121 lines
16 KiB
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121 lines
16 KiB
XML
<document id="18775BE589C743224025E96BC50B89D3" ID-CLB-Dataset="204277" ID-DOI="10.11646/zootaxa.5219.2.7" ID-GBIF-Dataset="84d3b99b-968c-4a1c-9975-a3d1cd882a24" ID-ISSN="1175-5326" ID-Zenodo-Dep="7413471" ID-ZooBank="B50DA711-9900-41BE-9CD9-36AD01B8C078" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670488696986" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2022" docId="03F587EAFF9CFFDDFF2144FFE98118B2" docLanguage="en" docName="zootaxa.5219.2.7.pdf" docOrigin="Zootaxa 5219 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liaoninosoma Mikhaljova 2022, gen. nov." docType="treatment" docVersion="6" lastPageNumber="186" masterDocId="FFCCFF92FF9DFFDCFFB64670EB111D09" masterDocTitle="The millipede family Diplomaragnidae Attems, 1907 confirmed for mainland China, with descriptions of a new genus and new species (Diplopoda, Chordeumatida)" masterLastPageNumber="192" masterPageNumber="185" pageNumber="186" updateTime="1699413216787" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="0828548B4E1D829D2AE51090E462A6AE">
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<mods:title id="388459BFA1B4AAB144554055187D09B8">The millipede family Diplomaragnidae Attems, 1907 confirmed for mainland China, with descriptions of a new genus and new species (Diplopoda, Chordeumatida)</mods:title>
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<mods:namePart id="D927B8D6D9FE8FC12801204DE0E84AB7">Mikhaljova, Elena V.</mods:namePart>
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<mods:title id="9E26F8BBE6E0A37F689B541E01E72B24">Zootaxa</mods:title>
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<mods:part id="2B5D262D547D861F76F1831DBF515E9A">
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<mods:date id="4A47300C40C3A6C1394253D8F53BC3B0">2022</mods:date>
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<mods:number id="D9696740FA81861663AE300BFD99CD08">2022-12-08</mods:number>
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<mods:number id="011D106727B46DA2932770B19F56A75B">5219</mods:number>
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<mods:number id="270285BBE9EFCB4ADD4A63C8208AEAD4">2</mods:number>
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<mods:identifier id="24C2F3C0DB931673D98173E2827CD8D4" type="DOI">10.11646/zootaxa.5219.2.7</mods:identifier>
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<treatment id="03F587EAFF9CFFDDFF2144FFE98118B2" ID-DOI="http://doi.org/10.5281/zenodo.7413473" ID-GBIF-Taxon="205936975" ID-Zenodo-Dep="7413473" LSID="urn:lsid:plazi:treatment:03F587EAFF9CFFDDFF2144FFE98118B2" httpUri="http://treatment.plazi.org/id/03F587EAFF9CFFDDFF2144FFE98118B2" lastPageNumber="186" pageId="1" pageNumber="186">
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<subSubSection id="C3466577FF9CFFDDFF2144FFEAAA1FA0" box="[151,443,655,681]" pageId="1" pageNumber="186" type="nomenclature">
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<paragraph id="8BE336FCFF9CFFDDFF2144FFEAAA1FA0" blockId="1.[151,443,655,681]" box="[151,443,655,681]" pageId="1" pageNumber="186">
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<heading id="D0AB8190FF9CFFDDFF2144FFEAAA1FA0" bold="true" box="[151,443,655,681]" fontSize="11" level="1" pageId="1" pageNumber="186" reason="1">
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<emphasis id="B928EAEEFF9CFFDDFF2144FFEAAA1FA0" bold="true" box="[151,443,655,681]" pageId="1" pageNumber="186">
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<taxonomicName id="4C5C4D7FFF9CFFDDFF2144FFEA541FA0" authority="Mikhaljova, 2022" authorityName="Mikhaljova" authorityYear="2022" box="[151,325,655,681]" class="Diplopoda" family="Diplomaragnidae" genus="Liaoninosoma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="186" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B928EAEEFF9CFFDDFF2144FFEA541FA0" bold="true" box="[151,325,655,681]" italics="true" pageId="1" pageNumber="186">Liaoninosoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21B5795FF9CFFDDFEFB44FFEAAA1FA0" box="[333,443,655,681]" pageId="1" pageNumber="186" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3466577FF9CFFDDFF2144A4E8D51FE2" box="[151,964,723,748]" pageId="1" pageNumber="186" type="materials_examined">
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<paragraph id="8BE336FCFF9CFFDDFF2144A4E8D51FE2" blockId="1.[151,964,723,748]" box="[151,964,723,748]" pageId="1" pageNumber="186">
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<typeStatus id="54E7885EFF9CFFDDFF2144A4EBD81FE5" box="[151,201,724,748]" pageId="1" pageNumber="186">Type</typeStatus>
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and only species:
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<taxonomicName id="4C5C4D7FFF9CFFDDFE3344A4E9641FE2" authority="Mikhaljova, 2022" authorityName="Mikhaljova" authorityYear="2022" box="[389,629,723,747]" class="Diplopoda" family="Diplomaragnidae" genus="Liaoninosoma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
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<emphasis id="B928EAEEFF9CFFDDFE3344A4E9641FE2" box="[389,629,723,747]" italics="true" pageId="1" pageNumber="186">Liaoninosoma marusiki</emphasis>
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</taxonomicName>
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<emphasis id="B928EAEEFF9CFFDDFDCD44A3E9C31FE2" bold="true" box="[635,722,723,747]" pageId="1" pageNumber="186">
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<taxonomicNameLabel id="A21B5795FF9CFFDDFDCD44A3E9DF1FE2" box="[635,718,723,747]" pageId="1" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
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,
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</emphasis>
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by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3466577FF9CFFDDFF214569EE491ECA" pageId="1" pageNumber="186" type="diagnosis">
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<paragraph id="8BE336FCFF9CFFDDFF214569EFCB1EA9" blockId="1.[151,1437,793,1467]" pageId="1" pageNumber="186">
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<emphasis id="B928EAEEFF9CFFDDFF214569EA031E3A" bold="true" box="[151,274,793,819]" pageId="1" pageNumber="186">Diagnosis.</emphasis>
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The genus differs from other genera of the family mainly by the posterior gonopod colpocoxite, which has an independent (separate), broad, longitudinal structure (“pad”) (
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<emphasis id="B928EAEEFF9CFFDDFC2D454DE8A61E5E" bold="true" box="[923,951,829,855]" pageId="1" pageNumber="186">cp</emphasis>
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), covered with dense spikes (
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<figureCitation id="13672A79FF9CFFDDFAB5454DEE441E5E" box="[1283,1365,829,855]" captionStart="FIGURES 15–20" captionStartId="4.[151,264,1511,1535]" captionTargetBox="[152,1435,181,1487]" captionTargetId="figure-20@4.[152,1435,181,1487]" captionTargetPageId="4" captionText="FIGURES 15–20. Liaoninosoma marusiki sp. nov., two male paratypes (FSCB). 15. Gonopods, posterior view (posterior angiocoxal process on the left removed). 16. Colpocoxite distal part without posterior angiocoxal process, posterior view. 17. Colpocoxite distal part, posterior view. 18. Gonopods, posterior view. 19. Spikes. 20. Gonopods without posterior gonopod telopodites, lateral view. Abbreviations: a, angiocoxite; at, anterior gonopod telopodite; ax, apex; b, mesal blade; ba, frontal blade; cp, colpocoxite “pad’’; ms, spear-shaped structure; p, conical process; pp, posterior angiocoxal process; sg, subglobule. Scales: 10 μm (Fig. 19), 20 μm (Figs 16, 17), 100 μm (Figs 15, 18, 20)." figureDoi="http://doi.org/10.5281/zenodo.7413481" httpUri="https://zenodo.org/record/7413481/files/figure.png" pageId="1" pageNumber="186">Fig. 16</figureCitation>
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), with a shallow, partly open-edged sheath groove for the anterior gonopod telopodite together with a rectangular laterally and slightly caudally curved distal part, in combination with a large posterior angiocoxal process.
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</paragraph>
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<paragraph id="8BE336FCFF9CFFDDFF7145DAEE491ECA" blockId="1.[151,1437,793,1467]" box="[199,1368,937,964]" pageId="1" pageNumber="186">A structure similar to the above-mentioned “pad” is not observed among all known genera of the family.</paragraph>
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</subSubSection>
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<subSubSection id="C3466577FF9CFFDDFF7145BDEE9C187A" pageId="1" pageNumber="186" type="description">
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<paragraph id="8BE336FCFF9CFFDDFF7145BDEBF9195A" blockId="1.[151,1437,793,1467]" pageId="1" pageNumber="186">
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<emphasis id="B928EAEEFF9CFFDDFF7145BDEA481EEE" bold="true" box="[199,345,973,999]" pageId="1" pageNumber="186">Description.</emphasis>
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Body with 30 rings (including telson). Ommatidia present, fields triangular. Medium-sized paraterga well-developed, horizontal. Male legs 3 and 4 not enlarged. Male legs 5–7 enlarged, coxae with low subconical outgrowths, femora swollen and curved, tarsi very long, sabre-shaped. Male legs 10 and 11 with coxal glands.
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</paragraph>
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<paragraph id="8BE336FCFF9CFFDDFF71422DEE9C187A" blockId="1.[151,1437,793,1467]" pageId="1" pageNumber="186">
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Telopodites of anterior gonopods flagelliform, 1-segmented, sheathed. Posterior gonopod colpocoxite with an independent (separate), broad, longitudinal spiky structure (“pad”) (
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<emphasis id="B928EAEEFF9CFFDDFC6D42F1E8E61992" bold="true" box="[987,1015,1153,1179]" pageId="1" pageNumber="186">cp</emphasis>
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) with a shallow, partly open-edged sheath groove for the anterior gonopod telopodite (
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<figureCitation id="13672A79FF9CFFDDFD5D42D5E86519B6" box="[747,884,1189,1216]" captionStart="FIGURES 15–20" captionStartId="4.[151,264,1511,1535]" captionTargetBox="[152,1435,181,1487]" captionTargetId="figure-20@4.[152,1435,181,1487]" captionTargetPageId="4" captionText="FIGURES 15–20. Liaoninosoma marusiki sp. nov., two male paratypes (FSCB). 15. Gonopods, posterior view (posterior angiocoxal process on the left removed). 16. Colpocoxite distal part without posterior angiocoxal process, posterior view. 17. Colpocoxite distal part, posterior view. 18. Gonopods, posterior view. 19. Spikes. 20. Gonopods without posterior gonopod telopodites, lateral view. Abbreviations: a, angiocoxite; at, anterior gonopod telopodite; ax, apex; b, mesal blade; ba, frontal blade; cp, colpocoxite “pad’’; ms, spear-shaped structure; p, conical process; pp, posterior angiocoxal process; sg, subglobule. Scales: 10 μm (Fig. 19), 20 μm (Figs 16, 17), 100 μm (Figs 15, 18, 20)." figureDoi="http://doi.org/10.5281/zenodo.7413481" httpUri="https://zenodo.org/record/7413481/files/figure.png" pageId="1" pageNumber="186">Figs 15, 16</figureCitation>
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). Colpocoxites fused medially in its basal half; their distal parts curved laterally. Each colpocoxite entire, undivided. Angiocoxites with a subglobule (
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<emphasis id="B928EAEEFF9CFFDDFAA142B9EE2119EA" bold="true" box="[1303,1328,1225,1251]" pageId="1" pageNumber="186">sg</emphasis>
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) strongly protruding caudad in posterior view (
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<figureCitation id="13672A79FF9CFFDDFD88429DE9D2180E" box="[574,707,1261,1288]" captionStart="FIGURES 15–20" captionStartId="4.[151,264,1511,1535]" captionTargetBox="[152,1435,181,1487]" captionTargetId="figure-20@4.[152,1435,181,1487]" captionTargetPageId="4" captionText="FIGURES 15–20. Liaoninosoma marusiki sp. nov., two male paratypes (FSCB). 15. Gonopods, posterior view (posterior angiocoxal process on the left removed). 16. Colpocoxite distal part without posterior angiocoxal process, posterior view. 17. Colpocoxite distal part, posterior view. 18. Gonopods, posterior view. 19. Spikes. 20. Gonopods without posterior gonopod telopodites, lateral view. Abbreviations: a, angiocoxite; at, anterior gonopod telopodite; ax, apex; b, mesal blade; ba, frontal blade; cp, colpocoxite “pad’’; ms, spear-shaped structure; p, conical process; pp, posterior angiocoxal process; sg, subglobule. Scales: 10 μm (Fig. 19), 20 μm (Figs 16, 17), 100 μm (Figs 15, 18, 20)." figureDoi="http://doi.org/10.5281/zenodo.7413481" httpUri="https://zenodo.org/record/7413481/files/figure.png" pageId="1" pageNumber="186">Figs 15, 18</figureCitation>
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). Angiocoxites (
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<emphasis id="B928EAEEFF9CFFDDFCC8429DE89D180E" bold="true" box="[894,908,1261,1287]" pageId="1" pageNumber="186">a</emphasis>
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) in anterior view strongly elongated ventrally, centrally convex along their length, however, they do not form independent, separate processes, but are fused with to colpocoxites (
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<figureCitation id="13672A79FF9CFFDDFEE24345EAB81846" box="[340,425,1333,1359]" captionStart="FIGURES 21–25" captionStartId="5.[151,266,1386,1410]" captionTargetBox="[151,1435,181,1362]" captionTargetId="figure-15@5.[151,1435,181,1362]" captionTargetPageId="5" captionText="FIGURES 21–25. Liaoninosoma marusiki sp. nov., two male paratypes (FSCB). 21. Gonopods, anterior view (slightly turned to the right). 22. Gonopods, anterior view (slightly skewed backwards). 23. Anterior part of colpocoxite “pad”. 24. Posterior gonopod colpocoxites, anterior view (slightly skewed backwards). 25. Central part of coxosternum of anterior gonopods, anterior view. Abbreviations: a, angiocoxite; bb, blade. Scales: 10 μm (Fig. 23), 20 μm (Fig. 25), 100 μm (Figs 21, 22, 24)." figureDoi="http://doi.org/10.5281/zenodo.7413483" httpUri="https://zenodo.org/record/7413483/files/figure.png" pageId="1" pageNumber="186">Fig. 21</figureCitation>
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). Anterior angiocoxal processes absent. Posterior angiocoxal processes present. Posterior gonopod telopodites 2-segmented, setose, telopoditomere 1 with a thin stem, telopoditomere 2 of medium length.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3466577FF9CFFDDFF71430DEE441891" box="[199,1365,1405,1432]" pageId="1" pageNumber="186" type="etymology">
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<paragraph id="8BE336FCFF9CFFDDFF71430DEE441891" blockId="1.[151,1437,793,1467]" box="[199,1365,1405,1432]" pageId="1" pageNumber="186">
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<emphasis id="B928EAEEFF9CFFDDFF71430DEA5F189E" bold="true" box="[199,334,1405,1431]" pageId="1" pageNumber="186">Etymology.</emphasis>
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The generic name refers to the
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<typeStatus id="54E7885EFF9CFFDDFD07430EE9F01891" box="[689,737,1406,1432]" pageId="1" pageNumber="186">type</typeStatus>
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locality,
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<collectingRegion id="4998F81EFF9CFFDDFCFE430DEF0A189E" box="[840,1051,1405,1431]" country="China" name="Liaoning" pageId="1" pageNumber="186">Liaoning Province</collectingRegion>
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. The name is a neuter noun.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3466577FF9CFFDDFF7143D1E98118B2" box="[199,656,1441,1467]" pageId="1" pageNumber="186" type="distribution">
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<paragraph id="8BE336FCFF9CFFDDFF7143D1E98118B2" blockId="1.[151,1437,793,1467]" box="[199,656,1441,1467]" pageId="1" pageNumber="186">
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<emphasis id="B928EAEEFF9CFFDDFF7143D1EA7018B2" bold="true" box="[199,353,1441,1467]" pageId="1" pageNumber="186">Distribution.</emphasis>
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<collectingCountry id="F34B766CFF9CFFDDFED143D1EABF18B2" box="[359,430,1441,1467]" name="China" pageId="1" pageNumber="186">China</collectingCountry>
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:
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<collectingRegion id="4998F81EFF9CFFDDFE0F43D1E99D18B2" box="[441,652,1441,1467]" country="China" name="Liaoning" pageId="1" pageNumber="186">Liaoning Province</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |