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<mods:title id="2812A5A4E8ED4EB7BDD22D870B9904A5">The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha)</mods:title>
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<mods:namePart id="D43882C64D30C7C4FAF67D3CCD2ADE61">Vignaud, Patrick</mods:namePart>
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Machimosaurini
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.,
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<emphasis id="B90DE802FF81730E7B680DC30FF4FA12" bold="true" box="[520,718,1491,1517]" pageId="49" pageNumber="596">(Zoological Code)</emphasis>
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<emphasis id="B90DE802FF81730E79D20E010C62F9D3" bold="true" box="[178,344,1553,1580]" italics="true" pageId="49" pageNumber="596">Machimosaurini</emphasis>
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.,
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<paragraph id="8BC63410FF81730E79110E290DA5F98F" blockId="49.[113,698,1553,1648]" pageId="49" pageNumber="596">RegNum registration number 798.</paragraph>
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genus:
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<emphasis id="B90DE802FF81730E79880E810CB9F956" box="[232,387,1681,1705]" italics="true" pageId="49" pageNumber="596">Machimosaurus</emphasis>
<bibRefCitation id="EFE849E1FF81730E78E80E810F14F956" author="von Meyer H" box="[392,558,1681,1705]" pageId="49" pageNumber="596" pagination="557 - 62" refId="ref56718" refString="von Meyer H. Mittheilungen, an Professor Bronn gerichtet. Neues Jahrbuch fur Mineralogie, Geognosie, Geologie und Petrefaktenkunde 1837; 1837: 557 - 62." type="journal article" year="1837">von Meyer 1837</bibRefCitation>
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(emend.
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).
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<emphasis id="B90DE802FF81730E78E30EF80CD3F8FD" box="[387,489,1768,1794]" italics="true" pageId="49" pageNumber="596">Etymology</emphasis>
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Pertaining to
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<emphasis id="B90DE802FF81730E78640F1E0C9BF8D9" box="[260,417,1806,1830]" italics="true" pageId="49" pageNumber="596">Machimosaurus</emphasis>
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. The stem
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- is based on the genus
<taxonomicName id="4C794F93FF81730E78660F3E0C99F8B9" authorityName="von Meyer" authorityYear="1837" box="[262,419,1838,1862]" class="Reptilia" family="Teleosauridae" genus="Machimosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E78660F3E0C99F8B9" box="[262,419,1838,1862]" italics="true" pageId="49" pageNumber="596">Machimosaurus</emphasis>
</taxonomicName>
. -
<emphasis id="B90DE802FF81730E78DB0F3E0CEFF8BA" box="[443,469,1838,1861]" italics="true" pageId="49" pageNumber="596">ini</emphasis>
is a Latin masculine plural suffix for pertaining to. Under Article 29.2 of the Zoological Code, the suffix -
<emphasis id="B90DE802FF81730E784F0F7D0C73F87B" box="[303,329,1901,1924]" italics="true" pageId="49" pageNumber="596">ini</emphasis>
denotes a tribe rank within the family-group.
</paragraph>
</subSubSection>
<subSubSection id="C363679BFF81730E7D4108800E52FDC7" pageId="49" pageNumber="596" type="description">
<paragraph id="8BC63410FF81730E7D4108800981FF55" blockId="49.[810,1459,144,269]" box="[1057,1211,144,170]" pageId="49" pageNumber="596">
<emphasis id="B90DE802FF81730E7D4108800981FF55" box="[1057,1211,144,170]" italics="true" pageId="49" pageNumber="596">Geological range</emphasis>
</paragraph>
<paragraph id="8BC63410FF81730E7A4A08A60889FEF2" blockId="49.[810,1459,144,269]" pageId="49" pageNumber="596">
Middle Jurassic (Bathonian) to Early Cretaceous (Hauterivian or possibly late Barremian?) (
<bibRefCitation id="EFE849E1FF81730E7D3908C609C1FF11" author="Fanti F &amp; Miyashita T &amp; Cantelli L" box="[1113,1275,214,238]" pageId="49" pageNumber="596" pagination="263 - 74" refId="ref51216" refString="Fanti F, Miyashita T, Cantelli L et al. The largest thalattosuchian (Crocodylomorpha) supports teleosaurid survival across the Jurassic-Cretaceous boundary. Cretaceous Research 2016; 61: 263 - 74. https: // doi. org / 10.1016 / j. cretres. 2015.11.011" type="journal article" year="2016">
Fanti
<emphasis id="B90DE802FF81730E7DF308C60986FF11" box="[1171,1212,214,238]" italics="true" pageId="49" pageNumber="596">et al</emphasis>
. 2016
</bibRefCitation>
,
<bibRefCitation id="EFE849E1FF81730E7C6608C60897FF11" author="Jouve S &amp; Mennecart B &amp; Douteau J" box="[1286,1453,214,238]" pageId="49" pageNumber="596" pagination="863 - 76" refId="ref55141" refString="Jouve S, Mennecart B, Douteau J et al. The oldest durophagous teleosauroid (Crocodylomorpha, Thalattosuchia) from the lower Bathonian of central High Atlas, Morocco. Palaeontology 2016; 59: 863 - 76. https: // doi. org / 10.1111 / pala. 12262" type="journal article" year="2016">
Jouve
<emphasis id="B90DE802FF81730E7C2508C60854FF11" box="[1349,1390,214,238]" italics="true" pageId="49" pageNumber="596">et al</emphasis>
. 2016
</bibRefCitation>
,
<bibRefCitation id="EFE849E1FF81730E7A4A08E50EE6FEF2" author="Cortes D &amp; Larsson HCE &amp; Maxwell EE" box="[810,988,245,269]" pageId="49" pageNumber="596" pagination="365 - 79" refId="ref50271" refString="Cortes D, Larsson HCE, Maxwell EE et al. An Early Cretaceous teleosauroid (Crocodylomorpha: Thalattosuchia) from Colombia. Ameghiniana 2019; 56: 365 - 79." type="journal article" year="2019">
Cortés
<emphasis id="B90DE802FF81730E7A1508E60EA7FEF2" box="[885,925,245,269]" italics="true" pageId="49" pageNumber="596">et al</emphasis>
. 2019
</bibRefCitation>
,
<bibRefCitation id="EFE849E1FF81730E7A8708E50989FEF2" author="Johnson MM &amp; Young MT &amp; Brusatte SL" box="[999,1203,245,269]" pageId="49" pageNumber="596" pagination="9808" refId="ref54662" refString="Johnson MM, Young MT, Brusatte SL. The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution. PeerJ 2020 a; 8: e 9808. https: // doi. org / 10.7717 / peerj. 9808" type="journal article" year="2020">
Johnson
<emphasis id="B90DE802FF81730E7D2108E60955FEF2" box="[1089,1135,245,269]" italics="true" pageId="49" pageNumber="596">et al.</emphasis>
2020
<bibRefCitation id="EFE849E1FF81730E7DC808E50989FEF2" author="Young MT &amp; Brignon A &amp; Sachs S" box="[1192,1203,245,269]" pageId="49" pageNumber="596" pagination="510 - 53" refId="ref61294" refString="Young MT, Brignon A, Sachs S et al. Cutting the Gordian knot: a historical and taxonomic revision of the Jurassic crocodylomorph Metriorhynchus. Zoological Journal of the Linnean Society 2021; 192: 510 - 53." type="journal article" year="2021">a</bibRefCitation>
</bibRefCitation>
, Young and Sachs 2021).
</paragraph>
<paragraph id="8BC63410FF81730E7AB109250836FEB0" blockId="49.[977,1292,309,335]" box="[977,1292,309,335]" pageId="49" pageNumber="596">
<emphasis id="B90DE802FF81730E7AB109250836FEB0" box="[977,1292,309,335]" italics="true" pageId="49" pageNumber="596">PhyloCode phylogenetic definition</emphasis>
</paragraph>
<paragraph id="8BC63410FF81730E7A4A094C0EBCFE2E" blockId="49.[810,1459,348,466]" pageId="49" pageNumber="596">
The largest clade within
<taxonomicName id="4C794F93FF81730E7D76094C099CFE8B" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[1046,1190,348,372]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="582" phylum="Chordata" rank="superFamily" superFamily="Teleosauroidea">
<emphasis id="B90DE802FF81730E7D76094C099CFE8B" box="[1046,1190,348,372]" italics="true" pageId="49" pageNumber="596">Teleosauroidea</emphasis>
</taxonomicName>
containing
<taxonomicName id="4C794F93FF81730E7C77094C0900FE6C" authority="(von Meyer 1837)" baseAuthorityName="von Meyer" baseAuthorityYear="1837" class="Reptilia" family="Teleosauridae" genus="Machimosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="596" phylum="Chordata" rank="species" species="hugii">
<emphasis id="B90DE802FF81730E7C77094C0E61FE6C" italics="true" pageId="49" pageNumber="596">Machimosaurus hugii</emphasis>
(
<bibRefCitation id="EFE849E1FF81730E7A14096B0915FE6C" author="von Meyer H" box="[884,1071,379,403]" pageId="49" pageNumber="596" pagination="557 - 62" refId="ref56718" refString="von Meyer H. Mittheilungen, an Professor Bronn gerichtet. Neues Jahrbuch fur Mineralogie, Geognosie, Geologie und Petrefaktenkunde 1837; 1837: 557 - 62." type="journal article" year="1837">von Meyer 1837</bibRefCitation>
)
</taxonomicName>
but not
<taxonomicName id="4C794F93FF81730E7DCD096C095DFE4D" authority="(Eudes-Deslongchamps 1868)" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1868" genus="Neosteneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="596" phylum="Chordata" rank="species" species="edwardsi">
<emphasis id="B90DE802FF81730E7DCD096C0889FE6C" box="[1197,1459,379,403]" italics="true" pageId="49" pageNumber="596">Neosteneosaurus edwardsi</emphasis>
(Eudes-Deslongchamps 1868)
</taxonomicName>
. Definition from
<bibRefCitation id="EFE849E1FF81730E7C43098A0EBBFE2D" author="Johnson MM &amp; Young MT &amp; Brignon A" pageId="49" pageNumber="596" pagination="1 - 7" refId="ref54877" refString="Johnson MM, Young MT, Brignon A et al. Addition to ' The phylogenetics of Teleosauroidea (Crocodylomorpha; Thalattosuchia) and implications for their ecology and evolution'. Bulletin of Phylogenetic Nomenclature 2022 b; 1: 1 - 7. https: // doi. org / 10.11646 / bpn. 1.1.1" type="journal article" year="2022">
Johnson
<emphasis id="B90DE802FF81730E7CE1098B0897FE4D" box="[1409,1453,410,434]" italics="true" pageId="49" pageNumber="596">et al</emphasis>
. (2022b)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BC63410FF81730E7D7209EA09F6FDEB" blockId="49.[1042,1228,507,531]" box="[1042,1228,506,532]" pageId="49" pageNumber="596">
<emphasis id="B90DE802FF81730E7D7209EA09F6FDEB" box="[1042,1228,506,532]" italics="true" pageId="49" pageNumber="596">Reference phylogeny</emphasis>
</paragraph>
<paragraph id="8BC63410FF81730E7A4A0A300E52FDC7" blockId="49.[810,872,545,568]" box="[810,872,544,568]" pageId="49" pageNumber="596">
<figureCitation id="13422895FF81730E7A4A0A300E58FDC7" box="[810,866,544,568]" captionStart="Figure 3" captionStartId="19.[113,178,1185,1209]" captionTargetBox="[117,1455,147,1154]" captionTargetId="figure-283@19.[114,1458,144,1157]" captionTargetPageId="19" captionText="Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/." figureDoi="http://doi.org/10.5281/zenodo.11241251" httpUri="https://zenodo.org/record/11241251/files/figure.png" pageId="49" pageNumber="596">Fig. 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C363679BFF81730E7A5B0A480EEDFAB7" pageId="49" pageNumber="596" type="diagnosis">
<paragraph id="8BC63410FF81730E7A5B0A480898FD8D" blockId="49.[810,1459,600,1352]" box="[827,1442,600,626]" pageId="49" pageNumber="596">
<emphasis id="B90DE802FF81730E7A5B0A480898FD8D" box="[827,1442,600,626]" italics="true" pageId="49" pageNumber="596">Zoological Code diagnosis and PhyloCode diagnostic apomorphies</emphasis>
</paragraph>
<paragraph id="8BC63410FF81730E7A4A0A6E0EEDFAB7" blockId="49.[810,1459,600,1352]" pageId="49" pageNumber="596">
Teleosauroid crocodylomorphs with the following unique combination of characters (13): expanded network of neurovascular foramina on the premaxillae, maxillae and dentaries (shared with the teleosaurids
<taxonomicName id="4C794F93FF81730E7D7B0ACC094CFD0B" authorityName="von Meyer" authorityYear="1832" box="[1051,1142,732,756]" class="Reptilia" family="Teleosauridae" genus="Aeolodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E7D7B0ACC094CFD0B" box="[1051,1142,732,756]" italics="true" pageId="49" pageNumber="596">Aeolodon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C794F93FF81730E7DD60ACC0879FD0B" authorityName="Andrews" authorityYear="1913" box="[1206,1347,732,756]" family="Teleosauridae" genus="Mycterosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E7DD60ACC0879FD0B" box="[1206,1347,732,756]" italics="true" pageId="49" pageNumber="596">Mycterosuchus</emphasis>
</taxonomicName>
; and also seen in extant crocodylians as housing the dome pressure receptors) (86.1); supratemporal fossae are parallelogram in shape (194.5); supratemporal fenestrae at least twice as long as the anterior width (196.1); supratemporal fenestrae at least twice as long as the orbits (shared with
<taxonomicName id="4C794F93FF81730E7DA10B690875FC6E" baseAuthorityName="de Stefano" baseAuthorityYear="1903" box="[1217,1359,889,913]" class="Reptilia" family="Dyrosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="596" phylum="Chordata" rank="family">Dyrosauridae</taxonomicName>
) (198.3); deep and pronounced reception pits conspicuous along the anterior and mid-maxilla (596.1); anterior maxillary tooth crowns have blunt and rounded apices (631.1); anterior-middle tooth crowns not recurved (647.0); presence of false ziphodonty across the dentition (652.2); apical tooth crowns have a strongly developed anastomosed enamel ornamentation (shared with the goniopholidids
<taxonomicName id="4C794F93FF81730E7A9A0C440955FB93" box="[1018,1135,1108,1132]" class="Reptilia" family="Goniopholididae" genus="Goniopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E7A9A0C440955FB93" box="[1018,1135,1108,1132]" italics="true" pageId="49" pageNumber="596">Goniopholis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C794F93FF81730E7DC80C440847FB93" authorityName="Salisbury &amp; Naish" authorityYear="2011" box="[1192,1405,1108,1132]" class="Reptilia" family="Goniopholididae" genus="Anteophthalmosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E7DC80C440847FB93" box="[1192,1405,1108,1132]" italics="true" pageId="49" pageNumber="596">Anteophthalmosuchus</emphasis>
</taxonomicName>
, and the metriorhynchid
<taxonomicName id="4C794F93FF81730E7D610C640944FB74" box="[1025,1150,1140,1163]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E7D610C640944FB74" box="[1025,1150,1140,1163]" italics="true" pageId="49" pageNumber="596">Torvoneustes</emphasis>
</taxonomicName>
) (663.1); three sacral vertebrae (708.1); sub-rectangular ischial plate (shared with some metriorhynchids) (804.1); tuberosity of the tibia is angled ventrally (824.1); dorsal osteoderm ornamentation composed of pits of highly variable size and length, and on keeled osteoderms the pits can become elongate pits, especially along the lateral margins (840.3).
</paragraph>
</subSubSection>
<subSubSection id="C363679BFF81730E7D520D780823FA5A" pageId="49" pageNumber="596" type="discussion">
<paragraph id="8BC63410FF81730E7D520D780991FA7D" blockId="49.[810,1305,1384,1446]" box="[1074,1195,1384,1410]" pageId="49" pageNumber="596">
<emphasis id="B90DE802FF81730E7D520D780991FA7D" box="[1074,1195,1384,1410]" italics="true" pageId="49" pageNumber="596">Composition</emphasis>
</paragraph>
<paragraph id="8BC63410FF81730E7A4A0D9E0823FA5A" blockId="49.[810,1305,1384,1446]" box="[810,1305,1422,1446]" pageId="49" pageNumber="596">
<taxonomicName id="4C794F93FF81730E7A4A0D9E0E88FA59" authorityName="Johnson, Young, Steel, Foffa, Smith, Hua, Havlik, Howlett &amp; Dyke" authorityYear="2017" box="[810,946,1422,1446]" genus="Lemmysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E7A4A0D9E0E88FA59" box="[810,946,1422,1446]" italics="true" pageId="49" pageNumber="596">Lemmysuchus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C794F93FF81730E7ADD0D9E0962FA59" authorityName="von Meyer" authorityYear="1837" box="[957,1112,1422,1446]" class="Reptilia" family="Teleosauridae" genus="Machimosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E7ADD0D9E0962FA59" box="[957,1112,1422,1446]" italics="true" pageId="49" pageNumber="596">Machimosaurus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C794F93FF81730E7DEE0D9E0828FA59" authorityName="Johnson, Young &amp; Brusatte" authorityYear="2019" box="[1166,1298,1422,1446]" genus="Yvridiosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF81730E7DEE0D9E0828FA59" box="[1166,1298,1422,1446]" italics="true" pageId="49" pageNumber="596">Yvridiosuchus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>