433 lines
61 KiB
XML
433 lines
61 KiB
XML
<document ID-DOI="10.37520/aemnp.2023.001" ID-ISSN="1804-6487" ID-Zenodo-Dep="7887421" ID-ZooBank="3F2C90B1-6EA1-4B38-A218-C314D09F6E00" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683048986818" checkinUser="felipe" docAuthor="Yasunaga, Tomohide, Schwartz, Michael D. & Chérot, Frédéric" docDate="2023" docId="03F587DFFFEEE1290DBD6FCFFDEFF9C4" docLanguage="en" docName="ActaEntMusNatPra.63.1.1-55.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2023.001" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Diognetus minusculus Yasunaga & Schwartz & Chérot 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="36" masterDocId="FFCCFFA7FFF3E10A0E7E6F72FFBDFFF1" masterDocTitle="Revision of the plant bug genus Diognetus, with descriptions of thirteen new species from the Oriental and Eastern Palearctic Regions (Hemiptera: Heteroptera: Miridae)" masterLastPageNumber="55" masterPageNumber="1" pageNumber="30" updateTime="1683049705842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Revision of the plant bug genus Diognetus, with descriptions of thirteen new species from the Oriental and Eastern Palearctic Regions (Hemiptera: Heteroptera: Miridae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Yasunaga, Tomohide</mods:namePart>
|
||
<mods:affiliation>Research Associate, Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024, USA, c / o Nameshi</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Schwartz, Michael D.</mods:namePart>
|
||
<mods:affiliation>- 33, Nagasaki 852 - 8061, Japan; e-mail: yasunagat. amnh @ gmail. com</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Chérot, Frédéric</mods:namePart>
|
||
<mods:affiliation>Département de l’Etude du Milieu Naturel et Agricole, Service Public de Wallonie, Gembloux, BE- 5030, Belgium; e-mail: frederic. cherot @ spw. wallonie. be</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Acta Entomologica Musei Nationalis Pragae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="series">
|
||
<mods:title>Acta. Ent. Mus. Natl. Pragae</mods:title>
|
||
</mods:detail>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-03-12</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>63</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>55</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.37520/aemnp.2023.001</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.37520/aemnp.2023.001</mods:identifier>
|
||
<mods:identifier type="ISSN">1804-6487</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7887421</mods:identifier>
|
||
<mods:identifier type="ZooBank">3F2C90B1-6EA1-4B38-A218-C314D09F6E00</mods:identifier>
|
||
</mods:mods>
|
||
<treatment LSID="urn:lsid:plazi:treatment:03F587DFFFEEE1290DBD6FCFFDEFF9C4" httpUri="http://treatment.plazi.org/id/03F587DFFFEEE1290DBD6FCFFDEFF9C4" lastPageId="35" lastPageNumber="36" pageId="29" pageNumber="30">
|
||
<subSubSection box="[963,1302,189,212]" pageId="29" pageNumber="30" type="nomenclature">
|
||
<paragraph blockId="29.[963,1302,189,212]" box="[963,1302,189,212]" pageId="29" pageNumber="30">
|
||
<heading box="[963,1302,189,212]" centered="true" fontSize="10" level="2" pageId="29" pageNumber="30" reason="3">
|
||
<emphasis bold="true" box="[963,1302,189,212]" pageId="29" pageNumber="30">
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[963,1209,189,212]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="minusculus" status="sp. nov.">
|
||
<emphasis bold="true" box="[963,1209,189,212]" italics="true" pageId="29" pageNumber="30">Diognetus minusculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1216,1302,189,212]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[1040,1223,220,239]" pageId="29" pageNumber="30" type="description">
|
||
<paragraph blockId="29.[828,1437,220,467]" box="[1040,1223,220,239]" pageId="29" pageNumber="30">
|
||
(
|
||
<figureCitation box="[1046,1154,220,239]" captionStart="Fig" captionStartId="28.[150,181,2034,2053]" captionTargetBox="[166,1407,841,1996]" captionTargetId="figure-338@28.[148,1439,833,2009]" captionTargetPageId="28" captionText="Fig. 17. Habitus images of Diognetus spp. A−C − D. minusculus sp. nov., J (A) and ♀ (B−C); D−E − D. pilosus, J (D) and ♀ (E); F − D. puspae sp. nov., J. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7887473" httpUri="https://zenodo.org/record/7887473/files/figure.png" pageId="29" pageNumber="30">Figs 17A− C</figureCitation>
|
||
,
|
||
<figureCitation box="[1163,1186,220,239]" captionStart="Fig" captionStartId="29.[151,182,2029,2048]" captionTargetBox="[159,1411,611,1983]" captionTargetId="figure-16@29.[151,1437,601,2008]" captionTargetPageId="29" captionText="Fig. 18. Male (A–D) and female (E–G) genitalia of Diognetus minusculus sp. nov. A − left paramere and apex of pygophore; B − right paramere; C−D − vesica (endosoma); E − ovipositor (gonapophysis I); F − genital chamber; G − posterior wall. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7887475" httpUri="https://zenodo.org/record/7887475/files/figure.png" pageId="29" pageNumber="30">18</figureCitation>
|
||
,
|
||
<figureCitation box="[1195,1217,220,239]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="29" pageNumber="30">33</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="29" pageNumber="30" type="materials_examined">
|
||
<paragraph blockId="29.[828,1437,220,467]" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[828,964,261,280]" pageId="29" pageNumber="30">Type material.</emphasis>
|
||
<materialsCitation collectingDate="1990-08-21" collectionCode="T, ZRC, AMNH" country="Malaysia" elevation="1100" latitude="4.8625" location="Bukit Larut" longLatPrecision="7" longitude="100.8" municipality="Taiping" pageId="29" pageNumber="30" specimenCode="PBI 00380752" specimenCount="1" stateProvince="Perak" typeStatus="holotype">
|
||
<typeStatus box="[972,1062,261,280]" pageId="29" pageNumber="30">HOLOTYPE</typeStatus>
|
||
: J,
|
||
<emphasis bold="true" box="[1106,1302,261,280]" pageId="29" pageNumber="30">
|
||
<collectingCountry box="[1106,1224,261,280]" name="Malaysia" pageId="29" pageNumber="30">MALAYSIA</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1236,1295,261,280]" country="Malaysia" name="Perak" pageId="29" pageNumber="30">PERAK</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<collectingMunicipality box="[1310,1378,261,280]" pageId="29" pageNumber="30">Taiping</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:03F587DFFFEEE1290DBD6FCFFDEFF9C4:8E836012FFEEE1170B126E77FCD7FEC2" country="Malaysia" latitude="4.8625" longLatPrecision="7" longitude="100.8" municipality="Taiping" name="Bukit Larut" pageId="29" pageNumber="30" stateProvince="Perak">Bukit Larut</location>
|
||
(Maxwell Hill),
|
||
<geoCoordinate box="[1026,1087,288,307]" degrees="4.8625" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="5" value="4.8625">4.8625</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1098,1180,288,307]" degrees="100.8000" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="5" value="100.8">100.8000</geoCoordinate>
|
||
,
|
||
<quantity box="[1192,1261,288,307]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="m" value="1100.0">
|
||
<elevation box="[1192,1261,288,307]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="m" value="1100.0">1100 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod box="[1272,1433,288,307]" pageId="29" pageNumber="30">mercury light trap</collectingMethod>
|
||
,
|
||
<date box="[828,940,315,334]" pageId="29" pageNumber="30" value="1990-08-21">
|
||
<collectingDate box="[828,940,315,334]" pageId="29" pageNumber="30" value="1990-08-21">21 Aug 1990</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode box="[948,962,315,334]" name="Tavera, Department of Geology and Geophysics" pageId="29" pageNumber="30">T</collectionCode>
|
||
. Yasunaga (
|
||
<collectionCode box="[1062,1106,315,334]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="29" pageNumber="30">ZRC</collectionCode>
|
||
) (
|
||
<collectionCode box="[1123,1189,315,334]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="29" pageNumber="30">AMNH</collectionCode>
|
||
_
|
||
<specimenCode box="[1200,1327,315,334]" collectionCode="PBI" pageId="29" pageNumber="30">PBI 00380752</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus box="[1341,1430,315,334]" pageId="29" pageNumber="30">PARATYPES</typeStatus>
|
||
:
|
||
<emphasis bold="true" box="[828,949,341,360]" pageId="29" pageNumber="30">
|
||
<collectingCountry box="[828,946,341,360]" name="Malaysia" pageId="29" pageNumber="30">MALAYSIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Same data as for
|
||
<typeStatus box="[1102,1179,341,360]" pageId="29" pageNumber="30">holotype</typeStatus>
|
||
, 2 JJ
|
||
<specimenCount box="[1240,1287,335,360]" pageId="29" pageNumber="30" type="female">3 ♀♀</specimenCount>
|
||
(
|
||
<collectionCode box="[1298,1357,341,360]" pageId="29" pageNumber="30">DOAT</collectionCode>
|
||
,
|
||
<collectionCode box="[1365,1427,341,360]" pageId="29" pageNumber="30">TYCN</collectionCode>
|
||
); same data, except for date
|
||
<date box="[1055,1167,368,387]" pageId="29" pageNumber="30" value="1989-12-29">29 Dec 1989</date>
|
||
,
|
||
<specimenCount box="[1175,1207,362,387]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode box="[1218,1281,368,387]" pageId="29" pageNumber="30">TYCN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="29.[828,1437,220,467]" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[828,1436,395,414]" pageId="29" pageNumber="30">
|
||
Additional material examined:
|
||
<collectingCountry box="[1132,1259,395,414]" name="Indonesia" pageId="29" pageNumber="30">INDONESIA</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1272,1429,395,414]" country="Indonesia" name="Sumatera Utara" pageId="29" pageNumber="30">NORTH SUMATRA</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
Brastagi, North of Toba Lake,
|
||
<geoCoordinate box="[1098,1161,421,440]" degrees="3.18" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="3.18">3.18°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1171,1243,421,440]" degrees="98.50" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="98.5">98.50°E</geoCoordinate>
|
||
,
|
||
<quantity box="[1254,1328,421,440]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="m" value="1400.0">1,400 m</quantity>
|
||
, at FL light,
|
||
<date box="[828,950,448,467]" pageId="29" pageNumber="30" value="1989-12-05" valueMax="1989-12-09" valueMin="1989-12-05">5–9 Dec 1989</date>
|
||
,
|
||
<collectionCode box="[958,972,448,467]" name="Tavera, Department of Geology and Geophysics" pageId="29" pageNumber="30">T</collectionCode>
|
||
. Yasunaga,
|
||
<specimenCount box="[1072,1104,442,467]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode box="[1115,1178,448,467]" pageId="29" pageNumber="30">TYCN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="35" lastPageNumber="36" pageId="29" pageNumber="30" type="description">
|
||
<paragraph blockId="29.[828,1436,492,547]" lastBlockId="35.[151,761,190,1589]" lastPageId="35" lastPageNumber="36" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[828,971,492,515]" pageId="29" pageNumber="30">Description.</emphasis>
|
||
Body ovoid, small,
|
||
<quantity box="[1208,1345,492,515]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.2" metricValueMin="3.5" pageId="29" pageNumber="30" unit="mm" value="3.85" valueMax="4.2" valueMin="3.5">3.5−4.2 mm</quantity>
|
||
in total length; smallest species among known congeners. COLORATION: Body generally brown to dark brown (
|
||
<figureCitation captionStart="Fig" captionStartId="28.[150,181,2034,2053]" captionTargetBox="[166,1407,841,1996]" captionTargetId="figure-338@28.[148,1439,833,2009]" captionTargetPageId="28" captionText="Fig. 17. Habitus images of Diognetus spp. A−C − D. minusculus sp. nov., J (A) and ♀ (B−C); D−E − D. pilosus, J (D) and ♀ (E); F − D. puspae sp. nov., J. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7887473" httpUri="https://zenodo.org/record/7887473/files/figure.png" pageId="30" pageNumber="31">Figs 17A− B</figureCitation>
|
||
); dorsum partly speckled with darker maculae. Head pale brown; vertex and frons somewhat darkened medially; clypeus sometimes suffused with red. Antennae pale brown; segment I partly darkened; apical 1/4−1/3 of segment II, and whole segment III (except for yellowish extreme base) and segment IV dark brown. Labium shiny pale reddish-brown; apex of segment IV narrowly dark brown (
|
||
<figureCitation box="[238,339,446,469]" captionStart="Fig" captionStartId="28.[150,181,2034,2053]" captionTargetBox="[166,1407,841,1996]" captionTargetId="figure-338@28.[148,1439,833,2009]" captionTargetPageId="28" captionText="Fig. 17. Habitus images of Diognetus spp. A−C − D. minusculus sp. nov., J (A) and ♀ (B−C); D−E − D. pilosus, J (D) and ♀ (E); F − D. puspae sp. nov., J. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7887473" httpUri="https://zenodo.org/record/7887473/files/figure.png" pageId="30" pageNumber="31">Fig. 17C</figureCitation>
|
||
). Pronotum relatively shining, brown, more or less darkened posterolaterally, with creamy yellow posterior margin; calli with a pair of dark, shiny spots; mesoscutum and scutellum grayish brown; pleura, except for brown propleuron, shiny black; scent efferent system creamy yellow, weakly tinged with red in some specimens. Hemelytron irregularly mottled with dark maculae; cuneus reddish-brown, with paler basal 2/3; membrane smoky brown, with pale veins and several semitransparent spots. Coxae and legs whitish brown; apical half of metafemur brown, with two faint pale rings subapically; apices of tibiae and tarsomeres III darkened. Ventral surface of abdomen almost uniformly pale brown. SURFACE AND VESTITURE: As in generic diagnosis; scutellum finely and uniformly punctate, less rugose (
|
||
<figureCitation box="[528,623,190,213]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Fig. 33A</figureCitation>
|
||
); hemelytron matte; punctures on corium partly sparse and shallow. STRUCTURE: Vertex wider than an eye in dorsal view, with narrow basal transverse carina (
|
||
<figureCitation box="[545,644,286,309]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Fig. 33D</figureCitation>
|
||
). Antennal segment II obviously shorter than labium. Labium long, reaching apex of mesocoxa but not exceeding apical metacoxa (
|
||
<figureCitation box="[213,310,382,405]" captionStart="Fig" captionStartId="28.[150,181,2034,2053]" captionTargetBox="[166,1407,841,1996]" captionTargetId="figure-338@28.[148,1439,833,2009]" captionTargetPageId="28" captionText="Fig. 17. Habitus images of Diognetus spp. A−C − D. minusculus sp. nov., J (A) and ♀ (B−C); D−E − D. pilosus, J (D) and ♀ (E); F − D. puspae sp. nov., J. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7887473" httpUri="https://zenodo.org/record/7887473/files/figure.png" pageId="35" pageNumber="36">Figs 17C</figureCitation>
|
||
,
|
||
<figureCitation box="[319,367,382,405]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">33C</figureCitation>
|
||
). Pronotum and propleuron uniformly punctate (
|
||
<figureCitation box="[253,383,414,437]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Figs 33B− C</figureCitation>
|
||
); scutellum rather flat (
|
||
<figureCitation box="[621,746,414,437]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Figs 33B, E</figureCitation>
|
||
); metathoracic scent efferent system as in
|
||
<figureCitation box="[569,659,446,469]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">
|
||
Fig.
|
||
<specimenCount box="[616,659,446,469]" pageId="35" pageNumber="36" type="generic">33F</specimenCount>
|
||
</figureCitation>
|
||
. Pretarsal structure as in
|
||
<figureCitation box="[303,389,478,501]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Fig. 33I</figureCitation>
|
||
; parempodia rather long and broad. MALE GENITALIA (
|
||
<figureCitation box="[385,520,510,533]" captionStart="Fig" captionStartId="29.[151,182,2029,2048]" captionTargetBox="[159,1411,611,1983]" captionTargetId="figure-16@29.[151,1437,601,2008]" captionTargetPageId="29" captionText="Fig. 18. Male (A–D) and female (E–G) genitalia of Diognetus minusculus sp. nov. A − left paramere and apex of pygophore; B − right paramere; C−D − vesica (endosoma); E − ovipositor (gonapophysis I); F − genital chamber; G − posterior wall. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7887475" httpUri="https://zenodo.org/record/7887475/files/figure.png" pageId="35" pageNumber="36">Figs 18A− D</figureCitation>
|
||
,
|
||
<figureCitation box="[530,637,510,533]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">33G, J− K</figureCitation>
|
||
): Left paramere modified in shape, with triangular protuberance of sensory lobe and somewhat flattened hypophysis that is hooked at apex (
|
||
<figureCitation box="[325,427,606,629]" captionStart="Fig" captionStartId="30.[152,183,2031,2050]" captionTargetBox="[215,1423,577,1980]" captionTargetId="figure-13@30.[151,1437,577,2004]" captionTargetPageId="30" captionText="Fig.19. Habitus images of Diognetus spp.A−B − D. schuhorum sp. nov., holotypeJ, dorsal (A) and ventral (B) views; C−D − D. styrax sp. nov., holotype J; E–F – D. yamato nom. nov., J. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7887479" httpUri="https://zenodo.org/record/7887479/files/figure.png" pageId="35" pageNumber="36">Figs 19A</figureCitation>
|
||
,
|
||
<figureCitation box="[438,478,606,629]" captionStart="Fig" captionStartId="49.[151,182,1892,1911]" captionTargetBox="[151,1436,189,1855]" captionTargetId="figure-16@49.[151,1436,189,1860]" captionTargetPageId="49" captionText="Fig. 36. Scanning electron micrographs for Diognetus yamato Yasunaga nom. nov. from Shikoku, Japan (A−I), J (A−F) and ♀ (G−I), and immature forms of D. cheimon sp. nov. from Nagasaki, Japan (J−L), D. styrax sp. nov. from Nantou, Taiwan (M−N) and D. vernus sp. nov. from Nagasaki (O). A − anterior body, left lateral view; B − apex of labium with mandibular and maxillary stylets; C − pleura and scent efferent system, left lateral view; D − metatarsus; E, L − pretarsal structure of metaleg; F − apex of pygophore with parameres, left lateral view; G − posterior wall; H − dorsal structure and interramal lobe; I − apex of ovipositor (gonapophysis I); J − 5th instar exuvia, right lateral view; K − 5th instar thorax and abdomen, dorsal view; M − 5th instar nymph, anterior body; N − same, dorsal habitus; O − 5th instar exuvia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7887519" httpUri="https://zenodo.org/record/7887519/files/figure.png" pageId="35" pageNumber="36">36J</figureCitation>
|
||
); right paramere relatively elongate (
|
||
<figureCitation box="[259,363,638,661]" captionStart="Fig" captionStartId="29.[151,182,2029,2048]" captionTargetBox="[159,1411,611,1983]" captionTargetId="figure-16@29.[151,1437,601,2008]" captionTargetPageId="29" captionText="Fig. 18. Male (A–D) and female (E–G) genitalia of Diognetus minusculus sp. nov. A − left paramere and apex of pygophore; B − right paramere; C−D − vesica (endosoma); E − ovipositor (gonapophysis I); F − genital chamber; G − posterior wall. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7887475" httpUri="https://zenodo.org/record/7887475/files/figure.png" pageId="35" pageNumber="36">Figs 18B</figureCitation>
|
||
,
|
||
<figureCitation box="[377,418,638,661]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">33J</figureCitation>
|
||
). Vesica (
|
||
<figureCitation box="[526,663,638,661]" captionStart="Fig" captionStartId="29.[151,182,2029,2048]" captionTargetBox="[159,1411,611,1983]" captionTargetId="figure-16@29.[151,1437,601,2008]" captionTargetPageId="29" captionText="Fig. 18. Male (A–D) and female (E–G) genitalia of Diognetus minusculus sp. nov. A − left paramere and apex of pygophore; B − right paramere; C−D − vesica (endosoma); E − ovipositor (gonapophysis I); F − genital chamber; G − posterior wall. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7887475" httpUri="https://zenodo.org/record/7887475/files/figure.png" pageId="35" pageNumber="36">Figs 18C− D</figureCitation>
|
||
) heavily sclerotized, with a distinct spicule (
|
||
<figureCitation box="[544,648,670,693]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Fig. 33K</figureCitation>
|
||
); RS with two apical pointed processes; secondary gonopore small. FEMALE GENITALIA (
|
||
<figureCitation box="[411,537,734,757]" captionStart="Fig" captionStartId="29.[151,182,2029,2048]" captionTargetBox="[159,1411,611,1983]" captionTargetId="figure-16@29.[151,1437,601,2008]" captionTargetPageId="29" captionText="Fig. 18. Male (A–D) and female (E–G) genitalia of Diognetus minusculus sp. nov. A − left paramere and apex of pygophore; B − right paramere; C−D − vesica (endosoma); E − ovipositor (gonapophysis I); F − genital chamber; G − posterior wall. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7887475" httpUri="https://zenodo.org/record/7887475/files/figure.png" pageId="35" pageNumber="36">Figs 18F–G</figureCitation>
|
||
,
|
||
<figureCitation box="[546,625,734,757]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">33L− O</figureCitation>
|
||
): Sclerotized ring comparatively large (
|
||
<figureCitation box="[432,530,766,789]" captionStart="Fig" captionStartId="29.[151,182,2029,2048]" captionTargetBox="[159,1411,611,1983]" captionTargetId="figure-16@29.[151,1437,601,2008]" captionTargetPageId="29" captionText="Fig. 18. Male (A–D) and female (E–G) genitalia of Diognetus minusculus sp. nov. A − left paramere and apex of pygophore; B − right paramere; C−D − vesica (endosoma); E − ovipositor (gonapophysis I); F − genital chamber; G − posterior wall. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7887475" httpUri="https://zenodo.org/record/7887475/files/figure.png" pageId="35" pageNumber="36">Figs 18F</figureCitation>
|
||
,
|
||
<figureCitation box="[543,590,766,789]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">33L</figureCitation>
|
||
); posterior wall (
|
||
<figureCitation box="[159,265,798,821]" captionStart="Fig" captionStartId="29.[151,182,2029,2048]" captionTargetBox="[159,1411,611,1983]" captionTargetId="figure-16@29.[151,1437,601,2008]" captionTargetPageId="29" captionText="Fig. 18. Male (A–D) and female (E–G) genitalia of Diognetus minusculus sp. nov. A − left paramere and apex of pygophore; B − right paramere; C−D − vesica (endosoma); E − ovipositor (gonapophysis I); F − genital chamber; G − posterior wall. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7887475" httpUri="https://zenodo.org/record/7887475/files/figure.png" pageId="35" pageNumber="36">Figs 18G</figureCitation>
|
||
,
|
||
<figureCitation box="[280,366,798,821]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">33M− O</figureCitation>
|
||
) with moderately developed dorsal structure that is covered with scaly microstructures (
|
||
<figureCitation captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Fig. 33N</figureCitation>
|
||
); interramal lobe wide, semi-circular (
|
||
<figureCitation box="[605,707,862,885]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Fig. 33M</figureCitation>
|
||
), not spinulate but densely furnished with scaly microstructures (
|
||
<figureCitation box="[159,259,926,949]" captionStart="Fig" captionStartId="46.[151,182,1924,1943]" captionTargetBox="[151,1436,189,1882]" captionTargetId="figure-13@46.[151,1436,189,1892]" captionTargetPageId="46" captionText="Fig. 33. Scanning electron micrographs for Diognetus minusculus sp. nov. from Malaysia (except for Sumatra, ♀ on C), J (A−B, D−E, G−K) and ♀ (C−F, L−O). A − dorsal habitus; B–C − left lateral habitus; D − head and anterior pronotum, dorsal view; E − scutellum and adjacent structures, left lateral view; F − pleura and scent efferent system, left lateral view; G − apex of pygophore with parameres, left lateral view; H − mesotarsus (lower) and metatarsus; I − pretarsal structure of metaleg; J − apex of pygophore with parameres, dorsal view; K − vesica; L − genital chamber, dorsal view; M − posterior wall; N − dorsal structure; O − interramal lobe." figureDoi="http://doi.org/10.5281/zenodo.7887513" httpUri="https://zenodo.org/record/7887513/files/figure.png" pageId="35" pageNumber="36">Fig. 33O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7887475" ID-Zenodo-Dep="7887475" httpUri="https://zenodo.org/record/7887475/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,182,2029,2048]" targetBox="[159,1411,611,1983]" targetPageId="29" targetType="figure">
|
||
<paragraph blockId="29.[151,1436,2029,2075]" pageId="29" pageNumber="30">
|
||
Fig. 18. Male (A–D) and female (E–G) genitalia of
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[593,782,2029,2048]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="minusculus" status="sp. nov.">
|
||
<emphasis box="[593,782,2029,2048]" italics="true" pageId="29" pageNumber="30">Diognetus minusculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[788,855,2029,2048]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A − left paramere and apex of pygophore; B − right paramere; C− D − vesica (endosoma); E − ovipositor (gonapophysis I); F − genital chamber; G − posterior wall. Scale bars 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7887479" ID-Zenodo-Dep="7887479" httpUri="https://zenodo.org/record/7887479/files/figure.png" pageId="30" pageNumber="31" startId="30.[152,183,2031,2050]" targetBox="[215,1423,577,1980]" targetPageId="30" targetType="figure">
|
||
<paragraph blockId="30.[149,1437,2026,2078]" pageId="30" pageNumber="31">
|
||
Fig.19. Habitus images of
|
||
<taxonomicName box="[376,503,2031,2050]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[376,464,2031,2050]" italics="true" pageId="30" pageNumber="31">Diognetus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
A− B −
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[567,686,2031,2050]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="schuhorum" status="sp. nov.">
|
||
<emphasis box="[567,686,2031,2050]" italics="true" pageId="30" pageNumber="31">D. schuhorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[690,757,2031,2050]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype J, dorsal (A) and ventral (B) views; C− D −
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[1208,1284,2031,2050]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="styrax" status="sp. nov.">
|
||
<emphasis box="[1208,1284,2031,2050]" italics="true" pageId="30" pageNumber="31">D. styrax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1288,1356,2031,2050]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype J; E–F –
|
||
<taxonomicName authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[235,324,2058,2077]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="yamato" status="nom. nov.">
|
||
<emphasis box="[235,324,2058,2077]" italics="true" pageId="30" pageNumber="31">D. yamato</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[329,417,2058,2077]" pageId="30" pageNumber="31">nom. nov.</taxonomicNameLabel>
|
||
, J. Scale bars 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7887485" ID-Zenodo-Dep="7887485" httpUri="https://zenodo.org/record/7887485/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,182,1743,1762]" targetBox="[173,1431,221,1690]" targetPageId="31" targetType="figure">
|
||
<paragraph blockId="31.[151,1436,1743,1789]" pageId="31" pageNumber="32">
|
||
Fig. 20. Male genitalia of
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[370,522,1743,1762]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="puspae" status="sp. nov.">
|
||
<emphasis box="[370,522,1743,1762]" italics="true" pageId="31" pageNumber="32">Diognetus puspae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[527,594,1743,1762]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A− C) and
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[693,813,1743,1762]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="schuhorum" status="sp. nov.">
|
||
<emphasis box="[693,813,1743,1762]" italics="true" pageId="31" pageNumber="32">D. schuhorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[818,884,1743,1762]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(D− F).A, E − left paramere; B, E − right paramere; C, F − vesica (endosoma). Scale bars 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7887487" ID-Zenodo-Dep="7887487" httpUri="https://zenodo.org/record/7887487/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,182,1913,1932]" targetBox="[173,1411,207,1875]" targetPageId="32" targetType="figure">
|
||
<paragraph blockId="32.[151,1437,1913,1959]" pageId="32" pageNumber="33">
|
||
Fig. 21. Male genitalia of
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[376,522,1913,1932]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="styrax" status="sp. nov.">
|
||
<emphasis box="[376,522,1913,1932]" italics="true" pageId="32" pageNumber="33">Diognetus styrax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[528,595,1913,1932]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A− C),
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[667,751,1913,1932]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="vernus" status="sp. nov.">
|
||
<emphasis box="[667,751,1913,1932]" italics="true" pageId="32" pageNumber="33">D. vernus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[756,824,1913,1932]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(D− G) and
|
||
<taxonomicName authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[928,1018,1913,1932]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="yamato" status="nom. nov.">
|
||
<emphasis box="[928,1018,1913,1932]" italics="true" pageId="32" pageNumber="33">D. yamato</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1023,1110,1913,1932]" pageId="32" pageNumber="33">nom. nov.</taxonomicNameLabel>
|
||
(H− J). A, D, H − left paramere; B, E, I − right paramere; C, F− G, J − vesica (endosoma). Scale bars 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7887489" ID-Zenodo-Dep="7887489" httpUri="https://zenodo.org/record/7887489/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,182,1886,1905]" targetBox="[151,1436,190,1848]" targetPageId="33" targetType="figure">
|
||
<paragraph blockId="33.[151,1437,1886,1958]" pageId="33" pageNumber="34">
|
||
Fig. 22. Female genitalia of
|
||
<taxonomicName authority="(Poppius, 1914)" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[395,692,1886,1905]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pilosus">
|
||
<emphasis box="[395,549,1886,1905]" italics="true" pageId="33" pageNumber="34">Diognetus pilosus</emphasis>
|
||
(
|
||
<bibRefCitation author="POPPIUS B." box="[561,687,1886,1905]" pageId="33" pageNumber="34" pagination="337 - 398" refId="ref30902" refString="POPPIUS B. 1914: Zur Kenntnis der Indo-Australischen Lygus - Arten. Annales Historico-Naturales Musei Nationalis Hungarici 12: 337 - 398." type="journal article" year="1914">Poppius, 1914</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(A–B – 00419636 from Sumatra, C–D – from Malaysia),
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[1196,1275,1886,1905]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="styrax" status="sp. nov.">
|
||
<emphasis box="[1196,1275,1886,1905]" italics="true" pageId="33" pageNumber="34">D. styrax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1281,1349,1886,1905]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(E− F),
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vernus" status="sp. nov.">
|
||
<emphasis italics="true" pageId="33" pageNumber="34">D. vernus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[214,281,1913,1932]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(G− I) and
|
||
<taxonomicName authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[377,466,1913,1932]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="yamato" status="nom. nov.">
|
||
<emphasis box="[377,466,1913,1932]" italics="true" pageId="33" pageNumber="34">D. yamato</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[472,559,1913,1932]" pageId="33" pageNumber="34">nom. nov.</taxonomicNameLabel>
|
||
(J− L). A, C, H, K − genital chamber. B, D, E, G, J − posterior wall; F, I, L − ovipositor (gonapophysis I); Scale bars 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7887493" ID-Zenodo-Dep="7887493" httpUri="https://zenodo.org/record/7887493/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,182,1842,1861]" targetBox="[151,1436,189,1820]" targetPageId="34" targetType="figure">
|
||
<paragraph blockId="34.[151,1436,1842,1888]" pageId="34" pageNumber="35">
|
||
Fig. 23. Dorsal vestiture pattern for 4 Japanese
|
||
<taxonomicName box="[549,706,1842,1861]" pageId="34" pageNumber="35">
|
||
<emphasis box="[549,637,1842,1861]" italics="true" pageId="34" pageNumber="35">Diognetus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[713,811,1842,1861]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cheimon" status="sp. nov.">
|
||
<emphasis box="[713,811,1842,1861]" italics="true" pageId="34" pageNumber="35">D. cheimon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[815,883,1842,1861]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="(Yasunaga, 1994)" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" baseAuthorityName="Yasunaga" baseAuthorityYear="1994" box="[890,1152,1842,1861]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="insulanus">
|
||
<emphasis box="[890,997,1842,1861]" italics="true" pageId="34" pageNumber="35">D. insulanus</emphasis>
|
||
(
|
||
<bibRefCitation author="YASUNAGA T." box="[1007,1146,1842,1861]" pageId="34" pageNumber="35" pagination="115 - 131" refId="ref31383" refString="YASUNAGA T. 1994: Pinalitus Kelton (Heteroptera, Miridae) and its allied genera of Japan, with descriptions of new genera and species. Japanese Journal of Entomology 62: 115 - 131." type="journal article" year="1994">Yasunaga, 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[1160,1249,1842,1861]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="laureus" status="sp. nov">
|
||
<emphasis box="[1160,1249,1842,1861]" italics="true" pageId="34" pageNumber="35">D. laureus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1253,1314,1842,1861]" pageId="34" pageNumber="35" rank="species">sp. nov</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[1354,1436,1842,1861]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="vernus" status="sp. nov.">
|
||
<emphasis box="[1354,1436,1842,1861]" italics="true" pageId="34" pageNumber="35">D. vernus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[151,218,1869,1888]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="35.[151,761,190,1589]" box="[181,487,958,981]" pageId="35" pageNumber="36">
|
||
<emphasis bold="true" box="[181,349,958,981]" italics="true" pageId="35" pageNumber="36">Measurements.</emphasis>
|
||
See
|
||
<tableCitation box="[401,481,958,981]" captionStart="Table 1" captionStartId="12.[151,198,190,209]" captionText="Table 1. Twelve measurements (in mm) for Diognetus spp. Abbreviations: FM – femur, HEM – hemelytra, L – length, LB – labium, PRN – pronotum, TB – tibia, VTX – vertex (interocular space), W – width." pageId="35" pageNumber="36">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="35" pageNumber="36" type="diagnosis">
|
||
<paragraph blockId="35.[151,761,190,1589]" pageId="35" pageNumber="36">
|
||
<emphasis bold="true" box="[151,399,990,1013]" pageId="35" pageNumber="36">Differential diagnosis.</emphasis>
|
||
Recognized by its tiny size; short antennal segment II with darkened apical 1/4−1/3; generally whitish brown femora; and unique shape of male and female genitalia as described above. At first sight similar to
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[180,336,1118,1141]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="duwalorum" status="sp. nov.">
|
||
<emphasis box="[180,336,1118,1141]" italics="true" pageId="35" pageNumber="36">D. duwalorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[345,430,1118,1141]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<collectingCountry box="[445,514,1118,1141]" name="Nepal" pageId="35" pageNumber="36">Nepal</collectingCountry>
|
||
), from which this new species can be distinguished by the darker dorsum, dark apical part of antennal segment II, and shorter labium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="35" pageNumber="36" type="etymology">
|
||
<paragraph blockId="35.[151,761,190,1589]" pageId="35" pageNumber="36">
|
||
<emphasis bold="true" box="[151,278,1214,1237]" pageId="35" pageNumber="36">Etymology.</emphasis>
|
||
From Latin,
|
||
<taxonomicName authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[410,528,1214,1237]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="minusculus">
|
||
<emphasis box="[410,528,1214,1237]" italics="true" pageId="35" pageNumber="36">minusculus</emphasis>
|
||
</taxonomicName>
|
||
(very small), referring to its smallest size among known congeners; adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="35" pageNumber="36" type="biology_ecology">
|
||
<paragraph blockId="35.[151,761,190,1589]" pageId="35" pageNumber="36">
|
||
<emphasis bold="true" box="[151,240,1278,1301]" pageId="35" pageNumber="36">Biology.</emphasis>
|
||
Unknown; all available individuals were collected at UV (mercury) lights, facing to a tropical rain forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,761,1342,1365]" pageId="35" pageNumber="36" type="distribution">
|
||
<paragraph blockId="35.[151,761,190,1589]" box="[151,761,1342,1365]" pageId="35" pageNumber="36">
|
||
<emphasis bold="true" box="[151,292,1342,1365]" pageId="35" pageNumber="36">Distribution.</emphasis>
|
||
<collectingCountry box="[296,391,1342,1365]" name="Malaysia" pageId="35" pageNumber="36">Malaysia</collectingCountry>
|
||
(
|
||
<collectingRegion box="[402,466,1342,1365]" country="Malaysia" name="Perak" pageId="35" pageNumber="36">Perak</collectingRegion>
|
||
),
|
||
<collectingCountry box="[481,581,1342,1365]" name="Indonesia" pageId="35" pageNumber="36">Indonesia</collectingCountry>
|
||
(
|
||
<collectingRegion box="[593,750,1342,1365]" country="Indonesia" name="Sumatera Utara" pageId="35" pageNumber="36">North Sumatra</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="35" pageNumber="36" type="discussion">
|
||
<paragraph blockId="35.[151,761,190,1589]" pageId="35" pageNumber="36">
|
||
<emphasis bold="true" box="[151,281,1374,1397]" pageId="35" pageNumber="36">Comments.</emphasis>
|
||
Based on the shape of the male genitalia, this new species is most closely related to
|
||
<taxonomicName authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" baseAuthorityName="Horvath" baseAuthorityYear="1905" box="[544,677,1406,1429]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="flavigenis">
|
||
<emphasis box="[544,677,1406,1429]" italics="true" pageId="35" pageNumber="36">D. flavigenis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry box="[689,753,1406,1429]" name="Japan" pageId="35" pageNumber="36">Japan</collectingCountry>
|
||
) and
|
||
<taxonomicName authority="Yasunaga & Schwartz & Chérot, 2023" authorityName="Yasunaga & Schwartz & Chérot" authorityYear="2023" box="[192,294,1438,1461]" class="Insecta" family="Miridae" genus="Diognetus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="puspae" status="sp. nov.">
|
||
<emphasis box="[192,294,1438,1461]" italics="true" pageId="35" pageNumber="36">D. puspae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[298,376,1438,1461]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<collectingCountry box="[386,453,1438,1461]" name="Nepal" pageId="35" pageNumber="36">Nepal</collectingCountry>
|
||
), although the external appearance of each species is distinctive (e.g., body size, color pattern of femora). The distribution pattern of the three related species may imply faunal relationships between the Himalayas and eastern (and SE) Asia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |