461 lines
58 KiB
XML
461 lines
58 KiB
XML
<document id="DE43945D6FA0C167D05502687B7F20CC" ID-CLB-Dataset="24900" ID-DOI="10.1093/isd/ixz015" ID-GBIF-Dataset="56172c62-3e7c-4ab7-83d4-1dd9f5c57eb5" ID-Zenodo-Dep="3542130" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573763013583" checkinUser="plazi" docAuthor="Hita-Garcia, Francisco, Lieberman, Ziv, Audisio, Tracy L., Liu, Cong & Economo, Evan P." docDate="2019" docId="03D9AC4AE579FF94FCC4F948BEB605B8" docLanguage="en" docName="insectSystematicsDiversity.ixz015.pdf" docOrigin="Insect Systematics and Diversity 5" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Discothyrea gryphon Hita-Garcia & Lieberman & Audisio & Liu & Economo 2019, sp. n." docType="treatment" docVersion="6" lastPageNumber="48" masterDocId="FFE0D432E555FFBBFFF6FF95BB120352" masterDocTitle="Revision of the Highly Specialized Ant Genus Discothyrea (Hymenoptera: Formicidae) in the Afrotropics with X-Ray Microtomography and 3 D Cybertaxonomy" masterLastPageNumber="84" masterPageNumber="1" pageNumber="45" updateTime="1698765580234" updateUser="ExternalLinkService">
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<mods:title id="9F134EADB0DB15EE9D4DFC5BAFD3D72C">Revision of the Highly Specialized Ant Genus Discothyrea (Hymenoptera: Formicidae) in the Afrotropics with X-Ray Microtomography and 3 D Cybertaxonomy</mods:title>
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<mods:namePart id="0A93A04F1EAC7FE1A6A8FA6F28D510DF">Hita-Garcia, Francisco</mods:namePart>
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<mods:namePart id="EBA79821FC1B8FA01F657517113055CF">Lieberman, Ziv</mods:namePart>
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<mods:namePart id="7D521C0B7F7A1CB4B320286E3A2F0890">Audisio, Tracy L.</mods:namePart>
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<mods:namePart id="E5A823555F11839FB1E39624A0D304A1">Liu, Cong</mods:namePart>
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<mods:title id="558B699346E0EF28F024A79AE4593134">Insect Systematics and Diversity</mods:title>
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<taxonomicName id="4C7066DFE579FF97FCC4F948BE2E05A6" authority="Hita Garcia & Lieberman" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[818,1340,1757,1780]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="gryphon" status="sp. nov.">
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<emphasis id="B904C14EE579FF97FCC4F948BF0D05A6" box="[818,1055,1757,1780]" italics="true" pageId="44" pageNumber="45">Discothyrea gryphon</emphasis>
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Hita Garcia & Lieberman
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(
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Figs.
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<date id="FFCE3B9CE579FF97FC80F895B8990445" box="[886,907,1792,1815]" pageId="44" pageNumber="45">4I</date>
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,
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<figureCitation id="134B01D9E579FF97FC61F895B8BE0445" box="[919,940,1792,1815]" captionStart="Fig" captionStartId="10.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@10.[154,1434,147,1419]" captionTargetPageId="10" captionText="Fig. 6. Diagnostic plate showing still images from surface volume renderings of the head in full-face view (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542144" httpUri="https://zenodo.org/record/3542144/files/figure.png" pageId="44" pageNumber="45">
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<date id="FFCE3B9CE579FF97FC61F895B8BE0445" box="[919,940,1792,1815]" pageId="44" pageNumber="45">6I</date>
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,
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<figureCitation id="134B01D9E579FF97FC4EF895B8DF0445" box="[952,973,1792,1815]" captionStart="Fig" captionStartId="11.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@11.[152,1435,147,1422]" captionTargetPageId="11" captionText="Fig. 7. Diagnostic plate showing still images from surface volume renderings of the head in profile (the remainder of the body virtually removed).All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542148" httpUri="https://zenodo.org/record/3542148/files/figure.png" pageId="44" pageNumber="45">
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<date id="FFCE3B9CE579FF97FC4EF895B8DF0445" box="[952,973,1792,1815]" pageId="44" pageNumber="45">7I</date>
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,
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<figureCitation id="134B01D9E579FF97FC2CF895B8FD0445" box="[986,1007,1792,1815]" captionStart="Fig" captionStartId="12.[146,172,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@12.[152,1435,147,1422]" captionTargetPageId="12" captionText="Fig. 8. Diagnostic plate showing still images from surface volume renderings of the right mandible in dorsal view (the remainder of the body virtually removed).All Afrotropical species are shown,species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542150" httpUri="https://zenodo.org/record/3542150/files/figure.png" pageId="44" pageNumber="45">
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<date id="FFCE3B9CE579FF97FC2CF895B8FD0445" box="[986,1007,1792,1815]" pageId="44" pageNumber="45">8I</date>
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,
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<figureCitation id="134B01D9E579FF97FC0DF895BF020445" box="[1019,1040,1792,1815]" captionStart="Fig" captionStartId="13.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@13.[152,1435,147,1422]" captionTargetPageId="13" captionText="Fig. 9. Diagnostic plate showing still images from surface volume renderings of the mesosoma in profile (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542152" httpUri="https://zenodo.org/record/3542152/files/figure.png" pageId="44" pageNumber="45">
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<figureCitation id="134B01D9E579FF97FBEBF895BF2F0445" box="[1053,1085,1792,1815]" captionStart="Fig" captionStartId="14.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@14.[152,1435,147,1422]" captionTargetPageId="14" captionText="Fig. 10. Diagnostic plate showing still images from surface volume renderings of the body in dorsal view with focus on mesosoma.All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542154" httpUri="https://zenodo.org/record/3542154/files/figure.png" pageId="44" pageNumber="45">
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<date id="FFCE3B9CE579FF97FBEBF895BF2F0445" box="[1053,1085,1792,1815]" pageId="44" pageNumber="45" value="2010-01">10I</date>
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<figureCitation id="134B01D9E579FF97FBBCF895BF740445" box="[1098,1126,1792,1815]" captionStart="Fig" captionStartId="15.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@15.[154,1434,147,1419]" captionTargetPageId="15" captionText="Fig. 11. Diagnostic plate showing still images from surface volume renderings of the posterior propodeum in posterior view (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542156" httpUri="https://zenodo.org/record/3542156/files/figure.png" pageId="44" pageNumber="45">
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<date id="FFCE3B9CE579FF97FBBCF895BF740445" box="[1098,1126,1792,1815]" pageId="44" pageNumber="45" value="2011-01">11I</date>
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<figureCitation id="134B01D9E579FF97FB83F895BF850445" box="[1141,1175,1792,1815]" captionStart="Fig" captionStartId="16.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@16.[154,1434,147,1419]" captionTargetPageId="16" captionText="Fig. 12. Diagnostic plate showing still images from surface volume renderings of the petiole and gaster in profile (the remainder of the body virtually removed).All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542158" httpUri="https://zenodo.org/record/3542158/files/figure.png" pageId="44" pageNumber="45">
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<date id="FFCE3B9CE579FF97FB83F895BF850445" box="[1141,1175,1792,1815]" pageId="44" pageNumber="45" value="2012-01">12I</date>
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,
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<figureCitation id="134B01D9E579FF97FB52F895BFD70445" box="[1188,1221,1792,1815]" captionStart="Fig" captionStartId="18.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1419]" captionTargetId="figure@18.[154,1434,147,1419]" captionTargetPageId="18" captionText="Fig. 14. Diagnostic plate showing differences in body pilosity. (A) D. mixta (CASENT0235473),(B) D. oculata (CASENT0235467),(C) D. aisnetu (CASENT0235475), (D) D.athene (CASENT0235476),(E) D.chimera (CASENT0235471),(F) D.damato (CASENT0247362),(G) D.dryad (CASENT0247371),(H) D.gaia (CASENT0247040), (I) D. gryphon (CASENT0247367), (J) D. hawkesi (CASENT0235470), (K) D. kalypso (CASENT0235468), (L) D. maia (CASENT0790541), (M) D. michelae (CASENT0235469), (N) D. patrizii (CASENT0235472), (O) D. penthos (CASENT0247383), (P) D. poweri (SAM-ENT-0011509), (Q) D. schulzei (CASENT0247370), (R) D. traegaordhi (CASENT0790122), (S) D. venus (CASENT0247017), (T) D. wakanda (CASENT0790326).With the exception of F, L, R,T, all other images are from https://www.antweb.org—photographers Michele Esposito and Will Ericson." figureDoi="http://doi.org/10.5281/zenodo.3542162" httpUri="https://zenodo.org/record/3542162/files/figure.png" pageId="44" pageNumber="45">
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<date id="FFCE3B9CE579FF97FB52F895BFD70445" box="[1188,1221,1792,1815]" pageId="44" pageNumber="45" value="2014-01">14I</date>
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,
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<figureCitation id="134B01D9E579FF97FB24F895BFE10445" box="[1234,1267,1792,1815]" captionStart="Fig" captionStartId="20.[146,173,1654,1671]" captionTargetBox="[249,1339,148,1619]" captionTargetId="figure@20.[248,1339,147,1619]" captionTargetPageId="20" captionText="Fig. 15. Antennal anatomy showing flagellar fusion in Discothyrea. (A) D. mixta (CASENT0285473)—light microscopy, (B) D. traegaordhi (CASENT0790122) —light microscopy, (C) D. traegaordhi (CASENT0790122)—equivalent view to 15B but performed with surface volume rendering, (D) Zasphinctus sarowiwai Hita Garcia, 2017 (CASENT0764654)—sagittal section of surface volume rendering, (E) D. traegaordhi (CASENT0790122)—sagittal section of surface volume rendering, (F) D. mixta (CASENT0790542)—surface volume rendering showing each antennomere in different color, (G) D. mixta (CASENT0790542)—equivalent view to 15F but sagittal section, (H) D. dryad (CASENT0247374)—sagittal section of surface volume rendering showing each antennomere in different color, (I) D. gryphon (CASENT0790103),sagittal section of surface volume rendering showing each antennomere in different color." figureDoi="http://doi.org/10.5281/zenodo.3542164" httpUri="https://zenodo.org/record/3542164/files/figure.png" pageId="44" pageNumber="45">
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<date id="FFCE3B9CE579FF97FB24F895BFE10445" box="[1234,1267,1792,1815]" pageId="44" pageNumber="45" value="2015-01">15I</date>
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,
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<figureCitation id="134B01D9E579FF97FAF6F895BE090445" box="[1280,1307,1792,1815]" captionStart="Fig" captionStartId="45.[146,173,1061,1078]" captionTargetBox="[147,768,147,1026]" captionTargetId="figure@45.[147,768,147,1026]" captionTargetPageId="45" captionText="Fig. 35. Stacked digital color images of D. gryphon sp. n. paratype (CASENT0247367—from https://www.antweb.org, photographerWill Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.3542204" httpUri="https://zenodo.org/record/3542204/files/figure.png" pageId="44" pageNumber="45">35</figureCitation>
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,
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<figureCitation id="134B01D9E579FF97FADEF895BE510445" box="[1320,1347,1792,1815]" captionStart="Fig" captionStartId="46.[146,173,1782,1799]" captionTargetBox="[202,1386,147,1747]" captionTargetId="figure@46.[202,1386,147,1747]" captionTargetPageId="46" captionText="Fig. 36. Still images from shaded surface display volume renderings of D. gryphon sp. n. holotype (CASENT0790103) showing virtually segmented body parts. (A) Body in dorsal view, (B) body in profile, (C) head in full-face view, (D) head in dorsal view, (E) head in anterodorsal view, (F) head in anterior view, (G) head in profile, (H) head in ventral view, (I) posterior propodeum in posterior view, (J) petiole in profile, (K) petiole and gaster in profile, (L) petiole in dorsal view, (M) petiole in anterior view, (N) petiole oblique anterior view, (O) gaster in dorsal view, (P) petiole in ventral view, (Q) abdominal sternite 3 in ventral view, (R) mesotibia and mesotarsus in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3542206" httpUri="https://zenodo.org/record/3542206/files/figure.png" pageId="44" pageNumber="45">36</figureCitation>
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;
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</paragraph>
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<paragraph id="8BCF1D5CE579FF97FCC4F8B5BF670465" blockId="44.[818,1442,1757,1937]" box="[818,1141,1824,1847]" pageId="44" pageNumber="45">Supp Video S9 [online only])</paragraph>
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<paragraph id="8BCF1D5CE579FF97FCC4F8D6B8A60404" blockId="44.[818,1442,1757,1937]" box="[818,948,1859,1878]" pageId="44" pageNumber="45">
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<emphasis id="B904C14EE579FF97FCC4F8D6B8A60404" bold="true" box="[818,948,1859,1878]" pageId="44" pageNumber="45">Type Material</emphasis>
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|
||
<materialsCitation id="3B181701E579FF97FCC4F8F5BEB004C3" ID-GBIF-Occurrence="2451560201" collectingDate="1973-08-12" collectorName="P. Werner" country="Rwanda" elevation="2100" latitude="-1.876" location="Kayove" longLatPrecision="78" longitude="29.357" pageId="44" pageNumber="45" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B904C14EE579FF97FCC4F8F5B8B90421" bold="true" box="[818,939,1888,1907]" pageId="44" pageNumber="45">
|
||
<typeStatus id="54CBA3FEE579FF97FCC4F8F5B8B90421" box="[818,939,1888,1907]" pageId="44" pageNumber="45" type="holotype">HOLOTYPE</typeStatus>
|
||
</emphasis>
|
||
, pinned worker,
|
||
<collectingCountry id="F3675DCCE579FF97FBBEF8F5BFB90421" box="[1096,1195,1888,1907]" name="Rwanda" pageId="44" pageNumber="45">RWANDA</collectingCountry>
|
||
,
|
||
<location id="8EAF4B87E579FF97FB42F8F4BFE90426" LSID="urn:lsid:plazi:treatment:03D9AC4AE579FF94FCC4F948BEB605B8:8EAF4B87E579FF97FB42F8F4BFE90426" box="[1204,1275,1889,1908]" country="Rwanda" latitude="-1.876" longLatPrecision="78" longitude="29.357" name="Kayove" pageId="44" pageNumber="45">Kayove</location>
|
||
, [
|
||
<geoCoordinate id="EE447B9BE579FF97FAFDF8F5BE580421" box="[1291,1354,1888,1907]" degrees="1.876" direction="south" orientation="latitude" pageId="44" pageNumber="45" precision="55" value="-1.876">-1.876</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE447B9BE579FF97FAA2F8F5BE8B0421" box="[1364,1433,1888,1907]" degrees="29.357" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="55" value="29.357">29.357</geoCoordinate>
|
||
],
|
||
<quantity id="4C88B0B9E579FF97FCC4F8EBB89204C3" box="[818,896,1918,1937]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="44" pageNumber="45" unit="m" value="2100.0">
|
||
<elevation id="005DFA6FE579FF97FCC4F8EBB89204C3" box="[818,896,1918,1937]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="44" pageNumber="45" unit="m" value="2100.0">2100 m</elevation>
|
||
</quantity>
|
||
, collection code ANTC37492,
|
||
<date id="FFCE3B9CE579FF97FB45F8EBBE3904C3" box="[1203,1323,1918,1937]" pageId="44" pageNumber="45" value="1973-08-12">
|
||
<collectingDate id="EF8AC274E579FF97FB45F8EBBE3904C3" box="[1203,1323,1918,1937]" pageId="44" pageNumber="45" value="1973-08-12">12.VIII.1973</collectingDate>
|
||
</date>
|
||
(
|
||
<emphasis id="B904C14EE579FF97FACBF8EBBE8904C2" box="[1341,1435,1918,1936]" italics="true" pageId="44" pageNumber="45">
|
||
<collectorName id="2685788AE579FF97FACBF8EBBE8904C2" box="[1341,1435,1918,1936]" pageId="44" pageNumber="45">P. Werner</collectorName>
|
||
</emphasis>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF0F4DD4E578FF96FF64FBB0B9D2073B" ID-DOI="http://doi.org/10.5281/zenodo.3542204" ID-Zenodo-Dep="3542204" httpUri="https://zenodo.org/record/3542204/files/figure.png" pageId="45" pageNumber="46" startId="45.[146,173,1061,1078]" targetBox="[147,768,147,1026]" targetPageId="45">
|
||
<paragraph id="8BCF1D5CE578FF96FF64FBB0B9D2073B" blockId="45.[146,772,1061,1129]" pageId="45" pageNumber="46">
|
||
<emphasis id="B904C14EE578FF96FF64FBB0BBC40764" bold="true" box="[146,214,1061,1078]" pageId="45" pageNumber="46">Fig. 35.</emphasis>
|
||
Stacked digital color images of
|
||
<taxonomicName id="4C7066DFE578FF96FDF1FBB0B9790764" authority="Hita-Garcia & Lieberman & Audisio & Liu & Economo, 2019" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[519,619,1061,1078]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="gryphon" status="sp. nov.">
|
||
<emphasis id="B904C14EE578FF96FDF1FBB0B9790764" box="[519,619,1061,1078]" italics="true" pageId="45" pageNumber="46">D. gryphon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377C35E578FF96FD8EFBB0B9BF0764" box="[632,685,1061,1078]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
|
||
paratype (CASENT0247367—from https://www.antweb.org, photographerWill Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF1D5CE578FF96FF64FB1AB8160645" blockId="45.[146,772,1167,1304]" pageId="45" pageNumber="46">
|
||
(
|
||
<collectionCode id="ED618599E578FF96FF6EFB1ABBF007F0" box="[152,226,1167,1186]" pageId="45" pageNumber="46">BMNH</collectionCode>
|
||
: CASENT0790103).
|
||
<materialsCitation id="3B181701E578FF96FE44FB1AB8120645" ID-GBIF-Occurrence="2451560184" collectingDate="1973-04-23" collectionCode="BMNH, SAMC, CASC, MHNG" collectorName="P. Werner" country="Rwanda" elevation="2100" latitude="-1.876" location="Kayove" longLatPrecision="78" longitude="29.357" pageId="45" pageNumber="46" specimenCode="CASENT0790104, CASENT0247366, CASENT0247367, CASENT0247368" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B904C14EE578FF96FE44FB1AB93E07F0" bold="true" box="[434,556,1167,1186]" pageId="45" pageNumber="46">
|
||
<typeStatus id="54CBA3FEE578FF96FE44FB1AB93E07F0" box="[434,556,1167,1186]" pageId="45" pageNumber="46" type="paratype">PARATYPES</typeStatus>
|
||
</emphasis>
|
||
, two pinned workers with same data as holotype (
|
||
<collectionCode id="ED618599E578FF96FE58FB38BAEA0792" box="[430,504,1197,1216]" pageId="45" pageNumber="46">BMNH</collectionCode>
|
||
:
|
||
<specimenCode id="DBD6B527E578FF96FDF3FB38B9A20792" box="[517,688,1197,1216]" collectionCode="BMNH" pageId="45" pageNumber="46">CASENT0790104</specimenCode>
|
||
;
|
||
<collectionCode id="ED618599E578FF96FD48FB38B9ED0792" box="[702,767,1197,1216]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="45" pageNumber="46" type="Museum">SAMC</collectionCode>
|
||
:
|
||
<specimenCode id="DBD6B527E578FF96FF64FB5FBA2C078F" box="[146,318,1226,1245]" collectionCode="SAMC" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="45" pageNumber="46" type="Museum">CASENT0247366</specimenCode>
|
||
); and two workers from
|
||
<collectingCountry id="F3675DCCE578FF96FDB3FB5FB9BA078F" box="[581,680,1226,1245]" name="Rwanda" pageId="45" pageNumber="46">RWANDA</collectingCountry>
|
||
,
|
||
<location id="8EAF4B87E578FF96FD4EFB5FB9ED078F" LSID="urn:lsid:plazi:treatment:03D9AC4AE579FF94FCC4F948BEB605B8:8EAF4B87E578FF96FD4EFB5FB9ED078F" box="[696,767,1226,1245]" country="Rwanda" latitude="-1.876" longLatPrecision="78" longitude="29.357" name="Kayove" pageId="45" pageNumber="46">Kayove</location>
|
||
, [
|
||
<geoCoordinate id="EE447B9BE578FF96FF61FB72BBC507A8" box="[151,215,1255,1274]" degrees="1.876" direction="south" orientation="latitude" pageId="45" pageNumber="46" precision="55" value="-1.876">-1.876</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE447B9BE578FF96FF17FB72BA3407A8" box="[225,294,1255,1274]" degrees="29.357" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="55" value="29.357">29.357</geoCoordinate>
|
||
],
|
||
<quantity id="4C88B0B9E578FF96FEC2FB72BA6F07A8" box="[308,381,1255,1274]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="45" pageNumber="46" unit="m" value="2100.0">
|
||
<elevation id="005DFA6FE578FF96FEC2FB72BA6F07A8" box="[308,381,1255,1274]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="45" pageNumber="46" unit="m" value="2100.0">2100 m</elevation>
|
||
</quantity>
|
||
, collection code ANTC42122,
|
||
<date id="FFCE3B9CE578FF96FD6DFB72B81307A8" box="[667,769,1255,1274]" pageId="45" pageNumber="46" value="1973-04-23">
|
||
<collectingDate id="EF8AC274E578FF96FD6DFB72B81307A8" box="[667,769,1255,1274]" pageId="45" pageNumber="46" value="1973-04-23">23.IV.1973</collectingDate>
|
||
</date>
|
||
(
|
||
<emphasis id="B904C14EE578FF96FF6FFA90BBE20645" box="[153,240,1285,1303]" italics="true" pageId="45" pageNumber="46">
|
||
<collectorName id="2685788AE578FF96FF6FFA90BBE20645" box="[153,240,1285,1303]" pageId="45" pageNumber="46">P.Werner</collectorName>
|
||
</emphasis>
|
||
) (
|
||
<collectionCode id="ED618599E578FF96FEF7FA90BA2F064A" box="[257,317,1285,1304]" pageId="45" pageNumber="46">CASC</collectionCode>
|
||
:
|
||
<specimenCode id="DBD6B527E578FF96FEB3FA91BAE30645" box="[325,497,1284,1303]" collectionCode="CASC" pageId="45" pageNumber="46">CASENT0247367</specimenCode>
|
||
;
|
||
<collectionCode id="ED618599E578FF96FE0FFA90B954064A" box="[505,582,1285,1304]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="45" pageNumber="46" type="Museum">MHNG</collectionCode>
|
||
:
|
||
<specimenCode id="DBD6B527E578FF96FDBBFA91B9E80645" box="[589,762,1284,1303]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="45" pageNumber="46" type="Museum">CASENT0247368</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FF64FAD7B811056D" blockId="45.[146,771,1346,1600]" pageId="45" pageNumber="46">
|
||
<emphasis id="B904C14EE578FF96FF64FAD7BBE50607" box="[146,247,1346,1365]" italics="true" pageId="45" pageNumber="46">Cybertype.</emphasis>
|
||
Volumetric raw data (in DICOM format), 3D rotation video, still images of surface volume rendering, and 3D surface (in PLY format) of the physical
|
||
<typeStatus id="54CBA3FEE578FF96FE52FAE9BAE706DD" box="[420,501,1404,1423]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
(CASENT0790103) in addition to stacked digital color images illustrating head in full-face view, profile and dorsal views of the body.The data are deposited at Dryad (Hita Garcia et al. 2019, http://doi.org/10.5061/dryad.3qm4183) and can be freely accessed as virtual representation of the type. In addition to the cybertype data at Dryad, we also provide a freely accessible 3D surface model of the
|
||
<typeStatus id="54CBA3FEE578FF96FE25F9B9B936056D" box="[467,548,1580,1599]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
at Sketchfab (Model 9).
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FF64F9FABA2505D0" blockId="45.[146,770,1647,1725]" box="[146,311,1647,1666]" pageId="45" pageNumber="46">
|
||
<heading id="D087AA30E578FF96FF64F9FABA2505D0" bold="true" box="[146,311,1647,1666]" fontSize="8" level="3" pageId="45" pageNumber="46" reason="6">
|
||
<emphasis id="B904C14EE578FF96FF64F9FABA2505D0" bold="true" box="[146,311,1647,1666]" pageId="45" pageNumber="46">Nontype Material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FF64F918B9E005EF" blockId="45.[146,770,1647,1725]" pageId="45" pageNumber="46">
|
||
<materialsCitation id="3B181701E578FF96FF64F918B9FC05EF" ID-GBIF-Occurrence="2451560220" collectingDate="1995-12-07" collectorName="H. G. Robertson" country="Tanzania" elevation="1600" latitude="-3.8833" location="Mkomazi Game Reserve" longLatPrecision="5" longitude="37.81667" municipality="Maji Kununua" pageId="45" pageNumber="46" specimenCount="1">
|
||
<emphasis id="B904C14EE578FF96FF64F918BA1C05F2" box="[146,270,1677,1696]" italics="true" pageId="45" pageNumber="46">
|
||
<collectingCountry id="F3675DCCE578FF96FF64F918BA1B05F2" box="[146,265,1677,1696]" name="Tanzania" pageId="45" pageNumber="46">TANZANIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<location id="8EAF4B87E578FF96FEEEF919B91405F2" LSID="urn:lsid:plazi:treatment:03D9AC4AE579FF94FCC4F948BEB605B8:8EAF4B87E578FF96FEEEF919B91405F2" box="[280,518,1676,1696]" country="Tanzania" latitude="-3.8833" longLatPrecision="5" longitude="37.81667" municipality="Maji Kununua" name="Mkomazi Game Reserve" pageId="45" pageNumber="46">Mkomazi Game Reserve</location>
|
||
,
|
||
<collectingMunicipality id="6BAB8726E578FF96FDE2F919B9B305F2" box="[532,673,1676,1696]" pageId="45" pageNumber="46">Maji Kununua</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE447B9BE578FF96FD59F918B9EC05F2" box="[687,766,1677,1696]" degrees="3.8833" direction="south" orientation="latitude" pageId="45" pageNumber="46" precision="5" value="-3.8833">−3.8833</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE447B9BE578FF96FF64F93FBBFF05EF" box="[146,237,1706,1725]" degrees="37.81667" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="1" value="37.81667">37.81667</geoCoordinate>
|
||
,
|
||
<quantity id="4C88B0B9E578FF96FF00F93FBA2C05EF" box="[246,318,1706,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="45" pageNumber="46" unit="m" value="1600.0">
|
||
<elevation id="005DFA6FE578FF96FF00F93FBA2C05EF" box="[246,318,1706,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="45" pageNumber="46" unit="m" value="1600.0">1600 m</elevation>
|
||
</quantity>
|
||
, montane forest,
|
||
<date id="FFCE3B9CE578FF96FE28F93FB95605EF" box="[478,580,1706,1725]" pageId="45" pageNumber="46" value="1995-12-07">
|
||
<collectingDate id="EF8AC274E578FF96FE28F93FB95605EF" box="[478,580,1706,1725]" pageId="45" pageNumber="46" value="1995-12-07">7.XII.1995</collectingDate>
|
||
</date>
|
||
(
|
||
<emphasis id="B904C14EE578FF96FDA7F93FB9F705EF" box="[593,741,1706,1725]" italics="true" pageId="45" pageNumber="46">
|
||
<collectorName id="2685788AE578FF96FDA7F93FB9F705EF" box="[593,741,1706,1725]" pageId="45" pageNumber="46">H.G. Robertson</collectorName>
|
||
</emphasis>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E578FF96FF64F97FBE8C03B6" pageId="45" pageNumber="46" type="diagnosis">
|
||
<paragraph id="8BCF1D5CE578FF96FF64F97FBBFE05AF" blockId="45.[146,770,1770,1936]" box="[146,236,1770,1789]" pageId="45" pageNumber="46">
|
||
<heading id="D087AA30E578FF96FF64F97FBBFE05AF" bold="true" box="[146,236,1770,1789]" fontSize="8" level="3" pageId="45" pageNumber="46" reason="6">
|
||
<emphasis id="B904C14EE578FF96FF64F97FBBFE05AF" bold="true" box="[146,236,1770,1789]" pageId="45" pageNumber="46">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FF64F892BE8C03B6" blockId="45.[146,770,1770,1936]" lastBlockId="45.[818,1442,149,228]" pageId="45" pageNumber="46">
|
||
The following character combination distinguishes
|
||
<taxonomicName id="4C7066DFE578FF96FD66F89DB8100448" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[656,770,1799,1818]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="gryphon">
|
||
<emphasis id="B904C14EE578FF96FD66F89DB8100448" box="[656,770,1799,1818]" italics="true" pageId="45" pageNumber="46">D. gryphon</emphasis>
|
||
</taxonomicName>
|
||
from the remainder of the complex: mandible with square apical tooth and subapical denticle; subquadrate head with anterolateral corner of gena sharply defined, nearly denticulate; anterior clypeal margin with long erect setae; mesosomal outline weakly convex; propodeum without strong angles or denticles; mesotibia without apicoventral spur; AT4 about 1.1 times longer than AT3 (ASI 108– 113); erect pilosity abundant on mesosoma and abdominal terga.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E578FF96FCC4FE9EB8C5053B" pageId="45" pageNumber="46" type="description">
|
||
<paragraph id="8BCF1D5CE578FF96FCC4FE9EBFA6024C" blockId="45.[818,1442,267,462]" box="[818,1204,267,286]" pageId="45" pageNumber="46">
|
||
<heading id="D087AA30E578FF96FCC4FE9EBFA6024C" bold="true" box="[818,1204,267,286]" fontSize="8" level="3" pageId="45" pageNumber="46" reason="6">
|
||
<emphasis id="B904C14EE578FF96FCC4FE9EBFA6024C" bold="true" box="[818,1204,267,286]" pageId="45" pageNumber="46">
|
||
Worker Measurements and Indices (
|
||
<emphasis id="B904C14EE578FF96FB8BFE9EBF9B024C" bold="true" box="[1149,1161,267,286]" italics="true" pageId="45" pageNumber="46">n</emphasis>
|
||
= 6)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FCC4FEBCBF83029C" blockId="45.[818,1442,267,462]" pageId="45" pageNumber="46">
|
||
EL 0.00–0.02; HL 0.51–0.53; HW 0.44–0.48; SL 0.30–0.32;
|
||
<collectingCountry id="F3675DCCE578FF96FA72FEBCBEB0026E" box="[1412,1442,297,316]" name="Philippines" pageId="45" pageNumber="46">PH</collectingCountry>
|
||
0.27–0.29; PW 0.30–0.33; DML 0.36–0.39; PrH 0.31–0.35; WL 0.56–0.58; HFL 0.29–0.31; PeL 0.08–0.11; PeW 0.20–0.22; PeH 0.20–0.23; LT3 0.30–0.33; LT4 0.33–0.36; OI 0–4; CI 87–92; SI 59–62; LMI 47–51; DMI 54–57; DMI2 82–86; ASI 108–113; HFI 52–54; DPeI 192–250; LPeI 200–275.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FCC4FE6EB8F5015C" blockId="45.[818,1443,507,1934]" box="[818,999,507,526]" pageId="45" pageNumber="46">
|
||
<heading id="D087AA30E578FF96FCC4FE6EB8F5015C" bold="true" box="[818,999,507,526]" fontSize="8" level="3" pageId="45" pageNumber="46" reason="6">
|
||
<emphasis id="B904C14EE578FF96FCC4FE6EB8F5015C" bold="true" box="[818,999,507,526]" pageId="45" pageNumber="46">Worker Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FCC4FD8DBF3C0674" blockId="45.[818,1443,507,1934]" pageId="45" pageNumber="46">
|
||
<emphasis id="B904C14EE578FF96FCC4FD8DB8760179" box="[818,868,536,555]" italics="true" pageId="45" pageNumber="46">Head</emphasis>
|
||
very broad (CI 87–92); posterior head margin straight, posterodorsal corners of head rounded; in frontal view sides of head slightly convex; head appearing subquadrate posterad antennal sockets; eyes absent or minute (OI 0–4), an asymmetrical pigmented spot, situated about one-third of the way between anterolateral corner of gena and posterior head margin; when present, eyes not visible in frontal view; anterolateral corner of gena sharply squared, appearing denticulate/dentate, somewhat projecting laterally; frontal lamella short and dentate in profile, apex acute; lamella translucent across entire disc, slightly more so basally but without distinct fenestra; medial clypeus broad, emarginate medially between antennae, lateral clypeus conspicuously narrowed, curving broadly between antennal sockets and anterolateral corners of head, entire clypeal margin bearing row of long erect setae.
|
||
<emphasis id="B904C14EE578FF96FB44FC03BE1200FB" box="[1202,1280,918,937]" italics="true" pageId="45" pageNumber="46">Antenna</emphasis>
|
||
with moderately long scape (SI 59–62), scape moderately incrassate, gently bent; pedicel subcylindrical, longer than broad; true antennomere count six; apparent antennomere count six to eight, flagellomeres basad apical club highly compressed, taken together only about as long as apical club.
|
||
<emphasis id="B904C14EE578FF96FC55FBBDBF0A0769" box="[931,1048,1064,1083]" italics="true" pageId="45" pageNumber="46">Ventral head</emphasis>
|
||
with poorly developed postoccipital ridge with very short but broad anteromedian carina; median region of hypostoma triangular, arms strongly narrowed and very weakly spatulate; palpal formula not examined.
|
||
<emphasis id="B904C14EE578FF96FB5EFB15BE1207C1" box="[1192,1280,1152,1171]" italics="true" pageId="45" pageNumber="46">Mandible</emphasis>
|
||
with squared apical tooth subtended by pointed preapical denticle; small prebasal angle present; basal angle rounded to somewhat truncate; ectal face with longitudinal carina confluent with masticatory margin for much of its length, leaving narrow strip of depressed region including prebasal denticle.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FCA4FAA4BFE1057C" blockId="45.[818,1443,507,1934]" pageId="45" pageNumber="46">
|
||
<emphasis id="B904C14EE578FF96FCA4FAA4B8A60611" box="[850,948,1329,1347]" italics="true" pageId="45" pageNumber="46">Mesosoma</emphasis>
|
||
in profile robust, dorsally feebly convex, sloping posteroventrally, pronotum distincly higher than propodeum; in dorsal view mesosoma moderately thick (DMI 54–57; DMI2 82–86) and narrowed posteriorly, pronotum wider than propodeum; posterior propodeal margin very weakly concave; posterodorsal corners of propodeum rounded, without teeth or angles; declivitous face of propodeum shallowly but clearly concave in profile and oblique posterior view; propodeal spiracle small and inconspicuous, directed posterolaterally; propodeal lobes short, truncate.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE578FF96FCA4F9ACB8C5053B" blockId="45.[818,1443,507,1934]" pageId="45" pageNumber="46">
|
||
<emphasis id="B904C14EE578FF96FCA4F9ACB86F0519" box="[850,893,1593,1611]" italics="true" pageId="45" pageNumber="46">Legs</emphasis>
|
||
short (HFI 52–54) and rather slender; mesotibia without apicoventral spur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E578FF94FCA4F9E6BA15012D" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46" type="materials_examined">
|
||
<paragraph id="8BCF1D5CE578FF94FCA4F9E6BA15012D" blockId="45.[818,1443,507,1934]" lastBlockId="47.[146,772,546,639]" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46">
|
||
<emphasis id="B904C14EE578FF96FCA4F9E6B8DF05D4" box="[850,973,1651,1670]" italics="true" pageId="45" pageNumber="46">Petiolar node</emphasis>
|
||
not strongly attenuated dorsally, rather robust in profile, about 2.0 to 2.8 times higher than broad (LPeI 200–275); in profile anterior face of node convex, apex blunt to rounded, posterior face convex; in dorsal view petiole rectangular, sides subparallel, about 1.9 to 2.5 times broader than long (DPeI 192–250); in anterior view, petiolar outline clearly pentagonal, edges well-defined, angles slightly rounded, apex somewhat broadly peaked; in oblique anterior view anterior face flat; in ventral view subrectangular to weakly campaniform, sides slightly divergent posteriorly to weakly convex; subpetiolar process large, lobate-triangular, apex rounded,
|
||
<emphasis id="B904C14EE57AFF94FF64FDB7BBCA0161" bold="true" box="[146,216,546,563]" pageId="47" pageNumber="48">Model 9.</emphasis>
|
||
3D surface model of
|
||
<taxonomicName id="4C7066DFE57AFF94FE7DFDB7BAF40161" authority="Hita-Garcia & Lieberman & Audisio & Liu & Economo, 2019" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[395,486,546,563]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gryphon" status="sp. nov.">
|
||
<emphasis id="B904C14EE57AFF94FE7DFDB7BAF40161" box="[395,486,546,563]" italics="true" pageId="47" pageNumber="48">D. gryphon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377C35E57AFF94FE1AFDB7B90B0161" box="[492,537,546,563]" pageId="47" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
|
||
<typeStatus id="54CBA3FEE57AFF94FDEBFDB7B9770161" box="[541,613,546,563]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
(CASENT0790103). An interactive version of this model is available in the HTML version of this article online and at https://sketchfab.com/3d-models/b89fada99bce4f169c7b 7d55f977326a.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0F4DD4E57BFF95FF64F963BAE4043F" ID-DOI="http://doi.org/10.5281/zenodo.3542206" ID-Zenodo-Dep="3542206" httpUri="https://zenodo.org/record/3542206/files/figure.png" pageId="46" pageNumber="47" startId="46.[146,173,1782,1799]" targetBox="[202,1386,147,1747]" targetPageId="46">
|
||
<paragraph id="8BCF1D5CE57BFF95FF64F963BAE4043F" blockId="46.[146,1444,1782,1901]" pageId="46" pageNumber="47">
|
||
<emphasis id="B904C14EE57BFF95FF64F963BBDC0455" bold="true" box="[146,206,1782,1799]" pageId="46" pageNumber="47">Fig. 36.</emphasis>
|
||
Still images from shaded surface display volume renderings of
|
||
<taxonomicName id="4C7066DFE57BFF95FD2BF963B82A0455" authority="Hita-Garcia & Lieberman & Audisio & Liu & Economo, 2019" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[733,824,1782,1799]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gryphon" status="sp. nov.">
|
||
<emphasis id="B904C14EE57BFF95FD2BF963B82A0455" box="[733,824,1782,1799]" italics="true" pageId="46" pageNumber="47">D. gryphon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377C35E57BFF95FCCBF963B8780455" box="[829,874,1782,1799]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
|
||
holotype (CASENT0790103) showing virtually segmented body parts. (A) Body in dorsal view, (B) body in profile, (C) head in full-face view, (D) head in dorsal view, (E) head in anterodorsal view, (F) head in anterior view, (G) head in profile, (H) head in ventral view, (I) posterior propodeum in posterior view, (J) petiole in profile, (K) petiole and gaster in profile, (L) petiole in dorsal view, (M) petiole in anterior view, (N) petiole oblique anterior view, (O) gaster in dorsal view, (P) petiole in ventral view, (Q) abdominal sternite 3 in ventral view, (R) mesotibia and mesotarsus in anterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36A4ED7E57AFF94FF64FD3BB9B905DB" pageId="47" pageNumber="48" type="description">
|
||
<paragraph id="8BCF1D5CE57AFF94FF64FD3BB900018C" blockId="47.[146,771,686,1673]" pageId="47" pageNumber="48">bearing numerous long, straight, white setae; petiolar spiracles large, round to slightly elliptical in ventral view.</paragraph>
|
||
<paragraph id="8BCF1D5CE57AFF94FF44FD7DB99A0773" blockId="47.[146,771,686,1673]" pageId="47" pageNumber="48">
|
||
<emphasis id="B904C14EE57AFF94FF44FD7DBA6E01A9" box="[178,380,744,763]" italics="true" pageId="47" pageNumber="48">Abdominal segment 3</emphasis>
|
||
asymmetrically campaniform, tergite prolonged anteriorly beyond anterior margin of sternite and widest point just anterad end of segment;
|
||
<collectingCountry id="F3675DCCE57AFF94FE16FCB6BAEF0064" box="[480,509,803,822]" name="American Samoa" pageId="47" pageNumber="48">AS</collectingCountry>
|
||
3 evenly rounded in profile, deepest point at about the longitudinal midline in profile, with thick median ridge;
|
||
<collectingCountry id="F3675DCCE57AFF94FEEAFCCBBA2B0023" box="[284,313,862,881]" name="American Samoa" pageId="47" pageNumber="48">AS</collectingCountry>
|
||
3 without carinate prora, but still with anterior face distinctly depressed and anterior margin of ventral face weakly concave in ventral view; AT4 about 1.1 times longer than AT3 (ASI 108–113); AT4 evenly rounded hemidemispherical;
|
||
<collectingCountry id="F3675DCCE57AFF94FD8CFC23B987009B" box="[634,661,950,969]" name="American Samoa" pageId="47" pageNumber="48">AS</collectingCountry>
|
||
4 with very well-developed anterior lip, overlapping nearly the entire width of
|
||
<collectingCountry id="F3675DCCE57AFF94FF64FC65BBA20751" box="[146,176,1008,1027]" name="American Samoa" pageId="47" pageNumber="48">AS</collectingCountry>
|
||
3, anterior margin very weakly concave in ventral view; successive abdominal segments short, telescopic, often concealed.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE57AFF94FF44FBBEB9F8067B" blockId="47.[146,771,686,1673]" pageId="47" pageNumber="48">
|
||
<emphasis id="B904C14EE57AFF94FF44FBBEBA1B076C" box="[178,265,1067,1086]" italics="true" pageId="47" pageNumber="48">Sculpture</emphasis>
|
||
on head foveolate-reticulate to areolate, foveolae smaller on front of head; mandible rather roughly sculptured with dense piligerous punctae; frontal lamella and medial clypeus distinctly punctate; mesosoma, petiole, and abdominal segment 3 foveolate-reticulate, foveolae smaller and more regular on dorsal than lateral mesosoma, with rugulae present between foveolae on lateral mesosoma, especially posteriorly; declivitous face of propodeum with narrow transverse rugulae and sometimes punctae; AT4 quite smooth and shiny despite numerous minute piligerous punctulae.
|
||
</paragraph>
|
||
<caption id="DF0F4DD4E57AFF94FCC4FBB0BE72073B" ID-DOI="http://doi.org/10.5281/zenodo.3542208" ID-Zenodo-Dep="3542208" httpUri="https://zenodo.org/record/3542208/files/figure.png" pageId="47" pageNumber="48" startId="47.[818,845,1061,1078]" targetBox="[819,1440,147,1026]" targetPageId="47">
|
||
<paragraph id="8BCF1D5CE57AFF94FCC4FBB0BE72073B" blockId="47.[818,1444,1061,1129]" pageId="47" pageNumber="48">
|
||
<emphasis id="B904C14EE57AFF94FCC4FBB0B8660764" bold="true" box="[818,884,1061,1078]" pageId="47" pageNumber="48">Fig. 37.</emphasis>
|
||
Stacked digital color images of
|
||
<taxonomicName id="4C7066DFE57AFF94FB51FBB0BE1A0764" authority="Hita-Garcia & Lieberman & Audisio & Liu & Economo, 2019" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[1191,1288,1061,1078]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="hawkesi" status="sp. nov.">
|
||
<emphasis id="B904C14EE57AFF94FB51FBB0BE1A0764" box="[1191,1288,1061,1078]" italics="true" pageId="47" pageNumber="48">D. hawkesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377C35E57AFF94FAE1FBB0BE5F0764" box="[1303,1357,1061,1078]" pageId="47" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
|
||
holotype (CASENT0235470—from https://www.antweb.org, photographerWill Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF1D5CE57AFF94FF44FAA6B9EC0539" blockId="47.[146,771,686,1673]" pageId="47" pageNumber="48">
|
||
<emphasis id="B904C14EE57AFF94FF44FAA6BBE90614" box="[178,251,1331,1350]" italics="true" pageId="47" pageNumber="48">Setation</emphasis>
|
||
consisting of distinct, erect, white pilosity on front of head, mesosomal, petiolar and abdominal dorsa, generally becoming longer and denser with each succesive tagma in the posterior direction; lateral mesosoma with mostly appressed pubescence; sides of abdominal segments 3 and 4, petiolar sternite, and abdominal sternite 3 also bearing long erect setae in addition to underlayer of appressed pubescence; abdominal segments five through seven with similar pilosity to AT4, setae not noticeably longer or more curved; scape and legs with long, dense pubescence, mostly appressed to decumbent, with scattered erect setae; ectal face of mandible with abundant, long, curved, erect to suberect setae; setation of masticatory margin undetermined.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE57AFF94FF44F9E3B9B905DB" blockId="47.[146,771,686,1673]" box="[178,683,1654,1673]" pageId="47" pageNumber="48">
|
||
<emphasis id="B904C14EE57AFF94FF44F9E3BBF505DB" box="[178,231,1654,1673]" italics="true" pageId="47" pageNumber="48">Color</emphasis>
|
||
uniformly dull testaceous-yellow to light brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E57AFF94FF64F95DBA080494" pageId="47" pageNumber="48" type="etymology">
|
||
<paragraph id="8BCF1D5CE57AFF94FF64F95DBBE70589" blockId="47.[146,770,1736,1990]" box="[146,245,1736,1755]" pageId="47" pageNumber="48">
|
||
<heading id="D087AA30E57AFF94FF64F95DBBE70589" bold="true" box="[146,245,1736,1755]" fontSize="8" level="3" pageId="47" pageNumber="48" reason="6">
|
||
<emphasis id="B904C14EE57AFF94FF64F95DBBE70589" bold="true" box="[146,245,1736,1755]" pageId="47" pageNumber="48">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE57AFF94FF64F973BA080494" blockId="47.[146,770,1736,1990]" pageId="47" pageNumber="48">
|
||
The gryphon of mythology was a legendary creature with a body composed of different animals.
|
||
<taxonomicName id="4C7066DFE57AFF94FE4CF896B9930444" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[442,641,1795,1814]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gryphon">
|
||
<emphasis id="B904C14EE57AFF94FE4CF896B9930444" box="[442,641,1795,1814]" italics="true" pageId="47" pageNumber="48">Discothyrea gryphon</emphasis>
|
||
</taxonomicName>
|
||
is named like for its strange combination of characters, which are approximated only by the aberrant
|
||
<taxonomicName id="4C7066DFE57AFF94FE9EF8ABBAC60403" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[360,468,1854,1873]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="chimera">
|
||
<emphasis id="B904C14EE57AFF94FE9EF8ABBAC60403" box="[360,468,1854,1873]" italics="true" pageId="47" pageNumber="48">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
. The archaic spelling
|
||
<taxonomicName id="4C7066DFE57AFF94FD45F8ABB8100403" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[691,770,1854,1873]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gryphon">
|
||
<emphasis id="B904C14EE57AFF94FD45F8ABB8100403" box="[691,770,1854,1873]" italics="true" pageId="47" pageNumber="48">gryphon</emphasis>
|
||
</taxonomicName>
|
||
is used in preference to the Latinate
|
||
<emphasis id="B904C14EE57AFF94FDF6F8CEB928043C" box="[512,570,1883,1902]" italics="true" pageId="47" pageNumber="48">griffin</emphasis>
|
||
as wordplay on the Greek root
|
||
<emphasis id="B904C14EE57AFF94FEF4F8ECBA5004D9" box="[258,322,1913,1931]" italics="true" pageId="47" pageNumber="48">grypos</emphasis>
|
||
, meaning hooked, in reference to the recurved abdomen of
|
||
<taxonomicName id="4C7066DFE57AFF94FEF9F803BA9304FB" authorityName="Roger" authorityYear="1863" box="[271,385,1942,1961]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904C14EE57AFF94FEF9F803BA9304FB" box="[271,385,1942,1961]" italics="true" pageId="47" pageNumber="48">Discothyrea</emphasis>
|
||
</taxonomicName>
|
||
. The specific epithet is given as an appositive noun.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E57AFF94FCC4FB0ABEB60658" pageId="47" pageNumber="48" type="distribution">
|
||
<paragraph id="8BCF1D5CE57AFF94FCC4FB0ABF0A07E0" blockId="47.[818,1444,1183,1290]" box="[818,1048,1183,1202]" pageId="47" pageNumber="48">
|
||
<heading id="D087AA30E57AFF94FCC4FB0ABF0A07E0" bold="true" box="[818,1048,1183,1202]" fontSize="8" level="3" pageId="47" pageNumber="48" reason="6">
|
||
<emphasis id="B904C14EE57AFF94FCC4FB0ABF0A07E0" bold="true" box="[818,1048,1183,1202]" pageId="47" pageNumber="48">Distribution and Biology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE57AFF94FCC4FB28BEB60658" blockId="47.[818,1444,1183,1290]" pageId="47" pageNumber="48">
|
||
At present, known only from the
|
||
<typeStatus id="54CBA3FEE57AFF94FB90FB28BF990782" box="[1126,1163,1213,1232]" pageId="47" pageNumber="48">type</typeStatus>
|
||
locality in Kayove in
|
||
<collectingCountry id="F3675DCCE57AFF94FAA1FB29BEB0079D" box="[1367,1442,1212,1231]" name="Rwanda" pageId="47" pageNumber="48">Rwanda</collectingCountry>
|
||
and Mkomazi in
|
||
<collectingCountry id="F3675DCCE57AFF94FC30FB4FBF0407BF" box="[966,1046,1242,1261]" name="Tanzania" pageId="47" pageNumber="48">Tanzania</collectingCountry>
|
||
(
|
||
<figureCitation id="134B01D9E57AFF94FBD4FB4FBF4B07BF" box="[1058,1113,1242,1261]" captionStart="Fig" captionStartId="5.[146,173,1283,1300]" captionTargetBox="[153,1435,148,1247]" captionTargetId="figure@5.[152,1435,147,1248]" captionTargetPageId="5" captionText="Fig. 4. Maps showing the known distribution ranges of Afrotropical Discothyrea species." figureDoi="http://doi.org/10.5281/zenodo.3542140" httpUri="https://zenodo.org/record/3542140/files/figure.png" pageId="47" pageNumber="48">
|
||
Fig.
|
||
<date id="FFCE3B9CE57AFF94FBB1FB4FBF4B07BF" box="[1095,1113,1242,1261]" pageId="47" pageNumber="48">4I</date>
|
||
</figureCitation>
|
||
). Both localities are montane forest at elevations from
|
||
<quantity id="4C88B0B9E57AFF94FC4BFB62BF5F0658" box="[957,1101,1271,1290]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="47" pageNumber="48" unit="m" value="1850.0" valueMax="2100.0" valueMin="1600.0">
|
||
<date id="FFCE3B9CE57AFF94FC4BFB62BF260658" box="[957,1076,1271,1290]" pageId="47" pageNumber="48" value="1600" valueMax="2100">1600 to 2100</date>
|
||
m
|
||
</quantity>
|
||
.
|
||
<taxonomicName id="4C7066DFE57AFF94FBA2FB62BE1F0658" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[1108,1293,1271,1290]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gryphon">
|
||
<emphasis id="B904C14EE57AFF94FBA2FB62BE1F0658" box="[1108,1293,1271,1290]" italics="true" pageId="47" pageNumber="48">Discothyrea gryphon</emphasis>
|
||
</taxonomicName>
|
||
lives in leaf litter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E57AFF94FCC4FAA2BEB605B8" pageId="47" pageNumber="48" type="discussion">
|
||
<paragraph id="8BCF1D5CE57AFF94FCC4FAA2B8840618" blockId="47.[818,1442,1335,1589]" box="[818,918,1335,1354]" pageId="47" pageNumber="48">
|
||
<heading id="D087AA30E57AFF94FCC4FAA2B8840618" bold="true" box="[818,918,1335,1354]" fontSize="8" level="3" pageId="47" pageNumber="48" reason="6">
|
||
<emphasis id="B904C14EE57AFF94FCC4FAA2B8840618" bold="true" box="[818,918,1335,1354]" pageId="47" pageNumber="48">Comments</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE57AFF94FCC4FAC1BE560567" blockId="47.[818,1442,1335,1589]" pageId="47" pageNumber="48">
|
||
<taxonomicName id="4C7066DFE57AFF94FCC4FAC1B8E40635" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[818,1014,1364,1383]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gryphon">
|
||
<emphasis id="B904C14EE57AFF94FCC4FAC1B8E40635" box="[818,1014,1364,1383]" italics="true" pageId="47" pageNumber="48">Discothyrea gryphon</emphasis>
|
||
</taxonomicName>
|
||
is easily recognizable on the basis of a combination of highly species-specific character states. The presence of a subapical tooth, dense standing pilosity, subquadrate head shape with sharply defined genal corners, and the row of long, straight setae on the anterior clypeal margin rapidly discriminate it from the remainder of the complex. As noted above, the similarity of
|
||
<taxonomicName id="4C7066DFE57AFF94FCC4F990B8890545" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[818,923,1540,1559]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gryphon">
|
||
<emphasis id="B904C14EE57AFF94FCC4F990B8890545" box="[818,923,1540,1559]" italics="true" pageId="47" pageNumber="48">D. gryphon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7066DFE57AFF94FC31F990BF390545" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[967,1067,1540,1559]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="chimera">
|
||
<emphasis id="B904C14EE57AFF94FC31F990BF390545" box="[967,1067,1540,1559]" italics="true" pageId="47" pageNumber="48">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
, in the characters of the head and general habitus, suggest that they may constitute a natural group.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE57AFF94FCC4F9F7B89A0527" blockId="47.[818,1444,1634,1770]" box="[818,904,1634,1653]" pageId="47" pageNumber="48">
|
||
<heading id="D087AA30E57AFF94FCC4F9F7B89A0527" bold="true" box="[818,904,1634,1653]" fontSize="8" level="3" pageId="47" pageNumber="48" reason="6">
|
||
<emphasis id="B904C14EE57AFF94FCC4F9F7B89A0527" bold="true" box="[818,904,1634,1653]" pageId="47" pageNumber="48">Variation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE57AFF94FCC4F9EABEB605B8" blockId="47.[818,1444,1634,1770]" pageId="47" pageNumber="48">All examined specimens are consistent in virtually all characters. However, the development of the subapical mandibular denticle ranges from an indistinct angle to a relatively distinct, small denticle. This variation may be due to the denticle being worn down with age.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |