261 lines
37 KiB
XML
261 lines
37 KiB
XML
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<mods:title id="274D92A9582E08FEB37285A08217BD47">Resolving taxonomic issues of cicadas (Hemiptera: Cicadidae) including new combinations, new synonymies, and revised status, with updates on the diversity of the Brazilian cicada fauna and new records for four South American countries</mods:title>
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<mods:namePart id="67C72BBF1A9A0410DDC4738C3401C4D3">Sanborn, Allen F.</mods:namePart>
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<mods:title id="30DB232B5E9609CB9AF8FF88DF1C36C3">Zootaxa</mods:title>
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<mods:part id="7848732AF1FEFA503FB392B02EEC1D6D">
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<mods:date id="A47EA0A674EB53CBB8086BA38C58BB48">2023</mods:date>
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<mods:number id="744A020C83FFB6B532CB1BE33D4D1258">2023-07-20</mods:number>
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<mods:identifier id="F4AAA092BE8B2122EA47530B2CC8A64D" type="DOI">10.11646/zootaxa.5318.3.2</mods:identifier>
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<treatment id="03E2878BFFE05921FF50F9ECFC5CFE33" ID-DOI="http://doi.org/10.5281/zenodo.8166924" ID-GBIF-Taxon="212427698" ID-Zenodo-Dep="8166924" LSID="urn:lsid:plazi:treatment:03E2878BFFE05921FF50F9ECFC5CFE33" httpUri="http://treatment.plazi.org/id/03E2878BFFE05921FF50F9ECFC5CFE33" lastPageId="12" lastPageNumber="351" pageId="10" pageNumber="349">
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<subSubSection id="C3516516FFE05927FF50F9ECFD17FC2A" box="[151,739,941,969]" pageId="10" pageNumber="349" type="nomenclature">
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<paragraph id="8BF4369DFFE05927FF50F9ECFD17FC2A" blockId="10.[151,739,941,969]" box="[151,739,941,969]" pageId="10" pageNumber="349">
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<heading id="D0BC81F1FFE05927FF50F9ECFD17FC2A" bold="true" box="[151,739,941,969]" fontSize="11" level="1" pageId="10" pageNumber="349" reason="1">
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<emphasis id="B93FEA8FFFE05927FF50F9ECFD17FC2A" bold="true" box="[151,739,941,969]" pageId="10" pageNumber="349">
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<taxonomicName id="4C4B4D1EFFE05927FF50F9ECFDF2FC2A" ID-CoL="659aca92-bea6-48f8-b5da-0961de0042b2" authority="(Stal, 1854)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Stal" baseAuthorityYear="1854" box="[151,518,941,969]" class="Insecta" family="Cicadidae" genus="Herrera" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="species" species="sahlbergi" status="comb. nov.">
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<emphasis id="B93FEA8FFFE05927FF50F9ECFE84FC2B" bold="true" box="[151,368,942,969]" italics="true" pageId="10" pageNumber="349">Herrera sahlbergi</emphasis>
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(
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<bibRefCitation id="EFDA4B6CFFE05927FE47F9EEFE09FC2A" author="Stal" box="[384,509,941,968]" firstAuthor="Stal" pageId="10" pageNumber="349" pagination="231 - 255" refId="ref16654" refString="Stal, C. (1854) Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 11, 231 - 255." type="journal article" year="1854">Stål, 1854</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A20C57F4FFE05927FDCAF9ECFD17FC2A" box="[525,739,942,969]" pageId="10" pageNumber="349" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516516FFE05927FF50F9B0FDA7FBE9" box="[151,595,1011,1035]" pageId="10" pageNumber="349" type="reference_group">
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<paragraph id="8BF4369DFFE05927FF50F9B0FDA7FBE9" blockId="10.[151,595,1011,1035]" box="[151,595,1011,1035]" pageId="10" pageNumber="349">
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<taxonomicName id="4C4B4D1EFFE05927FF50F9B0FDBBFBE9" ID-CoL="ab9b2838-dd69-4298-a022-a92b593c52c1" authority="Stal, 1854" authorityName="Stal" authorityYear="1854" box="[151,591,1011,1035]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="species" species="sahlbergi">
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<treatmentCitationGroup id="AB5B11B3FFE05927FF50F9B0FE12FBE9" box="[151,486,1011,1035]" pageId="10" pageNumber="349">
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<treatmentCitation id="0AEA108CFFE05927FF50F9B0FE12FBE9" author="Stal, C." box="[151,486,1011,1035]" page="243" pageId="10" pageNumber="349" year="1854">
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<emphasis id="B93FEA8FFFE05927FF50F9B0FEB2FBE9" box="[151,326,1011,1035]" italics="true" pageId="10" pageNumber="349">Cicada sahlbergi</emphasis>
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<bibRefCitation id="EFDA4B6CFFE05927FE8BF9B0FE12FBE9" author="Stal" box="[332,486,1011,1035]" firstAuthor="Stal" page="243" pageId="10" pageNumber="349" pagination="231 - 255" refId="ref16654" refString="Stal, C. (1854) Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 11, 231 - 255." type="journal article" year="1854">Stål, 1854: 243</bibRefCitation>
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</treatmentCitation>
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</treatmentCitationGroup>
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(Brasilia)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516516FFE05921FF50FE7AFAF7FE4F" lastPageId="12" lastPageNumber="351" pageId="10" pageNumber="349" type="discussion">
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<paragraph id="8BF4369DFFE05927FF50FE7AFE72FA5E" blockId="10.[151,1437,1081,2008]" pageId="10" pageNumber="349">
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<emphasis id="B93FEA8FFFE05927FF50FE7AFEFAFBB1" bold="true" box="[151,270,1081,1107]" pageId="10" pageNumber="349">REMARKS.</emphasis>
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A species missing from the Brazilian faunal list in
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<bibRefCitation id="EFDA4B6CFFE05927FC9FFE79FBC1FBB1" author="Nunes" box="[856,1077,1081,1108]" etAl="et al." firstAuthor="Nunes" pageId="10" pageNumber="349" refId="ref14077" refString="Nunes, V. L., Ruschel, T. P., Maccagnan, D. H. B., Simies, P. C. & Acosta, R. C. (2023) Cigarras do Brasil, Guia Fotographica. 1 a edicao. ebook. 46 pp. [https: // zenodo. org / record / 7712483]" type="book" year="2023">
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Nunes
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<emphasis id="B93FEA8FFFE05927FC6FFE79FC2FFBB1" box="[936,987,1081,1107]" italics="true" pageId="10" pageNumber="349">et al</emphasis>
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. (2023)
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</bibRefCitation>
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is
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<taxonomicName id="4C4B4D1EFFE05927FB9FFE7AFA6CFBB6" authority="Stal, 1854" authorityName="Stal" authorityYear="1854" box="[1112,1432,1081,1108]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="species" species="sahlbergi">
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<emphasis id="B93FEA8FFFE05927FB9FFE7AFAEEFBB1" box="[1112,1306,1081,1107]" italics="true" pageId="10" pageNumber="349">Cicada sahlbergi</emphasis>
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<bibRefCitation id="EFDA4B6CFFE05927FAE5FE7AFA6CFBB6" author="Stal" box="[1314,1432,1081,1108]" firstAuthor="Stal" pageId="10" pageNumber="349" pagination="231 - 255" refId="ref16654" refString="Stal, C. (1854) Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 11, 231 - 255." type="journal article" year="1854">Stål, 1854</bibRefCitation>
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</taxonomicName>
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.
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<bibRefCitation id="EFDA4B6CFFE05927FF50FE1EFEA5FB95" author="Metcalf" box="[151,337,1117,1143]" firstAuthor="Metcalf" pageId="10" pageNumber="349" pagination="587 - 919" refId="ref13934" refString="Metcalf, Z. P. (1963 b) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section II. Gaeninae and Cicadinae. North Carolina State College Contribution, 1502, 587 - 919." type="journal article" year="1963" yearSuffix="b">Metcalf (1963b)</bibRefCitation>
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placed
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<taxonomicName id="4C4B4D1EFFE05927FE6AFE1EFD1BFB9A" authority="Stal, 1854" authorityName="Stal" authorityYear="1854" box="[429,751,1117,1144]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="species" species="sahlbergi">
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<emphasis id="B93FEA8FFFE05927FE6AFE1EFD85FB95" box="[429,625,1117,1143]" italics="true" pageId="10" pageNumber="349">Cicada sahlbergi</emphasis>
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<bibRefCitation id="EFDA4B6CFFE05927FDBDFE1EFD1BFB9A" author="Stal" box="[634,751,1117,1144]" firstAuthor="Stal" pageId="10" pageNumber="349" pagination="231 - 255" refId="ref16654" refString="Stal, C. (1854) Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 11, 231 - 255." type="journal article" year="1854">
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Stål,
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<quantity id="4CB39B78FFE05927FD70FE1EFD1BFB9A" box="[695,751,1117,1144]" metricMagnitude="1" metricUnit="m" metricValue="4.70916" pageId="10" pageNumber="349" unit="in" value="1854.0">1854</quantity>
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</bibRefCitation>
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</taxonomicName>
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in his list of “Species of Uncertain Position” (Metcalf was suggesting the species actually belong in a different genus) under the genus
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<taxonomicName id="4C4B4D1EFFE05927FBD7FEC2FAEEFB7E" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[1040,1306,1153,1180]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA8FFFE05927FBD7FEC2FB95FB79" box="[1040,1121,1153,1179]" italics="true" pageId="10" pageNumber="349">Cicada</emphasis>
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<bibRefCitation id="EFDA4B6CFFE05927FBACFEC2FAEEFB7E" author="Linnaeus" box="[1131,1306,1153,1180]" firstAuthor="Linnaeus" pageId="10" pageNumber="349" refId="ref13514" refString="Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tomus I. Edito duodecima, reformata. Laurentius Salvius, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 157601" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
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</taxonomicName>
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suggesting he did not believe the species belonged in
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<taxonomicName id="4C4B4D1EFFE05927FDBBFEE6FD39FB5D" authorityName="Linnaeus" authorityYear="1758" box="[636,717,1189,1215]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA8FFFE05927FDBBFEE6FD39FB5D" box="[636,717,1189,1215]" italics="true" pageId="10" pageNumber="349">Cicada</emphasis>
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</taxonomicName>
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. The
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<typeStatus id="54F0883FFFE05927FCD7FEE6FC86FB5D" box="[784,882,1189,1215]" pageId="10" pageNumber="349" type="holotype">holotype</typeStatus>
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(“Typus”) was located in the NHRS and studied to determine its actual taxonomic position (
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<figureCitation id="13702A18FFE05927FD4DFE8AFD24FB01" box="[650,720,1225,1251]" captionStart="FIGURE 7" captionStartId="11.[152,255,1884,1908]" captionTargetBox="[151,1436,181,1832]" captionTargetId="figure-15@11.[151,1436,181,1860]" captionTargetPageId="11" captionText="FIGURE 7. Cicada sahlbergi Stål, 1854 type specimen (NHRM) dorsal (top) and ventral (middle) habitus. ©Naturhistoriska Riksmuseet, Stockholm. Photos by the author." figureDoi="http://doi.org/10.5281/zenodo.8166916" httpUri="https://zenodo.org/record/8166916/files/figure.png" pageId="10" pageNumber="349">Fig. 7</figureCitation>
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). The species was housed in the NHRS collection as
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<taxonomicName id="4C4B4D1EFFE05927FAFEFE89FEF4FAE5" class="Insecta" family="Cicadidae" genus="Prunasis" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="species" species="sahlbergi">
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<emphasis id="B93FEA8FFFE05927FAFEFE89FEF4FAE5" italics="true" pageId="10" pageNumber="349">Prunasis sahlbergi</emphasis>
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</taxonomicName>
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(http://www3.nrm.se/en/homoptera_nrm/s/homopts.html) although no official change to its generic assignment has been published since it was described as a species of
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<taxonomicName id="4C4B4D1EFFE05927FC6CFF52FC08FAC9" authorityName="Linnaeus" authorityYear="1758" box="[939,1020,1297,1323]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA8FFFE05927FC6CFF52FC08FAC9" box="[939,1020,1297,1323]" italics="true" pageId="10" pageNumber="349">Cicada</emphasis>
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</taxonomicName>
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. However, the species has six apical cells in the hindwings and all New World genera of Parnisini
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<bibRefCitation id="EFDA4B6CFFE05927FC94FF76FC08FAB2" author="Distant" box="[851,1020,1333,1360]" firstAuthor="Distant" pageId="10" pageNumber="349" pagination="203 - 216" refId="ref12367" refString="Distant, W. L. (1905 b) Rhynchotal notes XXXIV. Annals and Magazine of Natural History, Series 7, 16, 203 - 216. https: // doi. org / 10.1080 / 03745480509443671" type="journal article" year="1905" yearSuffix="b">Distant, 1905b</bibRefCitation>
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, including
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<taxonomicName id="4C4B4D1EFFE05927FBBDFF75FAAFFAB2" authority="Stal, 1854" authorityName="Stal" authorityYear="1854" box="[1146,1371,1333,1360]" class="Insecta" family="Cicadidae" genus="Prunasis" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA8FFFE05927FBBDFF75FB2AFAAD" box="[1146,1246,1334,1359]" italics="true" pageId="10" pageNumber="349">Prunasis</emphasis>
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<bibRefCitation id="EFDA4B6CFFE05927FB21FF76FAAFFAB2" author="Stal" box="[1254,1371,1333,1360]" firstAuthor="Stal" pageId="10" pageNumber="349" pagination="231 - 255" refId="ref16654" refString="Stal, C. (1854) Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 11, 231 - 255." type="journal article" year="1854">Stål, 1854</bibRefCitation>
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</taxonomicName>
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, have five hindwing apical cells with the exception of the two hindwing apical cells found in
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<taxonomicName id="4C4B4D1EFFE05927FBBBFF19FA6CFA96" authority="Torres, 1958" authorityName="Torres" authorityYear="1958" box="[1148,1432,1369,1396]" class="Insecta" family="Cicadidae" genus="Acyroneura" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA8FFFE05927FBBBFF19FB0BFA96" box="[1148,1279,1370,1396]" italics="true" pageId="10" pageNumber="349">Acyroneura</emphasis>
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<bibRefCitation id="EFDA4B6CFFE05927FAC0FF19FA6CFA96" author="Torres" box="[1287,1432,1369,1396]" firstAuthor="Torres" pageId="10" pageNumber="349" pagination="17 - 26" refId="ref16783" refString="Torres, B. A. (1958) Nuevos generos Acyroneura y Acuticephala, Guaranisaria bicolor nueva especie (Homoptera-Cicadidae). Neotropica, 4, 17 - 26." type="journal article" year="1958">Torres, 1958</bibRefCitation>
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</taxonomicName>
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. The six apical cells of the hindwing eliminate
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<taxonomicName id="4C4B4D1EFFE05927FD5BFF3DFCF4FA75" authorityName="Stal" authorityYear="1854" box="[668,768,1406,1431]" class="Insecta" family="Cicadidae" genus="Prunasis" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA8FFFE05927FD5BFF3DFCF4FA75" box="[668,768,1406,1431]" italics="true" pageId="10" pageNumber="349">Prunasis</emphasis>
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</taxonomicName>
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and the Parnisini as the correct generic or tribal assignment for
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<taxonomicName id="4C4B4D1EFFE05927FF78FFE2FE74FA59" authorityName="Stal" authorityYear="1854" box="[191,384,1441,1467]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="species" species="sahlbergi">
|
|
<emphasis id="B93FEA8FFFE05927FF78FFE2FE74FA59" box="[191,384,1441,1467]" italics="true" pageId="10" pageNumber="349">Cicada sahlbergi</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BF4369DFFE05927FF00FF86FB49F83A" blockId="10.[151,1437,1081,2008]" pageId="10" pageNumber="349">
|
|
The
|
|
<typeStatus id="54F0883FFFE05927FF38FF86FE95FA3D" box="[255,353,1477,1503]" pageId="10" pageNumber="349" type="holotype">holotype</typeStatus>
|
|
of
|
|
<taxonomicName id="4C4B4D1EFFE05927FE57FF86FDA2FA3D" authorityName="Stal" authorityYear="1854" box="[400,598,1477,1503]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="349" phylum="Arthropoda" rank="species" species="sahlbergi">
|
|
<emphasis id="B93FEA8FFFE05927FE57FF86FDA2FA3D" box="[400,598,1477,1503]" italics="true" pageId="10" pageNumber="349">Cicada sahlbergi</emphasis>
|
|
</taxonomicName>
|
|
is a female so many diagnostic structures are not available to help in determining the correct generic assignment. However, the metanotum is partially visible at the dorsal midline, fore wing veins CuP and 1A are fused in part, and hindwing veins RP and M are fused at their bases. In addition, the operculum is not S-shaped, the lateral margin is not concave, and it does not encapsulate the meracanthus. These characters eliminate the Tibicininae
|
|
<bibRefCitation id="EFDA4B6CFFE05927FDF6FC16FD21F992" author="Distant" box="[561,725,1621,1648]" firstAuthor="Distant" pageId="10" pageNumber="349" pagination="22 - 35" refId="ref12327" refString="Distant, W. L. (1905 a) Rhynchotal notes XXXIII. Annals and Magazine of Natural History, Series 7, 16, 22 - 35. https: // doi. org / 10.1080 / 03745480509443650" type="journal article" year="1905" yearSuffix="a">Distant, 1905a</bibRefCitation>
|
|
as a subfamily assignment for the species. The Tettigomyiinae
|
|
<bibRefCitation id="EFDA4B6CFFE05927FF50FC3AFECAF976" author="Distant" box="[151,318,1657,1684]" firstAuthor="Distant" pageId="10" pageNumber="349" pagination="265 - 281" refId="ref12407" refString="Distant, W. L. (1905 c) Rhynchotal notes XXXV. Annals and Magazine of Natural History, Series 7, 16, 265 - 281. https: // doi. org / 10.1080 / 03745480509442862" type="journal article" year="1905" yearSuffix="c">Distant, 1905c</bibRefCitation>
|
|
is an African subfamily (
|
|
<bibRefCitation id="EFDA4B6CFFE05927FDACFC3AFCAFF971" author="Marshall" box="[619,859,1657,1684]" etAl="et al." firstAuthor="Marshall" pageId="10" pageNumber="349" pagination="1 - 64" refId="ref13722" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. & Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
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|
Marshall
|
|
<emphasis id="B93FEA8FFFE05927FD1CFC39FCE4F971" box="[731,784,1657,1683]" italics="true" pageId="10" pageNumber="349">et al</emphasis>
|
|
. 2018
|
|
</bibRefCitation>
|
|
) and the Derotettinae Moulds, 2019 (in
|
|
<bibRefCitation id="EFDA4B6CFFE05927FAF3FC3AFF03F955" author="Simon" etAl="et al." firstAuthor="Simon" pageId="10" pageNumber="349" pagination="865 - 886" refId="ref16508" refString="Simon, C., Gordon, E. R., Moulds, M. S., Cole, J., Haji, D., Lemmon, A. R., Lemmon, E. M., Kortyna, M., Nazario, K., Wade, E. J., Meister, R., Goemans, G., Chiswell, S. M., Pessacq, P., Veloso, C., McCutchen, J. P. & Lukasik, P. (2019) Off-target capture data, endosymbiont genes and morphology reveal a relict lineage sister to all other singing cicadas. Biological Journal of the Linnean Society, 128, 865 - 886. https: // doi. org / 10.1093 / biolinnean / blz 120" type="journal article" year="2019">
|
|
Simon
|
|
<emphasis id="B93FEA8FFFE05927FA4FFC39FF59F955" italics="true" pageId="10" pageNumber="349">et al</emphasis>
|
|
. 2019
|
|
</bibRefCitation>
|
|
) contains a single genus of primitive cicadas found only in a limited region of
|
|
<collectingCountry id="F35C760DFFE05927FB4AFCDEFB08F955" box="[1165,1276,1693,1719]" name="Argentina" pageId="10" pageNumber="349">Argentina</collectingCountry>
|
|
(
|
|
<bibRefCitation id="EFDA4B6CFFE05927FACAFCDEFF25F939" author="Simon" etAl="et al." firstAuthor="Simon" pageId="10" pageNumber="349" pagination="865 - 886" refId="ref16508" refString="Simon, C., Gordon, E. R., Moulds, M. S., Cole, J., Haji, D., Lemmon, A. R., Lemmon, E. M., Kortyna, M., Nazario, K., Wade, E. J., Meister, R., Goemans, G., Chiswell, S. M., Pessacq, P., Veloso, C., McCutchen, J. P. & Lukasik, P. (2019) Off-target capture data, endosymbiont genes and morphology reveal a relict lineage sister to all other singing cicadas. Biological Journal of the Linnean Society, 128, 865 - 886. https: // doi. org / 10.1093 / biolinnean / blz 120" type="journal article" year="2019">
|
|
Simon
|
|
<emphasis id="B93FEA8FFFE05927FAA5FCDDFA61F955" box="[1378,1429,1693,1719]" italics="true" pageId="10" pageNumber="349">et al</emphasis>
|
|
. 2019
|
|
</bibRefCitation>
|
|
) so they can thus be eliminated for the Brazilian species. Of the Neotropical tribes remaining,
|
|
<taxonomicName id="4C4B4D1EFFE05927FAE7FC82FECFF8E2" authority="Distant, 1905d" authorityName="Distant" authorityYear="1905" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="359" phylum="Arthropoda" rank="tribe" tribe="Zammarini">
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Zammarini
|
|
<bibRefCitation id="EFDA4B6CFFE05927FF50FCA6FECFF8E2" author="Distant" box="[151,315,1765,1792]" firstAuthor="Distant" pageId="10" pageNumber="349" pagination="379 - 387" refId="ref12447" refString="Distant, W. L. (1905 d) Rhynchotal notes XXXI. Annals and Magazine of Natural History, Series 7, 15, 379 - 387. https: // doi. org / 10.1080 / 03745480509443064" type="journal article" year="1905" yearSuffix="d">Distant, 1905d</bibRefCitation>
|
|
</taxonomicName>
|
|
and Fidicinini can be eliminated by the hindcoxae with a large inner protuberance in species of these tribes, Plautillini
|
|
<bibRefCitation id="EFDA4B6CFFE05927FE9AFD4AFE01F8C1" author="Distant" box="[349,501,1801,1827]" firstAuthor="Distant" pageId="10" pageNumber="349" refId="ref12567" refString="Distant, W. L. (1906) A synonymic catalogue of Homoptera. Part I. Cicadidae. Trustees of the British Museum, London, 207 pp. https: // doi. org / 10.5962 / bhl. title. 8554" type="book" year="1906">Distant, 1906</bibRefCitation>
|
|
can be eliminated by the large fore wings and lateral expansion of the pronotum, Durangonini Moulds &
|
|
<bibRefCitation id="EFDA4B6CFFE05927FE66FD6EFDBDF8A5" author="Marshall" box="[417,585,1837,1863]" firstAuthor="Marshall" pageId="10" pageNumber="349" pagination="1 - 64" refId="ref13722" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. & Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">Marshall, 2018</bibRefCitation>
|
|
(in
|
|
<bibRefCitation id="EFDA4B6CFFE05927FDB4FD6EFCA7F8A5" author="Marshall" box="[627,851,1837,1864]" etAl="et al." firstAuthor="Marshall" pageId="10" pageNumber="349" pagination="1 - 64" refId="ref13722" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. & Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
|
|
Marshall
|
|
<emphasis id="B93FEA8FFFE05927FD1AFD6DFCF9F8A5" box="[733,781,1837,1863]" italics="true" pageId="10" pageNumber="349">et al</emphasis>
|
|
. 2018
|
|
</bibRefCitation>
|
|
) can be eliminated by the abutting RA1 and subcosta in the fore wing and the narrow anal cell of the hindwing, Chlorocystini
|
|
<bibRefCitation id="EFDA4B6CFFE05927FC11FD12FB88F88E" author="Distant" box="[982,1148,1873,1900]" firstAuthor="Distant" pageId="10" pageNumber="349" pagination="203 - 216" refId="ref12367" refString="Distant, W. L. (1905 b) Rhynchotal notes XXXIV. Annals and Magazine of Natural History, Series 7, 16, 203 - 216. https: // doi. org / 10.1080 / 03745480509443671" type="journal article" year="1905" yearSuffix="b">Distant, 1905b</bibRefCitation>
|
|
can be eliminated by the very narrow hind margin of the fore wing, narrow cruciform elevation, and angularly swollen ventral postclypeus, and Taphurini can be eliminated by the head being as wide or wider than the mesonotum (
|
|
<bibRefCitation id="EFDA4B6CFFE05927FBB8FDDAFAE0F851" author="Distant" box="[1151,1300,1945,1971]" firstAuthor="Distant" pageId="10" pageNumber="349" refId="ref12567" refString="Distant, W. L. (1906) A synonymic catalogue of Homoptera. Part I. Cicadidae. Trustees of the British Museum, London, 207 pp. https: // doi. org / 10.5962 / bhl. title. 8554" type="book" year="1906">Distant 1906</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFDA4B6CFFE05927FAD8FDD9FA63F856" author="Boer" box="[1311,1431,1945,1972]" firstAuthor="Boer" pageId="10" pageNumber="349" pagination="201 - 231" refId="ref11743" refString="Boer, A. J. de (1995) The phylogeny and taxonomy of the Chlorocystini (sensu stricto) (Homoptera, Tibicinidae). Contributions to Zoology, 65, 201 - 231. https: // doi. org / 10.1163 / 26660644 - 06504001" type="journal article" year="1995">Boer 1995</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFDA4B6CFFE05927FF50FDFEFEDBF83A" author="Moulds" box="[151,303,1981,2008]" firstAuthor="Moulds" pageId="10" pageNumber="349" pagination="375 - 446" refId="ref14021" refString="Moulds, M. S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375 - 446. https: // doi. org / 10.3853 / j. 0067 - 1975.57.2005.1447" type="journal article" year="2005">Moulds 2005</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFDA4B6CFFE05927FEFCFDFEFDD5F835" author="Marshall" box="[315,545,1981,2008]" etAl="et al." firstAuthor="Marshall" pageId="10" pageNumber="349" pagination="1 - 64" refId="ref13722" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. & Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
|
|
Marshall
|
|
<emphasis id="B93FEA8FFFE05927FE62FDFDFE22F835" box="[421,470,1981,2007]" italics="true" pageId="10" pageNumber="349">et al</emphasis>
|
|
. 2018
|
|
</bibRefCitation>
|
|
). The remaining available tribe is Carinetini
|
|
<bibRefCitation id="EFDA4B6CFFE05927FBD2FDFEFB4CF83A" author="Distant" box="[1045,1208,1981,2008]" firstAuthor="Distant" pageId="10" pageNumber="349" pagination="478 - 486" refId="ref12527" refString="Distant, W. L. (1905 f) Rhynchotal notes XXXII. Annals and Magazine of Natural History, Series 7, 15, 478 - 486. https: // doi. org / 10.1080 / 03745480509442837" type="journal article" year="1905" yearSuffix="f">Distant, 1905f</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
<caption id="DF346615FFE15926FF5FFD1FFD9FF87A" ID-DOI="http://doi.org/10.5281/zenodo.8166916" ID-Zenodo-Dep="8166916" httpUri="https://zenodo.org/record/8166916/files/figure.png" pageId="11" pageNumber="350" startId="11.[152,255,1884,1908]" targetBox="[151,1436,181,1832]" targetPageId="11" targetType="figure">
|
|
<paragraph id="8BF4369DFFE15926FF5FFD1FFD9FF87A" blockId="11.[151,1437,1884,1944]" pageId="11" pageNumber="350">
|
|
<emphasis id="B93FEA8FFFE15926FF5FFD1FFEEDF896" bold="true" box="[152,281,1884,1908]" pageId="11" pageNumber="350">FIGURE 7.</emphasis>
|
|
<taxonomicName id="4C4B4D1EFFE15926FEE7FD1FFDCAF896" authority="Stal, 1854" authorityName="Stal" authorityYear="1854" box="[288,574,1884,1908]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="350" phylum="Arthropoda" rank="species" species="sahlbergi">
|
|
<emphasis id="B93FEA8FFFE15926FEE7FD1FFE3BF896" box="[288,463,1884,1908]" italics="true" pageId="11" pageNumber="350">Cicada sahlbergi</emphasis>
|
|
<bibRefCitation id="EFDA4B6CFFE15926FE11FD1FFDCAF896" author="Stal" box="[470,574,1884,1908]" firstAuthor="Stal" pageId="11" pageNumber="350" pagination="231 - 255" refId="ref16654" refString="Stal, C. (1854) Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 11, 231 - 255." type="journal article" year="1854">Stål, 1854</bibRefCitation>
|
|
</taxonomicName>
|
|
type specimen (NHRM) dorsal (top) and ventral (middle) habitus. ©Naturhistoriska Riksmuseet, Stockholm. Photos by the author.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BF4369DFFE65921FF00FAD4FAF7FE4F" blockId="12.[151,1437,151,465]" pageId="12" pageNumber="351">
|
|
The body of
|
|
<taxonomicName id="4C4B4D1EFFE65921FE9DFAD4FDE8FF53" authorityName="Stal" authorityYear="1854" box="[346,540,151,177]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="species" species="sahlbergi">
|
|
<emphasis id="B93FEA8FFFE65921FE9DFAD4FDE8FF53" box="[346,540,151,177]" italics="true" pageId="12" pageNumber="351">Cicada sahlbergi</emphasis>
|
|
</taxonomicName>
|
|
tapers anteriorly and posteriorly, the pronotum has oblique lateral margins and is shorter than the mesonotum, and mainly hyaline wings as outlined by
|
|
<bibRefCitation id="EFDA4B6CFFE65921FC78FAF8FB9EFF37" author="Distant" box="[959,1130,186,213]" firstAuthor="Distant" pageId="12" pageNumber="351" pagination="478 - 486" refId="ref12527" refString="Distant, W. L. (1905 f) Rhynchotal notes XXXII. Annals and Magazine of Natural History, Series 7, 15, 478 - 486. https: // doi. org / 10.1080 / 03745480509442837" type="journal article" year="1905" yearSuffix="f">Distant (1905f)</bibRefCitation>
|
|
for species of the tribe (the tribal characteristics have not been updated with the addition of genera since the tribe was formed). The head being about as wide as the mesonotum, the vertex slightly longer than the front, the pronotum shorter than the mesonotum with the lateral margins slightly oblique, the abdomen about as long as the distance between the apex of the head and the cruciform elevation, the fore wings less than three times longer than broad, and the dorsal curvature of female abdominal segment 9 place the species in the genus
|
|
<taxonomicName id="4C4B4D1EFFE65921FD10FB2CFC2EFE6B" authority="Distant, 1905 f" authorityName="Distant" authorityYear="1905" box="[727,986,366,393]" class="Insecta" family="Cicadidae" genus="Herrera" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B93FEA8FFFE65921FD10FB2CFCC6FE6A" box="[727,818,367,392]" italics="true" pageId="12" pageNumber="351">Herrera</emphasis>
|
|
<bibRefCitation id="EFDA4B6CFFE65921FCFFFB2CFC2EFE6B" author="Distant" box="[824,986,366,393]" firstAuthor="Distant" pageId="12" pageNumber="351" pagination="478 - 486" refId="ref12527" refString="Distant, W. L. (1905 f) Rhynchotal notes XXXII. Annals and Magazine of Natural History, Series 7, 15, 478 - 486. https: // doi. org / 10.1080 / 03745480509442837" type="journal article" year="1905" yearSuffix="f">Distant, 1905f</bibRefCitation>
|
|
</taxonomicName>
|
|
. As a result,
|
|
<taxonomicName id="4C4B4D1EFFE65921FBA1FB2CFA68FE6B" authority="Stal, 1854" authorityName="Stal" authorityYear="1854" box="[1126,1436,366,393]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="species" species="sahlbergi">
|
|
<emphasis id="B93FEA8FFFE65921FBA1FB2CFAD2FE6B" box="[1126,1318,367,393]" italics="true" pageId="12" pageNumber="351">Cicada sahlbergi</emphasis>
|
|
<bibRefCitation id="EFDA4B6CFFE65921FAEBFB2DFA68FE6B" author="Stal" box="[1324,1436,366,393]" firstAuthor="Stal" pageId="12" pageNumber="351" pagination="231 - 255" refId="ref16654" refString="Stal, C. (1854) Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 11, 231 - 255." type="journal article" year="1854">Stål, 1854</bibRefCitation>
|
|
</taxonomicName>
|
|
is reassigned to the genus
|
|
<taxonomicName id="4C4B4D1EFFE65921FE7BFBD0FD4AFE4F" authority="Distant, 1905 f" authorityName="Distant" authorityYear="1905" box="[444,702,402,429]" class="Insecta" family="Cicadidae" genus="Herrera" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B93FEA8FFFE65921FE7BFBD0FDE3FE4E" box="[444,535,403,428]" italics="true" pageId="12" pageNumber="351">Herrera</emphasis>
|
|
<bibRefCitation id="EFDA4B6CFFE65921FDD9FBD0FD4AFE4F" author="Distant" box="[542,702,402,429]" firstAuthor="Distant" pageId="12" pageNumber="351" pagination="478 - 486" refId="ref12527" refString="Distant, W. L. (1905 f) Rhynchotal notes XXXII. Annals and Magazine of Natural History, Series 7, 15, 478 - 486. https: // doi. org / 10.1080 / 03745480509442837" type="journal article" year="1905" yearSuffix="f">Distant, 1905f</bibRefCitation>
|
|
</taxonomicName>
|
|
to become
|
|
<taxonomicName id="4C4B4D1EFFE65921FC87FBD0FB62FE4F" authority="(Stal, 1854)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Stal" baseAuthorityYear="1854" box="[832,1174,402,429]" class="Insecta" family="Cicadidae" genus="Herrera" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="species" species="sahlbergi" status="comb. nov.">
|
|
<emphasis id="B93FEA8FFFE65921FC87FBD0FBFFFE4F" box="[832,1035,403,429]" italics="true" pageId="12" pageNumber="351">Herrera sahlbergi</emphasis>
|
|
(
|
|
<bibRefCitation id="EFDA4B6CFFE65921FBDDFBD1FB79FE4F" author="Stal" box="[1050,1165,402,429]" firstAuthor="Stal" pageId="12" pageNumber="351" pagination="231 - 255" refId="ref16654" refString="Stal, C. (1854) Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 11, 231 - 255." type="journal article" year="1854">Stål, 1854</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
<emphasis id="B93FEA8FFFE65921FB5AFBD0FAF7FE4F" bold="true" box="[1181,1283,403,429]" pageId="12" pageNumber="351">
|
|
<taxonomicNameLabel id="A20C57F4FFE65921FB5AFBD0FAF7FE4F" box="[1181,1283,403,429]" pageId="12" pageNumber="351" rank="species">n. comb.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3516516FFE65921FF00FBF5FC5CFE33" box="[199,936,438,465]" pageId="12" pageNumber="351" type="distribution">
|
|
<paragraph id="8BF4369DFFE65921FF00FBF5FC5CFE33" blockId="12.[151,1437,151,465]" box="[199,936,438,465]" pageId="12" pageNumber="351">
|
|
<emphasis id="B93FEA8FFFE65921FF00FBF5FE85FE33" bold="true" box="[199,369,438,465]" pageId="12" pageNumber="351">DISTRIBUTION.</emphasis>
|
|
The species is endemic to
|
|
<collectingCountry id="F35C760DFFE65921FD58FBF4FD17FE33" box="[671,739,439,465]" name="Brazil" pageId="12" pageNumber="351">Brazil</collectingCountry>
|
|
(
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<bibRefCitation id="EFDA4B6CFFE65921FD36FBF4FC68FE33" author="Metcalf" box="[753,924,439,465]" firstAuthor="Metcalf" pageId="12" pageNumber="351" pagination="587 - 919" refId="ref13934" refString="Metcalf, Z. P. (1963 b) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section II. Gaeninae and Cicadinae. North Carolina State College Contribution, 1502, 587 - 919." type="journal article" year="1963" yearSuffix="b">Metcalf 1963b</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |