7" pageNumber="56">Microhabitat preference</th>
<td id="76BD5DC7FFCF00195AB8955FFDF3F946" box="[410,593,1652,1705]" gridcol="1" gridrow="17" pageId="7" pageNumber="56">Arboreal</td>
<td id="76BD5DC7FFCF001959A0955FFC86F946" box="[642,804,1652,1705]" gridcol="2" gridrow="17" pageId="7" pageNumber="56">Shrub/Arboreal</td>
<td id="76BD5DC7FFCF00195877955FFC56F946" box="[853,1012,1652,1705]" gridcol="3" gridrow="17" pageId="7" pageNumber="56">Arboreal</td>
<td id="76BD5DC7FFCF00195F32955FFB15F946" box="[1040,1207,1652,1705]" gridcol="4" gridrow="17" pageId="7" pageNumber="56">Arboreal</td>
<td id="76BD5DC7FFCF00195FC0955FFA23F946" box="[1250,1409,1652,1705]" gridcol="5" gridrow="17" pageId="7" pageNumber="56">Ground; cliff edges</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE336F9FFCFFFE15BE495C0FABAF839" blockId="7.[151,1437,1771,2006]" pageId="7" pageNumber="56">
<emphasis id="B928EAEBFFCFFFE15BE495C0FE07F8EA" bold="true" box="[198,421,1771,1797]" pageId="7" pageNumber="56">Coloration in life.</emphasis>
Dorsum and head mottled light green, dark green, and black (Fig. 4); interorbital bar distinct, black, and bordered anteriorly by lighter, green-brown border; lower surfaces of supratympanic fold black; black canthal blotch located just anterior to eye; black labial blotch located below eye, anterior to tympanic annulus; dorsal surfaces of limbs with alternating dark brown and green transverse bars, each bordered by thin cream transverse bars; dorsal surface of body with several large black blotches, irregularly distributed; area of black blotches and cream speckles on anterior portion of flanks; pupil bordered by copper ciliary ring of iris, sharply blending into predominant mottled gold and brown color pattern of iris.
</paragraph>
<caption id="DF236671FFC0FFEE5BB593BCFC0CFF00" pageId="8" pageNumber="57">
<paragraph id="8BE336F9FFC0FFEE5BB593BCFC0CFF00" blockId="8.[151,1436,151,513]" pageId="8" pageNumber="57">
<emphasis id="B928EAEBFFC0FFEE5BB593BCFEA9FF40" bold="true" box="[151,267,151,175]" pageId="8" pageNumber="57">TABLE 2.</emphasis>
Summary of mensural characters (in mm) and selected qualitative diagnostic characters (present, absent) in
<taxonomicName id="4C5C4D7AFFC0FFEE5BB59393FEEEFF20" box="[151,332,184,207]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC0FFEE5BB59393FEEEFF20" box="[151,332,184,207]" italics="true" pageId="8" pageNumber="57">Platymantis biak</emphasis>
</taxonomicName>
and other Philippine limestone forest frog species. Sample size, body size, species group, and general geographical distribution are included for reference (SVL given as range).
</paragraph>
</caption>
<paragraph id="8BE336F9FFC0FFEE5A8C9221FA88FECE" blockId="8.[151,1436,151,513]" box="[430,1322,266,290]" pageId="8" pageNumber="57">
<taxonomicName id="4C5C4D7AFFC0FFEE5A8C9221FE7EFECE" box="[430,476,266,289]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="biak" status="sp. nov.">
<emphasis id="B928EAEBFFC0FFEE5A8C9221FE7EFECE" box="[430,476,266,289]" italics="true" pageId="8" pageNumber="57">biak</emphasis>
</taxonomicName>
<emphasis id="B928EAEBFFC0FFEE5AC19221FD9AFECD" bold="true" box="[483,568,266,290]" pageId="8" pageNumber="57">
<taxonomicNameLabel id="A21B5790FFC0FFEE5AC19221FD9AFECD" box="[483,568,266,290]" pageId="8" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B928EAEBFFC0FFEE59B09221FA88FECE" box="[658,1322,266,289]" italics="true" pageId="8" pageNumber="57">
<taxonomicName id="4C5C4D7AFFC0FFEE59B09221FD79FECE" box="[658,731,266,289]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="bayani">bayani</taxonomicName>
<taxonomicName id="4C5C4D7AFFC0FFEE587A9221FC1AFECE" box="[856,952,266,289]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="insulatus">insulatus</taxonomicName>
<taxonomicName id="4C5C4D7AFFC0FFEE5F3F9221FBDBFECE" box="[1053,1145,266,289]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="spelaeus">spelaeus</taxonomicName>
<taxonomicName id="4C5C4D7AFFC0FFEE5FC09221FA88FECE" box="[1250,1322,266,289]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="paengi">paengi</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BE336F9FFC0FFEE5A8C9201FAB2FE63" blockId="8.[151,1436,151,513]" pageId="8" pageNumber="57">
(
<quantity id="4CA49B1CFFC0FFEE5A949201FE4BFEAE" box="[438,489,298,321]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="57" unit="m" value="12.0">12m</quantity>
, 4f) (
<quantity id="4CA49B1CFFC0FFEE59B89201FD62FEAE" box="[666,704,298,321]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="57" unit="m" value="9.0">9m</quantity>
, 9f) (
<quantity id="4CA49B1CFFC0FFEE58429201FC26FEAE" box="[864,900,298,321]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="57" unit="m" value="8.0">8m</quantity>
, 6f) (
<quantity id="4CA49B1CFFC0FFEE5F079201FBE8FEAE" box="[1061,1098,298,321]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="57" unit="m" value="4.0">4m</quantity>
, 3f) (
<quantity id="4CA49B1CFFC0FFEE5FC89201FABEFEAE" box="[1258,1308,298,321]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="57" unit="m" value="15.0">15m</quantity>
) SVL (m) 32.339.9 34.239.1 34.338.0 37.144.3 27.734.3 SVL (f) 37.442.4 44.349.8 40.242.9 53.5
<geoCoordinate id="EE68503EFFC0FFEE5F7B925EFB25FE63" box="[1113,1159,373,396]" direction="north" orientation="latitude" pageId="8" pageNumber="57" precision="5555" value="57.5">57.5</geoCoordinate>
N/A
</paragraph>
<paragraph id="8BE336F9FFC0FFEE5BBD928BFAF4FE58" blockId="8.[151,1436,151,513]" box="[159,1366,416,439]" pageId="8" pageNumber="57">Range Central Luzon Samar Island Gigantes Islands SW Negros NW Panay</paragraph>
<paragraph id="8BE336F9FFC0FFEE5A8C92EBFA97FDEE" blockId="8.[151,1436,151,513]" pageId="8" pageNumber="57">
Island Island Island Species Group
<taxonomicName id="4C5C4D7AFFC0FFEE5A8C92C1FC1FFDEE" box="[430,957,490,513]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="guentheri">guentheri guentheri guentheri</taxonomicName>
<taxonomicName id="4C5C4D7AFFC0FFEE5F3F92C1FA97FDEE" box="[1053,1333,490,513]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="dorsalis">dorsalis dorsalis</taxonomicName>
</paragraph>
<paragraph id="8BE336F9FFC0FFEE5BBD910FFADDFDD4" blockId="8.[159,1407,548,645]" box="[159,1407,548,571]" pageId="8" pageNumber="57">Digital disks Moderately Widely Widely Non-expanded Non-expanded</paragraph>
<paragraph id="8BE336F9FFC0FFEE5A8C916FFC1CFDB4" blockId="8.[159,1407,548,645]" box="[430,958,580,603]" pageId="8" pageNumber="57">expanded expanded expanded</paragraph>
<paragraph id="8BE336F9FFC0FFEE5BBD9145FB53FD6A" blockId="8.[159,1407,548,645]" box="[159,1265,622,645]" pageId="8" pageNumber="57">Dorsal skin rugosity to granular +, +</paragraph>
<paragraph id="8BE336F9FFC0FFEE5BBD9183FB53FD50" blockId="8.[159,1265,680,703]" box="[159,1265,680,703]" pageId="8" pageNumber="57">Dorsal dermal tubercles +, sparse +, small +</paragraph>
<paragraph id="8BE336F9FFC0FFEE5BBD91C9FE29FB5C" blockId="8.[159,1331,738,761]" lastBlockId="8.[151,1437,840,2024]" pageId="8" pageNumber="57">
Flanks Blotches anteriorly Spotted Spotted Immaculate Spotted
<emphasis id="B928EAEBFFC0FFEE5BE69063FEE3FC8D" bold="true" box="[196,321,840,866]" pageId="8" pageNumber="57">Variation.</emphasis>
Dominant color pattern consistently mottled in all observed specimens. Variation exists in the w-shaped pattern of pigmentation on the dorsal surface of the body in line with the forearms. Some individuals possess a sharply defined w-shaped pattern (KU 319976, 319980, 31998283, 319987,
<collectingCountry id="F34B7669FFC0FFEE5E7990B8FA3EFC42" box="[1371,1436,915,941]" name="Panama" pageId="8" pageNumber="57">PNM</collectingCountry>
968486, 968889, 9691, 9693, 9695, 9697, 9700), while the pattern on many other specimens is either obscured by the background body coloration or is vaguely defined (KU 31997375, 31997779, 319981,
<collectingCountry id="F34B7669FFC0FFEE5BB59728FF7AFBF2" box="[151,216,1027,1053]" name="Panama" pageId="8" pageNumber="57">PNM</collectingCountry>
969899, 968083, 967476, 9687, 9690, 9692, 9694, 9696, 967778, 9701, 970414). The outer metacarpal tubercle was fully divided in most individuals with the exception of a few individuals showing a single, large, connected outer metacarpal tubercle (KU 319974, 319980, 319981, 319983, 319987). Females are considerably larger and more robust. Summaries of univariate morphological variation in the series are presented in
<tableCitation id="C6DE0342FFC0FFEE5A0E97B2FE27FB5C" box="[300,389,1177,1203]" captionStart="TABLE 3" captionStartId="9.[151,239,151,175]" captionTargetBox="[151,1435,234,1360]" captionTargetPageId="9" captionText="TABLE 3. Summary of univariate morphological variation among mensural characters in the type series of Platymantis biak." httpUri="http://table.plazi.org/id/DF236671FFC1FFEF5BB593BCFF69FF20" pageId="8" pageNumber="57" tableUuid="DF236671FFC1FFEF5BB593BCFF69FF20">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3466572FFC0FFEE5BE69795FB58F898" pageId="8" pageNumber="57" type="distribution">
<paragraph id="8BE336F9FFC0FFEE5BE69795FBEBFB37" blockId="8.[151,1437,840,2024]" box="[196,1097,1214,1240]" pageId="8" pageNumber="57">
<emphasis id="B928EAEBFFC0FFEE5BE69795FEC6FB37" bold="true" box="[196,356,1214,1240]" pageId="8" pageNumber="57">Distribution.</emphasis>
<taxonomicName id="4C5C4D7AFFC0FFEE5A4E9795FD90FB38" box="[364,562,1214,1239]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC0FFEE5A4E9795FD90FB38" box="[364,562,1214,1239]" italics="true" pageId="8" pageNumber="57">Platymantis biak</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54E7885BFFC0FFEE58739795FC21FB37" box="[849,899,1214,1240]" pageId="8" pageNumber="57">type</typeStatus>
locality (Fig. 1).
</paragraph>
<paragraph id="8BE336F9FFC0FFEE5BE697C8FCA0FA57" blockId="8.[151,1437,840,2024]" pageId="8" pageNumber="57">
<emphasis id="B928EAEBFFC0FFEE5BE697C8FD97FB12" bold="true" box="[196,565,1251,1277]" pageId="8" pageNumber="57">Ecology and natural history.</emphasis>
<taxonomicName id="4C5C4D7AFFC0FFEE591C97CFFCAEFB12" box="[574,780,1252,1277]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC0FFEE591C97CFFCAEFB12" box="[574,780,1252,1277]" italics="true" pageId="8" pageNumber="57">Platymantis biak</emphasis>
</taxonomicName>
occurs in secondary-growth forest on karst substrate. Both males and females were observed perched on top of limestone rocks within large rock formations in the forest and around the entrances to cave systems. Additionally, males were observed up to 5 meters inside caves, some perched above the ground within depressions and holes in the karst rock walls (Fig. 2B, C). No specimens were observed perching in trees or shrubs, in contrast to the common behavior of most species in the
<taxonomicName id="4C5C4D7AFFC0FFEE5BE096B5FEEDFA58" box="[194,335,1438,1463]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B928EAEBFFC0FFEE5BE096B5FEEDFA58" box="[194,335,1438,1463]" italics="true" pageId="8" pageNumber="57">P. guentheri</emphasis>
</taxonomicName>
Group (
<bibRefCitation id="EFCD4B08FFC0FFEE5A9396B5FD54FA57" author="Brown" box="[433,758,1438,1464]" pageId="8" pageNumber="61" refString="Brown, R. M &amp; Gonzales, J. C. (2007) A new forest frog of the genus Platymantis (Amphibia; Anura; Ranidae) from the Bicol Peninsula of Luzon Island, Philippines. Copeia, 2007 (2), 251 - 266." type="journal article" year="2007">Brown and Gonzales, 2007</bibRefCitation>
).
</paragraph>
<paragraph id="8BE336F9FFC0FFEE5BE696E8FBFEF8E7" blockId="8.[151,1437,840,2024]" pageId="8" pageNumber="57">
Some individuals of
<taxonomicName id="4C5C4D7AFFC0FFEE5A9A96EFFDDDFA32" box="[440,639,1476,1501]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC0FFEE5A9A96EFFDDDFA32" box="[440,639,1476,1501]" italics="true" pageId="8" pageNumber="57">Platymantis biak</emphasis>
</taxonomicName>
were well camouflaged against the limestone (much like
<emphasis id="B928EAEBFFC0FFEE5E0596EFFE1AF9ED" italics="true" pageId="8" pageNumber="57">
<taxonomicName id="4C5C4D7AFFC0FFEE5E0596EFFA35FA32" box="[1319,1431,1476,1501]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="bayani">P. bayani</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC0FFEE5BB596C2FE82F9ED" box="[151,288,1513,1538]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="insulatus">P. insulatus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC0FFEE5A0C96C2FE11F9ED" box="[302,435,1513,1538]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="speleaus">P. speleaus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C5C4D7AFFC0FFEE5AD096C2FDC2F9ED" box="[498,608,1513,1538]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="paengi">
<emphasis id="B928EAEBFFC0FFEE5AD096C2FDC2F9ED" box="[498,608,1513,1538]" italics="true" pageId="8" pageNumber="57">P. paengi</emphasis>
</taxonomicName>
; Fig. 2B, C), and field workers found it difficult to locate and capture individuals against the limestone substrate. This was especially true for many of the females. However, males were often more conspicuous while calling atop rocky outcrops. When disturbed, the new species immediately jumped back into the rocks and escaped into limestone crevices. No egg masses or juveniles were found; other species of
<taxonomicName id="4C5C4D7AFFC0FFEE5ACE9555FDDAF978" box="[492,632,1662,1687]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="genus">
<emphasis id="B928EAEBFFC0FFEE5ACE9555FDDAF978" box="[492,632,1662,1687]" italics="true" pageId="8" pageNumber="57">Platymantis</emphasis>
</taxonomicName>
have direct development (
<bibRefCitation id="EFCD4B08FFC0FFEE58939555FB40F978" author="Brown" box="[945,1250,1662,1688]" pageId="8" pageNumber="61" refString="Brown, W. C. &amp; Alcala, A. C. (1982 a) A new cave Platymantis (Amphibia: Ranidae) from the Philippine Islands. Proceedings of the Biological Society of Washington, 95, 386 - 391." type="journal article" year="1982" yearSuffix="a">
Brown and Alcala, 1982
<emphasis id="B928EAEBFFC0FFEE5FF09555FB40F978" box="[1234,1250,1662,1687]" italics="true" pageId="8" pageNumber="57">a</emphasis>
</bibRefCitation>
,
<bibRefCitation id="EFCD4B08FFC0FFEE5FC59555FB55F978" author="Brown" box="[1255,1271,1662,1687]" pageId="8" pageNumber="61" refString="Brown, W. C. &amp; Alcala, A. C. (1982 b) Modes of reproduction of Philippine anurans. Pp. 416 - 428. In A. G. J. Rodin and K. Miyata (Eds.), Advances in Herpetology and Evolutionary Biology. Museum of Comparative Biology, Cambridge, Massachusetts, U. S. A." type="book chapter" year="1982" yearSuffix="b">b</bibRefCitation>
).
<taxonomicName id="4C5C4D7AFFC0FFEE5E329555FF5FF952" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="spelaeus">
<emphasis id="B928EAEBFFC0FFEE5E329555FF5FF952" italics="true" pageId="8" pageNumber="57">Platymantis spelaeus</emphasis>
</taxonomicName>
has been observed by one of us (CDS) to deposit direct-developing eggs in limestone crevices and the same is suspected for
<taxonomicName id="4C5C4D7AFFC0FFEE5AE995E2FC35F90C" authority="Brown and Alcala, 2000" authorityName="Brown and Alcala" authorityYear="2000" box="[459,919,1737,1763]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="insulatus">
<emphasis id="B928EAEBFFC0FFEE5AE995E2FDF6F90D" box="[459,596,1737,1762]" italics="true" pageId="8" pageNumber="57">P. insulatus</emphasis>
(
<bibRefCitation id="EFCD4B08FFC0FFEE594695E2FC2DF90C" author="Brown" box="[612,911,1737,1763]" pageId="8" pageNumber="60" refString="Brown, R. M &amp; Alcala, A. C. (2000) Geckos, cave frogs, and small land-bridge islands in the Visayan sea. Haring Ibon, 2, 19 - 22." type="journal article" year="2000">Brown and Alcala, 2000</bibRefCitation>
)
</taxonomicName>
. We assume that this species also breeds in limestone crevices and small caves (
<bibRefCitation id="EFCD4B08FFC0FFEE596295C5FCC1F8E7" author="Brown" box="[576,867,1774,1800]" pageId="8" pageNumber="60" refString="Brown, R. M &amp; Alcala, A. C. (2000) Geckos, cave frogs, and small land-bridge islands in the Visayan sea. Haring Ibon, 2, 19 - 22." type="journal article" year="2000">Brown and Alcala, 2000</bibRefCitation>
;
<bibRefCitation id="EFCD4B08FFC0FFEE585295C5FBF2F8E7" author="Brown" box="[880,1104,1774,1800]" pageId="8" pageNumber="60" refString="Brown, R. M., Dolino, C. N., Alcala, E., Diesmos, A. C. &amp; Alcala, A. C. (2003) The advertisement calls of two endangered species of endemic Philippine frogs: Platymantis spelaeus and P. insulatus (Anura; Ranidae). Haring Ibon, 43, 91 - 109." type="journal article" year="2003">
Brown
<emphasis id="B928EAEBFFC0FFEE58E595C5FBA3F8E8" box="[967,1025,1774,1799]" italics="true" pageId="8" pageNumber="57">et al.</emphasis>
, 2003
</bibRefCitation>
).
</paragraph>
<paragraph id="8BE336F9FFC0FFEE5BE69438FB58F898" blockId="8.[151,1437,840,2024]" pageId="8" pageNumber="57">
Sympatric anuran species include
<taxonomicName id="4C5C4D7AFFC0FFEE597A943FFCBBF8C2" box="[600,793,1812,1837]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="marina">
<emphasis id="B928EAEBFFC0FFEE597A943FFCBBF8C2" box="[600,793,1812,1837]" italics="true" pageId="8" pageNumber="57">Rhinella marina</emphasis>
</taxonomicName>
(introduced),
<taxonomicName id="4C5C4D7AFFC0FFEE58E3943FFB5EF8C2" box="[961,1276,1812,1837]" class="Amphibia" family="Dicroglossidae" genus="Hoplobatrachus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="rugulosus">
<emphasis id="B928EAEBFFC0FFEE58E3943FFB5EF8C2" box="[961,1276,1812,1837]" italics="true" pageId="8" pageNumber="57">Hoplobatrachus rugulosus</emphasis>
</taxonomicName>
(introduced),
<emphasis id="B928EAEBFFC0FFEE5BB59412FE71F8BD" box="[151,467,1849,1874]" italics="true" pageId="8" pageNumber="57">
<taxonomicName id="4C5C4D7AFFC0FFEE5BB59412FEE0F8BD" box="[151,322,1849,1874]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="picta">Kaloula picta</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC0FFEE5A729412FE71F8BD" box="[336,467,1849,1874]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="pulchra">K. pulchra</taxonomicName>
</emphasis>
(introduced),
<emphasis id="B928EAEBFFC0FFEE59A69412FC0FF898" italics="true" pageId="8" pageNumber="57">
<taxonomicName id="4C5C4D7AFFC0FFEE59A69412FC41F8BD" box="[644,995,1849,1874]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="macrocephalus">Limnonectes macrocephalus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC0FFEE58D39412FAE6F8BD" authority="Occidozyga" authorityName="Occidozyga" box="[1009,1348,1849,1874]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="woodworthi">L. woodworthi, Occidozyga</taxonomicName>
laevis,
<taxonomicName id="4C5C4D7AFFC0FFEE5BB59475FDEDF898" authority="Polypedates" authorityName="Polypedates" box="[151,591,1886,1911]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="mimulus">Platymantis cf. mimulus, Polypedates</taxonomicName>
leucomystax, Rana erythraea
</emphasis>
(introduced), and
<emphasis id="B928EAEBFFC0FFEE5FA79475FB58F898" box="[1157,1274,1886,1911]" italics="true" pageId="8" pageNumber="57">
<taxonomicName id="4C5C4D7AFFC0FFEE5FA79475FB54F898" box="[1157,1270,1886,1911]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="similis">R. similis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3466572FFC0FFEC5BE694A8FDC2FD0D" lastPageId="10" lastPageNumber="59" pageId="8" pageNumber="57" type="etymology">
<paragraph id="8BE336F9FFC0FFEE5BE694A8FDC8F807" blockId="8.[151,1437,840,2024]" pageId="8" pageNumber="57">
<emphasis id="B928EAEBFFC0FFEE5BE694A8FEF0F872" bold="true" box="[196,338,1923,1949]" pageId="8" pageNumber="57">Etymology.</emphasis>
We derive the specific epithet from the Tagalog term
<taxonomicName id="4C5C4D7AFFC0FFEE58F494A8FBAEF872" box="[982,1036,1923,1949]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="biak">biak</taxonomicName>
, meaning &quot;crevice&quot; or &quot;crack,&quot; in reference to the preferred limestone karst habitat at the
<typeStatus id="54E7885BFFC0FFEE583E9482FCEDF82C" box="[796,847,1961,1987]" pageId="8" pageNumber="57">type</typeStatus>
locality,
<taxonomicName id="4C5C4D7AFFC0FFEE58989482FC53F82C" box="[954,1009,1961,1987]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="biak">Biak</taxonomicName>
na Bato (meaning &quot;cracked rock&quot; or &quot;fragmented stone&quot;), Bulacan Province.
</paragraph>
<caption id="DF236671FFC1FFEF5BB593BCFF69FF20" ID-Table-UUID="DF236671FFC1FFEF5BB593BCFF69FF20" httpUri="http://table.plazi.org/id/DF236671FFC1FFEF5BB593BCFF69FF20" pageId="9" pageNumber="58" targetBox="[151,1435,234,1360]" targetIsTable="true" targetPageId="9">
<paragraph id="8BE336F9FFC1FFEF5BB593BCFF69FF20" blockId="9.[151,1435,151,207]" pageId="9" pageNumber="58">
<emphasis id="B928EAEBFFC1FFEF5BB593BCFEABFF40" bold="true" box="[151,265,151,175]" pageId="9" pageNumber="58">TABLE 3.</emphasis>
Summary of univariate morphological variation among mensural characters in the type series of
<taxonomicName id="4C5C4D7AFFC1FFEF5E3E93B3FF67FF20" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC1FFEF5E3E93B3FF67FF20" italics="true" pageId="9" pageNumber="58">Platymantis biak</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE336F9FFC1FFEF593B93C1FB06FABF" pageId="9" pageNumber="58">
<table id="F95CC459FFC100195BB593C1FA39FABF" box="[151,1435,234,1360]" gridcols="3" gridrows="18" pageId="9" pageNumber="58">
<tr id="356C34BBFFC100195BB593C1FA39FECE" box="[151,1435,234,289]" gridrow="0" pageId="9" pageNumber="58" rowspan-0="1">
<th id="76BD5DC7FFC10019593B93C1FCAAFECE" box="[537,776,234,289]" gridcol="1" gridrow="0" pageId="9" pageNumber="58">
Female
<emphasis id="B928EAEBFFC1FFEF593B9221FD84FECE" box="[537,550,266,289]" italics="true" pageId="9" pageNumber="58">n</emphasis>
= 8
</th>
<th id="76BD5DC7FFC1001958FF93C1FA39FECE" box="[989,1435,234,289]" gridcol="2" gridrow="0" pageId="9" pageNumber="58">
Male
<emphasis id="B928EAEBFFC1FFEF58FF9221FC48FECE" box="[989,1002,266,289]" italics="true" pageId="9" pageNumber="58">n</emphasis>
= 24
</th>
</tr>
<tr id="356C34BBFFC100195BB5921EFA39FEA3" box="[151,1435,309,332]" gridrow="1" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB5921EFF5CFEA3" box="[151,254,309,332]" gridcol="0" gridrow="1" pageId="9" pageNumber="58">SVL</th>
<td id="76BD5DC7FFC10019593B921EFCAAFEA3" box="[537,776,309,332]" gridcol="1" gridrow="1" pageId="9" pageNumber="58">37.442.4 (40.1 ± 1.5)</td>
<td id="76BD5DC7FFC1001958FF921EFA39FEA3" box="[989,1435,309,332]" gridcol="2" gridrow="1" pageId="9" pageNumber="58">32.339.9 (35.9 ± 1.8)</td>
</tr>
<tr id="356C34BBFFC100195BB5924FFA39FE4A" box="[151,1435,356,421]" gridrow="2" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB5924FFF5CFE4A" box="[151,254,356,421]" gridcol="0" gridrow="2" pageId="9" pageNumber="58">HL END</th>
<td id="76BD5DC7FFC10019593B924FFCAAFE4A" box="[537,776,356,421]" gridcol="1" gridrow="2" pageId="9" pageNumber="58">14.917.3 (15.7 ± 0.7) 3.84.4 (4.1 ± 0.2)</td>
<td id="76BD5DC7FFC1001958FF924FFA39FE4A" box="[989,1435,356,421]" gridcol="2" gridrow="2" pageId="9" pageNumber="58">11.816.0 (13.6 ± 0.9) 3.23.9 (3.6 ± 0.2)</td>
</tr>
<tr id="356C34BBFFC100195BB59292FA39FE3F" box="[151,1435,441,464]" gridrow="3" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB59292FF5CFE3F" box="[151,254,441,464]" gridcol="0" gridrow="3" pageId="9" pageNumber="58">SNL</th>
<td id="76BD5DC7FFC10019593B9292FCAAFE3F" box="[537,776,441,464]" gridcol="1" gridrow="3" pageId="9" pageNumber="58">5.97.2 (6.6 ± 0.6)</td>
<td id="76BD5DC7FFC1001958FF9292FA39FE3F" box="[989,1435,441,464]" gridcol="2" gridrow="3" pageId="9" pageNumber="58">5.16.4 (5.9 ± 0.4)</td>
</tr>
<tr id="356C34BBFFC100195BB592CFFA39FDCA" box="[151,1435,484,549]" gridrow="4" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB592CFFF5CFDCA" box="[151,254,484,549]" gridcol="0" gridrow="4" pageId="9" pageNumber="58">IOD IND</th>
<td id="76BD5DC7FFC10019593B92CFFCAAFDCA" box="[537,776,484,549]" gridcol="1" gridrow="4" pageId="9" pageNumber="58">3.14.3 (3.6 ± 0.4) 3.04.0 (3.6 ± 0.4)</td>
<td id="76BD5DC7FFC1001958FF92CFFA39FDCA" box="[989,1435,484,549]" gridcol="2" gridrow="4" pageId="9" pageNumber="58">2.64.2 (3.4 ± 0.4) 2.63.6 (3.2 ± 0.3)</td>
</tr>
<tr id="356C34BBFFC100195BB59112FA39FDBF" box="[151,1435,569,592]" gridrow="5" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB59112FF5CFDBF" box="[151,254,569,592]" gridcol="0" gridrow="5" pageId="9" pageNumber="58">ED</th>
<td id="76BD5DC7FFC10019593B9112FCAAFDBF" box="[537,776,569,592]" gridcol="1" gridrow="5" pageId="9" pageNumber="58">4.35.2 (4.9 ± 0.3)</td>
<td id="76BD5DC7FFC1001958FF9112FA39FDBF" box="[989,1435,569,592]" gridcol="2" gridrow="5" pageId="9" pageNumber="58">3.35.2 (4.3 ± 0.5)</td>
</tr>
<tr id="356C34BBFFC100195BB5914FFA39FD4A" box="[151,1435,612,677]" gridrow="6" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB5914FFF5CFD4A" box="[151,254,612,677]" gridcol="0" gridrow="6" pageId="9" pageNumber="58">TAD ETD</th>
<td id="76BD5DC7FFC10019593B914FFCAAFD4A" box="[537,776,612,677]" gridcol="1" gridrow="6" pageId="9" pageNumber="58">3.34.3 (3.8 ± 0.3) 0.51.5 (0.8 ± 0.3)</td>
<td id="76BD5DC7FFC1001958FF914FFA39FD4A" box="[989,1435,612,677]" gridcol="2" gridrow="6" pageId="9" pageNumber="58">2.74.0 (3.4 ± 0.3) 0.31.4 (0.7 ± 0.2)</td>
</tr>
<tr id="356C34BBFFC100195BB59192FA39FD3F" box="[151,1435,697,720]" gridrow="7" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB59192FF5CFD3F" box="[151,254,697,720]" gridcol="0" gridrow="7" pageId="9" pageNumber="58">HW</th>
<td id="76BD5DC7FFC10019593B9192FCAAFD3F" box="[537,776,697,720]" gridcol="1" gridrow="7" pageId="9" pageNumber="58">15.117.2 (16.1 ± 0.7)</td>
<td id="76BD5DC7FFC1001958FF9192FA39FD3F" box="[989,1435,697,720]" gridcol="2" gridrow="7" pageId="9" pageNumber="58">12.515.7 (14.1 ± 0.9)</td>
</tr>
<tr id="356C34BBFFC100195BB591CFFA39FCCA" box="[151,1435,740,805]" gridrow="8" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB591CFFF5CFCCA" box="[151,254,740,805]" gridcol="0" gridrow="8" pageId="9" pageNumber="58">UA FAL</th>
<td id="76BD5DC7FFC10019593B91CFFCAAFCCA" box="[537,776,740,805]" gridcol="1" gridrow="8" pageId="9" pageNumber="58">7.99.6 (8.6 ± 0.6) 9.411.2 (10.0 ± 0.6)</td>
<td id="76BD5DC7FFC1001958FF91CFFA39FCCA" box="[989,1435,740,805]" gridcol="2" gridrow="8" pageId="9" pageNumber="58">7.29.7 (8.5 ± 0.7) 7.910.3 (9.2 ± 0.7)</td>
</tr>
<tr id="356C34BBFFC100195BB59012FA39FCBF" box="[151,1435,825,848]" gridrow="9" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB59012FF5CFCBF" box="[151,254,825,848]" gridcol="0" gridrow="9" pageId="9" pageNumber="58">FL</th>
<td id="76BD5DC7FFC10019593B9012FCAAFCBF" box="[537,776,825,848]" gridcol="1" gridrow="9" pageId="9" pageNumber="58">18.822.0 (20.7 ± 0.9)</td>
<td id="76BD5DC7FFC1001958FF9012FA39FCBF" box="[989,1435,825,848]" gridcol="2" gridrow="9" pageId="9" pageNumber="58">16.721.0 (18.8 ± 1.2)</td>
</tr>
<tr id="356C34BBFFC100195BB5904FFA39FC4A" box="[151,1435,868,933]" gridrow="10" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB5904FFF5CFC4A" box="[151,254,868,933]" gridcol="0" gridrow="10" pageId="9" pageNumber="58">TBL TSL</th>
<td id="76BD5DC7FFC10019593B904FFCAAFC4A" box="[537,776,868,933]" gridcol="1" gridrow="10" pageId="9" pageNumber="58">20.023.6 (21.0 ± 1.1) 10.318.6 (12.3 ± 2.7)</td>
<td id="76BD5DC7FFC1001958FF904FFA39FC4A" box="[989,1435,868,933]" gridcol="2" gridrow="10" pageId="9" pageNumber="58">16.721.7 (19.0 ± 1.4) 9.412.5 (10.8 ± 0.9)</td>
</tr>
<tr id="356C34BBFFC100195BB59092FA39FC3F" box="[151,1435,953,976]" gridrow="11" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB59092FF5CFC3F" box="[151,254,953,976]" gridcol="0" gridrow="11" pageId="9" pageNumber="58">PL</th>
<td id="76BD5DC7FFC10019593B9092FCAAFC3F" box="[537,776,953,976]" gridcol="1" gridrow="11" pageId="9" pageNumber="58">11.319.4 (17.8 ± 2.7)</td>
<td id="76BD5DC7FFC1001958FF9092FA39FC3F" box="[989,1435,953,976]" gridcol="2" gridrow="11" pageId="9" pageNumber="58">14.720.4 (17.5 ± 1.5)</td>
</tr>
<tr id="356C34BBFFC100195BB590CFFA39FBCA" box="[151,1435,996,1061]" gridrow="12" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB590CFFF5CFBCA" box="[151,254,996,1061]" gridcol="0" gridrow="12" pageId="9" pageNumber="58">ML Toe4 L</th>
<td id="76BD5DC7FFC10019593B90CFFCAAFBCA" box="[537,776,996,1061]" gridcol="1" gridrow="12" pageId="9" pageNumber="58">10.312.1 (11.1 ± 0.6) 10.813.4 (12.1 ± 0.8)</td>
<td id="76BD5DC7FFC1001958FF90CFFA39FBCA" box="[989,1435,996,1061]" gridcol="2" gridrow="12" pageId="9" pageNumber="58">7.812.4 (10.2 ± 1.1) 9.612.8 (11.4 ± 1.0)</td>
</tr>
<tr id="356C34BBFFC100195BB59712FA39FBBF" box="[151,1435,1081,1104]" gridrow="13" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB59712FF5CFBBF" box="[151,254,1081,1104]" gridcol="0" gridrow="13" pageId="9" pageNumber="58">Fin1L</th>
<td id="76BD5DC7FFC10019593B9712FCAAFBBF" box="[537,776,1081,1104]" gridcol="1" gridrow="13" pageId="9" pageNumber="58">5.47.3 (6.3 ± 0.8)</td>
<td id="76BD5DC7FFC1001958FF9712FA39FBBF" box="[989,1435,1081,1104]" gridcol="2" gridrow="13" pageId="9" pageNumber="58">4.17.6 (5.5 ± 0.9)</td>
</tr>
<tr id="356C34BBFFC100195BB5974FFA39FB4A" box="[151,1435,1124,1189]" gridrow="14" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB5974FFF5CFB4A" box="[151,254,1124,1189]" gridcol="0" gridrow="14" pageId="9" pageNumber="58">Fin3L Fin1DW</th>
<td id="76BD5DC7FFC10019593B974FFCAAFB4A" box="[537,776,1124,1189]" gridcol="1" gridrow="14" pageId="9" pageNumber="58">7.68.8 (8.1 ± 0.5) 0.81.2 (1.1 ± 0.1)</td>
<td id="76BD5DC7FFC1001958FF974FFA39FB4A" box="[989,1435,1124,1189]" gridcol="2" gridrow="14" pageId="9" pageNumber="58">6.58.6 (7.6 ± 0.5) 0.81.2 (0.9 ± 0.1)</td>
</tr>
<tr id="356C34BBFFC100195BB59792FA39FB3F" box="[151,1435,1209,1232]" gridrow="15" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB59792FF5CFB3F" box="[151,254,1209,1232]" gridcol="0" gridrow="15" pageId="9" pageNumber="58">Fin3DW</th>
<td id="76BD5DC7FFC10019593B9792FCAAFB3F" box="[537,776,1209,1232]" gridcol="1" gridrow="15" pageId="9" pageNumber="58">1.31.9 (1.6 ± 0.2)</td>
<td id="76BD5DC7FFC1001958FF9792FA39FB3F" box="[989,1435,1209,1232]" gridcol="2" gridrow="15" pageId="9" pageNumber="58">0.11.2 (1.3 ± 0.2)</td>
</tr>
<tr id="356C34BBFFC100195BB597CFFA39FACA" box="[151,1435,1252,1317]" gridrow="16" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB597CFFF5CFACA" box="[151,254,1252,1317]" gridcol="0" gridrow="16" pageId="9" pageNumber="58">Toe4 DW PpFin3</th>
<td id="76BD5DC7FFC10019593B97CFFCAAFACA" box="[537,776,1252,1317]" gridcol="1" gridrow="16" pageId="9" pageNumber="58">1.01.5 (1.3 ± 0.2) 0.50.9 (0.7 ± 0.1)</td>
<td id="76BD5DC7FFC1001958FF97CFFA39FACA" box="[989,1435,1252,1317]" gridcol="2" gridrow="16" pageId="9" pageNumber="58">0.71.3 (1.1 ± 0.2) 0.40.9 (0.7 ± 0.1)</td>
</tr>
<tr id="356C34BBFFC100195BB59612FA39FABF" box="[151,1435,1337,1360]" gridrow="17" pageId="9" pageNumber="58">
<th id="76BD5DC7FFC100195BB59612FF5CFABF" box="[151,254,1337,1360]" gridcol="0" gridrow="17" pageId="9" pageNumber="58">PpToe4</th>
<td id="76BD5DC7FFC10019593B9612FCAAFABF" box="[537,776,1337,1360]" gridcol="1" gridrow="17" pageId="9" pageNumber="58">0.40.9 (0.7 ± 0.2)</td>
<td id="76BD5DC7FFC1001958FF9612FA39FABF" box="[989,1435,1337,1360]" gridcol="2" gridrow="17" pageId="9" pageNumber="58">0.50.8 (0.7 ± 0.1)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE336F9FFC1FFEF5BE696BDFD6CF9A9" blockId="9.[151,1437,1430,1979]" pageId="9" pageNumber="58">
<emphasis id="B928EAEBFFC1FFEF5BE696BDFEDEFA5F" bold="true" box="[196,380,1430,1456]" pageId="9" pageNumber="58">Comparisons.</emphasis>
The new species is assignable to the
<emphasis id="B928EAEBFFC1FFEF587996BCFC52FA5F" box="[859,1008,1431,1456]" italics="true" pageId="9" pageNumber="58">P. g u e n t h e r i</emphasis>
Group (
<emphasis id="B928EAEBFFC1FFEF5F4296BCFBEBFA3A" italics="true" pageId="9" pageNumber="58">
<taxonomicName id="4C5C4D7AFFC1FFEF5F4296BCFB53FA5F" box="[1120,1265,1431,1456]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="banahao">P. banahao</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC1FFEF5FDD96BCFADAFA5F" box="[1279,1400,1431,1456]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="bayani">P. bayani</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC1FFEF5EA796BCFEA6FA3A" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="cornutus">P. cornutus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC1FFEF5A339697FE3EFA3A" box="[273,412,1468,1493]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="diesmosi">P. diesmosi</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC1FFEF5A8B9697FD9DFA3A" box="[425,575,1468,1493]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="guentheri">P. guentheri</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC1FFEF596E9697FD78FA3A" box="[588,730,1468,1493]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="insulatus">P. insulatus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC1FFEF59CA9697FC2AFA3A" box="[744,904,1468,1493]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="luzonensis">P. luzonensis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC1FFEF58B79697FBE1FA3A" box="[917,1091,1468,1493]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="negrosensis">P. negrosensis</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C5C4D7AFFC1FFEF5FA79697FB53FA3A" box="[1157,1265,1468,1493]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="rabori">
<emphasis id="B928EAEBFFC1FFEF5FA79697FB53FA3A" box="[1157,1265,1468,1493]" italics="true" pageId="9" pageNumber="58">P. rabori</emphasis>
</taxonomicName>
) based on the following combination of characters: digital discs moderately expanded and larger in fingers than in toes, proximal portions of digits with narrow dermal folds in cross section, and subarticular tubercles large and highly protuberant (
<bibRefCitation id="EFCD4B08FFC1FFEF5A5D9507FDCBF9AA" author="Brown" box="[383,617,1580,1606]" pageId="9" pageNumber="61" refString="Brown, W. C., Alcala, A. C., Diesmos, A. C &amp; Alcala, E. (1997 c) Species of the guentheri group of Platymantis from the Philippines with descriptions of four new species. Proceedings of the California Academy of Sciences, 50, 1 - 20." type="journal article" year="1997" yearSuffix="c">
Brown
<emphasis id="B928EAEBFFC1FFEF5AFA9507FDB1F9AA" box="[472,531,1580,1605]" italics="true" pageId="9" pageNumber="58">et al.</emphasis>
, 1997
<emphasis id="B928EAEBFFC1FFEF597E9507FDCBF9AA" box="[604,617,1580,1605]" italics="true" pageId="9" pageNumber="58">c</emphasis>
</bibRefCitation>
; Fig. 3).
</paragraph>
<paragraph id="8BE336F9FFC1FFEF5BE6957AFB7BF854" blockId="9.[151,1437,1430,1979]" pageId="9" pageNumber="58">
The new species is morphologically most similar to
<taxonomicName id="4C5C4D7AFFC1FFEF581F957AFB85F985" box="[829,1063,1617,1642]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="bayani">
<emphasis id="B928EAEBFFC1FFEF581F957AFB85F985" box="[829,1063,1617,1642]" italics="true" pageId="9" pageNumber="58">Platymantis bayani</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5C4D7AFFC1FFEF5F41957AFAC5F985" box="[1123,1383,1617,1642]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="insulatus">
<emphasis id="B928EAEBFFC1FFEF5F41957AFAC5F985" box="[1123,1383,1617,1642]" italics="true" pageId="9" pageNumber="58">Platymantis insulatus</emphasis>
</taxonomicName>
, the other two members of the
<emphasis id="B928EAEBFFC1FFEF5AC6955CFDDAF97F" box="[484,632,1655,1680]" italics="true" pageId="9" pageNumber="58">P. g u e n t h e r i</emphasis>
Group with similar morphology and preference for karst forest microhabitats (
<tableCitation id="C6DE0342FFC1FFEF5A6D95B7FE1BF959" box="[335,441,1692,1718]" captionStart="TABLE 1" captionStartId="6.[151,240,151,175]" captionTargetBox="[165,1412,367,1936]" captionTargetPageId="6" captionText="TABLE 1. Summary of qualitative diagnostic characters (present, absent) in Platymantis biak and specimens of other members of the P. guentheri Species Group. Sample size for each sex, body size, eye diameter, digit disk proportions, and general geographical distribution (PAIC = Pleistocene Aggregate Island Complexes, sensu Brown and Diesmos, 2002) are included for reference (SVL and ED given as range over mean ± standard deviation; digit disk proportions given as percentage over mean ± standard deviation). Only adult specimens in good preservation condition were used for morphometric analyses." httpUri="http://table.plazi.org/id/DF236671FFCEFFE05BB593BCFE34FEA0" pageId="9" pageNumber="58" tableUuid="DF236671FFCEFFE05BB593BCFE34FEA0">Tables 1</tableCitation>
and
<tableCitation id="C6DE0342FFC1FFEF5AD495B7FDAAF959" box="[502,520,1692,1718]" captionStart="TABLE 2" captionStartId="8.[151,240,151,175]" captionText="TABLE 2. Summary of mensural characters (in mm) and selected qualitative diagnostic characters (present, absent) in Platymantis biak and other Philippine limestone forest frog species. Sample size, body size, species group, and general geographical distribution are included for reference (SVL given as range)." pageId="9" pageNumber="58">2</tableCitation>
).
<taxonomicName id="4C5C4D7AFFC1FFEF590395B7FD53F95A" box="[545,753,1692,1717]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC1FFEF590395B7FD53F95A" box="[545,753,1692,1717]" italics="true" pageId="9" pageNumber="58">Platymantis biak</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C5C4D7AFFC1FFEF5F2A95B7FBD8F95A" box="[1032,1146,1692,1717]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="bayani">
<emphasis id="B928EAEBFFC1FFEF5F2A95B7FBD8F95A" box="[1032,1146,1692,1717]" italics="true" pageId="9" pageNumber="58">P. bayani</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5C4D7AFFC1FFEF5F9A95B7FAE5F95A" box="[1208,1351,1692,1717]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="insulatus">
<emphasis id="B928EAEBFFC1FFEF5F9A95B7FAE5F95A" box="[1208,1351,1692,1717]" italics="true" pageId="9" pageNumber="58">P. insulatus</emphasis>
</taxonomicName>
by the presence of moderately (vs. widely) expanded digital discs and the absence of fully spotted flanks (vs. spotted). From
<taxonomicName id="4C5C4D7AFFC1FFEF5A6C95CCFD96F8EF" box="[334,564,1767,1792]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="bayani">
<emphasis id="B928EAEBFFC1FFEF5A6C95CCFD96F8EF" box="[334,564,1767,1792]" italics="true" pageId="9" pageNumber="58">Platymantis bayani</emphasis>
</taxonomicName>
, the new species differs by its smaller female body size (maximum SVL
<quantity id="4CA49B1CFFC1FFEF5BB59427FEA3F8C9" box="[151,257,1804,1830]" metricMagnitude="-2" metricUnit="m" metricValue="4.24" pageId="9" pageNumber="58" unit="mm" value="42.4">42.4 mm</quantity>
for
<taxonomicName id="4C5C4D7AFFC1FFEF5A119427FE26F8CA" box="[307,388,1804,1829]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC1FFEF5A119427FE26F8CA" box="[307,388,1804,1829]" italics="true" pageId="9" pageNumber="58">P. biak</emphasis>
</taxonomicName>
vs.
<quantity id="4CA49B1CFFC1FFEF5A949427FD82F8C9" box="[438,544,1804,1830]" metricMagnitude="-2" metricUnit="m" metricValue="4.9799999999999995" pageId="9" pageNumber="58" unit="mm" value="49.8">49.8 mm</quantity>
for
<taxonomicName id="4C5C4D7AFFC1FFEF59719427FD63F8CA" box="[595,705,1804,1829]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="bayani">
<emphasis id="B928EAEBFFC1FFEF59719427FD63F8CA" box="[595,705,1804,1829]" italics="true" pageId="9" pageNumber="58">P. bayani</emphasis>
</taxonomicName>
), smaller eye diameter (maximum ED
<quantity id="4CA49B1CFFC1FFEF5FB69427FB4DF8C9" box="[1172,1263,1804,1830]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="58" unit="mm" value="5.2">5.2 mm</quantity>
for
<emphasis id="B928EAEBFFC1FFEF5E039427FACDF8CA" box="[1313,1391,1804,1829]" italics="true" pageId="9" pageNumber="58">P. b i a k</emphasis>
vs.
<quantity id="4CA49B1CFFC1FFEF5BB5941AFF53F8A4" box="[151,241,1841,1867]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="58" unit="mm" value="6.8">6.8 mm</quantity>
for
<taxonomicName id="4C5C4D7AFFC1FFEF5A00941AFE2DF8A5" box="[290,399,1841,1866]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="bayani">
<emphasis id="B928EAEBFFC1FFEF5A00941AFE2DF8A5" box="[290,399,1841,1866]" italics="true" pageId="9" pageNumber="58">P. bayani</emphasis>
</taxonomicName>
), and the absence of granular skin (vs. absence or presence) and dorsal dermal tubercles (vs. presence). The new species further differs from
<taxonomicName id="4C5C4D7AFFC1FFEF5830947CFBB4F89F" box="[786,1046,1879,1904]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="insulatus">
<emphasis id="B928EAEBFFC1FFEF5830947CFBB4F89F" box="[786,1046,1879,1904]" italics="true" pageId="9" pageNumber="58">Platymantis insulatus</emphasis>
</taxonomicName>
by the presence of well-defined limb banding patterns (vs. absence or presence and poorly defined). Additional characters for distinguishing the new species from all species of the
<taxonomicName id="4C5C4D7AFFC1FFEF5940948AFD4DF855" box="[610,751,1953,1978]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B928EAEBFFC1FFEF5940948AFD4DF855" box="[610,751,1953,1978]" italics="true" pageId="9" pageNumber="58">P. guentheri</emphasis>
</taxonomicName>
Group are summarized in
<tableCitation id="C6DE0342FFC1FFEF5F0A948AFB2BF854" box="[1064,1161,1953,1979]" captionStart="TABLE 1" captionStartId="6.[151,240,151,175]" captionTargetBox="[165,1412,367,1936]" captionTargetPageId="6" captionText="TABLE 1. Summary of qualitative diagnostic characters (present, absent) in Platymantis biak and specimens of other members of the P. guentheri Species Group. Sample size for each sex, body size, eye diameter, digit disk proportions, and general geographical distribution (PAIC = Pleistocene Aggregate Island Complexes, sensu Brown and Diesmos, 2002) are included for reference (SVL and ED given as range over mean ± standard deviation; digit disk proportions given as percentage over mean ± standard deviation). Only adult specimens in good preservation condition were used for morphometric analyses." httpUri="http://table.plazi.org/id/DF236671FFCEFFE05BB593BCFE34FEA0" pageId="9" pageNumber="58" tableUuid="DF236671FFCEFFE05BB593BCFE34FEA0">Tables 1</tableCitation>
and
<tableCitation id="C6DE0342FFC1FFEF5FE1948AFB77F854" box="[1219,1237,1953,1979]" captionStart="TABLE 2" captionStartId="8.[151,240,151,175]" captionText="TABLE 2. Summary of mensural characters (in mm) and selected qualitative diagnostic characters (present, absent) in Platymantis biak and other Philippine limestone forest frog species. Sample size, body size, species group, and general geographical distribution are included for reference (SVL given as range)." pageId="9" pageNumber="58">2</tableCitation>
.
</paragraph>
<paragraph id="8BE336F9FFC2FFEC5BE693B3FC68FDC8" blockId="10.[151,1437,152,738]" pageId="10" pageNumber="59">
From all members of the
<emphasis id="B928EAEBFFC2FFEC5AD793B3FDD2FF5E" box="[501,624,152,177]" italics="true" pageId="10" pageNumber="59">P. d o r s a l i s</emphasis>
Group (
<emphasis id="B928EAEBFFC2FFEC59F893B3FB9DFF39" italics="true" pageId="10" pageNumber="59">
<taxonomicName id="4C5C4D7AFFC2FFEC59F893B3FCDCFF5E" box="[730,894,152,177]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="cagayensis">P. cagayensis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC58A993B3FB8DFF5E" box="[907,1071,152,177]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="corrugatus">P. corrugatus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5F1E93B3FB1FFF5E" box="[1084,1213,152,177]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="dorsalis">P. dorsalis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5FE893B3FAD8FF5E" box="[1226,1402,152,177]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="indeprensus">P. indeprensus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5EA493B3FEAAFF39" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="levigatus">P. levigatus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5A369396FE3FFF39" box="[276,413,189,214]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="mimulus">P. mimulus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5A8B9396FD91FF39" box="[425,563,189,214]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="naomiae">P. naomiae</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC59639396FD16FF39" box="[577,692,189,214]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="paengi">P. paengi</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC59E09396FC39FF39" box="[706,923,189,214]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="pseudodorsalis">P. pseudodorsalis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC588A9396FB9DFF39" box="[936,1087,189,214]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="pygmaeus">P. pygmaeus</taxonomicName>
</emphasis>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5F6D9396FB74FF39" box="[1103,1238,189,214]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="spelaeus">
<emphasis id="B928EAEBFFC2FFEC5F6D9396FB74FF39" box="[1103,1238,189,214]" italics="true" pageId="10" pageNumber="59">P. spelaeus</emphasis>
</taxonomicName>
, and
<emphasis id="B928EAEBFFC2FFEC5E399396FA25FF39" box="[1307,1415,189,214]" italics="true" pageId="10" pageNumber="59">P. t a y l o r i</emphasis>
),
<taxonomicName id="4C5C4D7AFFC2FFEC5BB593C9FEFCFF14" box="[151,350,226,251]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC2FFEC5BB593C9FEFCFF14" box="[151,350,226,251]" italics="true" pageId="10" pageNumber="59">Platymantis biak</emphasis>
</taxonomicName>
is distinguished by the presence (vs. absence) of moderately expanded digital discs (Fig. 3) and larger finger discs than toe discs.
<taxonomicName id="4C5C4D7AFFC2FFEC59469223FC8DFECE" box="[612,815,264,289]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="biak">
<emphasis id="B928EAEBFFC2FFEC59469223FC8DFECE" box="[612,815,264,289]" italics="true" pageId="10" pageNumber="59">Platymantis biak</emphasis>
</taxonomicName>
is further distinguished from all members of the
<taxonomicName id="4C5C4D7AFFC2FFEC5EA49223FF54FEA9" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="B928EAEBFFC2FFEC5EA49223FF54FEA9" italics="true" pageId="10" pageNumber="59">P. dorsalis</emphasis>
</taxonomicName>
Group, except
<taxonomicName id="4C5C4D7AFFC2FFEC5A8B9206FD86FEA9" box="[425,548,301,326]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="levigata">
<emphasis id="B928EAEBFFC2FFEC5A8B9206FD86FEA9" box="[425,548,301,326]" italics="true" pageId="10" pageNumber="59">P. levigata</emphasis>
</taxonomicName>
by the absence of dorsal folds and ridges, dermal tubercles, and rugose skin (vs. presence of one, two, or all characters), and from all members except
<taxonomicName id="4C5C4D7AFFC2FFEC5F3B9279FB2BFE84" box="[1049,1161,338,363]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="paengi">
<emphasis id="B928EAEBFFC2FFEC5F3B9279FB2BFE84" box="[1049,1161,338,363]" italics="true" pageId="10" pageNumber="59">P. paengi</emphasis>
</taxonomicName>
and
<emphasis id="B928EAEBFFC2FFEC5FE69279FAF3FE84" box="[1220,1361,338,363]" italics="true" pageId="10" pageNumber="59">
<taxonomicName id="4C5C4D7AFFC2FFEC5FE69279FAEFFE84" box="[1220,1357,338,363]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="spelaeus">P. spelaeus</taxonomicName>
,
</emphasis>
by its microhabitat preference (terrestrial substrates in limestone forest vs. leaf litter of forest floor and/or herb-layer vegetation in
<emphasis id="B928EAEBFFC2FFEC5A6E92B6FE4FFE34" italics="true" pageId="10" pageNumber="59">
<taxonomicName id="4C5C4D7AFFC2FFEC5A6E92B6FDA3FE59" box="[332,513,413,438]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="cagayensis">P. cagayensis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC592D92B6FD66FE59" box="[527,708,413,438]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="corrugatus">P. corrugatus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC59F092B6FCC3FE59" box="[722,865,413,438]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="dorsalis">P. dorsalis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC585292B6FB90FE59" box="[880,1074,413,438]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="indeprensus">P. indeprensus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5F6292B6FB71FE59" box="[1088,1235,413,438]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="mimulus">P. mimulus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5FC092B6FAD5FE59" box="[1250,1399,413,438]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="naomiae">P. naomiae</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5EA792B6FEEFFE34" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="pseudodorsalis">P. pseudodorsalis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5A7B92E9FE4FFE34" box="[345,493,450,475]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="pygmaeus">P. pygmaeus</taxonomicName>
</emphasis>
and
<emphasis id="B928EAEBFFC2FFEC590A92E9FD30FE34" box="[552,658,450,475]" italics="true" pageId="10" pageNumber="59">P. t a y l o r i</emphasis>
, or river banks and rocks in
<taxonomicName id="4C5C4D7AFFC2FFEC58CF92E9FBD4FE34" box="[1005,1142,450,475]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="levigatus">
<emphasis id="B928EAEBFFC2FFEC58CF92E9FBD4FE34" box="[1005,1142,450,475]" italics="true" pageId="10" pageNumber="59">P. levigatus</emphasis>
</taxonomicName>
). From
<emphasis id="B928EAEBFFC2FFEC5FF592E9FBB5FDEE" italics="true" pageId="10" pageNumber="59">
<taxonomicName id="4C5C4D7AFFC2FFEC5FF592E9FADBFE34" box="[1239,1401,450,475]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="cagayensis">P. cagayensis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5EA492E9FEB8FDEE" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="corrugatus">P. corrugatus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5A0492C3FE01FDEE" box="[294,419,488,513]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="dorsalis">P. dorsalis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5A8D92C3FDF9FDEE" box="[431,603,488,513]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="indeprensus">P. indeprensus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC594592C3FD48FDEE" box="[615,746,488,513]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="mimulus">P. mimulus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC59D792C3FCD8FDEE" box="[757,890,488,513]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="naomiae">P. naomiae</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC58A792C3FBB5FDEE" box="[901,1047,488,513]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="pygmaeus">P. pygmaeus</taxonomicName>
</emphasis>
and
<emphasis id="B928EAEBFFC2FFEC5F7292C3FB1DFDEE" box="[1104,1215,488,513]" italics="true" pageId="10" pageNumber="59">P. t a y l o r i,</emphasis>
the new species is distinguished by the absence (vs. presence) of dorsal folds and ridges.
</paragraph>
<paragraph id="8BE336F9FFC2FFEC5BE69119FDC2FD0D" blockId="10.[151,1437,152,738]" pageId="10" pageNumber="59">
The new species is distinguished from all
<taxonomicName id="4C5C4D7AFFC2FFEC599C9119FC9AFDA4" box="[702,824,562,587]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="hazelae">
<emphasis id="B928EAEBFFC2FFEC599C9119FC9AFDA4" box="[702,824,562,587]" italics="true" pageId="10" pageNumber="59">P. hazelae</emphasis>
</taxonomicName>
Group species (
<emphasis id="B928EAEBFFC2FFEC58DE9119FBB6FD9E" italics="true" pageId="10" pageNumber="59">
<taxonomicName id="4C5C4D7AFFC2FFEC58DE9119FBDBFDA4" box="[1020,1145,562,587]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="hazelae">P. hazelae</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5FA49119FB50FDA4" box="[1158,1266,562,587]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="isarog">P. isarog</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5FDD9119FAD8FDA4" box="[1279,1402,562,587]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="lawtoni">P. lawtoni</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5EA49119FEACFD9E" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="montanus">P. montanus</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5A389173FE1DFD9E" box="[282,447,600,625]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="panayensis">P. panayensis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC5AE99173FDC4FD9E" box="[459,614,600,625]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="polillensis">P. polillensis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC59519173FCF6FD9E" box="[627,852,600,625]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="sierramadrensis">P. sierramadrensis</taxonomicName>
,
<taxonomicName id="4C5C4D7AFFC2FFEC58439173FBB6FD9E" box="[865,1044,600,625]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="subterrestris">P. subterrestris</taxonomicName>
</emphasis>
) by its larger body size; the first finger slightly longer than the second; narrow dermal phalanges on the digits; protuberant (vs. flattened) subarticular tubercles, and a preferred microhabitat of terrestrial substrates in limestone forest (vs. arboreal habitat, and/or shrub-layer vegetation).
</paragraph>
</subSubSection>
</treatment>
</document>