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<document id="FFEA7F4FD607FBF4C5322948A5DF6BEF" ID-CLB-Dataset="50773" ID-DOI="10.5281/zenodo.172430" ID-GBIF-Dataset="06a9d708-5518-4797-8de3-595854d4f8b3" ID-ISSN="1175­5326" ID-Zenodo-Dep="172430" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459756796429" checkinUser="plazi" docAuthor="Hoch, Hannelore" docDate="2006" docId="03F58782E13CFFAFDB03FB71FE9F1633" docLanguage="en" docName="zt01209p047.pdf" docOrigin="Zootaxa 1209" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Oteana Hoch, 2006, gen. nov." docType="treatment" docVersion="10" lastPageNumber="7" masterDocId="FFCCFFFAE13EFFA9DA0BFFE8FFFD1572" masterDocTitle="New Cixiidae from Eastern Polynesia: Oteana gen. nov. and Manurevana gen. nov. (Hemiptera: Fulgoromorpha)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="3" updateTime="1698218175919" updateUser="ExternalLinkService">
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<mods:title id="51843FD817FAD284B9FAAE18C2BC4617">New Cixiidae from Eastern Polynesia: Oteana gen. nov. and Manurevana gen. nov. (Hemiptera: Fulgoromorpha)</mods:title>
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<mods:date id="B3A97CC25271AD2549FA4BE365FADEA0">2006</mods:date>
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<treatment id="03F58782E13CFFAFDB03FB71FE9F1633" ID-DOI="http://doi.org/10.5281/zenodo.6255917" ID-GBIF-Taxon="119343850" ID-Zenodo-Dep="6255917" LSID="urn:lsid:plazi:treatment:03F58782E13CFFAFDB03FB71FE9F1633" httpUri="http://treatment.plazi.org/id/03F58782E13CFFAFDB03FB71FE9F1633" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
<subSubSection id="C346651FE13CFFABDB03FB71FE2811C1" box="[264,469,1177,1203]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BE33694E13CFFABDB03FB71FE2811C1" blockId="2.[264,469,1177,1203]" box="[264,469,1177,1203]" pageId="2" pageNumber="3">
<heading id="D0AB81F8E13CFFABDB03FB71FE2811C1" bold="true" box="[264,469,1177,1203]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B928EA86E13CFFABDB03FB71FE2811C1" bold="true" box="[264,469,1177,1203]" pageId="2" pageNumber="3">
<taxonomicName id="4C5C4D17E13CFFABDB03FB71FE9D11C1" ID-CoL="63HVD" ID-ENA="1902045" box="[264,352,1177,1203]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA86E13CFFABDB03FB71FE9D11C1" bold="true" box="[264,352,1177,1203]" italics="true" pageId="2" pageNumber="3">Oteana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57FDE13CFFABDB6CFB71FE2811C1" box="[359,469,1177,1203]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C346651FE13CFFAFDB03FB31FE9F1633" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BE33694E13CFFABDB03FB31FAF21181" blockId="2.[264,1295,1241,1267]" box="[264,1295,1241,1267]" pageId="2" pageNumber="3">
<typeStatus id="54E78836E13CFFABDB03FB31FEBF1181" box="[264,322,1241,1267]" pageId="2" pageNumber="3">Type</typeStatus>
species:
<taxonomicName id="4C5C4D17E13CFFABDBA5FB31FC931181" authority="Fennah, 1958: 131" authorityName="Fennah" authorityPageNumber="131" authorityYear="1958" box="[430,878,1241,1267]" class="Insecta" family="Cixiidae" genus="Oliarus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="euphranor">
<emphasis id="B928EA86E13CFFABDBA5FB31FD771180" box="[430,650,1241,1266]" italics="true" pageId="2" pageNumber="3">Oliarus euphranor</emphasis>
<bibRefCitation id="EFCD4B65E13CFFABD899FB31FCC81181" author="Fennah" box="[658,821,1241,1267]" lastPageId="2" lastPageNumber="47" pageId="2" pageNumber="46" refString="Fennah, R. G. (1958) Fulgoroidea of South-Eastern Polynesia. The Transactions of the Royal Entomological Society of London, 110 (6), 117 - 220." type="journal article" year="1958">Fennah, 1958</bibRefCitation>
: 131
</taxonomicName>
, Society Islands: Tahiti (Mt.
<taxonomicName id="4C5C4D17E13CFFABDEC8FB31FAFA1181" box="[1219,1287,1241,1267]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="aorai">Aorai</taxonomicName>
)
</paragraph>
<paragraph id="8BE33694E13CFFABDB03FACCFE6D104C" blockId="2.[264,1323,1316,1905]" box="[264,400,1316,1342]" pageId="2" pageNumber="3">Description</paragraph>
<paragraph id="8BE33694E13CFFABDB33FAA4FDF610C4" blockId="2.[264,1323,1316,1905]" pageId="2" pageNumber="3">Habitus (fig. 1). Small to medium size cixiids (males 4.98.0 mm, females 5.59.0 mm) with moderately depressed body form, mesonotum pentecarinate, tegmina shallowly tectiform, translucent.</paragraph>
<paragraph id="8BE33694E13CFFABDB33FA2CFE1210AC" blockId="2.[264,1323,1316,1905]" box="[312,495,1476,1502]" pageId="2" pageNumber="3">Head (figs 24)</paragraph>
<paragraph id="8BE33694E13CFFABDB33FA04FB82126C" blockId="2.[264,1323,1316,1905]" pageId="2" pageNumber="3">Vertex long, narrow, ca. 1.5 times as long as posteriorly wide, in dorsal aspect surpassing the anterior margin of the compound eyes with ca. 1/3 its total length; lateral carinae ridged, converging anteriorly; median carina obsolete; posterior margin of vertex medially deeply incised. Areolets divided by a short robust median carina. Frons separated from vertex by a distinct transverse carina. Frons ca. 1.8 times as wide as medially long, widest at level of the antennae. Lateral ocelli distinct, median frontal ocellus present yet obsolete. Frontoclypeal suture parabolic. Frons and clypeus with a distinct median carina which forks towards the apex. Rostrum in repose attaining metatrochanters.</paragraph>
<paragraph id="8BE33694E13CFFABDB33F8C7FBA51203" blockId="2.[264,1323,1316,1905]" pageId="2" pageNumber="3">Antennae small, not visible in dorsal aspect. 1st antennal segment short, ring­like, 2nd antennal segment globose, irregularly beset with sense organs (plaques).</paragraph>
<caption id="DF23661CE13DFFAADB03F9F5FCEF1341" box="[264,786,1565,1587]" pageId="3" pageNumber="4">
<paragraph id="8BE33694E13DFFAADB03F9F5FCEF1341" blockId="3.[264,786,1565,1587]" box="[264,786,1565,1587]" pageId="3" pageNumber="4">
<emphasis id="B928EA86E13DFFAADB03F9F5FE811340" bold="true" box="[264,380,1565,1586]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="4C5C4D17E13DFFAADBE5F9F6FDC91341" box="[494,564,1566,1587]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA86E13DFFAADBE5F9F6FDC91341" box="[494,564,1566,1587]" italics="true" pageId="3" pageNumber="4">Oteana</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5C4D17E13DFFAAD84BF9F6FD7B1341" box="[576,646,1566,1587]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA86E13DFFAAD84BF9F6FD7B1341" box="[576,646,1566,1587]" italics="true" pageId="3" pageNumber="4">Oteana</emphasis>
</taxonomicName>
<taxonomicName id="4C5C4D17E13DFFAAD886F9F6FD401341" box="[653,701,1566,1587]" class="Magnoliopsida" family="Primulaceae" genus="Myrsine" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="spec">spec</taxonomicName>
., female.
</paragraph>
</caption>
<paragraph id="8BE33694E13DFFAADB33F98BFE70130F" blockId="3.[264,1321,1635,1781]" box="[312,397,1635,1661]" pageId="3" pageNumber="4">Thorax</paragraph>
<paragraph id="8BE33694E13DFFAADB33F963FE4C1387" blockId="3.[264,1321,1635,1781]" pageId="3" pageNumber="4">Pronotum short, posterior margin deeply incised, only slightly wider than head, with distinct median and lateral carinae. Mesonotum pentecarinate, ca. 1.2 times as wide as medially long.</paragraph>
<caption id="DF23661CE13AFFADDB03FC7DFAE716BA" httpUri="https://zenodo.org/record/172431/files/figure.png" pageId="4" pageNumber="5" targetBox="[495,1106,296,916]" targetPageId="4">
<paragraph id="8BE33694E13AFFADDB03FC7DFAE716BA" blockId="4.[264,1323,917,968]" pageId="4" pageNumber="5">
<emphasis id="B928EA86E13AFFADDB03FC7DFE5C16D8" bold="true" box="[264,417,917,938]" pageId="4" pageNumber="5">FIGURES 24.</emphasis>
<taxonomicName id="4C5C4D17E13AFFADDBA3FC7EFE1316D9" box="[424,494,918,939]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA86E13AFFADDBA3FC7EFE1316D9" box="[424,494,918,939]" italics="true" pageId="4" pageNumber="5">Oteana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57FDE13AFFADDBFFFC7EFDBB16D9" box="[500,582,918,939]" pageId="4" pageNumber="5" rank="genus">gen.nov.</taxonomicNameLabel>
(
<taxonomicName id="4C5C4D17E13AFFADD859FC7EFC9316D9" authority="Fennah, 1958" authorityName="Fennah" authorityYear="1958" box="[594,878,918,939]" class="Insecta" family="Cixiidae" genus="Oliarus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="euphranor">
<emphasis id="B928EA86E13AFFADD859FC7EFD2916D9" box="[594,724,918,939]" italics="true" pageId="4" pageNumber="5">O. euphranor</emphasis>
(Fennah, 1958)
</taxonomicName>
, male): head. 2. Vertex, dorsal aspect; 3. Frons and clypeus, ventral aspect; 4. transition vertex/frons with areolets, anterior aspect. Scale bar equals 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BE33694E13AFFADDB33FC1EFC1711D8" blockId="4.[264,1324,1014,1896]" pageId="4" pageNumber="5">Tegmina and wings well developed. Tegmina shallowly tectiform, in repose surpassing tip of abdomen with ca. ¼ their total length. Tegmen (fig. 5) ca. 3.2 times longer than maximally wide. Apex of outer subapical cell slightly distad of apex of inner subapical cell. Intercubital transverse veinlet entering margin of tegmen distinctly distad of claval suture. Longitudinal veins inconspicuously granulate.</paragraph>
<paragraph id="8BE33694E13AFFADDB33FB5FFCA610EA" blockId="4.[264,1324,1014,1896]" pageId="4" pageNumber="5">Legs (fig. 6) Metatibiae laterally generally with 3 small spines (except for the species from Moorea which display 1 and 23 spines, respectively), apically with 6 spines (5 arranged in an oblique row, lateral spine more strongly pronounced). Metabasitarsus ca. 1.5 times longer than 2nd and 3rd metatarsal joints together, distally with 7 spines in a slightly arched row (lateral spines are longest), 2nd metatarsal joint distally with 5 spines. Metatarsal joints without macrochaetae (platellae).</paragraph>
<paragraph id="8BE33694E13AFFADDB33FA4DFDCA10CD" blockId="4.[264,1324,1014,1896]" box="[312,567,1445,1471]" pageId="4" pageNumber="5">
<collectingCountry id="F34B7604E13AFFADDB33FA4DFE8910CD" box="[312,372,1445,1471]" name="Malta" pageId="4" pageNumber="5">Male</collectingCountry>
genital complex
</paragraph>
<paragraph id="8BE33694E13AFFACDB33FA23FCBE14D8" blockId="4.[264,1324,1014,1896]" lastBlockId="5.[264,1323,284,426]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Genital segment with laterodorsal margins strongly produced caudad in a subrectangular lobe with dorsal apical angles on the right or/and the left side more or less acute; thus, genital segment either bilaterally symmetrical or asymmetrical. Medioventral process in ventral aspect triangular, with fine lateral grooves converging ventrocaudally. Anal segment in dorsal aspect highly or broadly ovate, in most species bilaterally symmetrical. Parameres slender at base, with median portion bulging, apically produced dorsolaterally, in lateral aspect hook­shaped; mediodorsal margin more or less serrate. Aedeagus (fig. 7) with shaft and flagellum well developed. Shaft ventrally and apically with a characteristic set of spines. Inter­ and intraspecific variation is observed in the configuration of these spines. Flagellum long, in repose bent dorsobasad to the left side. Flagellum membranous, with a few more strongly sclerotized portions: at least one distal spine (fig. 7, b1) and a ventral, more or less triangular ridge near flagellum bend (fig 7, b2). Phallotreme at apex of flagellum (fig. 7, b3). To point out hypothesized homologous structures of the aedeagus in the species of
<taxonomicName id="4C5C4D17E13BFFACD90CFE81FCA014F0" box="[775,861,361,386]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA86E13BFFACD90CFE81FCA014F0" box="[775,861,361,386]" italics="true" pageId="5" pageNumber="6">Oteana</emphasis>
</taxonomicName>
, the same indications used in fig. 7 are referred to in the individual species descriptions.
</paragraph>
<caption id="DF23661CE13BFFACDB03F966FB7413AC" httpUri="https://zenodo.org/record/172432/files/figure.png" pageId="5" pageNumber="6" targetBox="[431,1162,450,1659]" targetPageId="5">
<paragraph id="8BE33694E13BFFACDB03F966FB7413AC" blockId="5.[264,1323,1678,1905]" pageId="5" pageNumber="6">
<emphasis id="B928EA86E13BFFACDB03F966FE5E13D1" bold="true" box="[264,419,1678,1699]" pageId="5" pageNumber="6">FIGURES 58.</emphasis>
<taxonomicName id="4C5C4D17E13BFFACDBA2F966FE1213D1" box="[425,495,1678,1699]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA86E13BFFACDBA2F966FE1213D1" box="[425,495,1678,1699]" italics="true" pageId="5" pageNumber="6">Oteana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57FDE13BFFACDBFEF966FDB013D1" box="[501,589,1678,1699]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
: 5. right tegmen (
<taxonomicName id="4C5C4D17E13BFFACD8F5F966FC7D13D1" box="[766,896,1678,1699]" class="Insecta" family="Cixiidae" genus="Oliarus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="euphranor">
<emphasis id="B928EA86E13BFFACD8F5F966FC7D13D1" box="[766,896,1678,1699]" italics="true" pageId="5" pageNumber="6">O. euphranor</emphasis>
</taxonomicName>
, female); 6. right hindleg, ventral aspect (
<taxonomicName id="4C5C4D17E13BFFACDF18F966FE9113B3" class="Insecta" family="Cixiidae" genus="Oliarus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="euphranor">
<emphasis id="B928EA86E13BFFACDF18F966FE9113B3" italics="true" pageId="5" pageNumber="6">O. euphranor</emphasis>
</taxonomicName>
, female); 7. configuration of the male copulatory organ, the aedeagus (
<taxonomicName id="4C5C4D17E13BFFACDE13F944FB9813B3" box="[1048,1125,1708,1729]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mato">
<emphasis id="B928EA86E13BFFACDE13F944FB9813B3" box="[1048,1125,1708,1729]" italics="true" pageId="5" pageNumber="6">O. mato</emphasis>
</taxonomicName>
); 8. external aspects of female genitalia (
<taxonomicName id="4C5C4D17E13BFFACDBC1F921FDED13AC" box="[458,528,1737,1758]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA86E13BFFACDBC1F921FDED13AC" box="[458,528,1737,1758]" italics="true" pageId="5" pageNumber="6">Oteana</emphasis>
</taxonomicName>
<taxonomicName id="4C5C4D17E13BFFACD81DF921FDBA13AC" box="[534,583,1737,1758]" class="Magnoliopsida" family="Primulaceae" genus="Myrsine" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="spec">spec</taxonomicName>
.). Scale bars equal 0.5 mm (figs 5, 6, 8) and 0.1 mm (fig. 7).
</paragraph>
</caption>
<paragraph id="8BE33694E13BFFACDB33F90EFE6E1203" blockId="5.[264,1323,1678,1905]" pageId="5" pageNumber="6">
Abbreviations: a, shaft (“periandrium” sensu
<bibRefCitation id="EFCD4B65E13BFFACD8E5F90EFC931389" author="Giffard" box="[750,878,1766,1787]" pageId="5" pageNumber="47" refString="Giffard, W. M. (1925) A review of the Hawaiian Cixiidae, with descriptions of species (Homoptera). Proceedings of the Hawaiian Entomological Society, 6, 1 - 121." type="journal article" year="1925">Giffard 1925</bibRefCitation>
); b, flagellum (“phallus” sensu
<bibRefCitation id="EFCD4B65E13BFFACDEABF90EFADD1389" author="Giffard" box="[1184,1312,1766,1787]" pageId="5" pageNumber="47" refString="Giffard, W. M. (1925) A review of the Hawaiian Cixiidae, with descriptions of species (Homoptera). Proceedings of the Hawaiian Entomological Society, 6, 1 - 121." type="journal article" year="1925">Giffard 1925</bibRefCitation>
); c, central part (“apodeme of phallus” sensu
<bibRefCitation id="EFCD4B65E13BFFACD8ACF8ECFCDB126B" author="Giffard" box="[679,806,1796,1817]" pageId="5" pageNumber="47" refString="Giffard, W. M. (1925) A review of the Hawaiian Cixiidae, with descriptions of species (Homoptera). Proceedings of the Hawaiian Entomological Society, 6, 1 - 121." type="journal article" year="1925">Giffard 1925</bibRefCitation>
); a1a4, ventral spines of shaft (a1, right lateral spine; a2, median right spine; a3, median left spine; a4, left lateral spine); a5, apical shaft spine; a6, subapical shaft spine; b1, distal flagellum spine; b2, sclerotized ridge of flagellum; b3, phallotreme (“functional orifice” sensu
<bibRefCitation id="EFCD4B65E13BFFACDB03F8B4FE751203" author="Giffard" box="[264,392,1884,1905]" pageId="5" pageNumber="47" refString="Giffard, W. M. (1925) A review of the Hawaiian Cixiidae, with descriptions of species (Homoptera). Proceedings of the Hawaiian Entomological Society, 6, 1 - 121." type="journal article" year="1925">Giffard 1925</bibRefCitation>
).
</paragraph>
<paragraph id="8BE33694E138FFAFDB33FEF4FDAE1444" blockId="6.[264,1323,284,473]" box="[312,595,284,310]" pageId="6" pageNumber="7">Female genitalia (fig. 8)</paragraph>
<paragraph id="8BE33694E138FFAFDB33FEACFBF514AB" blockId="6.[264,1323,284,473]" pageId="6" pageNumber="7">Caudal margin of 7th sternite shallowly sinuate, medially slightly concave. Ovipositor short, stout, in repose directed straight caudad. 9th tergite truncate, caudally concave, with wax producing area distinct, medially not divided. Anal segment plate­shaped, ventrally concave, wide at base. Ductus receptaculi tubular, not heliciform.</paragraph>
<paragraph id="8BE33694E138FFAFDB03FDE7FE76175B" blockId="6.[264,1321,527,593]" box="[264,395,527,553]" pageId="6" pageNumber="7">Etymology</paragraph>
<paragraph id="8BE33694E138FFAFDB33FDDFFAD41723" blockId="6.[264,1321,527,593]" box="[312,1321,567,593]" pageId="6" pageNumber="7">
Named after the traditional drum dance from Tahiti, the
<emphasis id="B928EA86E138FFAFD9C1FDDFFBFE1722" box="[970,1027,567,592]" italics="true" pageId="6" pageNumber="7">Otea</emphasis>
. The gender is feminine.
</paragraph>
<paragraph id="8BE33694E138FFAFDB03FD6FFE8D17D3" blockId="6.[264,1323,647,833]" box="[264,368,647,673]" pageId="6" pageNumber="7">Remarks</paragraph>
<paragraph id="8BE33694E138FFAFDB33FD47FE9F1633" blockId="6.[264,1323,647,833]" pageId="6" pageNumber="7">
<taxonomicName id="4C5C4D17E138FFAFDB33FD47FE7217BA" box="[312,399,687,712]" class="Insecta" family="Cixiidae" genus="Oteana" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA86E138FFAFDB33FD47FE7217BA" box="[312,399,687,712]" italics="true" pageId="6" pageNumber="7">Oteana</emphasis>
</taxonomicName>
can be distinguished from other pentastirine genera by bodily proportions, shape and arrangement of spines of the metatarsi (7, 5, no platellae), and especially by the configuration of the male genitalia (arrangement of ventral and apical aedeagal shaft spines).
</paragraph>
</subSubSection>
</treatment>
</document>