834 lines
103 KiB
XML
834 lines
103 KiB
XML
<document id="3AA84992B19D1BCC81E24C678DF85D0B" ID="10.11646/zootaxa.4196.1.2" ID-DOI="10.11646/zootaxa.4196.1.2" ID-GBIF-Dataset="fff68f44-8f36-4ff7-a6c1-5dcf5f201358" ID-ISSN="1175-5326" ID-Zenodo-Dep="167667" ID-ZooBank="42261121-4F80-4D02-96DE-77AD1CD912C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479712559727" checkinUser="plazi" docAuthor="Lavrenchenko, Leonid A., Voyta, Leonid L. & Hutterer, Rainer" docDate="2016" docId="03F58782FFE3684BFF1EF97F4E47DCF0" docLanguage="en" docName="zootaxa.4196.1.2.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crocidura yaldeni Lavrenchenko, Voyta & Hutterer, 2016, sp. nov." docType="treatment" docUuid="F4813A6D-1B58-4E0F-B02C-FF4D9FF69F55" docUuidSource="ZooBank" docVersion="9" masterDocId="FFCCFFFAFFE9685BFF89FFAE4C5DDF1D" masterDocTitle="Diversity of shrews in Ethiopia, with the description of two new species of Crocidura (Mammalia: Lipotyphla: Soricidae)" updateTime="1698393968789" updateUser="plazi">
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<mods:mods id="2EAB4C362AC2680DAE87053CDAC0549D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1F23E62441A052601C60C2616309A645">
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<mods:title id="B364063E297AC2EBA62C4104D22A9E24">Diversity of shrews in Ethiopia, with the description of two new species of Crocidura (Mammalia: Lipotyphla: Soricidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="98C866381E757FDB724E71911E7FB7EB" type="personal">
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<mods:role id="55CC7E3DA2F048055351C17ADFCF1641">
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<mods:roleTerm id="DCA5D6B3F8508AD04524109F2DF311FC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="160CFB415B8D40FAD14E56B98A95DABD">Lavrenchenko, Leonid A.</mods:namePart>
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<mods:name id="3A32C69E8D1197EF55D5754E0C39F4DF" type="personal">
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<mods:role id="D3217AC8D2ADD7B2438FF310DBFE3AD1">
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<mods:roleTerm id="B0EF1DA3F1A829A46A1565A21AB7EA47">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EDA0DE279644ACA250A01F316D532FD9">Voyta, Leonid L.</mods:namePart>
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</mods:name>
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<mods:name id="5B8ADA993D2C91476C704D55C3B195FC" type="personal">
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<mods:role id="5E8E7C77FAABCF3EBE9417266852B673">
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<mods:roleTerm id="3DA0C3106AA66304043570870A8F8F90">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6F4541FDABA9E9832DB649034043E901">Hutterer, Rainer</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="41164CDD8695DD55423D74BEEF0C45EB">text</mods:typeOfResource>
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<mods:titleInfo id="6FF84AAF7D20752F3F79591A48652DC9">
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<mods:title id="863500D8963DE5193074B9B08DB58119">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B7FD24A91DFFF7C4CA5E36DBFD72BC0F">
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<mods:date id="3B2B88B4BA95872B04D6D00D15D8863A">2016</mods:date>
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<mods:detail id="C493A5FA2B16F2F980226324BEBCED0D" type="volume">
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<mods:number id="14C1824BE382EF83F5AD8936ED96853E">4196</mods:number>
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<mods:detail id="BC81A87F86C75575EECDCB05A77D59A3" type="issue">
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<mods:number id="8876AA432169FE38AA9E2DFE666E3194">1</mods:number>
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</mods:detail>
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<mods:classification id="F0242766D6F777539D1AA2C46A9C1B43">journal volume</mods:classification>
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<mods:identifier id="3A4222D82B859D415AB82EA12B9890A9" type="DOI">10.11646/zootaxa.4196.1.2</mods:identifier>
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<mods:identifier id="0807EDDFAD5E2E00077489D59244FA27" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6D30F693A8EC41C74F4D41BB8B79CEC0" type="Zenodo-Dep">167667</mods:identifier>
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<mods:identifier id="1E1C3C5E1568B1F5BE0D5D9FCA5EE191" type="ZooBank">42261121-4F80-4D02-96DE-77AD1CD912C1</mods:identifier>
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</mods:mods>
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<treatment id="03F58782FFE3684BFF1EF97F4E47DCF0" ID-DOI="http://doi.org/10.5281/zenodo.5614915" ID-GBIF-Taxon="124814331" ID-Zenodo-Dep="5614915" LSID="urn:lsid:zoobank.org:act:F4813A6D-1B58-4E0F-B02C-FF4D9FF69F55" httpUri="http://treatment.plazi.org/id/03F58782FFE3684BFF1EF97F4E47DCF0" lastPageId="16" pageId="10">
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<subSubSection id="C346651FFFE36851FF1EF97F4DB7D810" pageId="10" type="nomenclature">
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<paragraph id="8BE33694FFE36851FF1EF97F4D8AD9F1" blockId="10.[151,490,1745,1805]" box="[151,471,1745,1772]" pageId="10">
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<heading id="D0AB81F8FFE36851FF1EF97F4D8AD9F1" bold="true" box="[151,471,1745,1772]" fontSize="11" level="1" pageId="10" reason="1">
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<emphasis id="B928EA86FFE36851FF1EF97F4D8AD9F1" bold="true" box="[151,471,1745,1772]" pageId="10">
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<taxonomicName id="4C5C4D17FFE36851FF1EF97F4D2FD9F6" box="[151,370,1745,1771]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="10" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE36851FF1EF97F4D2FD9F6" bold="true" box="[151,370,1745,1771]" italics="true" pageId="10">Crocidura yaldeni</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21B57FDFFE36851FEF0F97C4D8AD9F1" box="[377,471,1746,1772]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE36851FF1EF95A4DB7D810" blockId="10.[151,490,1745,1805]" box="[151,490,1780,1805]" pageId="10">
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<figureCitation id="13672A11FFE36851FF1EF95A4CBED810" box="[151,227,1780,1805]" captionStart="FIGURE 7" captionStartId="11.[151,250,1729,1751]" captionTargetBox="[192,1398,373,1707]" captionTargetId="figure@11.[188,1398,373,1707]" captionTargetPageId="11" captionText="FIGURE 7. Principal components plots: A—for all specimens of Crocidura (1 st versus 2 nd axis); B—for five middle-sized species (1 st versus 3 rd axis). Abbreviations: Af = C. afeworkbekelei sp. nov.; Bl = C. baileyi; G = C. glassi; H = C. harenna; L = C. lucina; M = C. macmillani; P = C. parvipes; T = C. thalia; Y = Crocidura yaldeni sp. nov. Below upper plot size ranges for small-, middle-, and large-sized groups shown by condylo-incisive length (CI) along PC 1." httpUri="https://zenodo.org/record/167674/files/figure.png" pageId="10">Figs. 7</figureCitation>
|
||
A, 8A, 9A, 10;
|
||
<tableCitation id="C6DE032FFFE36851FE19F95A4DBBD810" box="[400,486,1780,1805]" captionStart="TABLE 2" captionStartId="10.[151,239,152,175]" captionTargetBox="[159,1229,266,1517]" captionTargetPageId="10" captionText="TABLE 2. Comparison of the external and cranial measurements of C. yaldeni sp. nov. and C. thalia. Measurements in millimeters are presented as the mean, standard error of mean, standard deviation, range and number of specimens in parentheses." httpUri="http://table.plazi.org/id/DF23661CFFE36851FF1EFF364D46DFF2" pageId="10" tableUuid="DF23661CFFE36851FF1EFF364D46DFF2">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346651FFFE36851FF1EF8924E20D88D" pageId="10" type="description">
|
||
<paragraph id="8BE33694FFE36851FF1EF8924E06D84F" blockId="10.[151,637,1852,1936]" box="[151,603,1852,1874]" pageId="10">
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||
<treatmentCitationGroup id="AB4C11BAFFE36851FF1EF8924E06D84F" box="[151,603,1852,1874]" pageId="10">
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<taxonomicName id="4C5C4D17FFE36851FF1EF8924D5DD84F" ID-CoL="3W2V" box="[151,256,1852,1874]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFE36851FF1EF8924D5DD84F" box="[151,256,1852,1874]" italics="true" pageId="10">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
sp. B:
|
||
<treatmentCitation id="0AFD1085FFE36851FECEF8924E0BD84F" author="Bannikova" box="[327,598,1852,1874]" page="56" pageId="10" year="2001">
|
||
<bibRefCitation id="EFCD4B65FFE36851FECEF8924E6DD84F" author="Bannikova" box="[327,560,1852,1874]" pageId="10" refString="Bannikova, A. A., Lavrenchenko, L. A., Lomov A. A. & Mednikov, B. M. (2001) Molecular diversity of some Crocidura species (Insectivora, Soricidae) from Ethiopia. In: Denis, C., Granjon, L. & Poulet, A. (Eds.), African Small Mammals. IRD Editions, Paris, pp. 55 - 64." type="book chapter" year="2001">
|
||
Bannikova
|
||
<emphasis id="B928EA86FFE36851FE32F8934DBAD84F" box="[443,487,1852,1874]" italics="true" pageId="10">et al</emphasis>
|
||
., 2001
|
||
</bibRefCitation>
|
||
: 56
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE36851FF1EF8F54E06D86C" blockId="10.[151,637,1852,1936]" box="[151,603,1883,1905]" pageId="10">
|
||
<treatmentCitationGroup id="AB4C11BAFFE36851FF1EF8F54E06D86C" box="[151,603,1883,1905]" pageId="10">
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||
<taxonomicName id="4C5C4D17FFE36851FF1EF8F54D5DD86C" ID-CoL="3W2V" box="[151,256,1883,1905]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFE36851FF1EF8F54D5DD86C" box="[151,256,1883,1905]" italics="true" pageId="10">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
sp. B:
|
||
<treatmentCitation id="0AFD1085FFE36851FECEF8F54E0BD86C" author="Bannikova" box="[327,598,1883,1905]" page="47" pageId="10" year="2005">
|
||
<bibRefCitation id="EFCD4B65FFE36851FECEF8F54E6DD86C" author="Bannikova" box="[327,560,1883,1905]" pageId="10" refString="Bannikova, A. A., Lavrenchenko, L. A. & Kramerov, D. A. (2005) Phylogenetic relationships between Afrotropical and Palaearctic Crocidura species inferred from Inter-SINE-PCR. Biochemical Systematics and Ecology, 33, 45 - 59. http: // dx. doi. org / 10.1016 / j. bse. 2004.05.014" type="journal article" year="2005">
|
||
Bannikova
|
||
<emphasis id="B928EA86FFE36851FE32F8F24DBAD86C" box="[443,487,1883,1905]" italics="true" pageId="10">et al</emphasis>
|
||
., 2005
|
||
</bibRefCitation>
|
||
: 47
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE36851FF1EF8D44E20D88D" blockId="10.[151,637,1852,1936]" box="[151,637,1914,1936]" pageId="10">
|
||
<treatmentCitationGroup id="AB4C11BAFFE36851FF1EF8D44E20D88D" box="[151,637,1914,1936]" pageId="10">
|
||
<taxonomicName id="4C5C4D17FFE36851FF1EF8D44D5DD88D" ID-CoL="3W2V" box="[151,256,1914,1936]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFE36851FF1EF8D44D5DD88D" box="[151,256,1914,1936]" italics="true" pageId="10">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
sp. B:
|
||
<treatmentCitation id="0AFD1085FFE36851FECEF8D44E24D88D" author="Lavrenchenko" box="[327,633,1914,1936]" page="57" pageId="10" year="2009">
|
||
<bibRefCitation id="EFCD4B65FFE36851FECEF8D44E0ED88D" author="Lavrenchenko" box="[327,595,1914,1936]" pageId="10" refString="Lavrenchenko, L. A., Bannikova, A. A. & Lebedev, V. S. (2009) Shrews (Crocidura spp.) endemic to Ethiopia: recent adaptive radiation of an ancient lineage. Doklady Biological Sciences, 424, 57 - 60. http: // dx. doi. org / 10.1134 / S 0012496609010177" type="journal article" year="2009">
|
||
Lavrenchenko
|
||
<emphasis id="B928EA86FFE36851FE57F8D54E56D88D" box="[478,523,1914,1936]" italics="true" pageId="10">et al</emphasis>
|
||
., 2009
|
||
</bibRefCitation>
|
||
: 57
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346651FFFE36850FF1EF81B4DD3DE5D" lastPageId="11" pageId="10" type="materials_examined">
|
||
<paragraph id="8BE33694FFE36851FF1EF81B4DA1D8EF" blockId="10.[151,1436,1973,2034]" pageId="10">
|
||
<emphasis id="B928EA86FFE36851FF1EF81B4D59D8D3" bold="true" box="[151,260,1973,1998]" pageId="10">Holotype</emphasis>
|
||
. ZMMU S-165342; adult male, dry skin and skull; collector's number 30; collected by L.A. Lavrenchenko on
|
||
<date id="FFE21054FFE36851FED2F8774DA4D8EF" box="[347,505,2009,2034]" pageId="10" value="1997-04-16">16 April 1997</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE26850FF4EFF3948DFDFAD" blockId="11.[151,1436,151,320]" box="[199,1154,151,176]" pageId="11">
|
||
<materialsCitation id="3B343CC9FFE26850FF4EFF3948DFDFAD" ID-GBIF-Occurrence="1341356637" box="[199,1154,151,176]" country="Ethiopia" elevation="1900" latitude="7.5666666" location="Beletta Forest" longLatPrecision="1303" longitude="36.516666" pageId="11" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B928EA86FFE26850FF4EFF394D04DFAD" bold="true" box="[199,345,151,176]" pageId="11">
|
||
<typeStatus id="54E78836FFE26850FF4EFF394D5DDFAD" box="[199,256,151,176]" pageId="11">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<location id="8E83604FFFE26850FEEFFF394E5BDFB2" LSID="urn:lsid:plazi:treatment:03F58782FFE3684BFF1EF97F4E47DCF0:8E83604FFFE26850FEEFFF394E5BDFB2" box="[358,518,151,176]" country="Ethiopia" latitude="7.5666666" longLatPrecision="1303" longitude="36.516666" name="Beletta Forest" pageId="11">Beletta Forest</location>
|
||
, south-western
|
||
<collectingCountry id="F34B7604FFE26850FD3CFF394F4ADFAD" box="[693,791,151,176]" name="Ethiopia" pageId="11">Ethiopia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE685053FFE26850FCABFF364F26DFAD" box="[802,891,152,176]" direction="north" orientation="latitude" pageId="11" precision="925" value="7.5666666">07º34'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685053FFE26850FC0EFF394FB6DFAD" box="[903,1003,151,176]" direction="east" orientation="longitude" pageId="11" precision="925" value="36.516666">036º31'E</geoCoordinate>
|
||
,
|
||
<elevation id="0071D1A7FFE26850FC7FFF3648DFDFAD" box="[1014,1154,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" unit="m" value="1900.0">
|
||
<quantity id="4CA49B71FFE26850FC7FFF364815DFB2" box="[1014,1096,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" unit="m" value="1900.0">1900 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE26850FF4EFF124DD3DE5D" blockId="11.[151,1436,151,320]" pageId="11">
|
||
<emphasis id="B928EA86FFE26850FF4EFF124D62DFC8" bold="true" box="[199,319,188,213]" pageId="11">Paratypes</emphasis>
|
||
. ZMMU S-165340 (adult female, dry skin and skull, collector's number 20); ZMMU S-165341 (adult male, dry skin and skull, collector's number 28); ZMMU S-165343 (adult male, dry skin and skull, collector's number 35;
|
||
<figureCitation id="13672A11FFE26850FE1FFEAA4D85DE01" box="[406,472,260,285]" captionStart="FIGURE 8" captionStartId="12.[151,250,1480,1502]" captionTargetBox="[188,1398,338,1457]" captionTargetId="figure@12.[188,1398,337,1459]" captionTargetPageId="12" captionText="FIGURE 8. Dorsal, ventral, and lateral views of the cranium, and medial and lateral views of the mandible of Crocidura yaldeni sp. nov. S- 165343 (A) and C. thalia S- 164858 (B). Abbreviations: a = outline of lateral profile of the braincase; pp = paraoccipital process. Scale 5 mm." httpUri="https://zenodo.org/record/167675/files/figure.png" pageId="11">Fig. 8</figureCitation>
|
||
A); all three specimens from the type locality collected by L.A. Lavrenchenko between
|
||
<date id="FFE21054FFE26850FF1EFE864DD4DE5D" box="[151,393,295,320]" pageId="11" value="1997-04-13" valueMax="1997-04-19" valueMin="1997-04-13">13 and 19 April 1997</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF23661CFFE26850FF1EF96F480DD82E" httpUri="https://zenodo.org/record/167674/files/figure.png" pageId="11" targetBox="[192,1398,373,1707]" targetPageId="11">
|
||
<paragraph id="8BE33694FFE26850FF1EF96F480DD82E" blockId="11.[151,1436,1729,1843]" pageId="11">
|
||
<emphasis id="B928EA86FFE26850FF1EF96F4D49D9CA" bold="true" box="[151,276,1729,1751]" pageId="11">FIGURE 7.</emphasis>
|
||
Principal components plots: A—for all specimens of
|
||
<taxonomicName id="4C5C4D17FFE26850FCCDF96F4FF0D9CA" box="[836,941,1729,1751]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFE26850FCCDF96F4FF0D9CA" box="[836,941,1729,1751]" italics="true" pageId="11">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
(1st versus 2nd axis); B—for five middle-sized species (1st versus 3rd axis). Abbreviations: Af =
|
||
<taxonomicName id="4C5C4D17FFE26850FD18F94E4F1AD9E8" box="[657,839,1759,1781]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="afeworkbekelei" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE26850FD18F94E4F1AD9E8" box="[657,839,1759,1781]" italics="true" pageId="11">C. afeworkbekelei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE26850FCC7F9714FC3D9E8" bold="true" box="[846,926,1759,1781]" pageId="11">
|
||
<taxonomicNameLabel id="A21B57FDFFE26850FCC7F9714FC3D9E8" box="[846,926,1759,1781]" pageId="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; Bl =
|
||
<taxonomicName id="4C5C4D17FFE26850FC56F94E481CD9E8" box="[991,1089,1759,1781]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="baileyi">
|
||
<emphasis id="B928EA86FFE26850FC56F94E481CD9E8" box="[991,1089,1759,1781]" italics="true" pageId="11">C. baileyi</emphasis>
|
||
</taxonomicName>
|
||
; G =
|
||
<taxonomicName id="4C5C4D17FFE26850FBF5F94E4889D9E8" box="[1148,1236,1759,1781]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFE26850FBF5F94E4889D9E8" box="[1148,1236,1759,1781]" italics="true" pageId="11">C. glassi</emphasis>
|
||
</taxonomicName>
|
||
; H =
|
||
<taxonomicName id="4C5C4D17FFE26850FA87F94E4923D9E8" box="[1294,1406,1759,1781]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="harenna">
|
||
<emphasis id="B928EA86FFE26850FA87F94E4923D9E8" box="[1294,1406,1759,1781]" italics="true" pageId="11">C. harenna</emphasis>
|
||
</taxonomicName>
|
||
; L =
|
||
<taxonomicName id="4C5C4D17FFE26850FF25F9514D57D809" box="[172,266,1790,1812]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="lucina">
|
||
<emphasis id="B928EA86FFE26850FF25F9514D57D809" box="[172,266,1790,1812]" italics="true" pageId="11">C. lucina</emphasis>
|
||
</taxonomicName>
|
||
; M =
|
||
<taxonomicName id="4C5C4D17FFE26850FEC2F9514D86D809" box="[331,475,1790,1812]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="macmillani">
|
||
<emphasis id="B928EA86FFE26850FEC2F9514D86D809" box="[331,475,1790,1812]" italics="true" pageId="11">C. macmillani</emphasis>
|
||
</taxonomicName>
|
||
; P =
|
||
<taxonomicName id="4C5C4D17FFE26850FD9DF9514ED4D809" box="[532,649,1791,1812]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="parvipes">
|
||
<emphasis id="B928EA86FFE26850FD9DF9514ED4D809" box="[532,649,1791,1812]" italics="true" pageId="11">C. parvipes</emphasis>
|
||
</taxonomicName>
|
||
; T =
|
||
<taxonomicName id="4C5C4D17FFE26850FD4AF9514F41D809" box="[707,796,1790,1812]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE26850FD4AF9514F41D809" box="[707,796,1790,1812]" italics="true" pageId="11">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
; Y =
|
||
<taxonomicName id="4C5C4D17FFE26850FCD0F950484ED809" box="[857,1043,1790,1812]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE26850FCD0F950484ED809" box="[857,1043,1790,1812]" italics="true" pageId="11">Crocidura yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE26850FB93F9504831D809" bold="true" box="[1050,1132,1790,1812]" pageId="11">
|
||
<taxonomicNameLabel id="A21B57FDFFE26850FB93F9504831D809" box="[1050,1132,1790,1812]" pageId="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Below upper plot size ranges for small-, middle-, and large-sized groups shown by condylo-incisive length (CI) along PC 1.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C346651FFFE26857FF4EF8F4485DDE01" lastPageId="12" pageId="11" type="diagnosis">
|
||
<paragraph id="8BE33694FFE26857FF4EF8F4485DDE01" blockId="11.[151,1437,1882,2015]" lastBlockId="12.[151,1436,151,285]" lastPageId="12" pageId="11">
|
||
<emphasis id="B928EA86FFE26850FF4EF8F44D67D86E" bold="true" box="[199,314,1882,1907]" pageId="11">Diagnosis</emphasis>
|
||
. A large-sized, greyish-brown
|
||
<taxonomicName id="4C5C4D17FFE26850FD1AF8F44F5AD86E" box="[659,775,1882,1907]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFE26850FD1AF8F44F5AD86E" box="[659,775,1882,1907]" italics="true" pageId="11">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
similar in external measurements (HB, TL) to
|
||
<taxonomicName id="4C5C4D17FFE26850FA99F8F5492FD86E" box="[1296,1394,1882,1907]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="11" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE26850FA99F8F5492FD86E" box="[1296,1394,1882,1907]" italics="true" pageId="11">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
but slightly larger, on average, in cranial size (
|
||
<tableCitation id="C6DE032FFFE26850FDF9F8D04E95D88A" box="[624,712,1918,1943]" captionStart="TABLE 2" captionStartId="10.[151,239,152,175]" captionTargetBox="[159,1229,266,1517]" captionTargetPageId="10" captionText="TABLE 2. Comparison of the external and cranial measurements of C. yaldeni sp. nov. and C. thalia. Measurements in millimeters are presented as the mean, standard error of mean, standard deviation, range and number of specimens in parentheses." httpUri="http://table.plazi.org/id/DF23661CFFE36851FF1EFF364D46DFF2" pageId="11" tableUuid="DF23661CFFE36851FF1EFF364D46DFF2">Table 2</tableCitation>
|
||
). Distinguished from the latter species by the following features: moderately longer hindfoot and claws; relatively uniformly colored tail; bristle hairs of tail bicolored and longer; skull large; nasal aperture wide; dorsal profile of skull looks concave at the midpoint and, slightly convex above the upper tooth row (
|
||
<figureCitation id="13672A11FFE56857FED6FF394DFBDFAD" box="[351,422,151,176]" captionStart="FIGURE 8" captionStartId="12.[151,250,1480,1502]" captionTargetBox="[188,1398,338,1457]" captionTargetId="figure@12.[188,1398,337,1459]" captionTargetPageId="12" captionText="FIGURE 8. Dorsal, ventral, and lateral views of the cranium, and medial and lateral views of the mandible of Crocidura yaldeni sp. nov. S- 165343 (A) and C. thalia S- 164858 (B). Abbreviations: a = outline of lateral profile of the braincase; pp = paraoccipital process. Scale 5 mm." httpUri="https://zenodo.org/record/167675/files/figure.png" pageId="12">Fig. 8</figureCitation>
|
||
, A); upper first incisor with a hook-like apex; lingual outline of the second upper molar with a distinct symmetric incision (
|
||
<figureCitation id="13672A11FFE56857FDA5FF124E2EDFC9" box="[556,627,188,213]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="12">Fig. 9</figureCitation>
|
||
, A1-a); second and third upper molar in firmly contact (
|
||
<figureCitation id="13672A11FFE56857FB74FF12491EDFC9" box="[1277,1347,188,213]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="12">Fig. 9</figureCitation>
|
||
, A1-b); occlusal outline of the third upper molar more undulated (
|
||
<figureCitation id="13672A11FFE56857FC93FF714F03DFE5" box="[794,862,223,248]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="12">Fig. 9</figureCitation>
|
||
, A1-c); hypoconulids of the lower molars visible in medial view (
|
||
<figureCitation id="13672A11FFE56857FEA6FEAA4D30DE01" box="[303,365,260,285]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="12">Fig 9</figureCitation>
|
||
, A2-d); postcingulid of m3 forming a notch (
|
||
<figureCitation id="13672A11FFE56857FCE1FEAA4FF1DE01" box="[872,940,260,285]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="12">Fig. 9</figureCitation>
|
||
, A3-e).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF23661CFFE56857FF1EFA664DAAD906" httpUri="https://zenodo.org/record/167675/files/figure.png" pageId="12" targetBox="[188,1398,338,1457]" targetPageId="12">
|
||
<paragraph id="8BE33694FFE56857FF1EFA664DAAD906" blockId="12.[151,1436,1480,1563]" pageId="12">
|
||
<emphasis id="B928EA86FFE56857FF1EFA664D4BDAC3" bold="true" box="[151,278,1480,1502]" pageId="12">FIGURE 8.</emphasis>
|
||
Dorsal, ventral, and lateral views of the cranium, and medial and lateral views of the mandible of
|
||
<taxonomicName id="4C5C4D17FFE56857FABAFA664CBFDAE0" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="12" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE56857FABAFA664CBFDAE0" italics="true" pageId="12">Crocidura yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE56857FF60FA494D67DAE0" bold="true" box="[233,314,1511,1533]" pageId="12">
|
||
<taxonomicNameLabel id="A21B57FDFFE56857FF60FA494D67DAE0" box="[233,314,1511,1533]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
S-165343 (A) and
|
||
<taxonomicName id="4C5C4D17FFE56857FD8BFA464E06DAE0" box="[514,603,1511,1533]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="12" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE56857FD8BFA464E06DAE0" box="[514,603,1511,1533]" italics="true" pageId="12">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
S-164858 (B). Abbreviations: a = outline of lateral profile of the braincase; pp = paraoccipital process. Scale 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C346651FFFE5684BFF4EF9E64E1EDE01" lastPageId="16" pageId="12" type="description">
|
||
<paragraph id="8BE33694FFE56857FF4EF9E64D54D824" blockId="12.[151,1437,1608,2029]" pageId="12">
|
||
<emphasis id="B928EA86FFE56857FF4EF9E64D0CD97C" bold="true" box="[199,337,1608,1633]" pageId="12">Description</emphasis>
|
||
. Large-sized
|
||
<taxonomicName id="4C5C4D17FFE56857FE63F9E64E03D97C" box="[490,606,1608,1633]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFE56857FE63F9E64E03D97C" box="[490,606,1608,1633]" italics="true" pageId="12">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
(head and body length 84.0–
|
||
<quantity id="4CA49B71FFE56857FC2FF9E64850D97D" box="[934,1037,1608,1633]" metricMagnitude="-2" metricUnit="m" metricValue="9.95" pageId="12" unit="mm" value="99.5">99.5 mm</quantity>
|
||
; here regarded as large-sized shrew in comparison to examined species) with a moderately long tail, ranging between 62.2–77.6% of head and body length. Dorsal pelage grey-brown with pale ochre shades (differs from
|
||
<taxonomicName id="4C5C4D17FFE56857FC35F93F4843D9B4" box="[956,1054,1680,1705]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="12" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE56857FC35F93F4843D9B4" box="[956,1054,1680,1705]" italics="true" pageId="12">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
in slightly less intensive shades); dorsal hairs grey at base, brown at tip. Ventral pelage blackish-grey with pale ochre wash; ventral hairs dark grey at base, pale-yellowish at tip. Dorsal surface of fore- and hindfoot brownish. Tail uniformly colored, dark grey-brown above and brown below. Bristle hairs are long, dark-grey at base, pale-grey at tip, and present along the full length of the tail.
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE56856FF4EF8EA49DDDFE5" blockId="12.[151,1437,1608,2029]" lastBlockId="13.[151,1437,151,248]" lastPageId="13" pageId="12">
|
||
Skull (
|
||
<figureCitation id="13672A11FFE56857FE9AF8EA4D04D840" box="[275,345,1860,1885]" captionStart="FIGURE 8" captionStartId="12.[151,250,1480,1502]" captionTargetBox="[188,1398,338,1457]" captionTargetId="figure@12.[188,1398,337,1459]" captionTargetPageId="12" captionText="FIGURE 8. Dorsal, ventral, and lateral views of the cranium, and medial and lateral views of the mandible of Crocidura yaldeni sp. nov. S- 165343 (A) and C. thalia S- 164858 (B). Abbreviations: a = outline of lateral profile of the braincase; pp = paraoccipital process. Scale 5 mm." httpUri="https://zenodo.org/record/167675/files/figure.png" pageId="12">Fig. 8</figureCitation>
|
||
, A) with a long rostrum and wide braincase. The rostral part is distinctly inflated (similar to
|
||
<taxonomicName id="4C5C4D17FFE56857FA0BF8EB4C84D89C" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="12" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE56857FA0BF8EB4C84D89C" italics="true" pageId="12">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
and differs from others Ethiopian endemic and Afromontane-Afroalpine species). The nasal aperture is wide; the posterior margin of the aperture has no medial tip. The dorsal outline of the orbital parts is slightly flexed. The lateral profile of the braincase is slightly angulated (
|
||
<figureCitation id="13672A11FFE56857FC8AF81E4F11D8D4" box="[771,844,1968,1993]" captionStart="FIGURE 8" captionStartId="12.[151,250,1480,1502]" captionTargetBox="[188,1398,338,1457]" captionTargetId="figure@12.[188,1398,337,1459]" captionTargetPageId="12" captionText="FIGURE 8. Dorsal, ventral, and lateral views of the cranium, and medial and lateral views of the mandible of Crocidura yaldeni sp. nov. S- 165343 (A) and C. thalia S- 164858 (B). Abbreviations: a = outline of lateral profile of the braincase; pp = paraoccipital process. Scale 5 mm." httpUri="https://zenodo.org/record/167675/files/figure.png" pageId="12">Fig. 8</figureCitation>
|
||
, A-a). The nuchal crests are well-developed; their postero-lateral ends are prolonged and joint with noticeable paraoccipital processes (
|
||
<figureCitation id="13672A11FFE56857FBE2F87A48EFD8F0" box="[1131,1202,2004,2029]" captionStart="FIGURE 8" captionStartId="12.[151,250,1480,1502]" captionTargetBox="[188,1398,338,1457]" captionTargetId="figure@12.[188,1398,337,1459]" captionTargetPageId="12" captionText="FIGURE 8. Dorsal, ventral, and lateral views of the cranium, and medial and lateral views of the mandible of Crocidura yaldeni sp. nov. S- 165343 (A) and C. thalia S- 164858 (B). Abbreviations: a = outline of lateral profile of the braincase; pp = paraoccipital process. Scale 5 mm." httpUri="https://zenodo.org/record/167675/files/figure.png" pageId="12">Fig. 8</figureCitation>
|
||
, A-pp). The sagittal crest is faint (in adult specimens); the temporal line is clearly recognizable. Mandible with broad body; its lower margin running in a shallow convex inward curve; body achieving its maximum depth below p4–m3. Coronoid spicule massive. External temporal fossa well-developed and wide. Angular process with hooked tip (
|
||
<figureCitation id="13672A11FFE46856FA82FF714912DFE5" box="[1291,1359,223,248]" captionStart="FIGURE 8" captionStartId="12.[151,250,1480,1502]" captionTargetBox="[188,1398,338,1457]" captionTargetId="figure@12.[188,1398,337,1459]" captionTargetPageId="12" captionText="FIGURE 8. Dorsal, ventral, and lateral views of the cranium, and medial and lateral views of the mandible of Crocidura yaldeni sp. nov. S- 165343 (A) and C. thalia S- 164858 (B). Abbreviations: a = outline of lateral profile of the braincase; pp = paraoccipital process. Scale 5 mm." httpUri="https://zenodo.org/record/167675/files/figure.png" pageId="13">Fig. 8</figureCitation>
|
||
, A).
|
||
</paragraph>
|
||
<caption id="DF23661CFFE46856FF1EF94D4C9BD8AC" httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="13" targetBox="[188,1398,302,1739]" targetPageId="13">
|
||
<paragraph id="8BE33694FFE46856FF1EF94D4C9BD8AC" blockId="13.[151,1436,1763,1969]" pageId="13">
|
||
<emphasis id="B928EA86FFE46856FF1EF94D4D49D9E4" bold="true" box="[151,276,1763,1785]" pageId="13">FIGURE 9.</emphasis>
|
||
Specific features of
|
||
<taxonomicName id="4C5C4D17FFE46856FE65F94D4EF5D9E4" box="[492,680,1763,1785]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="13" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE46856FE65F94D4EF5D9E4" box="[492,680,1763,1785]" italics="true" pageId="13">Crocidura yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE46856FD26F94D4F5CD9E4" bold="true" box="[687,769,1763,1785]" pageId="13">
|
||
<taxonomicNameLabel id="A21B57FDFFE46856FD26F94D4F5CD9E4" box="[687,769,1763,1785]" pageId="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
S-165343 (A1–A4) and
|
||
<taxonomicName id="4C5C4D17FFE46856FB8DF94D4803D9E4" box="[1028,1118,1763,1785]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="13" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE46856FB8DF94D4803D9E4" box="[1028,1118,1763,1785]" italics="true" pageId="13">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
S-164858 (B1–B4). A1, B1— occlusal view of the M2 and M3; A2, B2—lingual side of the m1; A3, B3—occlusal view of the m3; A4, B4—lateral view of the P4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of
|
||
<taxonomicName id="4C5C4D17FFE46856FBFBF88F4887D82B" box="[1138,1242,1824,1846]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="13" phylum="Chordata" rank="species" species="yaldeni">
|
||
<emphasis id="B928EA86FFE46856FBFBF88F4887D82B" box="[1138,1242,1824,1846]" italics="true" pageId="13">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
: a—the symmetric and distinctly curved incision of the M2; b—postero-lingual part of the M2 in contact with M3; c—the shape of the outline of the posterior edge of M3; d—the hypoconulid shoulder of the m1 in medial view; e—postcingulid of m3 forming a notch; fthe undulated upper margin of P4 in lateral view; g—three-quarters of the crown base of P4 surrounded by a cingulum. Scale 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF23661CFFE76855FF1EFCF14EF0DC89" httpUri="https://zenodo.org/record/167677/files/figure.png" pageId="14" targetBox="[189,1398,194,841]" targetPageId="14">
|
||
<paragraph id="8BE33694FFE76855FF1EFCF14EF0DC89" blockId="14.[151,1436,863,916]" pageId="14">
|
||
<emphasis id="B928EA86FFE76855FF1EFCF14D7DDC68" bold="true" box="[151,288,863,885]" pageId="14">FIGURE 10.</emphasis>
|
||
Principal components plot for two large-sized species and
|
||
<taxonomicName id="4C5C4D17FFE76855FCF3FCCE4F8EDC68" box="[890,979,863,885]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFE76855FCF3FCCE4F8EDC68" box="[890,979,863,885]" italics="true" pageId="14">C. glassi</emphasis>
|
||
</taxonomicName>
|
||
(1st versus 2nd axis). Abbreviations: G =
|
||
<taxonomicName id="4C5C4D17FFE76855FA0DFCCE4C8FDC89" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFE76855FA0DFCCE4C8FDC89" italics="true" pageId="14">C. glassi</emphasis>
|
||
</taxonomicName>
|
||
; T =
|
||
<taxonomicName id="4C5C4D17FFE76855FE83FCD14D3FDC89" box="[266,354,894,916]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE76855FE83FCD14D3FDC89" box="[266,354,894,916]" italics="true" pageId="14">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
; Y =
|
||
<taxonomicName id="4C5C4D17FFE76855FE14FCD04E0ADC89" box="[413,599,894,916]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE76855FE14FCD04E0ADC89" box="[413,599,894,916]" italics="true" pageId="14">Crocidura yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21B57FDFFE76855FDD5FCD04EF0DC89" box="[604,685,894,916]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE33694FFE76855FF4EFC154D65DA99" blockId="14.[151,1437,955,2024]" pageId="14">
|
||
First upper incisor (
|
||
<date id="FFE21054FFE76855FE24FC124D9EDCC9" box="[429,451,956,980]" pageId="14">I1</date>
|
||
) is robust; the apex (partly worn) is comparably long and hooked. The talon is welldeveloped, with distinct lingual ridge (lingual cingulum). The posterior ridge of the talon apex is stronglydeveloped. Lateral cingulum is weak and extending dorsally about 3/4 of
|
||
<date id="FFE21054FFE76855FC50FBAA4FADDB01" box="[985,1008,1028,1052]" pageId="14">I1</date>
|
||
base height, with a faint bulge on the inferior part. Upper antemolars (A1–A3) are well-spaced, with a weak buccal cingulum and postero-buccal small cuspules. The upper first antemolar (A1) is very large; basal prominence of the anterior ridge is absent. The upper second antemolar (A2) is smallest, approximately 1/3 the height of A1, and 3/4 the height of A3. The upper third antemolar (A3) with straight buccal outline; the posterior edge is not strongly curved and connected with P4. The parastyle of the fourth upper premolar (P4) is well defined (tip partly worn); the upper margin (lateral view) is undulated and distinctly concave above parastyle and main cusp (
|
||
<figureCitation id="13672A11FFE76855FCF0FB754FE3DBE9" box="[889,958,1243,1268]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="14">Fig. 9</figureCitation>
|
||
, A4-f); three-quarters of the crown base is surrounded by a cingulum (
|
||
<figureCitation id="13672A11FFE76855FE58FB514E4ADA05" box="[465,535,1279,1304]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="14">Fig. 9</figureCitation>
|
||
, A4-g). The upper molars (M1–M2) are comparably wide. The lingual edge of the upper second molar (M2) has a well-developed and symmetrically curved incision (
|
||
<figureCitation id="13672A11FFE76855FB0BFA8D4894DA21" box="[1154,1225,1315,1340]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="14">Fig. 9</figureCitation>
|
||
, A1-a). The upper third molar (M3) is relatively long and wide, in close contact with M2 (
|
||
<figureCitation id="13672A11FFE76855FC4FFAE94856DA7D" box="[966,1035,1351,1376]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="14">Fig. 9</figureCitation>
|
||
, A1-b), posterior edge is undulated (
|
||
<figureCitation id="13672A11FFE76855FF16FAC54CB9DA99" box="[159,228,1387,1412]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="14">Fig. 9</figureCitation>
|
||
, A1-c).
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE76855FF4EFA214FE1D99D" blockId="14.[151,1437,955,2024]" pageId="14">
|
||
The first lower incisor (
|
||
<date id="FFE21054FFE76855FE57FA214DAFDAB5" box="[478,498,1423,1448]" pageId="14">i1</date>
|
||
) is wide at base, with a blunt tip; the tooth is partly worn in all specimens, and therefore the denticulation of the cutting ridge is unknown. The second incisor (a1) is elongated, approximately 1/3 of the lower border of the tooth is in contact with
|
||
<date id="FFE21054FFE76855FD4EFA794E82DAED" box="[711,735,1495,1520]" pageId="14">i1</date>
|
||
; its posterior border is considerably (about 1/4) overlapped by the fourth lower premolar (p4); the postero-lateral ridge has a bulge. The basal prominence on the anterior ridge of p4 is present but indistinct, the lingual cusp (metaconid) is small. The hypoconulids of the lower molars (m1-m3) are well-developed and visible from the medial side of the teeth (
|
||
<figureCitation id="13672A11FFE76855FCFBF9ED4FEAD941" box="[882,951,1603,1628]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="14">Fig. 9</figureCitation>
|
||
, A2-d). The postcingulid of the lower third molar (m3) forms a visible notch buccal to the entoconid (
|
||
<figureCitation id="13672A11FFE76855FCAAF9C94F3AD99D" box="[803,871,1639,1664]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="14">Fig. 9</figureCitation>
|
||
, A3-e).
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE76855FF4EF9254D22D9F1" blockId="14.[151,1437,955,2024]" pageId="14">
|
||
The chromosomal set of
|
||
<taxonomicName id="4C5C4D17FFE76855FE6AF9224E09D9B9" box="[483,596,1675,1700]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE76855FE6AF9224E09D9B9" box="[483,596,1675,1700]" italics="true" pageId="14">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE76855FDD5F9254EEAD9B9" bold="true" box="[604,695,1675,1700]" pageId="14">
|
||
<taxonomicNameLabel id="A21B57FDFFE76855FDD5F9254EEAD9B9" box="[604,695,1675,1700]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(2n = 36, NFa = 52) comprises 4 pairs of metacentric, 5 pairs of subtelocentric and 8 pairs of acrocentric autosomes. The X-chromosome is large metacentric; the Y-chromosome is small subtelocentric.
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE76855FF4EF9594D67D845" blockId="14.[151,1437,955,2024]" pageId="14">
|
||
<emphasis id="B928EA86FFE76855FF4EF9594D68D80D" bold="true" box="[199,309,1783,1808]" pageId="14">Variation</emphasis>
|
||
. Specimens of the
|
||
<typeStatus id="54E78836FFE76855FD98F9564E1CD80D" box="[529,577,1784,1808]" pageId="14">type</typeStatus>
|
||
series are uniformly colored. In external and skull measurements S-165340 (female) is slightly smaller (HB = 84.0 mm, vs. 94.0–
|
||
<quantity id="4CA49B71FFE76855FD73F8B54F03D82E" box="[762,862,1819,1844]" metricMagnitude="-2" metricUnit="m" metricValue="9.95" pageId="14" unit="mm" value="99.5">99.5 mm</quantity>
|
||
in males). The qualitative characters are relatively homogeneous.
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE76854FF4EF8CD4FB6DFE5" blockId="14.[151,1437,955,2024]" lastBlockId="15.[151,1437,151,680]" lastPageId="15" pageId="14">
|
||
<emphasis id="B928EA86FFE76855FF4EF8CD4D38D861" bold="true" box="[199,357,1891,1916]" pageId="14">Comparisons</emphasis>
|
||
. This is a large-sized shrew, comparable in external size (HB, TL) to
|
||
<taxonomicName id="4C5C4D17FFE76855FB13F8CA48A3D861" box="[1178,1278,1891,1916]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE76855FB13F8CA48A3D861" box="[1178,1278,1891,1916]" italics="true" pageId="14">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
, but slightly larger in cranial size (
|
||
<tableCitation id="C6DE032FFFE76855FE1EF8294DAED8BD" box="[407,499,1927,1952]" captionStart="TABLE 2" captionStartId="10.[151,239,152,175]" captionTargetBox="[159,1229,266,1517]" captionTargetPageId="10" captionText="TABLE 2. Comparison of the external and cranial measurements of C. yaldeni sp. nov. and C. thalia. Measurements in millimeters are presented as the mean, standard error of mean, standard deviation, range and number of specimens in parentheses." httpUri="http://table.plazi.org/id/DF23661CFFE36851FF1EFF364D46DFF2" pageId="14" tableUuid="DF23661CFFE36851FF1EFF364D46DFF2">Table 2</tableCitation>
|
||
). It is substantially larger than the following Ethiopian Afromontane–Afroalpine species:
|
||
<emphasis id="B928EA86FFE76855FF7FF8024DC2D8D9" box="[246,415,1963,1988]" italics="true" pageId="14">
|
||
<taxonomicName id="4C5C4D17FFE76855FF7FF8024DC6D8D9" box="[246,411,1963,1988]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="bottegoides">C. bottegoides</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="4C5C4D17FFE76855FE2FF8024E66D8D9" box="[422,571,1963,1988]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="bottegi">
|
||
<emphasis id="B928EA86FFE76855FE2FF8024D9DD8DE" box="[422,448,1964,1987]" italics="true" pageId="14">C.</emphasis>
|
||
cf.
|
||
<emphasis id="B928EA86FFE76855FE62F8054E66D8D9" box="[491,571,1963,1988]" italics="true" pageId="14">bottegi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE76855FDB2F8054E62D8D9" box="[571,575,1963,1988]" italics="true" pageId="14">,</emphasis>
|
||
<taxonomicName id="4C5C4D17FFE76855FDCFF8024F7DD8D9" box="[582,800,1963,1988]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="hildegardeae">
|
||
<emphasis id="B928EA86FFE76855FDCFF8024E3DD8DE" box="[582,608,1964,1987]" italics="true" pageId="14">C.</emphasis>
|
||
cf.
|
||
<emphasis id="B928EA86FFE76855FD02F8054F7DD8D9" box="[651,800,1963,1988]" italics="true" pageId="14">hildegardeae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE76855FCA9F80549C1D8D9" box="[800,1436,1963,1988]" italics="true" pageId="14">
|
||
,
|
||
<taxonomicName id="4C5C4D17FFE76855FCA5F8024FF6D8D9" box="[812,939,1963,1988]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="harenna">C. harenna</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFE76855FC3EF802486AD8D9" box="[951,1079,1963,1988]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="phaeura">C. phaeura</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFE76855FBCAF802489BD8D9" box="[1091,1222,1964,1988]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="parvipes">C. parvipes</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFE76855FB5BF80249C1D8D9" box="[1234,1436,1963,1988]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="afeworkbekelei" status="sp. nov.">C. afeworkbekelei</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B928EA86FFE76855FF1EF8614CA9D8F5" bold="true" box="[151,244,1999,2024]" pageId="14">
|
||
<taxonomicNameLabel id="A21B57FDFFE76855FF1EF8614CA9D8F5" box="[151,244,1999,2024]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B928EA86FFE76855FE8CF87E4EF6D8F5" box="[261,683,1999,2024]" italics="true" pageId="14">
|
||
<taxonomicName id="4C5C4D17FFE76855FE8CF87E4D2FD8F5" box="[261,370,1999,2024]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="lucina">C. lucina</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFE76855FE09F87E4E7BD8F5" box="[384,550,1999,2024]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="macmillani">C. macmillani</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFE76855FDBDF87E4EFAD8F5" box="[564,679,1999,2024]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="baileyi">C. baileyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B928EA86FFE76855FD60F87E4F0BD8F5" box="[745,854,1999,2024]" italics="true" pageId="14">
|
||
<taxonomicName id="4C5C4D17FFE76855FD60F87E4F0FD8F5" box="[745,850,1999,2024]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="14" phylum="Chordata" rank="species" species="glassi">C. glassi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The values of CI for all mentioned species range between
|
||
<quantity id="4CA49B71FFE66854FF76FF394DEADFB2" box="[255,439,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.9405" metricValueMax="2.451" metricValueMin="1.43" pageId="15" unit="mm" value="19.405" valueMax="24.51" valueMin="14.3">14.3–24.51 mm</quantity>
|
||
. The new species is considerably smaller than the other Ethiopian
|
||
<taxonomicName id="4C5C4D17FFE66854FB45FF39491DDFAD" box="[1228,1344,151,176]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFE66854FB45FF39491DDFAD" box="[1228,1344,151,176]" italics="true" pageId="15">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
species found in the Afromontane–Afroalpine biozones:
|
||
<taxonomicName id="4C5C4D17FFE66854FD48FF134F6FDFC8" box="[705,818,188,213]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="zaphiri">
|
||
<emphasis id="B928EA86FFE66854FD48FF134F6FDFC8" box="[705,818,188,213]" italics="true" pageId="15">C. zaphiri</emphasis>
|
||
</taxonomicName>
|
||
(HB =
|
||
<quantity id="4CA49B71FFE66854FC02FF124FB6DFC9" box="[907,1003,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="15" unit="mm" value="105.0">105 mm</quantity>
|
||
, CI unknown;
|
||
<bibRefCitation id="EFCD4B65FFE66854FB1CFF124C82DFE5" author="Churchfield" pageId="15" refString="Churchfield, S. & Jenkins, P. D. (2013 a) Crocidura zaphiri Zaphir's Shrew. In: Happold, M. & Happold, D. C. D. (Eds.), Mammals of Africa. Vol. IV: Hedgehogs, Shrews and Bats. Bloomsbury Publishing, London, pp. 148 - 148." type="book" year="2013" yearSuffix="a">Churchfield & Jenkins, 2013a</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="4C5C4D17FFE66854FEABFF4E4DC9DFE5" box="[290,404,223,248]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="olivieri">
|
||
<emphasis id="B928EA86FFE66854FEABFF4E4DC9DFE5" box="[290,404,223,248]" italics="true" pageId="15">C. olivieri</emphasis>
|
||
</taxonomicName>
|
||
(CI = 32.0–
|
||
<quantity id="4CA49B71FFE66854FD97FF4E4ED9DFEA" box="[542,644,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" pageId="15" unit="mm" value="34.3">34.3 mm</quantity>
|
||
;
|
||
<bibRefCitation id="EFCD4B65FFE66854FD06FF714F81DFE5" author="Churchfield" box="[655,988,223,248]" pageId="15" refString="Churchfield, S. & Hutterer, R. (2013) Crocidura olivieri African Giant Shrew. In: Happold, M. & Happold, D. C. D. (Eds.), Mammals of Africa. Vol. IV: Hedgehogs, Shrews and Bats. Bloomsbury Publishing, London, pp. 118 - 119." type="book chapter" year="2013">Churchfield & Hutterer, 2013</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE66854FF4EFEAA4DDADE79" blockId="15.[151,1437,151,680]" pageId="15">
|
||
The first PCA (see above, and
|
||
<figureCitation id="13672A11FFE66854FDCAFEAA4ED6DE01" box="[579,651,260,285]" captionStart="FIGURE 7" captionStartId="11.[151,250,1729,1751]" captionTargetBox="[192,1398,373,1707]" captionTargetId="figure@11.[188,1398,373,1707]" captionTargetPageId="11" captionText="FIGURE 7. Principal components plots: A—for all specimens of Crocidura (1 st versus 2 nd axis); B—for five middle-sized species (1 st versus 3 rd axis). Abbreviations: Af = C. afeworkbekelei sp. nov.; Bl = C. baileyi; G = C. glassi; H = C. harenna; L = C. lucina; M = C. macmillani; P = C. parvipes; T = C. thalia; Y = Crocidura yaldeni sp. nov. Below upper plot size ranges for small-, middle-, and large-sized groups shown by condylo-incisive length (CI) along PC 1." httpUri="https://zenodo.org/record/167674/files/figure.png" pageId="15">Fig. 7</figureCitation>
|
||
A) revealed significant differences between the groups of smaller, medium-sized and larger shrews along significant PC 1. Therefore, a further PCA was performed on two larger species and
|
||
<taxonomicName id="4C5C4D17FFE66854FE97FEE34DDDDE78" box="[286,384,332,357]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFE66854FE97FEE34DDDDE78" box="[286,384,332,357]" italics="true" pageId="15">C. glassi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE66854FF4EFEC14FF0DDB5" blockId="15.[151,1437,151,680]" pageId="15">
|
||
The third PCA (
|
||
<figureCitation id="13672A11FFE66854FEF3FEC14D96DE95" box="[378,459,367,392]" captionStart="FIGURE 10" captionStartId="14.[151,250,863,885]" captionTargetBox="[189,1398,194,841]" captionTargetId="figure@14.[188,1398,193,842]" captionTargetPageId="14" captionText="FIGURE 10. Principal components plot for two large-sized species and C. glassi (1 st versus 2 nd axis). Abbreviations: G = C. glassi; T = C. thalia; Y = Crocidura yaldeni sp. nov." httpUri="https://zenodo.org/record/167677/files/figure.png" pageId="15">Fig. 10</figureCitation>
|
||
) of all 25 cranial and mandibular linear measurements reveals that the smaller
|
||
<taxonomicName id="4C5C4D17FFE66854FAB3FEDE49C1DE95" box="[1338,1436,367,392]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFE66854FAB3FEDE49C1DE95" box="[1338,1436,367,392]" italics="true" pageId="15">C. glassi</emphasis>
|
||
</taxonomicName>
|
||
is clearly separated from the larger
|
||
<taxonomicName id="4C5C4D17FFE66854FDA1FE3B4ED6DEB0" box="[552,651,404,429]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE66854FDA1FE3B4ED6DEB0" box="[552,651,404,429]" italics="true" pageId="15">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5C4D17FFE66854FD4DFE3B4F6ADEB0" box="[708,823,404,429]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE66854FD4DFE3B4F6ADEB0" box="[708,823,404,429]" italics="true" pageId="15">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE66854FCC9FE3A4FC7DEB0" bold="true" box="[832,922,404,429]" pageId="15">
|
||
<taxonomicNameLabel id="A21B57FDFFE66854FCC9FE3A4FC7DEB0" box="[832,922,404,429]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
along PC 1. This statistically significant axis accounted for 91.49% of the total variance and is most correlated with measures of the general size of the skull and mandible, such as CI (
|
||
<emphasis id="B928EA86FFE66854FE1EFE734DFFDEE9" box="[407,418,477,500]" italics="true" pageId="15">r</emphasis>
|
||
= 0.674), PL (
|
||
<emphasis id="B928EA86FFE66854FDC2FE734E0BDEE9" box="[587,598,477,500]" italics="true" pageId="15">r</emphasis>
|
||
= 0.316), MBL (
|
||
<emphasis id="B928EA86FFE66854FC95FE734F7ADEE9" box="[796,807,477,500]" italics="true" pageId="15">r</emphasis>
|
||
= 0.297) and COR (
|
||
<emphasis id="B928EA86FFE66854FB9AFE734843DEE9" box="[1043,1054,477,500]" italics="true" pageId="15">r</emphasis>
|
||
= 2.788). The second component accounted only for a small percentage of the total variance (2.78%), and therefore was not significant. The PC 2 is most positively correlated with cranial width (PGW,
|
||
<emphasis id="B928EA86FFE66854FD7BFD8B4EA0DD21" box="[754,765,549,572]" italics="true" pageId="15">r</emphasis>
|
||
= 0.576; ZYG,
|
||
<emphasis id="B928EA86FFE66854FC3DFD8B4FE2DD21" box="[948,959,549,572]" italics="true" pageId="15">r</emphasis>
|
||
= 0.349; GW,
|
||
<emphasis id="B928EA86FFE66854FBE2FD8B482BDD21" box="[1131,1142,549,572]" italics="true" pageId="15">r</emphasis>
|
||
= 0.261). As can be seen from
|
||
<figureCitation id="13672A11FFE66854FF5CFDE94D75DD7D" box="[213,296,583,608]" captionStart="FIGURE 10" captionStartId="14.[151,250,863,885]" captionTargetBox="[189,1398,194,841]" captionTargetId="figure@14.[188,1398,193,842]" captionTargetPageId="14" captionText="FIGURE 10. Principal components plot for two large-sized species and C. glassi (1 st versus 2 nd axis). Abbreviations: G = C. glassi; T = C. thalia; Y = Crocidura yaldeni sp. nov." httpUri="https://zenodo.org/record/167677/files/figure.png" pageId="15">Fig. 10</figureCitation>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFE66854FEBDFDE64DCADD7D" box="[308,407,583,608]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE66854FEBDFDE64DCADD7D" box="[308,407,583,608]" italics="true" pageId="15">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5C4D17FFE66854FE59FDE64E18DD7D" box="[464,581,583,608]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE66854FE59FDE64E18DD7D" box="[464,581,583,608]" italics="true" pageId="15">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE66854FDC5FDE94EFADD7D" bold="true" box="[588,679,583,608]" pageId="15">
|
||
<taxonomicNameLabel id="A21B57FDFFE66854FDC5FDE94EFADD7D" box="[588,679,583,608]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are clearly distinguished along the first two principal components. The difference between the two species along the first axis reflects the larger average size of
|
||
<taxonomicName id="4C5C4D17FFE66854FB34FDC3496FDD98" box="[1213,1330,620,645]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE66854FB34FDC3496FDD98" box="[1213,1330,620,645]" italics="true" pageId="15">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE66854FAB3FDC249C8DD98" bold="true" box="[1338,1429,620,645]" pageId="15">
|
||
<taxonomicNameLabel id="A21B57FDFFE66854FAB3FDC249C8DD98" box="[1338,1429,620,645]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, while the second component is associated with changes in skull shape.
|
||
</paragraph>
|
||
<caption id="DF23661CFFE66854FF1EFA5749DBD933" httpUri="https://zenodo.org/record/167678/files/figure.png" pageId="15" targetBox="[200,1387,744,1496]" targetPageId="15">
|
||
<paragraph id="8BE33694FFE66854FF1EFA5749DBD933" blockId="15.[151,1436,1529,1582]" pageId="15">
|
||
<emphasis id="B928EA86FFE66854FF1EFA574D42D912" bold="true" box="[151,287,1529,1551]" pageId="15">FIGURE 11.</emphasis>
|
||
Afework Bekele (left, photograph taken by Viola Clausnitzer in March 2000) and the late Derek Yalden (right, photograph taken by Pat Morris), two biologists who greatly contributed to the knowledge of the mammal fauna of Ethiopia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE33694FFE66854FF4EF9F54862D819" blockId="15.[151,1437,1627,2012]" pageId="15">
|
||
Detailed comparison between similar in size species:
|
||
<taxonomicName id="4C5C4D17FFE66854FC97F9F24FCCD969" box="[798,913,1627,1652]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE66854FC97F9F24FCCD969" box="[798,913,1627,1652]" italics="true" pageId="15">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE66854FC11F9F54FACD969" bold="true" box="[920,1009,1627,1652]" pageId="15">
|
||
<taxonomicNameLabel id="A21B57FDFFE66854FC11F9F54FACD969" box="[920,1009,1627,1652]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C5C4D17FFE66854FB0CF9F248BAD969" box="[1157,1255,1627,1652]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE66854FB0CF9F248BAD969" box="[1157,1255,1627,1652]" italics="true" pageId="15">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
in: smaller P4s/ d, M3s/d; bigger M2L, M3W, M3L, UML, ZYG, PGW, LML; longer hindfoot and claws (
|
||
<tableCitation id="C6DE032FFFE66854FB1AF9D048B6D98A" box="[1171,1259,1662,1687]" captionStart="TABLE 2" captionStartId="10.[151,239,152,175]" captionTargetBox="[159,1229,266,1517]" captionTargetPageId="10" captionText="TABLE 2. Comparison of the external and cranial measurements of C. yaldeni sp. nov. and C. thalia. Measurements in millimeters are presented as the mean, standard error of mean, standard deviation, range and number of specimens in parentheses." httpUri="http://table.plazi.org/id/DF23661CFFE36851FF1EFF364D46DFF2" pageId="15" tableUuid="DF23661CFFE36851FF1EFF364D46DFF2">Table 2</tableCitation>
|
||
); the uniformly colored tail (tail of
|
||
<taxonomicName id="4C5C4D17FFE66854FEFCF90A4D85D9A1" box="[373,472,1699,1724]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE66854FEFCF90A4D85D9A1" box="[373,472,1699,1724]" italics="true" pageId="15">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
is mottled with brown and pale ochre); the bicolored and longer bristle hairs of tail; more massive
|
||
<date id="FFE21054FFE66854FEB1F9694D0CD9C2" box="[312,337,1735,1759]" pageId="15">I1</date>
|
||
; more compact upper antemolars row; stronger developed incision on lingual edge of the M2 (
|
||
<figureCitation id="13672A11FFE66854FAE6F9684CFAD81E" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="15">Fig. 9</figureCitation>
|
||
, A1-a vs. B1); the expression of hypoconulids on the lower molars (
|
||
<figureCitation id="13672A11FFE66854FC2DF9454FB4D81E" box="[932,1001,1771,1796]" captionStart="FIGURE 9" captionStartId="13.[151,250,1763,1785]" captionTargetBox="[188,1398,302,1739]" captionTargetId="figure@13.[188,1398,301,1741]" captionTargetPageId="13" captionText="FIGURE 9. Specific features of Crocidura yaldeni sp. nov. S- 165343 (A 1 – A 4) and C. thalia S- 164858 (B 1 – B 4). A 1, B 1 — occlusal view of the M 2 and M 3; A 2, B 2 — lingual side of the m 1; A 3, B 3 — occlusal view of the m 3; A 4, B 4 — lateral view of the P 4. Abbreviations: encd = entoconid; hc = hypocone; also see figure 5. Specific characters of C. yaldeni: a—the symmetric and distinctly curved incision of the M 2; b—postero-lingual part of the M 2 in contact with M 3; c—the shape of the outline of the posterior edge of M 3; d—the hypoconulid shoulder of the m 1 in medial view; e—postcingulid of m 3 forming a notch; fthe undulated upper margin of P 4 in lateral view; g—three-quarters of the crown base of P 4 surrounded by a cingulum. Scale 1 mm." httpUri="https://zenodo.org/record/167676/files/figure.png" pageId="15">Fig. 9</figureCitation>
|
||
, A2-d).
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFE66854FF4EF8A049C1D8C6" blockId="15.[151,1437,1627,2012]" pageId="15">
|
||
The standard karyotype is identical to that of two related species,
|
||
<taxonomicName id="4C5C4D17FFE66854FC20F8A14856D83A" box="[937,1035,1806,1831]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE66854FC20F8A14856D83A" box="[937,1035,1806,1831]" italics="true" pageId="15">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5C4D17FFE66854FBCBF8A14C86D856" authority="Lavrenchenko et al., 1997" authorityName="Lavrenchenko et al." authorityYear="1997" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFE66854FBCBF8A148F9D83A" box="[1090,1188,1806,1831]" italics="true" pageId="15">C. glassi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCD4B65FFE66854FB3AF8A04C8ED856" author="Lavrenchenko" pageId="15" refString="Lavrenchenko, L. A., Milishnikov, A. N., Aniskin, V. M., Warshavsky, A. A. & Gebrekidan, W. (1997) The genetic diversity of small mammals of the Bale Mountains, Ethiopia. SINET: Ethiopian Journal of Science, 20, 213 - 233. http: // dx. doi. org / 10.4314 / sinet. v 20 i 2.18102" type="journal article" year="1997">
|
||
Lavrenchenko
|
||
<emphasis id="B928EA86FFE66854FAD5F8BE49D3D83A" box="[1372,1422,1806,1831]" italics="true" pageId="15">et al</emphasis>
|
||
., 1997
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Phylogenetic analyses based upon repetitive DNA elements (taxonomic DNA fingerprint) and inter-SINE- PCR (IS-PCR) revealed that genetic distances between
|
||
<taxonomicName id="4C5C4D17FFE66854FC88F8F94F2CD872" box="[769,881,1878,1903]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE66854FC88F8F94F2CD872" box="[769,881,1878,1903]" italics="true" pageId="15">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE66854FCFEF8F84F8DD872" bold="true" box="[887,976,1878,1903]" pageId="15">
|
||
<taxonomicNameLabel id="A21B57FDFFE66854FCFEF8F84F8DD872" box="[887,976,1878,1903]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and its two closest relatives,
|
||
<taxonomicName id="4C5C4D17FFE66854FA82F8F94930D872" box="[1291,1389,1878,1903]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE66854FA82F8F94930D872" box="[1291,1389,1878,1903]" italics="true" pageId="15">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B928EA86FFE66854FF1EF8D24D55D889" box="[151,264,1915,1940]" italics="true" pageId="15">
|
||
<taxonomicName id="4C5C4D17FFE66854FF1EF8D24D59D889" box="[151,260,1915,1940]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="glassi">C. glassi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
fell within the range usually recorded for interspecific genetic differentiation within
|
||
<taxonomicName id="4C5C4D17FFE66854FAA1F8D549C1D889" box="[1320,1436,1915,1940]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFE66854FAA1F8D549C1D889" box="[1320,1436,1915,1940]" italics="true" pageId="15">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFCD4B65FFE66854FF17F8304DFFD8AA" author="Bannikova" box="[158,418,1950,1975]" pageId="15" refString="Bannikova, A. A., Lavrenchenko, L. A., Lomov A. A. & Mednikov, B. M. (2001) Molecular diversity of some Crocidura species (Insectivora, Soricidae) from Ethiopia. In: Denis, C., Granjon, L. & Poulet, A. (Eds.), African Small Mammals. IRD Editions, Paris, pp. 55 - 64." type="book chapter" year="2001">
|
||
Bannikova
|
||
<emphasis id="B928EA86FFE66854FEA9F80E4D0CD8AA" box="[288,337,1950,1975]" italics="true" pageId="15">et al</emphasis>
|
||
., 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCD4B65FFE66854FE24F8304EECD8AA" author="Bannikova" box="[429,689,1950,1975]" pageId="15" refString="Bannikova, A. A., Lavrenchenko, L. A. & Kramerov, D. A. (2005) Phylogenetic relationships between Afrotropical and Palaearctic Crocidura species inferred from Inter-SINE-PCR. Biochemical Systematics and Ecology, 33, 45 - 59. http: // dx. doi. org / 10.1016 / j. bse. 2004.05.014" type="journal article" year="2005">
|
||
Bannikova
|
||
<emphasis id="B928EA86FFE66854FDA7F80E4E02D8AA" box="[558,607,1950,1975]" italics="true" pageId="15">et al</emphasis>
|
||
., 2005
|
||
</bibRefCitation>
|
||
). The former analysis strongly supported that
|
||
<taxonomicName id="4C5C4D17FFE66854FB3AF831497ED8AA" box="[1203,1315,1950,1975]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFE66854FB3AF831497ED8AA" box="[1203,1315,1950,1975]" italics="true" pageId="15">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFE66854FAA3F83049DED8AA" bold="true" box="[1322,1411,1950,1975]" pageId="15">
|
||
<taxonomicNameLabel id="A21B57FDFFE66854FAA3F83049DED8AA" box="[1322,1411,1950,1975]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a sister species to
|
||
<taxonomicName id="4C5C4D17FFE66854FED7F86A4D9FD8C1" box="[350,450,1987,2012]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE66854FED7F86A4D9FD8C1" box="[350,450,1987,2012]" italics="true" pageId="15">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
; in the latter analysis
|
||
<taxonomicName id="4C5C4D17FFE66854FD37F86A4F7CD8C1" box="[702,801,1987,2012]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFE66854FD37F86A4F7CD8C1" box="[702,801,1987,2012]" italics="true" pageId="15">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5C4D17FFE66854FCD3F86A4FE1D8C1" box="[858,956,1987,2012]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="15" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFE66854FCD3F86A4FE1D8C1" box="[858,956,1987,2012]" italics="true" pageId="15">C. glassi</emphasis>
|
||
</taxonomicName>
|
||
had a tendency to form a clade against
|
||
<emphasis id="B928EA86FFE66854FA0AF86A49C1D8C6" box="[1411,1436,1988,2011]" italics="true" pageId="15">C.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFF9684BFF1EFF394E1EDE01" blockId="16.[151,1437,151,1005]" pageId="16">
|
||
<taxonomicName id="4C5C4D17FFF9684BFF1EFF394CBADFAD" box="[151,231,151,176]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFF9684BFF1EFF394CBADFAD" box="[151,231,151,176]" italics="true" pageId="16">yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFF9684BFF64FF394D1ADFAD" bold="true" box="[237,327,151,176]" pageId="16">
|
||
<taxonomicNameLabel id="A21B57FDFFF9684BFF64FF394D1ADFAD" box="[237,327,151,176]" pageId="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The phylogenetic analysis of an extended set of
|
||
<taxonomicName id="4C5C4D17FFF9684BFCEAFF394F89DFAD" box="[867,980,151,176]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EA86FFF9684BFCEAFF394F89DFAD" box="[867,980,151,176]" italics="true" pageId="16">Crocidura</emphasis>
|
||
</taxonomicName>
|
||
species using mitochondrial cytochrome b gene sequences (
|
||
<bibRefCitation id="EFCD4B65FFF9684BFEE4FF124EC7DFC9" author="Lavrenchenko" box="[365,666,188,213]" pageId="16" refString="Lavrenchenko, L. A., Bannikova, A. A. & Lebedev, V. S. (2009) Shrews (Crocidura spp.) endemic to Ethiopia: recent adaptive radiation of an ancient lineage. Doklady Biological Sciences, 424, 57 - 60. http: // dx. doi. org / 10.1134 / S 0012496609010177" type="journal article" year="2009">Lavrenchenko et al., 2009</bibRefCitation>
|
||
) revealed that
|
||
<taxonomicName id="4C5C4D17FFF9684BFCCDFF134FE8DFC8" box="[836,949,188,213]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFF9684BFCCDFF134FE8DFC8" box="[836,949,188,213]" italics="true" pageId="16">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFF9684BFC34FF124844DFC8" bold="true" box="[957,1049,188,213]" pageId="16">
|
||
<taxonomicNameLabel id="A21B57FDFFF9684BFC34FF124844DFC8" box="[957,1049,188,213]" pageId="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C5C4D17FFF9684BFBDBFF1348AEDFC8" box="[1106,1267,188,213]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="macmillani">
|
||
<emphasis id="B928EA86FFF9684BFBDBFF1348AEDFC8" box="[1106,1267,188,213]" italics="true" pageId="16">C. macmillani</emphasis>
|
||
</taxonomicName>
|
||
form the most basal branch of the group of Ethiopian endemics (including
|
||
<taxonomicName id="4C5C4D17FFF9684BFCB2FF4E4FC0DFE5" box="[827,925,223,248]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFF9684BFCB2FF4E4FC0DFE5" box="[827,925,223,248]" italics="true" pageId="16">C. glassi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFC25FF4E4853DFE5" box="[940,1038,223,248]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFF9684BFC25FF4E4853DFE5" box="[940,1038,223,248]" italics="true" pageId="16">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFB94FF4E48D9DFE5" box="[1053,1156,223,248]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="lucina">
|
||
<emphasis id="B928EA86FFF9684BFB94FF4E48D9DFE5" box="[1053,1156,223,248]" italics="true" pageId="16">C. lucina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5C4D17FFF9684BFB32FF4E4975DFE5" box="[1211,1320,223,248]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="baileyi">
|
||
<emphasis id="B928EA86FFF9684BFB32FF4E4975DFE5" box="[1211,1320,223,248]" italics="true" pageId="16">C. baileyi</emphasis>
|
||
</taxonomicName>
|
||
), whereas
|
||
<taxonomicName id="4C5C4D17FFF9684BFF1EFEAB4CA4DE00" box="[151,249,260,285]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="thalia">
|
||
<emphasis id="B928EA86FFF9684BFF1EFEAB4CA4DE00" box="[151,249,260,285]" italics="true" pageId="16">C. thalia</emphasis>
|
||
</taxonomicName>
|
||
appears as sister to
|
||
<taxonomicName id="4C5C4D17FFF9684BFE53FEAB4E66DE00" box="[474,571,260,285]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="glassi">
|
||
<emphasis id="B928EA86FFF9684BFE53FEAB4E66DE00" box="[474,571,260,285]" italics="true" pageId="16">C. glassi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346651FFFF9684BFF4EFE894E92DC9D" pageId="16" type="distribution">
|
||
<paragraph id="8BE33694FFF9684BFF4EFE894D0FDEE8" blockId="16.[151,1437,151,1005]" pageId="16">
|
||
<emphasis id="B928EA86FFF9684BFF4EFE894D04DE5D" bold="true" box="[199,345,295,320]" pageId="16">Distribution</emphasis>
|
||
. The new species has been found only in the Beletta Forest (
|
||
<geoCoordinate id="EE685053FFF9684BFB83FE86483BDE5D" box="[1034,1126,296,320]" direction="north" orientation="latitude" pageId="16" precision="925" value="7.5666666">07º34'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685053FFF9684BFBF8FE89489ADE5D" box="[1137,1223,295,320]" direction="east" orientation="longitude" pageId="16" precision="925" value="36.516666">36º31'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CA49B71FFF9684BFB5DFE86497ADE22" box="[1236,1319,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="16" unit="m" value="1900.0">1900 m</quantity>
|
||
a.s.l.). We failed to trap
|
||
<taxonomicName id="4C5C4D17FFF9684BFEA4FEE34DC2DE78" box="[301,415,332,357]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFF9684BFEA4FEE34DC2DE78" box="[301,415,332,357]" italics="true" pageId="16">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFF9684BFE21FEE24E5EDE78" bold="true" box="[424,515,332,357]" pageId="16">
|
||
<taxonomicNameLabel id="A21B57FDFFF9684BFE21FEE24E5EDE78" box="[424,515,332,357]" pageId="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in any other site of this forest and other humid Afromontane forest blocks of SW
|
||
<collectingCountry id="F34B7604FFF9684BFF1EFEC14CA7DE95" box="[151,250,367,392]" name="Ethiopia" pageId="16">Ethiopia</collectingCountry>
|
||
: the Sheko Forest (
|
||
<geoCoordinate id="EE685053FFF9684BFE6BFEDE4E1FDE95" box="[482,578,368,392]" direction="north" orientation="latitude" pageId="16" precision="925" value="7.0666666">07°04'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685053FFF9684BFDDBFEC14EF0DE95" box="[594,685,367,392]" direction="east" orientation="longitude" pageId="16" precision="925" value="35.5">35°30'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CA49B71FFF9684BFD34FEDE4F4ADE9A" box="[701,791,368,392]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="16" unit="m" value="1930.0">1930 m</quantity>
|
||
a.s.l.), the Dushi Area of the Godare Forest (
|
||
<geoCoordinate id="EE685053FFF9684BFAB1FEDE49CADE95" box="[1336,1431,368,392]" direction="north" orientation="latitude" pageId="16" precision="925" value="7.35">07°21'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685053FFF9684BFF1EFE3A4CAFDEB0" box="[151,242,404,429]" direction="east" orientation="longitude" pageId="16" precision="925" value="35.216667">35°13'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CA49B71FFF9684BFE88FE3A4D04DEB1" box="[257,345,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="16" unit="m" value="1200.0">1200 m</quantity>
|
||
a.s.l.), the Meti Area of the Godare Forest (
|
||
<geoCoordinate id="EE685053FFF9684BFCD7FE3A4FE0DEB1" box="[862,957,404,428]" direction="north" orientation="latitude" pageId="16" precision="925" value="7.2833333">07°17'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685053FFF9684BFC45FE3A4875DEB0" box="[972,1064,404,429]" direction="east" orientation="longitude" pageId="16" precision="925" value="35.266666">35°16'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CA49B71FFF9684BFBBFFE3A48D3DEB1" box="[1078,1166,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="16" unit="m" value="1370.0">1370 m</quantity>
|
||
a.s.l.) and the Inegawa Forest (
|
||
<geoCoordinate id="EE685053FFF9684BFF67FE194D10DECD" box="[238,333,439,464]" direction="north" orientation="latitude" pageId="16" precision="925" value="7.4166665">07°25'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685053FFF9684BFED3FE194DEBDECD" box="[346,438,439,464]" direction="east" orientation="longitude" pageId="16" precision="925" value="35.4">35°24'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CA49B71FFF9684BFE4AFE164E44DED2" box="[451,537,440,464]" metricMagnitude="3" metricUnit="m" metricValue="2.34" pageId="16" unit="m" value="2340.0">2340 m</quantity>
|
||
a.s.l.). Therefore, the currently known distribution range of this new species is extremely small.
|
||
</paragraph>
|
||
<paragraph id="8BE33694FFF9684BFF4EFE514E92DC9D" blockId="16.[151,1437,151,1005]" pageId="16">
|
||
<emphasis id="B928EA86FFF9684BFF4EFE514D7DDD05" bold="true" box="[199,288,511,536]" pageId="16">Habitat</emphasis>
|
||
. All four specimens of
|
||
<taxonomicName id="4C5C4D17FFF9684BFDA1FDAE4EC4DD05" box="[552,665,511,536]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFF9684BFDA1FDAE4EC4DD05" box="[552,665,511,536]" italics="true" pageId="16">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFF9684BFD28FE514EA1DD05" bold="true" box="[673,764,511,536]" pageId="16">
|
||
<taxonomicNameLabel id="A21B57FDFFF9684BFD28FE514EA1DD05" box="[673,764,511,536]" pageId="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were captured in the riverine variant of humid Afromontane forest on the bank of the small river (trees:
|
||
<taxonomicName id="4C5C4D17FFF9684BFD2CFD8A4FC9DD20" box="[677,916,548,573]" class="Magnoliopsida" family="Araliaceae" genus="Schefflera" kingdom="Plantae" order="Apiales" pageId="16" phylum="Tracheophyta" rank="species" species="abyssinica">
|
||
<emphasis id="B928EA86FFF9684BFD2CFD8A4FC9DD20" box="[677,916,548,573]" italics="true" pageId="16">Schefflera abyssinica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFC2EFD8B48DEDD21" box="[935,1155,549,572]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="16" phylum="Tracheophyta" rank="species" species="macrostacis">
|
||
<emphasis id="B928EA86FFF9684BFC2EFD8B48DEDD21" box="[935,1155,549,572]" italics="true" pageId="16">Croton macrostacis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFB1FFD8A49C8DD20" box="[1174,1429,548,573]" class="Magnoliopsida" family="Sapindaceae" genus="Allophylus" kingdom="Animalia" order="Sapindales" pageId="16" phylum="Tracheophyta" rank="species" species="abyssinicus">
|
||
<emphasis id="B928EA86FFF9684BFB1FFD8A49C8DD20" box="[1174,1429,548,573]" italics="true" pageId="16">Allophylus abyssinicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFF1EFDE64DA7DD7D" authority="Malacanta" authorityName="Malacanta" box="[151,506,583,608]" class="Magnoliopsida" family="Sapotaceae" genus="Aningeria" higherTaxonomySource="CoL" kingdom="Plantae" order="Ericales" pageId="16" phylum="Tracheophyta" rank="species" species="altissima">
|
||
<emphasis id="B928EA86FFF9684BFF1EFDE64D2CDD7D" box="[151,369,583,608]" italics="true" pageId="16">Aningeria altissima</emphasis>
|
||
,
|
||
<emphasis id="B928EA86FFF9684BFE0DFDE94DA7DD7D" box="[388,506,583,608]" italics="true" pageId="16">Malacanta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFF9684BFD8FFDE94E3FDD7D" box="[518,610,583,608]" italics="true" pageId="16">alnifolia</emphasis>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFDFDFDE94F67DD7D" box="[628,826,583,608]" class="Liliopsida" family="Arecaceae" genus="Phoenix" kingdom="Plantae" order="Arecales" pageId="16" phylum="Tracheophyta" rank="species" species="reclinata">
|
||
<emphasis id="B928EA86FFF9684BFDFDFDE94F67DD7D" box="[628,826,583,608]" italics="true" pageId="16">Phoenix reclinata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFCC4FDE74817DD7D" box="[845,1098,583,608]" class="Magnoliopsida" family="Simaroubaceae" genus="Brucea" kingdom="Plantae" order="Sapindales" pageId="16" phylum="Tracheophyta" rank="species" species="antiderinterica">
|
||
<emphasis id="B928EA86FFF9684BFCC4FDE74817DD7D" box="[845,1098,583,608]" italics="true" pageId="16">Brucea antiderinterica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFBD4FDE9495ADD7D" box="[1117,1287,583,608]" class="Magnoliopsida" family="Araliaceae" genus="Polyscias" kingdom="Plantae" order="Apiales" pageId="16" phylum="Tracheophyta" rank="species" species="fulva">
|
||
<emphasis id="B928EA86FFF9684BFBD4FDE9495ADD7D" box="[1117,1287,583,608]" italics="true" pageId="16">Polyscias fulva</emphasis>
|
||
</taxonomicName>
|
||
; small trees:
|
||
<taxonomicName id="4C5C4D17FFF9684BFF1EFDC34D6BDD98" box="[151,310,620,645]" class="Magnoliopsida" family="Rutaceae" genus="Vepris" kingdom="Plantae" order="Sapindales" pageId="16" phylum="Tracheophyta" rank="species" species="dainelii">
|
||
<emphasis id="B928EA86FFF9684BFF1EFDC34D6BDD98" box="[151,310,620,645]" italics="true" pageId="16">Vepris dainelii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFECDFDC24D80DD98" box="[324,477,620,645]" class="Magnoliopsida" family="Rutaceae" genus="Teclea" higherTaxonomySource="CoL" kingdom="Plantae" order="Sapindales" pageId="16" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B928EA86FFF9684BFECDFDC24D80DD98" box="[324,477,620,645]" italics="true" pageId="16">Teclea nobilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFE62FDC34EB0DD99" box="[491,749,620,644]" class="Reptilia" family="Teiidae" genus="Dracaena" kingdom="Animalia" order="Squamata" pageId="16" phylum="Chordata" rank="species" species="afromontana">
|
||
<emphasis id="B928EA86FFF9684BFE62FDC34EB0DD99" box="[491,749,620,644]" italics="true" pageId="16">Dracaena afromontana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFD72FDC34F26DD99" box="[763,891,620,644]" class="Reptilia" family="Teiidae" genus="Dracaena" kingdom="Animalia" order="Squamata" pageId="16" phylum="Chordata" rank="species" species="fragrans">
|
||
<emphasis id="B928EA86FFF9684BFD72FDC34F26DD99" box="[763,891,620,644]" italics="true" pageId="16">D. fragrans</emphasis>
|
||
</taxonomicName>
|
||
; shrubs:
|
||
<taxonomicName id="4C5C4D17FFF9684BFC6BFDC248D5DD98" box="[994,1160,620,645]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="16" phylum="Tracheophyta" rank="species" species="arabica">
|
||
<emphasis id="B928EA86FFF9684BFC6BFDC248D5DD98" box="[994,1160,620,645]" italics="true" pageId="16">Coffea arabica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFB1FFDC249C8DD98" box="[1174,1429,620,645]" class="Magnoliopsida" family="Rubiaceae" genus="Canthium" kingdom="Plantae" order="Gentianales" pageId="16" phylum="Tracheophyta" rank="species" species="oligocarpum">
|
||
<emphasis id="B928EA86FFF9684BFB1FFDC249C8DD98" box="[1174,1429,620,645]" italics="true" pageId="16">Canthium oligocarpum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFF1EFD214DDCDDB5" box="[151,385,655,680]" class="Magnoliopsida" family="Rubiaceae" genus="Galiniera" kingdom="Plantae" order="Gentianales" pageId="16" phylum="Tracheophyta" rank="species" species="coffeoides">
|
||
<emphasis id="B928EA86FFF9684BFF1EFD214DDCDDB5" box="[151,385,655,680]" italics="true" pageId="16">Galiniera coffeoides</emphasis>
|
||
</taxonomicName>
|
||
; climbers:
|
||
<taxonomicName id="4C5C4D17FFF9684BFD9AFD214EB6DDB5" box="[531,747,655,680]" class="Magnoliopsida" family="Primulaceae" genus="Embelia" kingdom="Plantae" order="Ericales" pageId="16" phylum="Tracheophyta" rank="species" species="schimperi">
|
||
<emphasis id="B928EA86FFF9684BFD9AFD214EB6DDB5" box="[531,747,655,680]" italics="true" pageId="16">Embelia schimperi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFC8CFD214FB1DDB5" box="[773,1004,655,680]" class="Mammalia" family="Soricidae" genus="Phychotria" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="neglecta">
|
||
<emphasis id="B928EA86FFF9684BFC8CFD214FB1DDB5" box="[773,1004,655,680]" italics="true" pageId="16">Phychotria neglecta</emphasis>
|
||
</taxonomicName>
|
||
; ferns:
|
||
<taxonomicName id="4C5C4D17FFF9684BFBDEFD3F4952DDB5" box="[1111,1295,655,680]" class="Polypodiopsida" family="Pteridaceae" genus="Pteris" kingdom="Plantae" order="Polypodiales" pageId="16" phylum="Tracheophyta" rank="species" species="dentatum">
|
||
<emphasis id="B928EA86FFF9684BFBDEFD3F4952DDB5" box="[1111,1295,655,680]" italics="true" pageId="16">Pteris dentatum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFAA0FD214D53DDD0" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="16" phylum="Tracheophyta" rank="species" species="sandersoni">
|
||
<emphasis id="B928EA86FFF9684BFAA0FD214D53DDD0" italics="true" pageId="16">Asplenium sandersoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFEABFD1A4DD3DDD0" box="[290,398,692,717]" class="Polypodiopsida" family="Polypodiaceae" genus="Pleopeltis" kingdom="Plantae" order="Polypodiales" pageId="16" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B928EA86FFF9684BFEABFD1A4DD3DDD0" box="[290,398,692,717]" italics="true" pageId="16">Pleopeltis</emphasis>
|
||
</taxonomicName>
|
||
sp.; herbs:
|
||
<emphasis id="B928EA86FFF9684BFDA9FD1A4EFBDDD1" box="[544,678,692,716]" italics="true" pageId="16">Afromomum</emphasis>
|
||
sp.). Three rodent species,
|
||
<taxonomicName id="4C5C4D17FFF9684BFC7AFD1A4907DDD0" authority="Osgood" authorityName="Osgood" box="[1011,1370,692,717]" class="Mammalia" family="Muridae" genus="Lophuromys" kingdom="Animalia" order="Rodentia" pageId="16" phylum="Chordata" rank="species" species="chrysopus">
|
||
<emphasis id="B928EA86FFF9684BFC7AFD1A48ABDDD0" box="[1011,1270,692,717]" italics="true" pageId="16">Lophuromys chrysopus</emphasis>
|
||
Osgood
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D17FFF9684BFAE4FD1B4D03DDED" authority="Rhoads" authorityName="Rhoads" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="16" phylum="Chordata" rank="species" species="mahomet">
|
||
<emphasis id="B928EA86FFF9684BFAE4FD1B4CA6DDED" italics="true" pageId="16">Mus mahomet</emphasis>
|
||
Rhoads
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5C4D17FFF9684BFE2FFD794F1DDDED" authority="Rueppell" authorityName="Rueppell" box="[422,832,727,752]" class="Mammalia" family="Muridae" genus="Stenocephalemys" kingdom="Animalia" order="Rodentia" pageId="16" phylum="Chordata" rank="species" species="albipes">
|
||
<emphasis id="B928EA86FFF9684BFE2FFD794EE0DDED" box="[422,701,727,752]" italics="true" pageId="16">Stenocephalemys albipes</emphasis>
|
||
(Rueppell)
|
||
</taxonomicName>
|
||
were also collected at the same trapping site. All individuals of the new shrew were caught in Sherman live traps placed on the ground no more than
|
||
<quantity id="4CA49B71FFF9684BFB5FFD5248A2DC09" box="[1238,1279,764,788]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="16" unit="m" value="1.0">1 m</quantity>
|
||
from the river bank. Probably the very restricted range of
|
||
<taxonomicName id="4C5C4D17FFF9684BFDFEFC8E4EB4DC25" box="[631,745,799,824]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="16" phylum="Chordata" rank="species" species="yaldeni" status="sp. nov.">
|
||
<emphasis id="B928EA86FFF9684BFDFEFC8E4EB4DC25" box="[631,745,799,824]" italics="true" pageId="16">C. yaldeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA86FFF9684BFD78FCB14F11DC25" bold="true" box="[753,844,799,824]" pageId="16">
|
||
<taxonomicNameLabel id="A21B57FDFFF9684BFD78FCB14F11DC25" box="[753,844,799,824]" pageId="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is associated with yet unknown habitat requirements, presumably more specific than 'river edge'. However, no morphological adaptations to some specific life style (including semi-aquatic adaptations) were observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346651FFFF9684BFF4EFC224E47DCF0" pageId="16" type="etymology">
|
||
<paragraph id="8BE33694FFF9684BFF4EFC224E47DCF0" blockId="16.[151,1437,151,1005]" pageId="16">
|
||
<emphasis id="B928EA86FFF9684BFF4EFC224D1BDCB8" bold="true" box="[199,326,908,933]" pageId="16">Etymology</emphasis>
|
||
. The new species is named in honor of the late Dr. Derek W. Yalden (
|
||
<date id="FFE21054FFF9684BFB2AFC22497FDCB9" box="[1187,1314,908,932]" pageId="16" value="1940" valueMax="2013">1940-2013</date>
|
||
), who has contributed greatly to our knowledge on Ethiopian small mammals (
|
||
<figureCitation id="13672A11FFF9684BFC14FC014FADDCD5" box="[925,1008,943,968]" captionStart="FIGURE 11" captionStartId="15.[151,250,1529,1551]" captionTargetBox="[200,1387,744,1496]" captionTargetId="figure@15.[188,1398,733,1508]" captionTargetPageId="15" captionText="FIGURE 11. Afework Bekele (left, photograph taken by Viola Clausnitzer in March 2000) and the late Derek Yalden (right, photograph taken by Pat Morris), two biologists who greatly contributed to the knowledge of the mammal fauna of Ethiopia." httpUri="https://zenodo.org/record/167678/files/figure.png" pageId="16">Fig. 11</figureCitation>
|
||
). As the vernacular name for the new species we propose Beletta Shrew.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |