428 lines
42 KiB
XML
428 lines
42 KiB
XML
<document id="E44C8F76D2BA77286E9BED6E778027EA" ID-DOI="http://dx.doi.org/10.3897/zse.93.11576" ID-GBIF-Dataset="24bdcd60-a1f2-4701-a3e7-1632d38d6e2c" ID-Pensoft-Pub="1860-0743-2-265" ID-ZooBank="4483DB5A3DD14AB685F7BA9A077878A5" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1860-0743-2-265" ModsDocOrigin="Zoosystematics and Evolution 93 (2)" ModsDocTitle="Three new species of the microhylid frog genus Choerophryne (Amphibia, Anura, Microhylidae) from Papua New Guinea" checkinTime="1493983377162" checkinUser="pensoft" docAuthor="Guenther, Rainer & Richards, Stephen" docDate="2017" docId="2C45F1FC83C8D250251FC0C7FB1567E2" docLanguage="en" docName="ZoosystEvol 93(2): 265-279" docOrigin="Zoosystematics and Evolution 93 (2)" docSource="http://dx.doi.org/10.3897/zse.93.11576" docTitle="Choerophryne crucifer Guenther & Richards, 2017, sp. n." docType="treatment" docUuid="31112467-259E-401A-ACB6-59E0B0960FEA" docUuidSource="ZooBank" docVersion="6" lastPageNumber="268" masterDocId="FFB9FFE3B2325519853A0564215BFFDF" masterDocTitle="Three new species of the microhylid frog genus Choerophryne (Amphibia, Anura, Microhylidae) from Papua New Guinea" masterLastPageNumber="279" masterPageNumber="265" pageNumber="265" updateTime="1701317639626" updateUser="plazi">
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<mods:title id="259203AD7703189CFA01C9D66AD64D22">Three new species of the microhylid frog genus Choerophryne (Amphibia, Anura, Microhylidae) from Papua New Guinea</mods:title>
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<mods:name id="204E6CB062C9FD3C1C719FA2D5F2059F" type="personal">
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<mods:namePart id="B6E1A25AE04EDF7A0C841D111D429A55">Guenther, Rainer</mods:namePart>
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<mods:namePart id="A70F1BB2DC19E7924326D5D69732E9DB">Richards, Stephen</mods:namePart>
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<mods:title id="5594DC83577C61BED548899416FF6B5B">Zoosystematics and Evolution</mods:title>
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<mods:part id="85E6E5F07E8CC67DA1C7A702E936DD3A">
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<mods:date id="EB43A591C85B8B1E1028A85C989AEA06">2017</mods:date>
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<mods:number id="797E4DAE8AD620233E7F838D768B6E1B">93</mods:number>
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<mods:url id="AD7FE4BF7111FE08D8B06A2A89FE77B3">http://dx.doi.org/10.3897/zse.93.11576</mods:url>
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<mods:classification id="BF525133F4F2C9587404CC34D43CB0CD">journal article</mods:classification>
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<mods:identifier id="F4F488DACE906632EE2E555370028A40" type="DOI">http://dx.doi.org/10.3897/zse.93.11576</mods:identifier>
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<treatment id="2C45F1FC83C8D250251FC0C7FB1567E2" ID-GBIF-Taxon="128848772" LSID="urn:lsid:zoobank.org:act:31112467-259E-401A-ACB6-59E0B0960FEA" httpUri="http://treatment.plazi.org/id/2C45F1FC83C8D250251FC0C7FB1567E2" lastPageId="3" lastPageNumber="268" pageId="0" pageNumber="265">
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<subSubSection id="2AE2457114D0EBF3AA76838B077D4BDB" pageId="0" pageNumber="265" type="nomenclature">
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<paragraph id="4B7C6FA199F12B9D7180C5F891932897" pageId="0" pageNumber="265">
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<taxonomicName id="BD4ED156710CCE163425590978668307" ID-CoL="69Z45" LSID="http://zoobank.org/31112467-259E-401A-ACB6-59E0B0960FEA" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="0" pageNumber="265" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
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<taxonomicNameLabel id="15E332B9FD805DEC1400E9E108B10C19" pageId="0" pageNumber="265">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="9DD5D2D8B18E62924BF85BB62F29EA81" pageId="0" pageNumber="265" type="holotype">
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<paragraph id="1A7654B4526811F2FA42A5AB1744FAF1" pageId="0" pageNumber="265">Holotype.</paragraph>
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<paragraph id="6917FDBE9A60902C2892DF5B06D20142" pageId="0" pageNumber="265">
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SAMA R69448 (Field number: FN SJR 8623), adult male, Iagifu Ridge near Moro, Southern Highlands Province, Papua New Guinea (
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<geoCoordinate id="4A5A64D16E9BD3933734721BB2D68C8C" direction="south" orientation="latitude" precision="1" value="-6.3683167">06°22.099'S</geoCoordinate>
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,
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<geoCoordinate id="91D26171E37342B8ADE9EAB48E8A2439" direction="east" orientation="longitude" precision="1" value="143.2229">143°13.374'E</geoCoordinate>
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; 900 m asl) collected on 21-11-2004 by S.J. Richards.
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</paragraph>
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</subSubSection>
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<subSubSection id="8722AD237B2867B971E6CC9C212E4AF9" pageId="0" pageNumber="265" type="paratypes">
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<paragraph id="C0E7E946E4B49B600F5E3A00088D9D03" pageId="0" pageNumber="265">Paratypes.</paragraph>
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<paragraph id="72CFE12C7DF1558C97B945EE3063959D" pageId="0" pageNumber="265">
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ZMB 84336 (FN SJR 8622), same data as holotype; SAMA R69440-69442 (FN SJR 3147-3148, 3151), Darai Plateau, Gulf Province, Papua New Guinea (
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<geoCoordinate id="9D7629F2813E6E18753BEAD6441E2B51" direction="south" orientation="latitude" precision="1" value="-7.1295166">07°07.771'S</geoCoordinate>
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,
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<geoCoordinate id="28020F47AE87E6943A070FDF555C5587" direction="east" orientation="longitude" precision="1" value="143.61343">143°36.806'E</geoCoordinate>
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; 400 m asl) collected on 23-07-2003 by S.J. Richards.
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</paragraph>
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</subSubSection>
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<subSubSection id="E726436E325D034B175EEF2642D782F5" pageId="0" pageNumber="265" type="diagnosis">
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<paragraph id="F174A6CD28C8F5E56660150601EAFED7" pageId="0" pageNumber="265">Diagnosis.</paragraph>
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<paragraph id="572F8014709DE1385BC488DE52623819" pageId="0" pageNumber="265">
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A species of the genus
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<taxonomicName id="3D0B02C8D3FDD77157F5F6F8DB8C3287" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne" order="Anura" pageId="0" pageNumber="265" phylum="Chordata" rank="genus">Choerophryne</taxonomicName>
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lacking an elongated snout. Snout-urostyle length in males (n=5) from 13.4-17.3 mm (mean 14.7
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<normalizedToken id="55FA73C6EC2A6EF653CCF3C769139F9E" originalValue="±">+/-</normalizedToken>
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1.58 mm). No webs between fingers or toes; fifth toe longer than third; finger discs wider than toe discs (ratio T4D/F3D 0.78-0.88); shanks short (TL/SUL 0.38-0.44). Eyes medium sized (ED/SUL 0.110-0.127), eye-naris distance greater than internarial distance (END/IND 1.00-1.25). Dorsum with a brown hour-glass mark that has an approximately median constriction; and a pale cross with a definite posterior
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<normalizedToken id="E9A7A6714B5B2F66B07FF6B4311700B8" originalValue="‘extension’">'extension'</normalizedToken>
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on head. Dorsal surfaces covered with tubercles in life, lower surface smooth and brown with numerous whitish dots, inguinal region yellowish. Advertisement call a series of musical clicks sounding like
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<normalizedToken id="B782B460946D232C6D409568B4EB1BBB" originalValue="‘tink-tink-tink…’">'tink-tink-tink...'</normalizedToken>
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lasting 1.01-4.75 s and containing 5-22 clicks (notes) per call at a repetition rate of 4.39-5.18 notes/s. Dominant frequency is at 5.5 kHz.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6D4BCF584013253A1F730A8DAB2AAB2" lastPageId="1" lastPageNumber="266" pageId="0" pageNumber="265" type="description">
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<paragraph id="14676BD4CECB062C5A0FB34B4B841AC6" pageId="0" pageNumber="265">Description of the holotype.</paragraph>
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<paragraph id="809F0EF356D721DA2E3D657569DC08A4" lastPageId="1" lastPageNumber="266" pageId="0" pageNumber="265">
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Adult male with a SUL of 14.9 mm. Additional measurements and ratios are listed in Table 1. Head broader than long (HL/HW 0.73); tip of snout rounded in dorsal view and truncate in lateral view; nostrils near tip of snout, directed laterally and not visible from above, distance between nares less than distance between eye and naris (END/IND 1.25); canthus rostralis in dorsal view straight and rounded; loreal region slightly sloped; tongue oval, narrower anteriorly than posteriorly, posterior margin without indentation; anterior prepharyngeal ridge well developed and smooth, posterior
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<normalizedToken id="5E47FDF0FDFC7121E93F230D9EB38D1D" originalValue="“ridge”">"ridge"</normalizedToken>
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a plaque with many longitudinal furrows; fairly long vocal slits on both sides of mouth floor; tympanum small (about one-third of eye diameter) and hardly visible; no supratympanic fold. Shanks of medium length (TL/SUL 0.44). Fingers unwebbed with broad, grooved terminal discs, their relative lengths 3>4>2>1 (Fig. 1a, b); disc of third finger slightly more than twice width of penultimate phalanx, no prominent metacarpal or subarticular tubercles. All toes with wide, grooved terminal discs, those of fourth toe slightly narrower than those of third finger; no webs between toes, no metatarsal tubercles, subarticular tubercles weakly developed; relative lengths of toes 4>5>3>2>1 (Fig. 1a, b). Skin smooth with small
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<pageBreakToken id="840FC04F5260E0039C88A973D8C50CA8" pageId="1" pageNumber="266" start="start">but</pageBreakToken>
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distinct tubercles dorsally in life, less conspicuous in preservative. No distinct tubercles on ventral surfaces in preservative.
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</paragraph>
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<caption id="5D7FD5C51141FF37865C9A353409EA21" pageId="1" pageNumber="266">
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<paragraph id="DAFBFB3313FA212F0C1080C5CFD78D0A" pageId="1" pageNumber="266">
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Figure 1a. Preserved holotype of
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<taxonomicName id="86185B7F262FE33BA037D11DE86B2167" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
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sp. n. in dorsal view.
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</paragraph>
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</caption>
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<caption id="77FE5EA4A8F83118F194B990BF08DBE4" pageId="1" pageNumber="266">
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<paragraph id="59C575F4A5E4AA3A30A1974B3EC7CFBC" pageId="1" pageNumber="266">
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Figure 1b. Preserved holotype of
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<taxonomicName id="37633C77583E27BEFDE17BE69430A38E" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
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sp. n. in ventral view.
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</paragraph>
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</caption>
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<caption id="FFC3B2114577603FBE952335C3F71176" pageId="1" pageNumber="266">
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<paragraph id="A35968D142FA2FD97160DF5A45E9A677" pageId="1" pageNumber="266">
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Table 1. Body measurements and body ratios of the type series of
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<taxonomicName id="578398F7EFFDC5A398DAD374E4EF87A5" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
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sp. n. SAMA R69448 is the holotype, all five types are adult males; all measurements in mm. Reg.-No = Registration number; SD = standard deviation; for explanations of abbreviations for measurements see "Material and methods".
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</paragraph>
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</caption>
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<paragraph id="5EFC678651E06C61D55384EF0419BF01" pageId="1" pageNumber="266">
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<table id="79D8447BC33FD9911819AA2E2A8C1F09" pageId="1" pageNumber="266">
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<tr id="5D598C2A2C926057E4BE4E1C27003968" pageId="1" pageNumber="266">
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<th id="26D0629359281D2497D6683ED01E72B0" colspan="1" pageId="1" pageNumber="266" rowspan="1">Reg.-No.</th>
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<th id="ACD744D4B46F85A63B00E0DF441BA793" colspan="1" pageId="1" pageNumber="266" rowspan="1">
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<named-content id="F7862E40DBF0607062DC5D91F6417262" content-type="dwc:institutional_code" pageId="1" pageNumber="266">SAMA</named-content>
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R69440
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</th>
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<th id="3248C8CB1111FB52DB31C3D2C2460D01" colspan="1" pageId="1" pageNumber="266" rowspan="1">
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<named-content id="3BB435C1DC73AAB87C4F98206B4739BD" content-type="dwc:institutional_code" pageId="1" pageNumber="266">SAMA</named-content>
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R69441
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</th>
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<th id="2EBABB667AC918255E387A6310C83BB6" colspan="1" pageId="1" pageNumber="266" rowspan="1">
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<named-content id="BE0358797C4470DC6287E0C722B60FFE" content-type="dwc:institutional_code" pageId="1" pageNumber="266">SAMA</named-content>
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R69442
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</th>
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<th id="7AA47E9D3F6EC5A54E2271F39D71584A" colspan="1" pageId="1" pageNumber="266" rowspan="1">
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<named-content id="5F0D74A130992CB8EBAB9FA596D5A856" content-type="dwc:institutional_code" pageId="1" pageNumber="266">ZMB</named-content>
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84336
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</th>
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<th id="373B21B0F6CD76D1EFB562B8654D26E8" colspan="1" pageId="1" pageNumber="266" rowspan="1">
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<named-content id="BF334266F737886297C33983E9E3B36E" content-type="dwc:institutional_code" pageId="1" pageNumber="266">SAMA</named-content>
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R69448
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</th>
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<th id="90C048C5D821A7E93D01C9A4C6580152" colspan="1" pageId="1" pageNumber="266" rowspan="1">
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Mean
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<normalizedToken id="BA743CC584E6964AC03B603C375D33EB" originalValue="±">+/-</normalizedToken>
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SD
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</th>
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||
</tr>
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||
<tr id="396E6810EC68527053CBC630884359E3" pageId="1" pageNumber="266">
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<td id="11D48D8CB07E672B90F2FB40122EF5E5" colspan="1" pageId="1" pageNumber="266" rowspan="1">SUL</td>
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</tr>
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||
<tr id="AF1C4720C1E8BB3A2DC59FE9CBB71FCC" pageId="1" pageNumber="266">
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<td id="07CDD8E1FA48669783E34F9261BE8F94" colspan="1" pageId="1" pageNumber="266" rowspan="1">TL</td>
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</tr>
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<tr id="A4DB027596DF5B1A79CB4BB51F89B13D" pageId="1" pageNumber="266">
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<td id="7801356C95A316B17931E6FD29FBC6F1" colspan="1" pageId="1" pageNumber="266" rowspan="1">TaL</td>
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</tr>
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<tr id="30533D6BB0CA81AFE81E3ACD25C217C7" pageId="1" pageNumber="266">
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<td id="261F9B48A1E54A445DC9882711919C2D" colspan="1" pageId="1" pageNumber="266" rowspan="1">T4L</td>
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</tr>
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||
<tr id="370B140195754E1E0C3EE4FA5D5BD8F8" pageId="1" pageNumber="266">
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<td id="2B7C349EF809E32217BF16D58793DA16" colspan="1" pageId="1" pageNumber="266" rowspan="1">T4D</td>
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</tr>
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||
<tr id="7E3DCBF505E51056D4F5BBF9C96E7DE3" pageId="1" pageNumber="266">
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||
<td id="9F699B4DC61F07E526DB23D29B2E8221" colspan="1" pageId="1" pageNumber="266" rowspan="1">T1D</td>
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</tr>
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||
<tr id="0BA2BEEE5A517350756F77D91969039B" pageId="1" pageNumber="266">
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||
<td id="D0818F8ACBC062B7F62F15CF945D6164" colspan="1" pageId="1" pageNumber="266" rowspan="1">F3L</td>
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||
</tr>
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||
<tr id="574EFD96BF12F0543F4A3C65D240B4BC" pageId="1" pageNumber="266">
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||
<td id="8EFEA77FB780E90D5EA76B45CD9B4E9C" colspan="1" pageId="1" pageNumber="266" rowspan="1">F3D</td>
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||
</tr>
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||
<tr id="CE762BC2D2920B36EDA4177DF8A0EBE6" pageId="1" pageNumber="266">
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||
<td id="080B77F1255DAC6CDA9D95554E909130" colspan="1" pageId="1" pageNumber="266" rowspan="1">F1D</td>
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||
</tr>
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||
<tr id="274C15912443D79DE134587FBC1D3F43" pageId="1" pageNumber="266">
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||
<td id="B212B5E2BE36342CEA67EBB0A7CCF12F" colspan="1" pageId="1" pageNumber="266" rowspan="1">HL</td>
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||
</tr>
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||
<tr id="05CC7D8D148A4742CD5840A3A368950B" pageId="1" pageNumber="266">
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||
<td id="CA15D130F839E03929ED39738F6EF057" colspan="1" pageId="1" pageNumber="266" rowspan="1">HW</td>
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</tr>
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||
<tr id="718EA3D295BAC13356258C64B981A101" pageId="1" pageNumber="266">
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<td id="E1F9883AF856D4D2973A630BBEE1AC12" colspan="1" pageId="1" pageNumber="266" rowspan="1">END</td>
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||
</tr>
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||
<tr id="9C9A062A4DFE82151F7F55DB3BE74AF8" pageId="1" pageNumber="266">
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<td id="FFDCF47E14B0EF528A0CA5179185E87B" colspan="1" pageId="1" pageNumber="266" rowspan="1">IND</td>
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||
</tr>
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||
<tr id="7B39D9319A514565CD25D02649CA7E54" pageId="1" pageNumber="266">
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||
<td id="CE669041781543ECDC69EC4CE467C461" colspan="1" pageId="1" pageNumber="266" rowspan="1">SL</td>
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||
</tr>
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||
<tr id="ACC6BA7BF26AB7BD73C65BC7143EE28C" pageId="1" pageNumber="266">
|
||
<td id="5E1BB52B6665A46EC42EC652B6EF9646" colspan="1" pageId="1" pageNumber="266" rowspan="1">ED</td>
|
||
</tr>
|
||
<tr id="35831BAF2DB33211FB1474080F6EFCAF" pageId="1" pageNumber="266">
|
||
<td id="F7AE13EDA20FB006DDA56BAD0C967261" colspan="1" pageId="1" pageNumber="266" rowspan="1">TyD</td>
|
||
</tr>
|
||
<tr id="8967E6AF293A3DF7F0D2F55CA4CB39E0" pageId="1" pageNumber="266">
|
||
<td id="7F50A4E2727CEBFFE71162978F93D9A5" colspan="1" pageId="1" pageNumber="266" rowspan="1">TLSUL</td>
|
||
</tr>
|
||
<tr id="67F3DAB84EA262E44CEB3CBFE64162DD" pageId="1" pageNumber="266">
|
||
<td id="E90691AFCB846589AD473AEA749311BD" colspan="1" pageId="1" pageNumber="266" rowspan="1">TaLSUL</td>
|
||
</tr>
|
||
<tr id="090322807C13BC71A680A01EC267F881" pageId="1" pageNumber="266">
|
||
<td id="3DF797E915E05C75A93AE2FA1F64A60F" colspan="1" pageId="1" pageNumber="266" rowspan="1">T4LSUL</td>
|
||
</tr>
|
||
<tr id="1AA0A361CD31D99959FDBFB339B2F940" pageId="1" pageNumber="266">
|
||
<td id="B2C264B1189976FDCE91158008AE91C4" colspan="1" pageId="1" pageNumber="266" rowspan="1">T4DSUL</td>
|
||
</tr>
|
||
<tr id="0FD18570C50B65022026306914A88FB2" pageId="1" pageNumber="266">
|
||
<td id="4ECD695E837EFE810B37607464A192C0" colspan="1" pageId="1" pageNumber="266" rowspan="1">F3LSUL</td>
|
||
</tr>
|
||
<tr id="407074AA590C16389632FF3E9B0F2A0F" pageId="1" pageNumber="266">
|
||
<td id="A498A9189A95A54AF7EC44F10AA625AD" colspan="1" pageId="1" pageNumber="266" rowspan="1">F3DSUL</td>
|
||
</tr>
|
||
<tr id="59F91B0C5CEE6BF336D28DA5CC217AB0" pageId="1" pageNumber="266">
|
||
<td id="1B0FFF7C828A554F39DCF01C1E15AAF8" colspan="1" pageId="1" pageNumber="266" rowspan="1">T4DF3D</td>
|
||
</tr>
|
||
<tr id="A15C53BECFADAF57444C46EB97BFE196" pageId="1" pageNumber="266">
|
||
<td id="47DD6042AE2487F4152A0CFFF7E334D5" colspan="1" pageId="1" pageNumber="266" rowspan="1">T1DF1D</td>
|
||
</tr>
|
||
<tr id="85D1972C897AC03A2388AE9198A1BEE9" pageId="1" pageNumber="266">
|
||
<td id="DABB8FD7ECB9A6CCB0882F70D1587E40" colspan="1" pageId="1" pageNumber="266" rowspan="1">HLSUL</td>
|
||
</tr>
|
||
<tr id="04AA2031CAA92A5EA82547B9CF882E79" pageId="1" pageNumber="266">
|
||
<td id="99C6D5089998F55D89C557EFAFBC8AFF" colspan="1" pageId="1" pageNumber="266" rowspan="1">HWSUL</td>
|
||
</tr>
|
||
<tr id="5E224A322BA503BBD188D57DDF72AC44" pageId="1" pageNumber="266">
|
||
<td id="EECE239D6C8683E09B61108D110D70A6" colspan="1" pageId="1" pageNumber="266" rowspan="1">HLHW</td>
|
||
</tr>
|
||
<tr id="9B32A8436DE313BE78F074584ACA21F3" pageId="1" pageNumber="266">
|
||
<td id="85AA3FFE6B26AB06B320A2C9D9B1C895" colspan="1" pageId="1" pageNumber="266" rowspan="1">ENDIND</td>
|
||
</tr>
|
||
<tr id="64635548ADD5C38A70880C89319BDD5F" pageId="1" pageNumber="266">
|
||
<td id="7B875BD67B7190B67997E67403264AFF" colspan="1" pageId="1" pageNumber="266" rowspan="1">EDSUL</td>
|
||
</tr>
|
||
<tr id="A99DFD1B75D7573A93789D6D10C2F013" pageId="1" pageNumber="266">
|
||
<td id="9F3DABB2172635D0AAC429948BB312E7" colspan="1" pageId="1" pageNumber="266" rowspan="1">TyDSUL</td>
|
||
</tr>
|
||
<tr id="9AFD1EA73227F02746F6E1E9021F77CD" pageId="1" pageNumber="266">
|
||
<td id="42A275A6A03D8CF3FF9EB147271E0B25" colspan="1" pageId="1" pageNumber="266" rowspan="1">TyDED</td>
|
||
</tr>
|
||
<tr id="72A2070E85D3D823EC5ECDCAFDCA5000" pageId="1" pageNumber="266">
|
||
<td id="1CD818EBD6571A3848A6539E305A0469" colspan="1" pageId="1" pageNumber="266" rowspan="1">SLSUL</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8F8705CD00AC95C1C43A64259C3FAC57" pageId="1" pageNumber="266" type="colour of the holotype in preservative">
|
||
<paragraph id="BDA08E88ED319BFB3978AA8A5C3EF43A" pageId="1" pageNumber="266">Colour of the holotype in preservative</paragraph>
|
||
<paragraph id="614B59D9B63A4996C25E66B9D9971C36" pageId="1" pageNumber="266">
|
||
(Fig. 1a, b). Colour of holotype in life unknown. In preservative ground colour of dorsal surfaces of head, body and hind limbs light-grey, that of fore limbs yellowish; a dun hour-glass shaped patch from eyes to inguinal region, this patch constricted at mid-dorsum and
|
||
<normalizedToken id="0EA3CEADEB620F679D0A0CB5848412D7" originalValue="“split”">"split"</normalizedToken>
|
||
anteriorly by posterior arm of pale cross on head; some diffuse smaller brownish spots on head and dorsal surfaces of extremities. Inguinal region with conspicuous dark brown areas bordered anteriorly by yellow flecks; dark brown inguinal spots merge ventrally into brown colour of abdomen. Underside of thighs, chest and throat also brown and interspersed with small whitish spots. An irregularly shaped dark brown band across anal region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11B0E2ECC4193C9965605FB4AD2FDCFA" pageId="1" pageNumber="266" type="colour in life">
|
||
<paragraph id="A50B57C635A9D476D2F35E5459B6773D" pageId="1" pageNumber="266">Colour in life.</paragraph>
|
||
<paragraph id="77FF2895BE5ACD33ECEA116627DADEE5" pageId="1" pageNumber="266">Based on paratype SAMA R69440, mid-dorsum covered by a large brown patch, dorsolaterally bordered by an irregular off-white longitudinal stripe. Flanks reddish-brown with a dark grey reticulum, inguinal region with a yellowish area, dorsal surfaces of extremities grey-yellowish with irregular dark grey, brownish and reddish spots; ventral surfaces grey with whitish mottles (Fig. 2).</paragraph>
|
||
<caption id="732F48E462C246BF108C7C32AD7C32C9" pageId="1" pageNumber="266">
|
||
<paragraph id="E81C63B1BCB3896F5C755067C8E3F21A" pageId="1" pageNumber="266">
|
||
Figure 2. Paratype SAMA R69440 of
|
||
<taxonomicName id="9E418366AFD618926B37D46118FBA075" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
|
||
sp. n. in life.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="1BD3D8307B8FDD90CE911847D1965F32" pageId="1" pageNumber="266" type="morphological variation">
|
||
<paragraph id="67D38D8E3EA192CF64D8BF49F384FBEB" pageId="1" pageNumber="266">Morphological variation.</paragraph>
|
||
<paragraph id="105AB90F03CD6132861ECA2C3B9BA8B9" pageId="1" pageNumber="266">
|
||
Measurements and body ratios of the type specimens are presented in Table 1. While four males had a SUL between 13.8 and 14.9 mm, one male measured 17.3 mm. This difference of 3.5 mm between adult males is fairly large for frogs of this small size. All five specimens exhibit a dark brown mid-dorsal patch with a pair of anterior and a pair of posterior
|
||
<normalizedToken id="9579E5996C72F30BD4AE12DC3CF53976" originalValue="“wings”">"wings"</normalizedToken>
|
||
, the posterior ones in almost all cases more strongly expanded than the anterior ones. This patch is bordered dorsolaterally by an irregular whitish stripe in four specimens and by a light brown stripe in one specimen. A complete cross-shaped figure on the head is evident in four specimens; it is incomplete in ZMB 84336 where only the posterior section is pronounced. All specimens exhibit the large, pale semicircular patch on the posterior end of the body and pale spots or bands proximally on posterior thighs and distally on dorsal shanks. Throat, chest, abdomen and inferior surface of thighs are brown and more or less densely speckled with whitish spots in all specimens. Ventral surfaces of extremities are off-white and mottled with brownish and/or whitish spots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1160412C85372AB11CF5D3A57061CABD" pageId="1" pageNumber="266" type="distribution">
|
||
<paragraph id="342D849BF97C2E74A3C8062C8314F5D2" pageId="1" pageNumber="266">Distribution and ecological notes.</paragraph>
|
||
<paragraph id="2299C565106CF094899C095C3401E9C2" pageId="1" pageNumber="266">
|
||
All records of
|
||
<taxonomicName id="0B96D0EDD145C8F78C9864CFC4E67710" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="1" pageNumber="266" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
|
||
are from lowland and foothill forest in south-central Papua New Guinea, at altitudes ranging from ~100 m in the lowlands and foothills of Gulf Province to nearly 1,000 m asl on Iagifu Ridge in Southern Highlands Province. Males called at night from the upper surfaces of leaves, generally between 2-10 m above the forest floor.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DDFB8A160CB5438E784CA7A8C9F1692" lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266" type="vocalisation">
|
||
<paragraph id="F52F755A11CA2D66D29B56FEB37C85D9" pageId="1" pageNumber="266">Vocalisation.</paragraph>
|
||
<paragraph id="32C045F26B00D7FA569ADB5E3EBE59B0" lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266">
|
||
The advertisement call of the new species, recorded at air temperatures of 23.2-24.0 °C, consists of a series, or
|
||
<normalizedToken id="E4CD2723260FC664373621D31AAE95E7" originalValue="‘train’">'train'</normalizedToken>
|
||
of rather musical clicks (Fig. 3). Calls
|
||
<pageBreakToken id="07C753993E37C8F74A60A61C197BD624" pageId="2" pageNumber="267" start="start">follow</pageBreakToken>
|
||
one another at variable intervals (a few seconds to half a minute and more). Eleven calls were analysed from ZMB 84336 and 16 from SAMA R69442. Because characteristics of both call trains are similar, they are treated together here. 27 calls had a mean duration of 2.75
|
||
<normalizedToken id="FD96A9CD7CF9ED079BAFCEC6D5A51C3E" originalValue="±">+/-</normalizedToken>
|
||
0.82 s, range 1.01-4.75 s. Mean number of clicks (notes) per call 13.2
|
||
<normalizedToken id="BECA07AA90229E1B1B8FD4499DA8E926" originalValue="±">+/-</normalizedToken>
|
||
3.81, range 5-22. Notes had a mean duration of 20.6
|
||
<normalizedToken id="E5579F920C48AD3A82C27383437D4679" originalValue="±">+/-</normalizedToken>
|
||
2.91 ms, range 13-25 ms, n=93. Inter-note intervals lasted 200.6
|
||
<normalizedToken id="1BCDA74AC9510D583F5C8C6FFA85D6BE" originalValue="±">+/-</normalizedToken>
|
||
13.4 ms, range 176-238 ms, n=92. Mean note repetition rate 4.82
|
||
<normalizedToken id="E39CC10814929FDFC7EAE1ADCA7403AF" originalValue="±">+/-</normalizedToken>
|
||
0.21, range 4.39-5.18 notes/s, n=27. Frequencies scatter mostly between 5 and 6 kHz; the dominant frequency is at 5.5 kHz (Fig. 4). Notes start at maximum sound amplitude and, after a short time at this level, amplitude drops quickly. All notes have a similar maximum amplitude.
|
||
</paragraph>
|
||
<caption id="F19C67BE66CA8D406308C4588D477BAB" pageId="2" pageNumber="267">
|
||
<paragraph id="BC669F2E77FFF5984096B3C9A8AE0D19" pageId="2" pageNumber="267">
|
||
Figure 3. Wave form (above) and spectrogram (below) of an advertisement call from
|
||
<taxonomicName id="67FD6318FDEC82C5315DB30A84C23635" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
|
||
with 12 notes.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BFFC0560B5C9E62706429D65D15DFBDF" pageId="2" pageNumber="267">
|
||
<paragraph id="75DFB7E4572C337F57FED5BABA9B6ADB" pageId="2" pageNumber="267">
|
||
Figure 4. Power spectrum of an advertisement call of
|
||
<taxonomicName id="6BA9A440DAACAA893A80AD89E969C403" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="2" pageNumber="267" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
|
||
sp. n.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="FAF10C6638C584B66AFA514623DF75CF" pageId="2" pageNumber="267" type="etymology">
|
||
<paragraph id="6DB7CD97523A2C517929AE835BD68A63" pageId="2" pageNumber="267">Etymology.</paragraph>
|
||
<paragraph id="38FFE46727F616D4043EBC06CFDA3DAB" pageId="2" pageNumber="267">
|
||
The specific epithet
|
||
<taxonomicName id="0244F2B2B18617B2C97C750F8FE200CB" lsidName="crucifer" pageId="2" pageNumber="267" rank="species" species="crucifer">crucifer</taxonomicName>
|
||
is a Latin substantive in apposition and means carrier (porter) of a cross. It refers to the conspicuous yellowish cross on the head of most specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B298CDC784E8BEA064F16DEBBAF362C7" pageId="3" pageNumber="268" type="comparisons">
|
||
<paragraph id="C2E7A48462AAF8F8A30D6B148D3947F9" pageId="3" pageNumber="268">
|
||
<pageBreakToken id="5838447F7FD87F97FF232172B1735A6B" pageId="3" pageNumber="268" start="start">Comparisons</pageBreakToken>
|
||
with other species.
|
||
</paragraph>
|
||
<paragraph id="05E2115C35FA1D6811F30E15F7594390" pageId="3" pageNumber="268">
|
||
With its short snout
|
||
<taxonomicName id="7206129768436A3F18C69765F14360CB" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne crucifer" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="crucifer">Choerophryne crucifer</taxonomicName>
|
||
differs from all twelve
|
||
<taxonomicName id="28A7CB42D9448D4977A3038643CADFA9" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="genus">Choerophryne</taxonomicName>
|
||
with an elongated snout and would have been placed in the genus
|
||
<taxonomicName id="465B760E453BE56DB67B0B0DDEA5822B" class="Amphibia" family="Microhylidae" genus="Albericus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Albericus" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="genus">Albericus</taxonomicName>
|
||
in the former sense. According to
|
||
<bibRefCitation id="10E90043F0C45EF99195B2B074A1052A" author="Frost, DR" journalOrPublisher="American Museum of Natural History, New York" pageId="9" pageNumber="274" title="Amphibian Species of the World: an online reference, version 6.0" url="http://research.amnh.org/herpetology/amphibia/index.php" year="2016">Frost (2016)</bibRefCitation>
|
||
18 species of short-snouted
|
||
<taxonomicName id="CDCF539FA2675DEF45973AB1D21B6518" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="genus">Choerophryne</taxonomicName>
|
||
are recognized at present. The advertisement call of most short-snouted species is described as a buzz. Apart from
|
||
<taxonomicName id="BB64A3A72C7B7D9196D52195253EF148" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
, calls consisting of clicking notes are produced only by
|
||
<taxonomicName id="A6D3262B27B2E1606E9BA97B53CF1494" lsidName="C. gudrunae" pageId="3" pageNumber="268" rank="species" species="gudrunae">C. gudrunae</taxonomicName>
|
||
,
|
||
<taxonomicName id="67DE0400415C76913BADAF343C5EBB9D" lsidName="C. gunnari" pageId="3" pageNumber="268" rank="species" species="gunnari">C. gunnari</taxonomicName>
|
||
, C.
|
||
<taxonomicName id="428485923D52E10EAE503EC80F856EB7" lsidName="sanguinopicta" pageId="3" pageNumber="268" rank="species" species="sanguinopicta">sanguinopicta</taxonomicName>
|
||
and
|
||
<taxonomicName id="406E8006B2C1CABEE4A06954A7803B96" lsidName="C. valkuriarum" pageId="3" pageNumber="268" rank="species" species="valkuriarum">C. valkuriarum</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="ED07F6B2281B5939399F3BE594CD1711" pageId="3" pageNumber="268">
|
||
<taxonomicName id="88EEAE83FB4DDB33C38012E5D33AC004" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne gudrunae" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="gudrunae">Choerophryne gudrunae</taxonomicName>
|
||
has longer legs (TL/SVL 0.44-0.46 vs. 0.38-0.44) and a higher ratio of END/IND (1.20-1.50 vs. 1.00-1.25) than
|
||
<taxonomicName id="02A37C74ABED379AC982C71D0BA4ED65" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
. The advertisement calls of the former consist of about 10 clicks, those of
|
||
<taxonomicName id="4EBA8B1A25EA9F87CC32F68A244A1AD7" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
of 13 on average. Dominant frequency of
|
||
<taxonomicName id="73D77E17779547FD5554D0D3EC23C627" lsidName="gudrunae" pageId="3" pageNumber="268" rank="species" species="gudrunae">gudrunae</taxonomicName>
|
||
-calls is at 3.5 kHz, of
|
||
<taxonomicName id="60D22E12F1194E65C7471B1AE742667F" lsidName="crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">crucifer</taxonomicName>
|
||
-calls at 5.5 kHz.
|
||
</paragraph>
|
||
<paragraph id="027A15AC25C1F9D6B619069D1A68C355" pageId="3" pageNumber="268">
|
||
<taxonomicName id="0764AD7578A333A575CDDD4D8A990895" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne gunnari" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="gunnari">Choerophryne gunnari</taxonomicName>
|
||
has longer legs (TL/SVL 0.44-0.51), a higher ratio END/IND (1.25-1.45) and conspicuous blue pigmentation on the belly and hind limbs (absent in
|
||
<taxonomicName id="23992FC6450BF92B334CA924D8D8D84C" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
). Note repetition rate of the advertisement call of
|
||
<taxonomicName id="EA921D30E363A6118EDD0C214E19FAD3" lsidName="C. gunnari" pageId="3" pageNumber="268" rank="species" species="gunnari">C. gunnari</taxonomicName>
|
||
is, according to the spectrograms published by
|
||
<bibRefCitation id="41815CDE52E8007EE23332ADD0464200" author="Menzies, JI" journalOrPublisher="Australian Journal of Zoology" pageId="10" pageNumber="275" pagination="327 - 360" title="A study of Albericus (Anura: Microhylidae) of New Guinea." url="https://doi.org/10.1071/ZO99003" volume="47" year="1999">Menzies (1999</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="330D305EFA3A685B8027233400827608" author="Menzies, J" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="10" pageNumber="275" title="The frogs of New Guinea and the Solomon Islands." year="2006">2006</bibRefCitation>
|
||
), about two clicks/s (erroneously reported as about 4 clicks/s in the 1999 paper) vs. 4.4-5.2 clicks/s in
|
||
<taxonomicName id="05D94F9A0DD235C09A161464161E1E42" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
. Figure 17 in
|
||
<bibRefCitation id="364DA262D10D024266D70E1B6B164BCF" author="Menzies, JI" journalOrPublisher="Australian Journal of Zoology" pageId="10" pageNumber="275" pagination="327 - 360" title="A study of Albericus (Anura: Microhylidae) of New Guinea." url="https://doi.org/10.1071/ZO99003" volume="47" year="1999">Menzies (1999)</bibRefCitation>
|
||
shows a dominant frequency of about 3.3 kHz for this species, which is much lower than in calls from
|
||
<taxonomicName id="864350DF892599B7BDAF6931FA041D69" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
(5.5 kHz).
|
||
<bibRefCitation id="5CE2403B9C045D3A1DD4E0A3D638D680" author="Menzies, JI" journalOrPublisher="Australian Journal of Zoology" pageId="10" pageNumber="275" pagination="327 - 360" title="A study of Albericus (Anura: Microhylidae) of New Guinea." url="https://doi.org/10.1071/ZO99003" volume="47" year="1999">Menzies (1999)</bibRefCitation>
|
||
also reports that the clicks uttered by
|
||
<taxonomicName id="8C3A944055C46918F6399B6184B83129" lsidName="C. gunnari" pageId="3" pageNumber="268" rank="species" species="gunnari">C. gunnari</taxonomicName>
|
||
have 'no musical
|
||
<normalizedToken id="4E0EE5A9F5AD07DB08C9452AD832A255" originalValue="quality’">quality'</normalizedToken>
|
||
whereas those produced by
|
||
<taxonomicName id="6509DEB6A7C0F4E543798D6D25064224" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
are distinctly musical, sounding to the ear like
|
||
<normalizedToken id="EA010299E05CD12A3D0E968CC4F3D300" originalValue="‘tink..tink..tink’">'tink..tink..tink'</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="83068553B5AADAFC53D1333B13E47895" pageId="3" pageNumber="268">
|
||
<taxonomicName id="2CF3CEA787944F520BDEA22D8B9ACCC0" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne sanguinopicta" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="sanguinopicta">Choerophryne sanguinopicta</taxonomicName>
|
||
occurs at elevations of>1,400 m asl (vs. lowlands and foothills <1000 m asl) and exhibits an extraordinary polychromatic colouration (absent in
|
||
<taxonomicName id="8801DC3178C7DFE6E5211EFD3FC8BF60" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
). Moreover its advertisement calls are very long series (several minutes) of clicks with a note (= call) repetition rate of less than three calls/s and a dominant frequency of around 4.0 kHz (vs. shorter series of clicks, each series lasting less than 4 seconds and with a repetition rate of more than 4.4 per second and a higher dominant frequency (at 5.5 kHz) in
|
||
<taxonomicName id="7F2CCF43C528F7C656D606986C5D2AAE" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="13FC3EF727F108FC773D4A7B04CFC43D" pageId="3" pageNumber="268">
|
||
<taxonomicName id="37E0BA3D7DD829E37E6EE23DB510125F" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne valkuriarum" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="valkuriarum">Choerophryne valkuriarum</taxonomicName>
|
||
generally occurs at elevations over 2000 m asl (vs. <1000 m asl in
|
||
<taxonomicName id="6502F6BFC60CD20B23A714BFEAE40F67" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
). According to
|
||
<bibRefCitation id="C1B671199AF988F010CB783B163187AB" author="Kraus, F" journalOrPublisher="Pacific Science" pageId="10" pageNumber="275" pagination="43 - 53" title="A colorful new species of Albericus (Anura: Microhylidae) from southeastern New Guinea." url="https://doi.org/10.1353/psc.2005.0008" volume="59" year="2005 a">Kraus and Allison (2005a)</bibRefCitation>
|
||
HW/SVL in
|
||
<taxonomicName id="98B68299C7370BB683BF46E5BBA9309C" lsidName="C. valkuriarum" pageId="3" pageNumber="268" rank="species" species="valkuriarum">C. valkuriarum</taxonomicName>
|
||
is 0.37-0.43 and F3D/SVL is 0.067-0.088, we found values of 0.33-0.35 and 0.054-0.064 in
|
||
<taxonomicName id="7886F3BE114A5AAB89E10F41692BD565" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
. There are also differences in the advertisement calls.
|
||
<bibRefCitation id="2936CC36C7F470BA988618A9CABDB69F" author="Menzies, JI" journalOrPublisher="Australian Journal of Zoology" pageId="10" pageNumber="275" pagination="327 - 360" title="A study of Albericus (Anura: Microhylidae) of New Guinea." url="https://doi.org/10.1071/ZO99003" volume="47" year="1999">Menzies (1999)</bibRefCitation>
|
||
notes a maximum repetition rate for
|
||
<taxonomicName id="80E8C7B03A6FBC94A456C6EE0F3ABD56" lsidName="C. valkuriarum" pageId="3" pageNumber="268" rank="species" species="valkuriarum">C. valkuriarum</taxonomicName>
|
||
of two notes/s and reports that notes are uttered "once or in irregular series". In
|
||
<taxonomicName id="091841965536ED0CC0D444CE4FF96AD3" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
clicks are uttered at regular intervals and at a rate of 4.4-5.2 per s.
|
||
</paragraph>
|
||
<paragraph id="F8A4F1C3862310DD96B7DFB5CF7ED54C" pageId="3" pageNumber="268">
|
||
<taxonomicName id="2BADC5A968C6A61AB4A1F713ED61E6F9" class="Amphibia" family="Microhylidae" genus="Choerophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Choerophryne variegata" order="Anura" pageId="3" pageNumber="268" phylum="Chordata" rank="species" species="variegata">Choerophryne variegata</taxonomicName>
|
||
is known from a single specimen, and its advertisement call is unknown. According to measurements by
|
||
<bibRefCitation id="A30BE733484929A01F62CB7027205BC4" author="Menzies, JI" journalOrPublisher="Australian Journal of Zoology" pageId="10" pageNumber="275" pagination="327 - 360" title="A study of Albericus (Anura: Microhylidae) of New Guinea." url="https://doi.org/10.1071/ZO99003" volume="47" year="1999">Menzies (1999)</bibRefCitation>
|
||
the holotype of this species differs from the new species described here by the ratios TL/SVL (0.47 vs. 0.38-0.44), F3D/SVL (0.071 vs. 0.054-0.065), END/IND (1.38 vs. 1.00-1.25) and ED/SVL (0.133 vs. 0.110-0.127). Moreover, toes 4 and 5 are connected by webbing in
|
||
<taxonomicName id="B05247E80BFAA924ED094A94F8484568" lsidName="C. variegata" pageId="3" pageNumber="268" rank="species" species="variegata">C. variegata</taxonomicName>
|
||
and not so in
|
||
<taxonomicName id="660FEB8E8468DEC35F8C6000C612736E" lsidName="C. crucifer" pageId="3" pageNumber="268" rank="species" species="crucifer">C. crucifer</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |