237 lines
33 KiB
XML
237 lines
33 KiB
XML
<document id="4C9E2011D9B9631387C309FDCC517FE6" ID-CLB-Dataset="20637" ID-DOI="10.5252/geodiversitas2022v44a6" ID-GBIF-Dataset="1f85fdb9-78c1-4927-96c3-0dfafdf1daf6" ID-ISSN="1638-9395" ID-Zenodo-Dep="6079765" ID-ZooBank="urn:lsid:zoobank.org:pub:30BBF1E1-A978-4DD1-8C1A-43B23A6BD474" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1644843376998" checkinUser="carolina" docAuthor="Ossó, Àlex, Gagnaison, Cyril & Gain, Olivier" docDate="2022" docId="03E18F24A010B54618B0FE53022294A1" docLanguage="en" docName="Geodiversitas.44.6.207-228.pdf" docOrigin="Geodiversitas 44 (6)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Liocarcinus kuehni" docType="treatment" docVersion="9" lastPageNumber="220" masterDocId="FFD8F75CA01FB5491844FFE50723910C" masterDocTitle="A re-appraisal of the middle-late Miocene fossil decapod crustaceans of the ‘ Faluns’ (Anjou-Touraine, France)" masterLastPageNumber="228" masterPageNumber="207" pageNumber="220" updateTime="1699311364356" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="0CD7F30C578EAF88E49282AF0B412F71">A re-appraisal of the middle-late Miocene fossil decapod crustaceans of the ‘ Faluns’ (Anjou-Touraine, France)</mods:title>
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<mods:namePart id="B43D884B614AF9E3FE1963861AAD80D8">Ossó, Àlex</mods:namePart>
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<mods:namePart id="C1B456EFFF019042C32A600D70680F03">Gagnaison, Cyril</mods:namePart>
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<mods:namePart id="F51ECD0E4BD4FED7DB10D74E5CF64558">Gain, Olivier</mods:namePart>
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<mods:title id="B81D3647017EA91098714A0D7E3A218B">Geodiversitas</mods:title>
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<mods:part id="33D8D86C8356B8837BB4C9F3E412D463">
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<mods:date id="76C0C7A2F6A41DF3CB770DF408B4CC87">2022</mods:date>
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<mods:number id="D70C93EC4DC773735621DF21A570CAD3">44</mods:number>
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<treatment id="03E18F24A010B54618B0FE53022294A1" ID-DOI="http://doi.org/10.5281/zenodo.6310575" ID-GBIF-Taxon="193376739" ID-Zenodo-Dep="6310575" LSID="urn:lsid:plazi:treatment:03E18F24A010B54618B0FE53022294A1" httpUri="http://treatment.plazi.org/id/03E18F24A010B54618B0FE53022294A1" lastPageNumber="220" pageId="15" pageNumber="220">
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<subSubSection id="C3526DB9A010B54618B0FE5305B490DD" box="[244,663,438,465]" pageId="15" pageNumber="220" type="nomenclature">
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<paragraph id="8BF73E32A010B54618B0FE5305B490DD" blockId="15.[244,663,438,465]" box="[244,663,438,465]" pageId="15" pageNumber="220">
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<heading id="D0BF895EA010B54618B0FE5305B490DD" box="[244,663,438,465]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="220" reason="2">
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<taxonomicName id="4C4845B1A010B54618B0FE5305B490DD" ID-CoL="36d9afed-246b-4a01-9912-6303d775c4a7" authority="(Bachmayer, 1953)" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" box="[244,663,438,465]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
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<emphasis id="B93CE220A010B54618B0FE5306E090DD" bold="true" box="[244,451,438,465]" italics="true" pageId="15" pageNumber="220">Liocarcinus kuehni</emphasis>
|
||
(
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||
<bibRefCitation id="EFD943C3A010B5461995FE5305AC90DD" author="BACHMAYER F." box="[465,655,438,465]" firstAuthor="Bachmayer" journalOrPublisher="Mitteilungen der Geologischen Gesellschaft in Wien" pageId="15" pageNumber="220" pagination="237 - 262" part="44" refId="ref14972" refString="BACHMAYER F. 1953 a. - Die Dekapodenfauna des tortonischen Leithakalkes von Deutsch-Altenburg (Niederosterreich). Mitteilungen der Geologischen Gesellschaft in Wien 44: 237 - 262." title="Die Dekapodenfauna des tortonischen Leithakalkes von Deutsch-Altenburg (Niederosterreich)" type="journal article" year="1953">Bachmayer, 1953</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3526DB9A010B54619C7FE330634934E" pageId="15" pageNumber="220" type="description">
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<paragraph id="8BF73E32A010B54619C7FE33052B90FC" blockId="15.[387,520,470,497]" box="[387,520,470,497]" pageId="15" pageNumber="220">
|
||
(
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<figureCitation id="137322B7A010B54619C8FE33052390FC" box="[396,512,470,497]" captionStart="FIG" captionStartId="14.[132,142,1691,1708]" captionTargetBox="[132,1455,215,1649]" captionTargetId="figure-131@14.[132,1455,183,1649]" captionTargetPageId="14" captionText="FIG. 5. — Decapods from the ‘Savignean facies’ (Langhian) of the ‘Blandinerie’ quarry,Breil (Maine-et-Loire):A, indeterminate majoid,ULB-IV-A (26): dorsal view;B-E, Derilambrus cf.szaboi (Müller,1974):B, ULB-IV-A (33),right propodus,outer margin;C, inner margin;D, ULB-IV-A (35),right propodus,outer margin;E, right propodus, inner margin;F-K, Pilumnus mediterraneus (Lőrenthey, 1897):F, ULB-IV-A (14),dorsal view;G, frontal view;H, ULB-IV-A (18),dorsal view;I, frontal view;J, ULB-IV-A (20), left chela,outer margin;K, inner margin;L-M, undeterminated Carcinidae:L, ULB-IV-A (52),right chela;M, ULB-IV-A (50) left chela;N-Q, Liocarcinus kuehni (Bachmayer, 1953a):N, ULB-IV-A (10),dorsal view;O, frontal view;P, ULB-IV-A (13), dorsal view;Q, frontal view.Scale bars:A-I, N-Q, 5 mm;J-M, 10 mm. Photographs by À. Ossó." figureDoi="http://doi.org/10.5281/zenodo.6079787" httpUri="https://zenodo.org/record/6079787/files/figure.png" pageId="15" pageNumber="220">Fig. 5N-Q</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF73E32A010B54618C0FDF50634934E" blockId="15.[132,775,527,578]" pageId="15" pageNumber="220">
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<treatmentCitationGroup id="AB58191CA010B54618C0FDF50634934E" pageId="15" pageNumber="220">
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<treatmentCitation id="0AE91823A010B54618C0FDF5058D932B" box="[132,686,527,551]" pageId="15" pageNumber="220">
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<taxonomicName id="4C4845B1A010B54618C0FDF5051B932B" box="[132,568,527,551]" pageId="15" pageNumber="220">
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<emphasis id="B93CE220A010B54618C0FDF5063C932B" box="[132,287,527,551]" italics="true" pageId="15" pageNumber="220">Portunites kühni</emphasis>
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<bibRefCitation id="EFD943C3A010B5461960FDEA051B932B" author="BACHMAYER F." box="[292,568,527,551]" firstAuthor="Bachmayer" journalOrPublisher="Mitteilungen der Geologischen Gesellschaft in Wien" page="249 - 251" pageId="15" pageNumber="220" pagination="237 - 262" part="44" refId="ref14972" refString="BACHMAYER F. 1953 a. - Die Dekapodenfauna des tortonischen Leithakalkes von Deutsch-Altenburg (Niederosterreich). Mitteilungen der Geologischen Gesellschaft in Wien 44: 237 - 262." title="Die Dekapodenfauna des tortonischen Leithakalkes von Deutsch-Altenburg (Niederosterreich)" type="journal article" year="1953" yearSuffix="a">Bachmayer, 1953a: 249-251</bibRefCitation>
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</taxonomicName>
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, pl. 2, fig. 2.
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</treatmentCitation>
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—
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<treatmentCitation id="0AE91823A010B5461A8BFDF50634934E" pageId="15" pageNumber="220">
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||
<bibRefCitation id="EFD943C3A010B5461A8BFDF50630934E" author="KARASAWA H." firstAuthor="Karasawa" journalOrPublisher="Bulletin of the Mizunami Fossil Museum" page="17" pageId="15" pageNumber="220" pagination="1 - 33" part="17" refId="ref17951" refString="KARASAWA H. 1990. - Decapod Crustaceans from the Miocene Mizunami Group, Central Japan Part 2 Section Oxyrhyncha, Cancridea and Brachyrhyncha. Bulletin of the Mizunami Fossil Museum 17: 1 - 33." title="Decapod Crustaceans from the Miocene Mizunami Group, Central Japan Part 2 Section Oxyrhyncha, Cancridea and Brachyrhyncha" type="journal article" year="1990">Karasawa 1990: 17</bibRefCitation>
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.
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</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3526DB9A010B54618C0FDBF05E19380" pageId="15" pageNumber="220" type="reference_group">
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||
<paragraph id="8BF73E32A010B54618C0FDBF05E19380" blockId="15.[132,776,601,652]" pageId="15" pageNumber="220">
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||
<treatmentCitationGroup id="AB58191CA010B54618C0FDBF05E19380" pageId="15" pageNumber="220">
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||
<taxonomicName id="4C4845B1A010B54618C0FDBF0615937D" ID-CoL="36d9afed-246b-4a01-9912-6303d775c4a7" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" box="[132,310,601,625]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
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<emphasis id="B93CE220A010B54618C0FDBF0615937D" box="[132,310,601,625]" italics="true" pageId="15" pageNumber="220">Liocarcinus kuehni</emphasis>
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</taxonomicName>
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||
–
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<treatmentCitation id="0AE91823A010B5461916FDBC05B9937D" author="MULLER P." box="[338,666,601,625]" page="84" pageId="15" pageNumber="220" year="1984">
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||
<bibRefCitation id="EFD943C3A010B5461916FDBC06D5937D" author="MULLER P." box="[338,502,601,625]" firstAuthor="Muller" journalOrPublisher="Geologica Hungarica, Series Palaeontologica" page="84" pageId="15" pageNumber="220" pagination="25 - 317" part="42" refId="ref19937" refString="MULLER P. 1984. - Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica 42: 25 - 317." title="Decapod Crustacea of the Badenian" type="journal article" year="1984">Müller 1984: 84</bibRefCitation>
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, pl. 71, figs 2, 3
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</treatmentCitation>
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||
. —
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<treatmentCitation id="0AE91823A010B5461A83FDBC06029380" author="HYZNY M. & HARZHAUSER M. & DANNINGER W." page="226" pageId="15" pageNumber="220" year="2015">
|
||
<bibRefCitation id="EFD943C3A010B5461A83FDBC06029380" author="HYZNY M. & HARZHAUSER M. & DANNINGER W." etAl="et al." firstAuthor="Hyzny" journalOrPublisher="Geologica Carpathica" page="226" pageId="15" pageNumber="220" pagination="217 - 233" part="66" refId="ref17657" refString="HYZNY M., HARZHAUSER M. & DANNINGER W. 2015. - Decapod Crustacea of the Central Paratethyan Ottnangian Stage (middle Burdigalian): implications for systematics and biogeography. Geologica Carpathica 66 (3): 217 - 233. https: // doi. org / 10.1515 / geoca- 2015 - 0021" title="Decapod Crustacea of the Central Paratethyan Ottnangian Stage (middle Burdigalian): implications for systematics and biogeography" type="journal article" year="2015">
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Hyžný
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<emphasis id="B93CE220A010B54618C0FD9007909380" box="[132,179,628,652]" italics="true" pageId="15" pageNumber="220">et al.</emphasis>
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2015: 226
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
. —
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||
<treatmentCitation id="0AE91823A010B546190EFD91059D9380" author="HYZNY M. & DULAI A." box="[330,702,628,652]" page="208" pageId="15" pageNumber="220" year="2021">
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||
<bibRefCitation id="EFD943C3A010B546190EFD9105719380" author="HYZNY M. & DULAI A." box="[330,594,628,652]" firstAuthor="Hyzny" journalOrPublisher="GeoLitera Publishing House, Institute of Geosciences, University of Szeged, Hungary" page="208" pageId="15" pageNumber="220" pagination="296 p" refId="ref17569" refString="HYZNY M. & DULAI A. 2021. - Badenian Decapods of Hungary. GeoLitera Publishing House, Institute of Geosciences, University of Szeged, Hungary, 296 p." title="Badenian Decapods of Hungary" type="book" year="2021">Hyžný & Dulai 2021: 208</bibRefCitation>
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, fig. 80.14
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3526DB9A010B54618C0FD4105B99236" pageId="15" pageNumber="220" type="materials_examined">
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<paragraph id="8BF73E32A010B54618C0FD4105A393FC" blockId="15.[130,775,675,753]" pageId="15" pageNumber="220">MATERIAL EXAMINED AND MEASUREMENTS (in mm). — Two dorsal carapaces, ULB-IV-A (10): L = 15, W = 17, F = 7, FOW = 12. — ULB-IV-A (13): L = 6, W = 6.5, F = 3, FOW = 5,5.</paragraph>
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<paragraph id="8BF73E32A010B54618C0FCED05B99236" blockId="15.[132,774,776,826]" pageId="15" pageNumber="220">
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LOCALITY AND HORIZON. — ‘Blandinerie’ quarry, Breil (
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<collectingRegion id="498CF0D0A010B5461AE0FCED079F9236" country="France" name="Pays de la Loire" pageId="15" pageNumber="220">Maine-et-Loire</collectingRegion>
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). ‘Savignean facies’, Langhian (middle Miocene).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3526DB9A010B54618C0FC90057D9600" pageId="15" pageNumber="220" type="description">
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||
<paragraph id="8BF73E32A010B54618C0FC9006369281" blockId="15.[130,778,885,1804]" box="[132,277,885,911]" pageId="15" pageNumber="220">DESCRIPTION</paragraph>
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<paragraph id="8BF73E32A010B54618C0FC70057D9600" blockId="15.[130,778,885,1804]" pageId="15" pageNumber="220">Small sized carapace, subhexagonal, slightly wider than long; carapace surface covered by transverse rows of minuscule granules forwardly directed, forming terraces of different sizes. Regions weakly defined. Front gently trilobate, edge finely serrate, medial tooth slightly more advanced than the lateral teeth. Orbits relatively large; supraorbital margin arched, finely serrate, with two closed fissures, first at mid length second near outer orbital tooth; inner orbital tooth blunt; outer orbital tooth broad subtriangular, forwardly directed; infraorbital margin not preserved.Anterolateral margin slightly arcuate, with four subtriangular teeth (excluding the outer orbital tooth), the latest (epibranchial tooth) the sharper. Posterolateral margin slightly concave; anterior half sharp edged, with serrate keel; posterior half blunt edged; reentrant of fifth pereiopod well marked, gently concave. Posterior margin slightly convex, finely rimmed. Frontal region flattened.Gastric process poorly defined. Epigastric lobes slightly marked. Protogastric lobes slightly swollen. Mesogastric region weakly marked, very narrow anteriorly, subrhomboidal posteriorly, slightly elevated above the rest of regions. Metagastric region faintly defined. Urogastric region slightly depressed. Cardiac region broad, swollen anteriorly. Intestinal region depressed. Hepatic region depressed. Epibranchial region sigmoidal, interrupted medially, defined by long terraces, and a sharp ridge distally that ends in the epibranchial tooth. Mesobranchial region slightly swollen internally. Metabranchial region depressed. Cervical groove faintly marked.Branchiocardiac groove weakly marked. Ventral parts and appendages not preserved.</paragraph>
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</subSubSection>
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<subSubSection id="C3526DB9A010B54618C0F8D7022294A1" pageId="15" pageNumber="220" type="discussion">
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<paragraph id="8BF73E32A010B54618C0F8D707CB9646" blockId="15.[130,776,1842,2028]" box="[132,232,1842,1868]" pageId="15" pageNumber="220">REMARKS</paragraph>
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<paragraph id="8BF73E32A010B54618C0F8B40294909D" blockId="15.[130,776,1842,2028]" lastBlockId="15.[811,1463,215,1454]" pageId="15" pageNumber="220">
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||
The dorsal morphology of the two samples of decapods described herein correspond exactly with the
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<typeStatus id="54F38090A010B5461A4CF89405449687" box="[520,615,1905,1931]" pageId="15" pageNumber="220" type="holotype">holotype</typeStatus>
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||
of
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||
<taxonomicName id="4C4845B1A010B5461AD6F89707EF96A7" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
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<emphasis id="B93CE220A010B5461AD6F89707EF96A7" italics="true" pageId="15" pageNumber="220">Liocarcinus kuehni</emphasis>
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||
</taxonomicName>
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||
from the upper Miocene of
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||
<collectingCountry id="F35F7EA2A010B5461A54F874054296A7" box="[528,609,1937,1963]" name="Austria" pageId="15" pageNumber="220">Austria</collectingCountry>
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||
, described and depicted as
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||
<emphasis id="B93CE220A010B54618B9F857068B96C7" box="[253,424,1969,1995]" italics="true" pageId="15" pageNumber="220">Portunites kühni</emphasis>
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by
|
||
<bibRefCitation id="EFD943C3A010B5461988F854042096C0" author="BACHMAYER F." box="[460,771,1969,1996]" firstAuthor="Bachmayer" journalOrPublisher="Mitteilungen der Geologischen Gesellschaft in Wien" page="249 - 251" pageId="15" pageNumber="220" pagination="237 - 262" part="44" refId="ref14972" refString="BACHMAYER F. 1953 a. - Die Dekapodenfauna des tortonischen Leithakalkes von Deutsch-Altenburg (Niederosterreich). Mitteilungen der Geologischen Gesellschaft in Wien 44: 237 - 262." title="Die Dekapodenfauna des tortonischen Leithakalkes von Deutsch-Altenburg (Niederosterreich)" type="journal article" year="1953" yearSuffix="a">Bachmayer (1953a: 249-251</bibRefCitation>
|
||
, pl. 2, fig. 2) (see also
|
||
<bibRefCitation id="EFD943C3A010B5461923F834053896E7" author="MULLER P." box="[359,539,2001,2028]" firstAuthor="Muller" journalOrPublisher="Geologica Hungarica, Series Palaeontologica" page="84" pageId="15" pageNumber="220" pagination="25 - 317" part="42" refId="ref19937" refString="MULLER P. 1984. - Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica 42: 25 - 317." title="Decapod Crustacea of the Badenian" type="journal article" year="1984">Müller 1984: 84</bibRefCitation>
|
||
, pl. 71, figs 3, 4). The carapace outline, same number and shape of anterolateral teeth, shape of the orbits, similar size, and in particular the squamous carapace surface formed by transverse terraces of different sizes, in despite that the
|
||
<typeStatus id="54F38090A010B5461CE8FED2022E905D" box="[1196,1293,311,337]" pageId="15" pageNumber="220" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54F38090A010B5461D0CFED2028C905D" box="[1352,1455,311,337]" pageId="15" pageNumber="220" type="paratype">paratypes</typeStatus>
|
||
does not preserve the front, allows to consider the ‘Faluns’ specimens conspecific of
|
||
<taxonomicName id="4C4845B1A010B5461C70FE9203D5909D" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" box="[1076,1270,375,401]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
|
||
<emphasis id="B93CE220A010B5461C70FE9203D5909D" box="[1076,1270,375,401]" italics="true" pageId="15" pageNumber="220">Liocarcinus kuehni</emphasis>
|
||
</taxonomicName>
|
||
with confidence.
|
||
</paragraph>
|
||
<paragraph id="8BF73E32A010B5461B07FE72023092C3" blockId="15.[811,1463,215,1454]" pageId="15" pageNumber="220">
|
||
Strikingly, the dorsal carapace, the only part preserved of our specimens and of Bachmayer’s
|
||
<typeStatus id="54F38090A010B5461C2EFE52038090DD" box="[1130,1187,439,465]" pageId="15" pageNumber="220">types</typeStatus>
|
||
, seems identical to that of the extant
|
||
<taxonomicName id="4C4845B1A010B5461BD9FE32021D90FD" authority="(Pennant, 1777)" baseAuthorityName="Pennant" baseAuthorityYear="1777" box="[925,1342,471,497]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="corrugatus">
|
||
<emphasis id="B93CE220A010B5461BD9FE3203A790FC" box="[925,1156,471,496]" italics="true" pageId="15" pageNumber="220">Liocarcinus corrugatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD943C3A010B5461CD6FE32021590FD" author="PENNANT T." box="[1170,1334,471,497]" firstAuthor="Pennant" journalOrPublisher="White, London" pageId="15" pageNumber="220" pagination="157 p" refId="ref20414" refString="PENNANT T. 1777. - British Zoology. Vol. IV. Crustacea. Mollusca. Testacea: i-viii. White, London: 157 p. https: // doi. org / 10.5962 / bhl. title. 62481" title="British Zoology. Vol. IV. Crustacea. Mollusca. Testacea: i-viii" type="book" year="1777">Pennant, 1777</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However, they apparently differ in having a much smaller size than the extant species. In this respect, it is interesting to know that molecular analyses byPlagge
|
||
<emphasis id="B93CE220A010B5461C1BFDD203B3935C" box="[1119,1168,566,592]" italics="true" pageId="15" pageNumber="220">et al.</emphasis>
|
||
(2016), concluded that the European and the Asian populations of
|
||
<taxonomicName id="4C4845B1A010B5461C82FDB20285937C" baseAuthorityName="Pennant" baseAuthorityYear="1777" box="[1222,1446,599,624]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="corrugatus">
|
||
<emphasis id="B93CE220A010B5461C82FDB20285937C" box="[1222,1446,599,624]" italics="true" pageId="15" pageNumber="220">Liocarcinus corrugatus</emphasis>
|
||
</taxonomicName>
|
||
, only separated morphologically by subtle dorsal and penial differences, are in fact two genetically separated species, as the results of their analyses demonstrated, thus recognizing the Asian population as
|
||
<taxonomicName id="4C4845B1A010B5461C71FD33027093FC" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[1077,1363,725,752]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="strigilis">
|
||
<emphasis id="B93CE220A010B5461C71FD3303BF93E3" box="[1077,1180,725,751]" italics="true" pageId="15" pageNumber="220">L. strigilis</emphasis>
|
||
<bibRefCitation id="EFD943C3A010B5461CE7FD30027093FC" author="STIMPSON W." box="[1187,1363,725,752]" firstAuthor="Stimpson" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="15" pageNumber="220" pagination="225 - 252" part="10" refId="ref21541" refString="STIMPSON W. 1858. - Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. Pars VII. Crustacea Anomura. Proceedings of the Academy of Natural Sciences of Philadelphia 10: 225 - 252 [pages 63 - 90 on separate]. https: // doi. org / 10.5962 / bhl. title. 51447" title="Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. Pars VII. Crustacea Anomura" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
. In view of that, and given the scarce number of specimens of the fossil form, the two samples herein studied, and the three Bachmayer’s samples seen by Müller, is not sufficient to infer that
|
||
<taxonomicName id="4C4845B1A010B5461B1AFCB303029263" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" box="[862,1057,853,879]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
|
||
<emphasis id="B93CE220A010B5461B1AFCB303029263" box="[862,1057,853,879]" italics="true" pageId="15" pageNumber="220">Liocarcinus kuehni</emphasis>
|
||
</taxonomicName>
|
||
is the same species that
|
||
<taxonomicName id="4C4845B1A010B5461D64FCB302849263" baseAuthorityName="Pennant" baseAuthorityYear="1777" box="[1312,1447,854,879]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="corrugatus">
|
||
<emphasis id="B93CE220A010B5461D64FCB302849263" box="[1312,1447,854,879]" italics="true" pageId="15" pageNumber="220">L. corrugatus</emphasis>
|
||
</taxonomicName>
|
||
. In addition,
|
||
<bibRefCitation id="EFD943C3A010B5461BF8FC9003ED929C" author="PLAGGE C. & SON N. T. & NG P. K. L. & TURKAY M. & STREIT B & KLAUS S." box="[956,1230,885,912]" etAl="et al." firstAuthor="Plagge" journalOrPublisher="Raffles Bulletin of Zoology" page="386" pageId="15" pageNumber="220" pagination="374 - 388" part="64" refId="ref20456" refString="PLAGGE C., SON N. T., NG P. K. L., TURKAY M., STREIT B & KLAUS S. 2016. - Liocarcinus corrugatus (Pennant, 1777) (Crustacea: Brachyura: Portunidae): a cosmopolitan brachyuran species? Raffles Bulletin of Zoology 64: 374 - 388." title="Liocarcinus corrugatus (Pennant, 1777) (Crustacea: Brachyura: Portunidae): a cosmopolitan brachyuran species?" type="journal article" year="2016">
|
||
Plagge
|
||
<emphasis id="B93CE220A010B5461C4EFC9303639283" box="[1034,1088,885,911]" italics="true" pageId="15" pageNumber="220">et al.</emphasis>
|
||
(2016: 386)
|
||
</bibRefCitation>
|
||
suggested that both extant species are relict of a former range of the eastern Tethys Ocean”,
|
||
<taxonomicName id="4C4845B1A010B5461BCEFC53036D92C3" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" box="[906,1102,949,975]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
|
||
<emphasis id="B93CE220A010B5461BCEFC53036D92C3" box="[906,1102,949,975]" italics="true" pageId="15" pageNumber="220">Liocarcinus kuehni</emphasis>
|
||
</taxonomicName>
|
||
would confirm it.
|
||
</paragraph>
|
||
<paragraph id="8BF73E32A010B5461B07FC30022294A1" blockId="15.[811,1463,215,1454]" pageId="15" pageNumber="220">
|
||
<bibRefCitation id="EFD943C3A010B5461B07FC30039592E3" author="OSSO A. & GAGNAISON C." box="[835,1206,981,1007]" firstAuthor="Osso" journalOrPublisher="Geodiversitas" page="374 - 376" pageId="15" pageNumber="220" pagination="367 - 383" part="41" refId="ref20274" refString="OSSO A. & GAGNAISON C. 2019. - An appraisal of the Middle- Late Miocene fossil decapod crustaceans of the ' Faluns' (Anjou- Touraine, France). Geodiversitas 41 (9): 367 - 383. https: // doi. org / 10.5252 / geodiversitas 2019 v 41 a 9. http: // geodiversitas. com / 41 / 9" title="An appraisal of the Middle- Late Miocene fossil decapod crustaceans of the ' Faluns' (Anjou- Touraine, France)" type="journal article" year="2019">Ossó & Gagnaison (2019: 374-376</bibRefCitation>
|
||
, fig.5A-B) reported and described a specimen from the ‘Faluns’ as
|
||
<taxonomicName id="4C4845B1A010B5461CA6FC10025E9503" box="[1250,1405,1013,1039]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B93CE220A010B5461CA6FC1002759502" box="[1250,1366,1013,1038]" italics="true" pageId="15" pageNumber="220">Liocarcinus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, and compared it with all the known fossil forms of
|
||
<taxonomicName id="4C4845B1A010B5461D71FBF0028B9522" authorityName="Stimpson" authorityYear="1871" box="[1333,1448,1045,1070]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CE220A010B5461D71FBF0028B9522" box="[1333,1448,1045,1070]" italics="true" pageId="15" pageNumber="220">Liocarcinus</emphasis>
|
||
</taxonomicName>
|
||
. However, it was left in open nomenclature due to its poor preservation and the sole available specimen. Now, compared with the two new specimens of
|
||
<taxonomicName id="4C4845B1A010B5461C7EFB9003DB9582" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" box="[1082,1272,1140,1166]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
|
||
<emphasis id="B93CE220A010B5461C7EFB9003DB9582" box="[1082,1272,1140,1166]" italics="true" pageId="15" pageNumber="220">Liocarcinus kuehni</emphasis>
|
||
</taxonomicName>
|
||
, it still appears to be different. Its smooth carapace surface could be due to the reworking, but the specimen of
|
||
<taxonomicName id="4C4845B1A010B5461C32FB50021595C2" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" box="[1142,1334,1204,1230]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
|
||
<emphasis id="B93CE220A010B5461C32FB50021595C2" box="[1142,1334,1204,1230]" italics="true" pageId="15" pageNumber="220">Liocarcinus kuehni</emphasis>
|
||
</taxonomicName>
|
||
ULB-IV-A (13) (
|
||
<figureCitation id="137322B7A010B5461B23FB3104EF95E2" box="[871,972,1236,1263]" captionStart="FIG" captionStartId="14.[132,142,1691,1708]" captionTargetBox="[132,1455,215,1649]" captionTargetId="figure-131@14.[132,1455,183,1649]" captionTargetPageId="14" captionText="FIG. 5. — Decapods from the ‘Savignean facies’ (Langhian) of the ‘Blandinerie’ quarry,Breil (Maine-et-Loire):A, indeterminate majoid,ULB-IV-A (26): dorsal view;B-E, Derilambrus cf.szaboi (Müller,1974):B, ULB-IV-A (33),right propodus,outer margin;C, inner margin;D, ULB-IV-A (35),right propodus,outer margin;E, right propodus, inner margin;F-K, Pilumnus mediterraneus (Lőrenthey, 1897):F, ULB-IV-A (14),dorsal view;G, frontal view;H, ULB-IV-A (18),dorsal view;I, frontal view;J, ULB-IV-A (20), left chela,outer margin;K, inner margin;L-M, undeterminated Carcinidae:L, ULB-IV-A (52),right chela;M, ULB-IV-A (50) left chela;N-Q, Liocarcinus kuehni (Bachmayer, 1953a):N, ULB-IV-A (10),dorsal view;O, frontal view;P, ULB-IV-A (13), dorsal view;Q, frontal view.Scale bars:A-I, N-Q, 5 mm;J-M, 10 mm. Photographs by À. Ossó." figureDoi="http://doi.org/10.5281/zenodo.6079787" httpUri="https://zenodo.org/record/6079787/files/figure.png" pageId="15" pageNumber="220">Fig. 5P, Q</figureCitation>
|
||
) although it is also strongly worn, it retains still the terraces in the carapace, absent in
|
||
<taxonomicName id="4C4845B1A010B5461CFFFB1102779402" box="[1211,1364,1268,1294]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B93CE220A010B5461CFFFB11020C9401" box="[1211,1327,1268,1293]" italics="true" pageId="15" pageNumber="220">Liocarcinus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
In addition, the three characteristic tubercles, longitudinally aligned three tubercles at both sides of carapace that
|
||
<taxonomicName id="4C4845B1A010B5461D52FAD102939442" box="[1302,1456,1332,1358]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B93CE220A010B5461D52FAD102A99441" box="[1302,1418,1332,1357]" italics="true" pageId="15" pageNumber="220">Liocarcinus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
exhibits, are not present, or at least not interpreted as such, in the specimens of
|
||
<taxonomicName id="4C4845B1A010B5461BAEFA9103739481" baseAuthorityName="Bachmayer" baseAuthorityYear="1953" box="[1002,1104,1395,1421]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kuehni">
|
||
<emphasis id="B93CE220A010B5461BAEFA9103739481" box="[1002,1104,1395,1421]" italics="true" pageId="15" pageNumber="220">L. kuehni</emphasis>
|
||
</taxonomicName>
|
||
described herein. Consequently, we retain the former in open nomenclature.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |