treatments-xml/data/86/43/87/864387EE1127724EF8A10F68815222F9.xml
2024-06-21 12:42:39 +02:00

292 lines
33 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="08BBBF4B604801803B352B7248706764" ID-CLB-Dataset="3175" ID-DOI="10.11646/zootaxa.5072.3.1" ID-GBIF-Dataset="a5454a8b-04cd-4cc6-ae0b-f13f31a7af34" ID-ISSN="1175-5326" ID-Zenodo-Dep="5744549" ID-ZooBank="CAB2C8C4-E0A4-4805-9DEB-94769C2F90EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638283643950" checkinUser="plazi" docAuthor="Mahecha-J, Oscar, Florczyk, Klaudia, Willmott, Keith, Cerdeña, José, Zubek, Anna, Boyer, Pierre, Farfán, Jackie, Lachowska-Cierlik, Dorota &amp; Pyrcz, Tomasz W." docDate="2021" docId="864387EE1127724EF8A10F68815222F9" docLanguage="en" docName="zootaxa.5072.3.1.pdf" docOrigin="Zootaxa 5072 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Manerebia pauperata Pyrcz &amp; Willmott 2006, n. stat." docType="treatment" docVersion="6" lastPageNumber="230" masterDocId="7A7AFF9611307253F836080285142354" masterDocTitle="Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina)" masterLastPageNumber="237" masterPageNumber="201" pageNumber="224" updateTime="1699471567071" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="637DF702E42565ED8297C0C5D73857C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC4FB314165E27BEA67C163145F99F8A">
<mods:title id="B06CAE5C0FD9A6D3B1890F728CA7BEB9">Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina)</mods:title>
</mods:titleInfo>
<mods:name id="649F56F2376A8D8D75B1372DC144539D" type="personal">
<mods:role id="2435BCEE4B726489879F8EE5E44E9CE4">
<mods:roleTerm id="FE7C345EC9C9E4C97B8DAD4E54D5BD08">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0633D8DD7DB86099E69FAF72556E6E9A">Mahecha-J, Oscar</mods:namePart>
<mods:affiliation id="08B5A26E54B44D5F34B7CBD8F9B56C4E">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland. &amp; Instituto de Ciencias Naturales ICN / Laboratorio de Ecología del Paisaje y Modelación de Ecosistemas ECOLMOD, Universidad Nacional de Colombia. Carrera 30 No. 45 - 03, Bogotá D. C., Colombia</mods:affiliation>
</mods:name>
<mods:name id="C8D354745A8653BCBCCE442D58539EC2" type="personal">
<mods:role id="67AC10D94618CEB3E046C72DA9BA9953">
<mods:roleTerm id="EBFDC2D0501B4DAB6BCB11B30FFF0E87">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6B1E5B125288091E2AEC3482097ABD2">Florczyk, Klaudia</mods:namePart>
<mods:nameIdentifier id="89A665E8A52DF069D7CEE78CECD33BC9" type="ORCID">0000-0002-5731-8605</mods:nameIdentifier>
<mods:affiliation id="64007AF1CA300A0DF98CAB4832630502">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland. &amp; klaudia. florczyk @ uj. edu. pl; https: // orcid. org / 0000 - 0002 - 5731 - 8605</mods:affiliation>
<mods:nameIdentifier id="5D7562DE80C14BFA0045111E913F1B39" type="email">klaudia.florczyk@uj.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="F863D8DB74ADDB56415C3F0A20B1BF70" type="personal">
<mods:role id="58A4F9ADD4C15F9842DB2B378177FB2E">
<mods:roleTerm id="8B770A2ED8ACC7824E1103DE97205A0C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="12EC3D29645E8551A9E2AB2667290430">Willmott, Keith</mods:namePart>
<mods:affiliation id="059A27327C67E34BD78B054938778370">Florida Natural History Museum, University of Florida, Gainesville, USA.</mods:affiliation>
</mods:name>
<mods:name id="A9AA084F5F334D21E71FA46BF2382C32" type="personal">
<mods:role id="9071748FFF2ECDBFBAFC92E5E5437A50">
<mods:roleTerm id="DE1A2DEB4A298C7EDBFB99FE9F80B182">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97243302228BC7E55226684824F9B829">Cerdeña, José</mods:namePart>
<mods:nameIdentifier id="83BD1F972F24A755D01A6821EE927270" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="772DFCE0AEB340596185EC0A5A97DB1F">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Escuela de Biología UNSA, Arequipa, Perú. &amp; cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480</mods:affiliation>
<mods:nameIdentifier id="8FE289074C70577EB75093D4CF84B6A2" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
</mods:name>
<mods:name id="52D4CE708B32B7D81E5B72DBADD5040C" type="personal">
<mods:role id="5B4E9CABBCF477A628008CCC51B43F25">
<mods:roleTerm id="B64F1618A4E323BF0D9CB3A9D1CC1DCA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7120EABD22A96A7E2E1AF0DDCA0B22A">Zubek, Anna</mods:namePart>
<mods:nameIdentifier id="007F695C7A283384BCC2D9E3122CC2AA" type="ORCID">0000-0003-3663-522X</mods:nameIdentifier>
<mods:affiliation id="357AFB2918D558570D1D11107BB3B77B">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland. &amp; anna. zubek @ uj. edu. pl; https: // orcid. org / 0000 - 0003 - 3663 - 522 X</mods:affiliation>
<mods:nameIdentifier id="C05EBBB366661E1E9586E2C64FEFB526" type="email">anna.zubek@uj.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="082734E74187E8D07DDC6B7C061F5AA4" type="personal">
<mods:role id="7802343305C3DC5F4D7932C6D69320C0">
<mods:roleTerm id="944DB93865DC0720C034A77D2BBA87D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="764CFF084A31F54CB284900732D8181B">Boyer, Pierre</mods:namePart>
<mods:affiliation id="FE0E464663856C7CAA75E09EAFDBB859">Lotissement lHorizon, 13610, Le Puy Sainte Réparade, France. pierdom @ aliceadsl. fr</mods:affiliation>
</mods:name>
<mods:name id="B840B7CD1AB0415BD54AEEEA74020542" type="personal">
<mods:role id="AC0350AE461394DAABE5E74B5B402408">
<mods:roleTerm id="02F249A895DE49B3D270BDB0809D0129">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C3E4D56BF8812E0731D090D36DF30C13">Farfán, Jackie</mods:namePart>
<mods:nameIdentifier id="A729624850C72854BBCA9C2F67C94762" type="ORCID">0000-0001-5913-0756</mods:nameIdentifier>
<mods:affiliation id="614EC520B78CB22BC50C62119A37F4CA">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Escuela de Biología UNSA, Arequipa, Perú. &amp; jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="9F95D4B22AFA0E3C6265AC9CD8053AFA" type="email">jjackie4u@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="57BB05A7FDC6F2C8C0DA6199D179E2F0" type="personal">
<mods:role id="FAB888E090DFD47A027FEA692483B19D">
<mods:roleTerm id="90914701E272992AC4271E826C40F1D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C167D53545563D37125B343681FA79E1">Lachowska-Cierlik, Dorota</mods:namePart>
<mods:nameIdentifier id="565C28952424E1463E3638C37626D724" type="ORCID">0000-0001-9072-1677</mods:nameIdentifier>
<mods:affiliation id="CAF85EC90B9AC9B6B4FEF5D9640451C6">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland &amp; dorota. lachowska-cierlik @ uj. edu. pl; https: // orcid. org / 0000 - 0001 - 9072 - 1677</mods:affiliation>
<mods:nameIdentifier id="1E286BBE581E1413B7B0F82226D8294C" type="email">dorota.lachowska-cierlik@uj.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="300BDD83CA5F4A4491CD9F7523F8C26F" type="personal">
<mods:role id="A00630B6F77E8658B914BEB8D10E7DC7">
<mods:roleTerm id="4D01610B01408D1C120F56A95A4E960B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FA97AD1432320E76F896407D093B3F0">Pyrcz, Tomasz W.</mods:namePart>
<mods:nameIdentifier id="8512F38EA968D14245588ED49383DCF5" type="ORCID">0000-0003-4822-0670</mods:nameIdentifier>
<mods:affiliation id="86337908375FB755711CC5AA404C01A1">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland. &amp; Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland &amp; tomasz. pyrcz @ uj. edu. pl; https: // orcid. org / 0000 - 0003 - 4822 - 0670</mods:affiliation>
<mods:nameIdentifier id="3701B758DA88E24EEDD075654983B704" type="email">tomasz.pyrcz@uj.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="EE214FE10866B21C41571A8E8144967C">text</mods:typeOfResource>
<mods:relatedItem id="9483DC41CD03023275625F4C4A975612" type="host">
<mods:titleInfo id="025961A1E088D13C7749D5CEDBBAEAEE">
<mods:title id="4D3909EC127005EACCD2A472F6D6EEB8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B15F51DAD92B0C45A5E62615C3360FE4">
<mods:date id="3E5B1BDC1D4E4A2875B49650FD1A8C3B">2021</mods:date>
<mods:detail id="75AC41486788D4DD67D25E9C86A1942E" type="pubDate">
<mods:number id="98004C60710C64F238437E8F807CFCAE">2021-11-30</mods:number>
</mods:detail>
<mods:detail id="742CFF07B62E40BA2F8CD6B8755DD4A3" type="volume">
<mods:number id="290D2B4C1348B9304625D2C6CF4E4A48">5072</mods:number>
</mods:detail>
<mods:detail id="71DE2C13A448E007BD030D02C26ACBEB" type="issue">
<mods:number id="C2C5FD8A175B5136CB090A4814A176F0">3</mods:number>
</mods:detail>
<mods:extent id="B1958E2C63AB454959881B09AE85B707" unit="page">
<mods:start id="E50462845D23A53A2C34C301AD456EF9">201</mods:start>
<mods:end id="0AE50EFCEEA083AD3B29C6A3C2512386">237</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="24145C55DC1532C177E7227209576819">journal article</mods:classification>
<mods:identifier id="CB2924A0CE7928D6B06ABDC921853C74" type="CLB-Dataset">3175</mods:identifier>
<mods:identifier id="F92B17DB13920A03C5B19FB9434E2187" type="DOI">10.11646/zootaxa.5072.3.1</mods:identifier>
<mods:identifier id="31BAA07DEBE87712F56CCA0F40176AFC" type="GBIF-Dataset">a5454a8b-04cd-4cc6-ae0b-f13f31a7af34</mods:identifier>
<mods:identifier id="EE911198797EB63E88A0A3AAD58ABA5D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="983D21686218E50DD36D08A682739FB7" type="Zenodo-Dep">5744549</mods:identifier>
<mods:identifier id="58657463002026011459AC08176C1325" type="ZooBank">CAB2C8C4-E0A4-4805-9DEB-94769C2F90EB</mods:identifier>
</mods:mods>
<treatment id="864387EE1127724EF8A10F68815222F9" ID-DOI="http://doi.org/10.5281/zenodo.5744593" ID-GBIF-Taxon="190874999" ID-Zenodo-Dep="5744593" LSID="urn:lsid:plazi:treatment:864387EE1127724EF8A10F68815222F9" httpUri="http://treatment.plazi.org/id/864387EE1127724EF8A10F68815222F9" lastPageId="29" lastPageNumber="230" pageId="23" pageNumber="224">
<subSubSection id="46F0657311277244F8A10F68863924D1" box="[151,813,1898,1925]" pageId="23" pageNumber="224" type="nomenclature">
<paragraph id="0E5536F811277244F8A10F68863924D1" blockId="23.[151,813,1898,1925]" box="[151,813,1898,1925]" pageId="23" pageNumber="224">
<heading id="551D819411277244F8A10F68863924D1" bold="true" box="[151,813,1898,1925]" fontSize="11" level="1" pageId="23" pageNumber="224" reason="1">
<emphasis id="3C9EEAEA11277244F8A10F68863924D1" bold="true" box="[151,813,1898,1925]" pageId="23" pageNumber="224">
<taxonomicName id="C9EA4D7B11277244F8A10F6887D824D0" authority="Pyrcz &amp; Willmott, 2006" authorityName="Pyrcz &amp; Willmott" authorityYear="2006" box="[151,716,1898,1925]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="pauperata" status="stat. nov.">
<emphasis id="3C9EEAEA11277244F8A10F68848924D1" bold="true" box="[151,413,1898,1925]" italics="true" pageId="23" pageNumber="224">Manerebia pauperata</emphasis>
<bibRefCitation id="6A7B4B0911277244F9920F6987D824D0" author="Pyrcz, T. W. &amp; Willmott, K. R. &amp; Hall, J. P. W. &amp; Viloria, A. L." box="[420,716,1898,1925]" pageId="23" pageNumber="224" pagination="37 - 79" refId="ref22357" refString="Pyrcz, T. W., Willmott, K. R., Hall, J. P. W. &amp; Viloria, A. L. (2006) A review of the genus Manerebia Staudinger (Lepidoptera: Nymphalidae: Satyrinae) in the northern Andes. Journal of Research on the Lepidoptera, 39, 37 - 79." type="journal article" year="2006">Pyrcz &amp; Willmott, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="27AD579111277244FAEC0F69863924D1" box="[730,813,1899,1925]" pageId="23" pageNumber="224">n. stat.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="46F0657311277244F8A10FAD878224B2" pageId="23" pageNumber="224" type="reference_group">
<paragraph id="0E5536F811277244F8A10FAD878224B2" blockId="23.[151,1437,1967,2022]" pageId="23" pageNumber="224">
<treatmentCitationGroup id="2EFA11D611277244F8A10FAD878224B2" pageId="23" pageNumber="224">
<treatmentCitation id="8F4B10E911277244F8A10FAD87E12493" author="Pyrcz, T. W. &amp; Willmott, K. R. &amp; Hall, J. P. W. &amp; Viloria, A. L." box="[151,757,1967,1992]" page="47" pageId="23" pageNumber="224" year="2006">
<taxonomicName id="C9EA4D7B11277244F8A10FAD87E12493" authority="Pyrcz &amp; Willmott, 2006: 47" authorityName="Pyrcz &amp; Willmott" authorityPageNumber="47" authorityYear="2006" box="[151,757,1967,1992]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="subSpecies" species="satura" subSpecies="pauperata">
<emphasis id="3C9EEAEA11277244F8A10FAD84D52493" box="[151,449,1967,1991]" italics="true" pageId="23" pageNumber="224">Manerebia satura pauperata</emphasis>
<materialsCitation id="BE823CA511277244F9FD0FB287E12493" ID-GBIF-Occurrence="3413048346" box="[459,757,1967,1992]" collectionCode="CEPUJ" country="Ecuador" location="Km" municipality="Zamora" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Loja" typeStatus="holotype">
<bibRefCitation id="6A7B4B0911277244F9FD0FB287E12493" author="Pyrcz, T. W. &amp; Willmott, K. R. &amp; Hall, J. P. W. &amp; Viloria, A. L." box="[459,757,1967,1992]" pageId="23" pageNumber="224" pagination="37 - 79" refId="ref22357" refString="Pyrcz, T. W., Willmott, K. R., Hall, J. P. W. &amp; Viloria, A. L. (2006) A review of the genus Manerebia Staudinger (Lepidoptera: Nymphalidae: Satyrinae) in the northern Andes. Journal of Research on the Lepidoptera, 39, 37 - 79." type="journal article" year="2006">Pyrcz &amp; Willmott, 2006: 47</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. Type locality:
<location id="0B35602311277244FBAC0FB286D42493" LSID="urn:lsid:plazi:treatment:864387EE1127724EF8A10F68815222F9:0B35602311277244FBAC0FB286D42493" box="[922,960,1968,1991]" country="Ecuador" municipality="Zamora" name="Km" pageId="23" pageNumber="224" stateProvince="Loja">Km</location>
40, road
<collectingRegion id="CC2EF81A11277244FC1C0FAD814C2493" box="[1066,1112,1967,1991]" country="Ecuador" name="Loja" pageId="23" pageNumber="224">Loja</collectingRegion>
<collectingMunicipality id="EE31AC8211277244FC4E0FB281DD2493" box="[1144,1225,1968,1991]" pageId="23" pageNumber="224">Zamora</collectingMunicipality>
,
<collectingRegion id="CC2EF81A11277244FCE00FAD808C2493" box="[1238,1432,1967,1991]" country="Ecuador" name="Zamora-Chinchipe" pageId="23" pageNumber="224">Zamora-Chinchipe</collectingRegion>
,
<collectingCountry id="76FD766811277244F8F10FCC840924B2" box="[199,285,1998,2022]" name="Ecuador" pageId="23" pageNumber="224">Ecuador</collectingCountry>
.
<typeStatus id="D151885A11277244F9110FCC849124B2" box="[295,389,1998,2022]" pageId="23" pageNumber="224" type="holotype">Holotype</typeStatus>
male: CEPUJ [examined].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="5A9566701128724BF8A10F70812824A2" ID-DOI="http://doi.org/10.5281/zenodo.5744579" ID-Zenodo-Dep="5744579" httpUri="https://zenodo.org/record/5744579/files/figure.png" pageId="24" pageNumber="225" startId="24.[151,250,1906,1930]" targetBox="[301,1286,181,1882]" targetPageId="24">
<paragraph id="0E5536F81128724BF8A10F70870724DE" blockId="24.[151,1084,1906,2038]" box="[151,531,1906,1930]" pageId="24" pageNumber="225">
<emphasis id="3C9EEAEA1128724BF8A10F70840A24DE" bold="true" box="[151,286,1906,1930]" pageId="24" pageNumber="225">FIGURE 11.</emphasis>
Adults live and habitats
</paragraph>
<paragraph id="0E5536F81128724BF8A10F94813F24FA" blockId="24.[151,1084,1906,2038]" box="[151,1067,1942,1966]" pageId="24" pageNumber="225">
A.
<taxonomicName id="C9EA4D7B1128724BF8F10F95848124FA" authorityName="Pyrcz &amp; Boyer" authorityYear="2021" box="[199,405,1942,1966]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subSpecies" species="prattorum" subSpecies="udima">
<emphasis id="3C9EEAEA1128724BF8F10F9585C824FA" box="[199,220,1943,1966]" italics="true" pageId="24" pageNumber="225">M</emphasis>
.
<emphasis id="3C9EEAEA1128724BF8DF0F95848124FA" box="[233,405,1942,1966]" italics="true" pageId="24" pageNumber="225">prattorum udima</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA1128724BF9AA0F9484A424FA" bold="true" box="[412,432,1942,1966]" pageId="24" pageNumber="225">n.</emphasis>
<taxonomicName id="C9EA4D7B1128724BF9800F94873924FA" box="[438,557,1942,1966]" pageId="24" pageNumber="225" rank="subSpecies" subSpecies="female">
<emphasis id="3C9EEAEA1128724BF9800F9484CB24FA" bold="true" box="[438,479,1942,1966]" pageId="24" pageNumber="225">ssp.</emphasis>
female
</taxonomicName>
, Peru, La Florida —via La Udima. Photo P. Boyer
</paragraph>
<paragraph id="0E5536F81128724BF8A10FB881032486" blockId="24.[151,1084,1906,2038]" box="[151,1047,1978,2002]" pageId="24" pageNumber="225">
B.
<taxonomicName id="C9EA4D7B1128724BF8F10FB984812486" authorityName="Pyrcz &amp; Boyer" authorityYear="2021" box="[199,405,1978,2002]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subSpecies" species="prattorum" subSpecies="udima">
<emphasis id="3C9EEAEA1128724BF8F10FB985C82486" box="[199,220,1979,2002]" italics="true" pageId="24" pageNumber="225">M</emphasis>
.
<emphasis id="3C9EEAEA1128724BF8DF0FB984812486" box="[233,405,1978,2002]" italics="true" pageId="24" pageNumber="225">prattorum udima</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA1128724BF9AA0FB884A42486" bold="true" box="[412,432,1978,2002]" pageId="24" pageNumber="225">n.</emphasis>
<taxonomicName id="C9EA4D7B1128724BF9800FB8870D2486" box="[438,537,1978,2002]" pageId="24" pageNumber="225" rank="subSpecies" subSpecies="male">
<emphasis id="3C9EEAEA1128724BF9800FB884CB2486" bold="true" box="[438,479,1978,2002]" pageId="24" pageNumber="225">ssp.</emphasis>
male
</taxonomicName>
, Peru, La Florida —via La Udima. Photo P. Boyer
</paragraph>
<paragraph id="0E5536F81128724BF8A10FDC812824A2" blockId="24.[151,1084,1906,2038]" box="[151,1084,2014,2038]" pageId="24" pageNumber="225">
C. Abra de Porculla, view towards Pacific coast, type locality of
<taxonomicName id="C9EA4D7B1128724BFB0D0FDD868E24A2" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[827,922,2014,2038]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="punku">
<emphasis id="3C9EEAEA1128724BFB0D0FDD864424A2" box="[827,848,2015,2038]" italics="true" pageId="24" pageNumber="225">M</emphasis>
.
<emphasis id="3C9EEAEA1128724BFB6A0FDC868E24A2" box="[860,922,2014,2038]" italics="true" pageId="24" pageNumber="225">punku</emphasis>
</taxonomicName>
. Photo T. Pyrcz
</paragraph>
</caption>
<caption id="5A9566701129724AF8A10F9584B72480" ID-DOI="http://doi.org/10.5281/zenodo.5744583" ID-Zenodo-Dep="5744583" httpUri="https://zenodo.org/record/5744583/files/figure.png" pageId="25" pageNumber="226" startId="25.[151,250,1943,1967]" targetBox="[572,1012,184,1914]" targetPageId="25">
<paragraph id="0E5536F81129724AF8A10F9584B72480" blockId="25.[151,1436,1943,2004]" pageId="25" pageNumber="226">
<emphasis id="3C9EEAEA1129724AF8A10F95843624FB" bold="true" box="[151,290,1943,1967]" pageId="25" pageNumber="226">FIGURE 12.</emphasis>
Bayesian Inference tree (BI) reconstructed from a fragment of COI mitochondrial gene. Posterior probability values indicated on nodes.
</paragraph>
</caption>
<caption id="5A956670112A7249F8A10F9584E22480" ID-DOI="http://doi.org/10.5281/zenodo.5744585" ID-Zenodo-Dep="5744585" httpUri="https://zenodo.org/record/5744585/files/figure.png" pageId="26" pageNumber="227" startId="26.[151,250,1943,1967]" targetBox="[575,1017,184,1919]" targetPageId="26">
<paragraph id="0E5536F8112A7249F8A10F9584E22480" blockId="26.[151,1436,1943,2004]" pageId="26" pageNumber="227">
<emphasis id="3C9EEAEA112A7249F8A10F95843524FB" bold="true" box="[151,289,1943,1967]" pageId="26" pageNumber="227">FIGURE 13.</emphasis>
Maximum likelihood tree (ML) reconstructed from a fragment of COI mitochondrial gene. Ultrafast bootstrap support values indicated on nodes.
</paragraph>
</caption>
<caption id="5A956670112B7248F8A10F9584A42480" ID-DOI="http://doi.org/10.5281/zenodo.5744589" ID-Zenodo-Dep="5744589" httpUri="https://zenodo.org/record/5744589/files/figure.png" pageId="27" pageNumber="228" startId="27.[151,250,1943,1967]" targetBox="[591,998,182,1916]" targetPageId="27">
<paragraph id="0E5536F8112B7248F8A10F9584A42480" blockId="27.[151,1436,1943,2004]" pageId="27" pageNumber="228">
<emphasis id="3C9EEAEA112B7248F8A10F95843524FB" bold="true" box="[151,289,1943,1967]" pageId="27" pageNumber="228">FIGURE 14.</emphasis>
The Bayesian implementation of the Poisson Tree Processes (bPTP) for delimiting species based on a fragment of COI mitochondrial gene.
</paragraph>
</caption>
<caption id="5A956670112C724FF8A10F9A85DA2481" ID-DOI="http://doi.org/10.5281/zenodo.5744591" ID-Zenodo-Dep="5744591" httpUri="https://zenodo.org/record/5744591/files/figure.png" pageId="28" pageNumber="229" startId="28.[151,250,1944,1968]" targetBox="[591,997,185,1917]" targetPageId="28">
<paragraph id="0E5536F8112C724FF8A10F9A85DA2481" blockId="28.[151,1436,1944,2005]" pageId="28" pageNumber="229">
<emphasis id="3C9EEAEA112C724FF8A10F9A840B24E4" bold="true" box="[151,287,1944,1968]" pageId="28" pageNumber="229">FIGURE 15.</emphasis>
The generalized mixed Yule-coalescent (GMYC) for delimiting species based on a fragment of COI mitochondrial gene.
</paragraph>
</caption>
<subSubSection id="46F06573112D724EF8A10895815222F9" pageId="29" pageNumber="230" type="discussion">
<paragraph id="0E5536F8112D724EF8A10895815222F9" blockId="29.[151,1437,151,429]" pageId="29" pageNumber="230">
<emphasis id="3C9EEAEA112D724EF8A10895841F23E5" bold="true" box="[151,267,151,177]" pageId="29" pageNumber="230">Remarks.</emphasis>
In both BI (
<figureCitation id="96D12A7D112D724EF9A7089584CF23E6" box="[401,475,151,178]" captionStart="FIGURE 12" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[572,1012,184,1914]" captionTargetId="figure-21@25.[572,1014,181,1919]" captionTargetPageId="25" captionText="FIGURE 12. Bayesian Inference tree (BI) reconstructed from a fragment of COI mitochondrial gene. Posterior probability values indicated on nodes." figureDoi="http://doi.org/10.5281/zenodo.5744583" httpUri="https://zenodo.org/record/5744583/files/figure.png" pageId="29" pageNumber="230">Fig.12</figureCitation>
) and ML (
<figureCitation id="96D12A7D112D724EFA660895878B23E5" box="[592,671,151,177]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[575,1017,184,1919]" captionTargetId="figure-23@26.[570,1017,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Maximum likelihood tree (ML) reconstructed from a fragment of COI mitochondrial gene. Ultrafast bootstrap support values indicated on nodes." figureDoi="http://doi.org/10.5281/zenodo.5744585" httpUri="https://zenodo.org/record/5744585/files/figure.png" pageId="29" pageNumber="230">Fig. 13</figureCitation>
) trees
<taxonomicName id="C9EA4D7B112D724EFADF0895869523E4" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" baseAuthorityName="Pyrcz &amp; Willmott" baseAuthorityYear="2006" box="[745,897,151,176]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="pauperata" status="stat. nov.">
<emphasis id="3C9EEAEA112D724EFADF0895869523E4" box="[745,897,151,176]" italics="true" pageId="29" pageNumber="230">M. pauperata</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA112D724EFBB1089586C223E5" bold="true" box="[903,982,151,177]" pageId="29" pageNumber="230">
<taxonomicNameLabel id="27AD5791112D724EFBB1089586C223E5" box="[903,982,151,177]" pageId="29" pageNumber="230">n. stat.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="C9EA4D7B112D724EFC3D0895816323E4" box="[1035,1143,151,176]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="satura">
<emphasis id="3C9EEAEA112D724EFC3D0895816323E4" box="[1035,1143,151,176]" italics="true" pageId="29" pageNumber="230">M. satura</emphasis>
</taxonomicName>
situate in distant branches of the tree and cluster in highly resolved clades with other congeners. Both
<taxonomicName id="C9EA4D7B112D724EFBD508B981472381" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" baseAuthorityName="Pyrcz &amp; Willmott" baseAuthorityYear="2006" box="[995,1107,187,213]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="lamasi" status="stat. nov.">
<emphasis id="3C9EEAEA112D724EFBD508B986EE2380" box="[995,1018,187,212]" italics="true" pageId="29" pageNumber="230">M</emphasis>
.
<emphasis id="3C9EEAEA112D724EFC3E08B981472381" box="[1032,1107,187,213]" italics="true" pageId="29" pageNumber="230">lamasi</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA112D724EFC6C08B981BD2381" bold="true" box="[1114,1193,187,213]" pageId="29" pageNumber="230">
<taxonomicNameLabel id="27AD5791112D724EFC6C08B981BD2381" box="[1114,1193,187,213]" pageId="29" pageNumber="230">n. stat.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="C9EA4D7B112D724EFCD108B9806B2380" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" baseAuthorityName="Pyrcz &amp; Willmott" baseAuthorityYear="2006" box="[1255,1407,187,213]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="pauperata" status="stat. nov.">
<emphasis id="3C9EEAEA112D724EFCD108B981EA2380" box="[1255,1278,187,212]" italics="true" pageId="29" pageNumber="230">M</emphasis>
.
<emphasis id="3C9EEAEA112D724EFD3A08B9806B2380" box="[1292,1407,187,212]" italics="true" pageId="29" pageNumber="230">pauperata</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA112D724EFDB008B985DE23AD" bold="true" pageId="29" pageNumber="230">
<taxonomicNameLabel id="27AD5791112D724EFDB008B985DE23AD" pageId="29" pageNumber="230">n. stat.</taxonomicNameLabel>
</emphasis>
have also been collected in sympatry in the Cordillera del Cóndor in southern
<collectingCountry id="76FD7668112D724EFC7308DD81B123AD" box="[1093,1189,223,249]" name="Ecuador" pageId="29" pageNumber="230">Ecuador</collectingCountry>
. Additionally, the two taxa also consistently differ in wing colour patterns (
<figureCitation id="96D12A7D112D724EFAEC0901865B2249" box="[730,847,258,285]" captionStart="FIGURE 5" captionStartId="11.[151,250,1849,1873]" captionTargetBox="[264,1317,190,1798]" captionTargetId="figure-21@11.[240,1348,181,1825]" captionTargetPageId="11" captionText="FIGURE 5. Adults (left—dorsum, right—venter) A. M. benigni tessmanni ♂ Ecuador, Río Troya B. M. benigni tessmanni ♀, Ecuador, Jimbura —San Andrés C. M. pauperata n. stat. ♂ Ecuador, Zamora —Chinchipe D. M. lamasi n. stat. ♂, Ecuador, Morona—Santiago" figureDoi="http://doi.org/10.5281/zenodo.5744559" httpUri="https://zenodo.org/record/5744559/files/figure.png" pageId="29" pageNumber="230">Fig. 5C, D</figureCitation>
) and male genitalia. We therefore formally recognize
<taxonomicName id="C9EA4D7B112D724EF8A1092584132215" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" baseAuthorityName="Pyrcz &amp; Willmott" baseAuthorityYear="2006" box="[151,263,295,321]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="lamasi" status="stat. nov.">
<emphasis id="3C9EEAEA112D724EF8A1092584132215" box="[151,263,295,321]" italics="true" pageId="29" pageNumber="230">M. lamasi</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA112D724EF93B092584492215" bold="true" box="[269,349,295,321]" pageId="29" pageNumber="230">
<taxonomicNameLabel id="27AD5791112D724EF93B092584492215" box="[269,349,295,321]" pageId="29" pageNumber="230">n. stat.</taxonomicNameLabel>
</emphasis>
as a distinct species. Despite an overall similar colour pattern in comparison with
<taxonomicName id="C9EA4D7B112D724EFCC5092580742214" box="[1267,1376,295,321]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="satura">
<emphasis id="3C9EEAEA112D724EFCC50925801E2214" box="[1267,1290,295,320]" italics="true" pageId="29" pageNumber="230">M</emphasis>
.
<emphasis id="3C9EEAEA112D724EFD2E092580742214" box="[1304,1376,295,320]" italics="true" pageId="29" pageNumber="230">satura</emphasis>
</taxonomicName>
from central and southern
<collectingCountry id="76FD7668112D724EF948094984A02231" box="[382,436,331,357]" name="Peru" pageId="29" pageNumber="230">Peru</collectingCountry>
, with the exception of the absence of the HWV yellow median band,
<taxonomicName id="C9EA4D7B112D724EFC86094980532230" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" baseAuthorityName="Pyrcz &amp; Willmott" baseAuthorityYear="2006" box="[1200,1351,331,357]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="pauperata" status="stat. nov.">
<emphasis id="3C9EEAEA112D724EFC86094981D32230" box="[1200,1223,331,356]" italics="true" pageId="29" pageNumber="230">M</emphasis>
.
<emphasis id="3C9EEAEA112D724EFCE2094980532230" box="[1236,1351,331,356]" italics="true" pageId="29" pageNumber="230">pauperata</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA112D724EFD7B094980882231" bold="true" box="[1357,1436,331,357]" pageId="29" pageNumber="230">
<taxonomicNameLabel id="27AD5791112D724EFD7B094980882231" box="[1357,1436,331,357]" pageId="29" pageNumber="230">n. stat.</taxonomicNameLabel>
</emphasis>
shows some genitalic differences. Moreover, the species-delimitation methods (
<figureCitation id="96D12A7D112D724EFC20096D816022DD" box="[1046,1140,367,393]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[591,998,182,1916]" captionTargetId="figure-21@27.[589,998,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. The Bayesian implementation of the Poisson Tree Processes (bPTP) for delimiting species based on a fragment of COI mitochondrial gene." figureDoi="http://doi.org/10.5281/zenodo.5744589" httpUri="https://zenodo.org/record/5744589/files/figure.png" pageId="29" pageNumber="230">Figs. 14</figureCitation>
,
<figureCitation id="96D12A7D112D724EFCB7096C818A22DD" box="[1153,1182,366,393]" captionStart="FIGURE 15" captionStartId="28.[151,250,1944,1968]" captionTargetBox="[591,997,185,1917]" captionTargetId="figure-23@28.[589,998,181,1920]" captionTargetPageId="28" captionText="FIGURE 15. The generalized mixed Yule-coalescent (GMYC) for delimiting species based on a fragment of COI mitochondrial gene." figureDoi="http://doi.org/10.5281/zenodo.5744591" httpUri="https://zenodo.org/record/5744591/files/figure.png" pageId="29" pageNumber="230">15</figureCitation>
) and genetic distances (Supplementary material 2) clearly support treating the two taxa as separate species.
</paragraph>
</subSubSection>
</treatment>
</document>