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<mods:title id="35AAA3FCD806EF91C59BA9A7620A8422">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<taxonomicName id="51AA4D67FFEFFFC544A8FABB92B17049" ID-CoL="98YJ5" authority="(Meyrick, 1918)" baseAuthorityName="Meyrick" baseAuthorityYear="1918" box="[151,644,1338,1365]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="nephodesma">
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<emphasis id="A4DEEAF6FFEFFFC544A8FABB91837048" bold="true" box="[151,438,1338,1365]" italics="true" pageId="5" pageNumber="80">Tenupalpa nephodesma</emphasis>
(
<bibRefCitation id="F23B4B15FFEFFFC545FAFABB924E7049" author="Meyrick, E." box="[453,635,1338,1364]" pageId="5" pageNumber="80" pagination="97 - 160" refId="ref23843" refString="Meyrick, E. (1918) Exotic Microlepidoptera II. Thornhanger, Marlborough, Wilts, pp. 97 - 160." type="book chapter" year="1918">Meyrick, 1918</bibRefCitation>
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<figureCitation id="0E912A61FFEFFFC544A0FADE90DD7064" box="[159,232,1375,1401]" captionStart="FIGURES 36" captionStartId="4.[151,264,1894,1918]" captionTargetBox="[161,1422,453,1848]" captionTargetId="figure-154@4.[151,1435,453,1871]" captionTargetPageId="4" captionText="FIGURES 36. Abdominal and genital structures of Tenupalpa spp. 3, distal segments of the abdomen and male genitalia of T. angustella in external view; 4, denuded male 8th abdominal segment of T. angustella (image from Lee et al., 2021: Fig. 7I), slide no. LGE18719; 5, male genitalia of T. angustella, slide no. LGE16107; 6, female genitalia of T. nephodesma, slide no. LGE19022 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819798" httpUri="https://zenodo.org/record/5819798/files/figure.png" pageId="5" pageNumber="80">Figs 6</figureCitation>
,
<figureCitation id="0E912A61FFEFFFC544CBFADF91277064" box="[244,274,1374,1401]" captionStart="FIGURES 1525" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 1525. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="5" pageNumber="80">15</figureCitation>
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<figureCitation id="0E912A61FFEFFFC54521FADE91097064" box="[286,316,1375,1401]" captionStart="FIGURES 2645" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 2645. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="5" pageNumber="80">27</figureCitation>
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<figureCitation id="0E912A61FFEFFFC54577FADE91537064" box="[328,358,1375,1401]" captionStart="FIGURES 4653" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 4653. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="5" pageNumber="80">47</figureCitation>
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<figureCitation id="0E912A61FFEFFFC5454DFADE91A57064" box="[370,400,1375,1401]" captionStart="FIGURES 6671" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 6671. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="5" pageNumber="80">66</figureCitation>
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<figureCitation id="0E912A61FFEFFFC545A3FADF91D67064" box="[412,483,1374,1401]" captionStart="FIGURES 9597" captionStartId="31.[151,264,1868,1892]" captionTargetBox="[151,1435,344,1845]" captionTargetId="figure-90@31.[151,1435,344,1845]" captionTargetPageId="31" captionText="FIGURES 9597. Type specimen of Tenupalpa nephodesma deposited in NHMUK. 95. Holotype, female; 96. Label data; 97. Female genitalia (holotype), slide no. J.F.G.C. 8367, 97a, detail of signum." figureDoi="http://doi.org/10.5281/zenodo.5819838" httpUri="https://zenodo.org/record/5819838/files/figure.png" pageId="5" pageNumber="80">9597</figureCitation>
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<emphasis id="A4DEEAF6FFEFFFC544A8FA2291BD70A6" box="[151,392,1443,1467]" italics="true" pageId="5" pageNumber="80">Thiotricha nephodesma</emphasis>
<bibRefCitation id="F23B4B15FFEFFFC545B1FA22926C70A6" author="Meyrick, E." box="[398,601,1443,1467]" pageId="5" pageNumber="80" pagination="97 - 160" refId="ref23843" refString="Meyrick, E. (1918) Exotic Microlepidoptera II. Thornhanger, Marlborough, Wilts, pp. 97 - 160." type="book chapter" year="1918">Meyrick, 1918: 103</bibRefCitation>
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<bibRefCitation id="F23B4B15FFEFFFC5471AFA2293EB70A6" author="Clarke, J. F. G." box="[805,990,1443,1467]" pageId="5" pageNumber="80" refId="ref23257" refString="Clarke, J. F. G. (1969) Catalogue of the type specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. 7. Trustees of the British Museum, London, 531 pp., 265 pls." type="book" year="1969">Clarke, 1969: 463</bibRefCitation>
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<bibRefCitation id="F23B4B15FFEFFFC547D6FA2294EE70A6" author="Ivinskis P. P. &amp; Piskunov, V. I. &amp; Emeliyanov, I. M." box="[1001,1243,1443,1467]" pageId="5" pageNumber="80" pagination="37 - 44" refId="ref23428" refString="Ivinskis P. P., Piskunov, V. I. &amp; Emeliyanov, I. M. (1984) [Review of the fauna of Gelechiid moths of the Thiotricha Meyrick, 1886 genus and its place in the system of Lepidoptera: Gelechiidae, Dichomerinae]. Trudy AN Lithuanian SSR, 2 (86), 37 - 44. [in Russian]" type="journal article" year="1984">
Ivinskis
<emphasis id="A4DEEAF6FFEFFFC5407FFA25945970A6" box="[1088,1132,1443,1467]" italics="true" pageId="5" pageNumber="80">et al</emphasis>
., 1984: 39
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<collectingCountry id="EEBD7674FFEFFFC5415DFA2295AC70A6" box="[1378,1433,1443,1467]" name="India" pageId="5" pageNumber="80">India</collectingCountry>
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<treatmentCitation id="170B10F5FFEFFFC544A8FA61925470E4" author="Lee, G. - E. &amp; Han, T. M. &amp; Park, H. C. &amp; Qi, M. J. &amp; Li, H. H." box="[151,609,1504,1529]" page="371" pageId="5" pageNumber="80" year="2021">
<taxonomicName id="51AA4D67FFEFFFC544A8FA61925470E4" ID-CoL="98YJ5" authority="Lee et al., 2021: 371" authorityName="Lee" authorityPageNumber="371" authorityYear="2021" box="[151,609,1504,1529]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC544A8FA6191B270E5" box="[151,391,1504,1528]" italics="true" pageId="5" pageNumber="80">Tenupalpa nephodesma</emphasis>
<bibRefCitation id="F23B4B15FFEFFFC545B2FA60925470E4" author="Lee, G. - E. &amp; Han, T. M. &amp; Park, H. C. &amp; Qi, M. J. &amp; Li, H. H." box="[397,609,1504,1529]" pageId="5" pageNumber="80" pagination="357 - 379" refId="ref23692" refString="Lee, G. - E., Han, T. M., Park, H. C., Qi, M. J. &amp; Li, H. H. (2021) A phylogeny of the subfamily Thiotrichinae (Lepidoptera: Gel- echiidae) with a revision of the generic classification based on molecular and morphological analyses. Systematic Entomology, 46, 357 - 379. https: // doi. org / 10.1111 / syen. 12466" type="journal article" year="2021">
Lee
<emphasis id="A4DEEAF6FFEFFFC54586FA6091D970E5" box="[441,492,1504,1528]" italics="true" pageId="5" pageNumber="80">et al.</emphasis>
, 2021: 371
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<emphasis id="A4DEEAF6FFEFFFC544A8F9A6917C735D" bold="true" box="[151,329,1574,1601]" pageId="5" pageNumber="80">Type material.</emphasis>
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<location id="9375603FFFEFFFC5466EF9A692E2735C" LSID="urn:lsid:plazi:treatment:1E0387F2FFEFFFC644A8FABB93A07608:9375603FFFEFFFC5466EF9A692E2735C" box="[593,727,1575,1601]" country="India" name="Khasi Hills" pageId="5" pageNumber="80">Khasi Hills</location>
,
<date id="E2141024FFEFFFC546D8F9A6937F735C" box="[743,842,1575,1601]" pageId="5" pageNumber="80" value="1908-09">
<collectingDate id="F250E9CCFFEFFFC546D8F9A6937F735C" box="[743,842,1575,1601]" pageId="5" pageNumber="80" value="1908-09">ix. 1908</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFEFFFC547B1F9A69421735C" box="[910,1044,1575,1601]" pageId="5" pageNumber="80">E. Meyrick</collectorName>
, genitalia slide no. J.
<collectorName id="3B5F5332FFEFFFC54131F9A6956C735C" box="[1294,1369,1575,1601]" name="J.F.G. Clarke" pageId="5" pageNumber="80">F.G.C.</collectorName>
8367 (
<collectionCode id="F0BBAE21FFEFFFC544A0F9CA91257378" box="[159,272,1611,1637]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="80" type="Museum">NHMUK</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="961536E4FFEFFFC544F8F9EF93A173CC" blockId="5.[151,1437,1574,2033]" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544F8F9EF92007395" bold="true" box="[199,565,1646,1672]" pageId="5" pageNumber="80">Additional material examined:</emphasis>
<materialsCitation id="26C23CB9FFEFFFC54604F9EF93DF73B0" ID-GBIF-Occurrence="3427696331" accessionNumber="ZJL05343" collectingDate="2001-07-31" collectorName="Houhun Li &amp; Xinpu Wang" country="China" elevation="2100" latitude="27.55" location="Mt. Fanjing" longLatPrecision="742" longitude="108.41" pageId="5" pageNumber="80" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou">
<emphasis id="A4DEEAF6FFEFFFC54604F9EF93B27395" bold="true" box="[571,903,1646,1672]" pageId="5" pageNumber="80">
<collectingCountry id="EEBD7674FFEFFFC54604F9EF92A87395" box="[571,669,1646,1672]" name="China" pageId="5" pageNumber="80">CHINA</collectingCountry>
:
<collectingRegion id="546EF806FFEFFFC54698F9EF93B77395" box="[679,898,1646,1672]" country="China" name="Guizhou" pageId="5" pageNumber="80">Guizhou Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFEFFFC547B2F9EE938D7394" box="[909,952,1647,1673]" pageId="5" pageNumber="80" type="male">1 ♂</specimenCount>
,
<location id="9375603FFFEFFFC547FCF9EE94727394" LSID="urn:lsid:plazi:treatment:1E0387F2FFEFFFC644A8FABB93A07608:9375603FFFEFFFC547FCF9EE94727394" box="[963,1095,1647,1673]" country="China" latitude="27.55" longLatPrecision="742" longitude="108.41" name="Mt. Fanjing" pageId="5" pageNumber="80" stateProvince="Guizhou">Mt. Fanjing</location>
(
<geoCoordinate id="F39E5023FFEFFFC5406AF9EF94827394" box="[1109,1207,1646,1673]" degrees="27.55" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="555" value="27.55">27.55°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFEFFFC540FDF9EE951A7394" box="[1218,1327,1647,1673]" degrees="108.41" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="555" value="108.41">108.41°E</geoCoordinate>
),
<quantity id="51529B01FFEFFFC5417EF9EE95A27394" box="[1345,1431,1647,1673]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="80" unit="m" value="2100.0">
<elevation id="1D87D1D7FFEFFFC5417EF9EE95A27394" box="[1345,1431,1647,1673]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="80" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="E2141024FFEFFFC544A8F912912F73B0" box="[151,282,1683,1709]" pageId="5" pageNumber="80" value="2001-07-31">
<collectingDate id="F250E9CCFFEFFFC544A8F912912F73B0" box="[151,282,1683,1709]" pageId="5" pageNumber="80" value="2001-07-31">31.vii.2001</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFEFFFC54569F91291E473B0" box="[342,465,1683,1709]" pageId="5" pageNumber="80">Houhun Li</collectorName>
and
<collectorName id="3B5F5332FFEFFFC54637F91292AC73B0" box="[520,665,1683,1709]" pageId="5" pageNumber="80">Xinpu Wang</collectorName>
, genitalia slide no. ZJL05343
</materialsCitation>
;
<materialsCitation id="26C23CB9FFEFFFC547CAF91293A573CC" ID-GBIF-Occurrence="3427696313" accessionNumber="LGE19022" collectingDate="2004-08-19" collectorName="Yunli Xiao" country="China" county="Daozhen County" elevation="1300" location="Chengjiashan" pageId="5" pageNumber="80" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou">
<specimenCount id="80ACFD6DFFEFFFC547CAF912941573B0" box="[1013,1056,1683,1709]" pageId="5" pageNumber="80" type="female">1 ♀</specimenCount>
,
<location id="9375603FFFEFFFC54011F91294FC73B0" LSID="urn:lsid:plazi:treatment:1E0387F2FFEFFFC644A8FABB93A07608:9375603FFFEFFFC54011F91294FC73B0" box="[1070,1225,1683,1709]" country="China" county="Daozhen County" name="Chengjiashan" pageId="5" pageNumber="80" stateProvince="Guizhou">Chengjiashan</location>
,
<quantity id="51529B01FFEFFFC540EAF912951973B0" box="[1237,1324,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="80" unit="m" value="1300.0">
<elevation id="1D87D1D7FFEFFFC540EAF912951973B0" box="[1237,1324,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="80" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFEFFFC54106F91290DE73CC" pageId="5" pageNumber="80">Daozhen County</collectingCounty>
,
<date id="E2141024FFEFFFC544C9F93691B573CC" box="[246,384,1719,1745]" pageId="5" pageNumber="80" value="2004-08-19">
<collectingDate id="F250E9CCFFEFFFC544C9F93691B573CC" box="[246,384,1719,1745]" pageId="5" pageNumber="80" value="2004-08-19">19.viii.2004</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFEFFFC54585F936920273CC" box="[442,567,1719,1745]" pageId="5" pageNumber="80">Yunli Xiao</collectorName>
, genitalia slide no. LGE19022
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFEFFFC644F8F95B916D77D0" lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80" type="diagnosis">
<paragraph id="961536E4FFEFFFC544F8F95B93A27298" blockId="5.[151,1437,1574,2033]" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544F8F95B917773E9" bold="true" box="[199,322,1754,1780]" pageId="5" pageNumber="80">Diagnosis.</emphasis>
The species is distinguished by the following characteristics: a broad transverse fascia before middle of the forewing, a fine longitudinal line arising from the fascia toward the apex, a small trapezoidal projection on the valva, and a conical-shaped anellus lobe. The female genitalia of
<taxonomicName id="51AA4D67FFEFFFC54751F8A294207220" box="[878,1045,1827,1853]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC54751F8A294207220" box="[878,1045,1827,1853]" italics="true" pageId="5" pageNumber="80">T. nephodesma</emphasis>
</taxonomicName>
can be distinguished by a long and somewhat boomerang-shaped signum in lateral view. Externally,
<taxonomicName id="51AA4D67FFEFFFC54752F8C69421727C" box="[877,1044,1863,1889]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC54752F8C69421727C" box="[877,1044,1863,1889]" italics="true" pageId="5" pageNumber="80">T. nephodesma</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="51AA4D67FFEFFFC540E3F8C69541727C" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1244,1396,1863,1889]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="latifasciata" status="sp. nov.">
<emphasis id="A4DEEAF6FFEFFFC540E3F8C69541727C" box="[1244,1396,1863,1889]" italics="true" pageId="5" pageNumber="80">T. latifasciata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFEFFFC54144F8C690FD7298" bold="true" pageId="5" pageNumber="80">
<taxonomicNameLabel id="BFED578DFFEFFFC54144F8C690FD7298" pageId="5" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but the apex of the forewing is not darkened in
<taxonomicName id="51AA4D67FFEFFFC546D7F8EA93A57298" box="[744,912,1899,1925]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC546D7F8EA93A57298" box="[744,912,1899,1925]" italics="true" pageId="5" pageNumber="80">T. nephodesma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="961536E4FFEFFFC544F8F80F956172D0" blockId="5.[151,1437,1574,2033]" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544F8F80F914B72B5" bold="true" box="[199,382,1934,1960]" pageId="5" pageNumber="80">Re-description.</emphasis>
Adult (
<figureCitation id="0E912A61FFEFFFC545EFF80E921072B4" box="[464,549,1935,1961]" captionStart="FIGURES 4653" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 4653. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="5" pageNumber="80">Figs 47</figureCitation>
,
<figureCitation id="0E912A61FFEFFFC54610F80F927A72B4" box="[559,591,1934,1961]" captionStart="FIGURES 9597" captionStartId="31.[151,264,1868,1892]" captionTargetBox="[151,1435,344,1845]" captionTargetId="figure-90@31.[151,1435,344,1845]" captionTargetPageId="31" captionText="FIGURES 9597. Type specimen of Tenupalpa nephodesma deposited in NHMUK. 95. Holotype, female; 96. Label data; 97. Female genitalia (holotype), slide no. J.F.G.C. 8367, 97a, detail of signum." figureDoi="http://doi.org/10.5281/zenodo.5819838" httpUri="https://zenodo.org/record/5819838/files/figure.png" pageId="5" pageNumber="80">95</figureCitation>
).
<emphasis id="A4DEEAF6FFEFFFC5465EF80E92AB72B4" box="[609,670,1935,1961]" italics="true" pageId="5" pageNumber="80">Head</emphasis>
: Silvery white. Labial palpus dirty white, except palpomere III mixed with grey. Antenna with scape dirty white; flagellum dark fuscous, except base of dorsal surface dirty white.
</paragraph>
<paragraph id="961536E4FFEFFFC644F8F85691EC7400" blockId="5.[151,1437,1574,2033]" lastBlockId="6.[151,1437,151,789]" lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544F8F856912D72EC" box="[199,280,2007,2033]" italics="true" pageId="5" pageNumber="80">Thorax</emphasis>
: Dorsum, tegula, and forewing dirty white mixed and pale brown; wingspan 9.0 mm; forewing with anterior edge of costa black in basal half, a very broad fuscous fascia between basal 1/3 and 2/3 of wing, a fine fuscous median line from outer margin of former fascia, running longitudinally toward apex; fringe scales grey. Hindwing and fringe scales grey. Legs dirty white and pale brown mixed, hind tibia broadly dark fuscous after middle and shortly dark fuscous at apex.
</paragraph>
<paragraph id="961536E4FFECFFC644F8FEA6916D77D0" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FEA69104745C" box="[199,305,295,321]" italics="true" pageId="6" pageNumber="81">Abdomen</emphasis>
: Male with coremata slightly shorter than sternum VIII; sternum VIII sub-pentagonal, rounded laterally (
<figureCitation id="0E912A61FFECFFC644EEFECA91107478" box="[209,293,330,357]" captionStart="FIGURES 1525" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 1525. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="6" pageNumber="81">Fig. 15</figureCitation>
). Male genitalia (
<figureCitation id="0E912A61FFECFFC645D3FECA920B747B" box="[492,574,331,358]" captionStart="FIGURES 6671" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 6671. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="6" pageNumber="81">Fig. 66</figureCitation>
) with uncus small, sub-pentagonal, protruded medially. Gnathos hook short, gradually narrowing to apex. Tegumen with a set of papilliform tubercles at middle (
<figureCitation id="0E912A61FFECFFC64061FEEE94817494" box="[1118,1204,367,393]" captionStart="FIGURES 2645" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 2645. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="6" pageNumber="81">Fig. 27</figureCitation>
). Valva with a small trapezoidal projection near middle of costa, apex rounded. Anellus lobe elongate-conical with a short subapical bristle, slightly curved, pointed apically. Juxta convex in basal 1/3 of inner margin. Vinculum simple. Saccus broad, slightly exceeding tegumen pedunculus. Aedeagus with basal half dilated, distal part slender with apex rounded. Female genitalia (
<figureCitation id="0E912A61FFECFFC6455DFE7E91997704" box="[354,428,511,537]" captionStart="FIGURES 36" captionStartId="4.[151,264,1894,1918]" captionTargetBox="[161,1422,453,1848]" captionTargetId="figure-154@4.[151,1435,453,1871]" captionTargetPageId="4" captionText="FIGURES 36. Abdominal and genital structures of Tenupalpa spp. 3, distal segments of the abdomen and male genitalia of T. angustella in external view; 4, denuded male 8th abdominal segment of T. angustella (image from Lee et al., 2021: Fig. 7I), slide no. LGE18719; 5, male genitalia of T. angustella, slide no. LGE16107; 6, female genitalia of T. nephodesma, slide no. LGE19022 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819798" httpUri="https://zenodo.org/record/5819798/files/figure.png" pageId="6" pageNumber="81">Figs 6</figureCitation>
,
<figureCitation id="0E912A61FFECFFC64586FE7E91E37704" box="[441,470,511,537]" captionStart="FIGURES 9597" captionStartId="31.[151,264,1868,1892]" captionTargetBox="[151,1435,344,1845]" captionTargetId="figure-90@31.[151,1435,344,1845]" captionTargetPageId="31" captionText="FIGURES 9597. Type specimen of Tenupalpa nephodesma deposited in NHMUK. 95. Holotype, female; 96. Label data; 97. Female genitalia (holotype), slide no. J.F.G.C. 8367, 97a, detail of signum." figureDoi="http://doi.org/10.5281/zenodo.5819838" httpUri="https://zenodo.org/record/5819838/files/figure.png" pageId="6" pageNumber="81">97</figureCitation>
) with papillae anales simple. Apophyses anteriores slightly shorter than 1/2 length of apophyses posteriores. Tergum VIII with posterior margin deeply concave at middle in V-shape, bearing a bundle of setae at the cleft; sternum VIII broadly convex anteriorly. Ostium bursae round, opening at middle of sternum VIII. Ductus bursae broad at base, constricted after base then gradually broadening toward corpus bursae; colliculum situated near base. Corpus bursae ovate, slightly shorter than ductus bursae; signum linear, situated in posterior half of corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644F8FD57945577EC" box="[199,1120,726,754]" pageId="6" pageNumber="81" type="distribution">
<paragraph id="961536E4FFECFFC644F8FD57945577EC" blockId="6.[151,1437,151,789]" box="[199,1120,726,754]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FD57916C77ED" bold="true" box="[199,345,726,752]" pageId="6" pageNumber="81">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFECFFC64558FD56919F77EC" box="[359,426,727,753]" name="China" pageId="6" pageNumber="81">China</collectingCountry>
(
<collectingRegion id="546EF806FFECFFC64586FD56922A77EF" box="[441,543,727,754]" country="China" name="Guizhou" pageId="6" pageNumber="81">Guizhou</collectingRegion>
),
<collectingCountry id="EEBD7674FFECFFC6460DFD56925A77EC" box="[562,623,727,753]" name="India" pageId="6" pageNumber="81">India</collectingCountry>
. This species is newly recorded from
<collectingCountry id="EEBD7674FFECFFC64029FD56946877EC" box="[1046,1117,727,753]" name="China" pageId="6" pageNumber="81">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644F8FD7A93A07608" box="[199,917,763,789]" pageId="6" pageNumber="81" type="discussion">
<paragraph id="961536E4FFECFFC644F8FD7A93A07608" blockId="6.[151,1437,151,789]" box="[199,917,763,789]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FD7A910E7608" bold="true" box="[199,315,763,789]" pageId="6" pageNumber="81">Remarks.</emphasis>
The male genitalia are newly illustrated in this paper.
</paragraph>
</subSubSection>
</treatment>
</document>