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<document id="CB1ADA5BA59A8FA5117766FEF38FA027" ID-DOI="10.11646/zootaxa.3857.1.6" ID-GBIF-Dataset="f134bbeb-a658-4640-9687-04a86526fb23" ID-ISSN="1175-5326" ID-Zenodo-Dep="227424" ID-ZooBank="C11922A0-154E-4026-BEA2-261358622FD0" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460897283549" checkinUser="plazi" docAuthor="Skibińska, Kornelia, Krzemiński, Wiesław &amp; Coram, Robert A." docDate="2014" docId="1A02878D524D5746FF7D0F239BDD8C90" docLanguage="en" docName="zt03857p130.pdf" docOrigin="Zootaxa 3857 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nannotanyderus oliviae Skibinska, Krzeminski &amp; Coram, sp. nov." docType="treatment" docVersion="7" lastPageNumber="127" masterDocId="E63BFFF5524C5744FFEA093C9A5D8D01" masterDocTitle="Discovery of the most ancient member of family Tanyderidae (Diptera) from the Lower Jurassic (Sinemurian) of England" masterLastPageNumber="130" masterPageNumber="125" pageNumber="126" updateTime="1698360182778" updateUser="plazi">
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<mods:title id="A642E3705A2041CCAD168BA01A8F1C29">Discovery of the most ancient member of family Tanyderidae (Diptera) from the Lower Jurassic (Sinemurian) of England</mods:title>
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<mods:namePart id="733DC7DF4606E4EB6DCF146F18FC64E5">Skibińska, Kornelia</mods:namePart>
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<mods:namePart id="6ADB4EBF196D96AE221DB57286AD1B7A">Krzemiński, Wiesław</mods:namePart>
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<mods:namePart id="23D23D7E67B5BEC08982C93C26FCE940">Coram, Robert A.</mods:namePart>
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<mods:title id="BC98D51E7A9FAA7665FBD652061C7730">Zootaxa</mods:title>
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<mods:date id="0A4B26B1D3EA2D78E71A287AFBAD71CA">2014</mods:date>
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<treatment id="1A02878D524D5746FF7D0F239BDD8C90" ID-DOI="http://doi.org/10.5281/zenodo.6140760" ID-GBIF-Taxon="119587531" ID-Zenodo-Dep="6140760" LSID="urn:lsid:plazi:treatment:1A02878D524D5746FF7D0F239BDD8C90" httpUri="http://treatment.plazi.org/id/1A02878D524D5746FF7D0F239BDD8C90" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126">
<subSubSection id="DAB16510524D5745FF7D0F239B4C8B5A" pageId="1" pageNumber="126" type="nomenclature">
<paragraph id="9214369B524D5745FF7D0F2399E28B38" blockId="1.[151,959,1567,1627]" box="[151,959,1567,1593]" pageId="1" pageNumber="126">
<heading id="C95C81F7524D5745FF7D0F2399E28B38" bold="true" box="[151,959,1567,1593]" fontSize="11" level="1" pageId="1" pageNumber="126" reason="1">
<emphasis id="A0DFEA89524D5745FF7D0F2399E28B38" bold="true" box="[151,959,1567,1593]" pageId="1" pageNumber="126">
<taxonomicName id="55AB4D18524D5745FF7D0F2399068B38" authority="Skibinska, Krzeminski &amp; Coram" authorityName="Skibinska, Krzeminski &amp; Coram" box="[151,859,1567,1593]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="oliviae" status="sp. nov.">
<emphasis id="A0DFEA89524D5745FF7D0F239BE58B38" bold="true" box="[151,440,1567,1593]" italics="true" pageId="1" pageNumber="126">Nannotanyderus oliviae</emphasis>
Skibińska, Krzemiński &amp; Coram
</taxonomicName>
<taxonomicNameLabel id="BBEC57F2524D5745FC880F2399E28B38" box="[866,959,1567,1593]" pageId="1" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9214369B524D5745FF7D0F7E9B4C8B5A" blockId="1.[151,959,1567,1627]" box="[151,273,1602,1627]" pageId="1" pageNumber="126">
(
<figureCitation id="0A902A1E524D5745FF750F7E9B578B5A" box="[159,266,1602,1627]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="2.[151,250,1072,1094]" captionStartId-1="2.[151,250,1821,1843]" captionStartId-2="3.[151,250,703,725]" captionStartId-3="3.[151,250,1383,1405]" captionTargetBox-0="[151,1436,476,1034]" captionTargetBox-1="[151,1436,1190,1783]" captionTargetBox-2="[283,1318,223,620]" captionTargetBox-3="[220,1347,840,1320]" captionTargetId-0="figure@2.[151,1436,476,1035]" captionTargetId-1="figure@2.[151,1436,1190,1783]" captionTargetId-2="figure@3.[236,1350,193,682]" captionTargetId-3="figure@3.[171,1415,806,1362]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionText-0="FIGURE 2. Photography of the wing venation of Nannotanyderus oliviae sp. nov.; holotype (Lower Jurassic, Sinemurian). Scales 0.5 mm." captionText-1="FIGURE 3. Photography of the wing venation of Nannotanyderus oliviae sp. nov.; paratype Scales 0.5 mm." captionText-2="FIGURE 4. Reconstruction of the wing venation of Nannotanyderus oliviae sp. nov.; holotype (Lower Jurassic, Sinemurian). Scale 0.5 mm." captionText-3="FIGURE 5. Reconstruction of the wing venation of Nannotanyderus oliviae sp. nov.; paratype (Lower Jurassic, Sinemurian). Scale 0.5 mm." httpUri-0="https://zenodo.org/record/227426/files/figure.png" httpUri-1="https://zenodo.org/record/227427/files/figure.png" httpUri-2="https://zenodo.org/record/227428/files/figure.png" httpUri-3="https://zenodo.org/record/227429/files/figure.png" pageId="1" pageNumber="126">Figs. 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DAB16510524D5745FF7D0FB69E0E8BA3" box="[151,1107,1674,1699]" pageId="1" pageNumber="126" type="etymology">
<paragraph id="9214369B524D5745FF7D0FB69E0E8BA3" blockId="1.[151,1437,1674,2037]" box="[151,1107,1674,1699]" pageId="1" pageNumber="126">
<emphasis id="A0DFEA89524D5745FF7D0FB69B4B8BA2" bold="true" box="[151,278,1674,1699]" pageId="1" pageNumber="126">Etymology</emphasis>
. The new species name is dedicated to Olivia, daughter of Robert Coram.
</paragraph>
</subSubSection>
<subSubSection id="DAB16510524D5745FF2D0F91989E8A34" pageId="1" pageNumber="126" type="materials_examined">
<paragraph id="9214369B524D5745FF2D0F91989E8A34" blockId="1.[151,1437,1674,2037]" pageId="1" pageNumber="126">
<emphasis id="A0DFEA89524D5745FF2D0F919B218BC7" bold="true" box="[199,380,1709,1734]" pageId="1" pageNumber="126">
<typeStatus id="4D108839524D5745FF2D0F919B5F8BC7" box="[199,258,1709,1734]" pageId="1" pageNumber="126">Type</typeStatus>
material.
</emphasis>
<typeStatus id="4D108839524D5745FE600F919BAF8BC7" box="[394,498,1709,1734]" pageId="1" pageNumber="126" type="holotype">Holotype</typeStatus>
No. I-F/MP/
<date id="E615105B524D5745FD720F91995B8BC7" box="[664,774,1709,1734]" pageId="1" pageNumber="126" value="1621-02-13">2/1621/13</date>
, complete wing;
<typeStatus id="4D108839524D5745FC360F939E638BC6" box="[988,1086,1711,1735]" pageId="1" pageNumber="126" type="paratype">Paratype</typeStatus>
No. I-F/MP/
<date id="E615105B524D5745FB0E0F919F0F8BC7" box="[1252,1362,1709,1734]" pageId="1" pageNumber="126" value="1600-01-12">1/1600/12</date>
, wing lacking anal lobe below A1 vein. Both from Brooki Bed, Shales-with-Beef Member, Charmouth Mudstone Formation, Lias Group (Lower Sinemurian, Turneri zone, Brooki subzone), Monmouth Beach, Lyme Regis, Dorset,
<collectingCountry id="EABC760B524D5745FF050E219B458A35" box="[239,280,1821,1844]" name="United Kingdom" pageId="1" pageNumber="126">UK</collectingCountry>
(national grid reference SY 334 914).
</paragraph>
</subSubSection>
<subSubSection id="DAB16510524D5745FF2D0E7C9FCE8AF4" pageId="1" pageNumber="126" type="diagnosis">
<paragraph id="9214369B524D5745FF2D0E7C9FCE8AF4" blockId="1.[151,1437,1674,2037]" pageId="1" pageNumber="126">
<emphasis id="A0DFEA89524D5745FF2D0E7C9B1C8A58" bold="true" box="[199,321,1856,1881]" pageId="1" pageNumber="126">Diagnosis.</emphasis>
The new species differs from all other members of the genus in that veins Rs, R4+5 and R4 form a nearly straight line, and vein R4+5 is shorter than in other described species. It also differs from
<taxonomicName id="55AB4D18524D5745FB580E549F338A7E" box="[1202,1390,1896,1919]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="grimmenensis">
<emphasis id="A0DFEA89524D5745FB580E549F338A7E" box="[1202,1390,1896,1919]" italics="true" pageId="1" pageNumber="126">N. grimmenensis</emphasis>
</taxonomicName>
and
<taxonomicName id="55AB4D18524D5745FF7D0EB39B458AA7" box="[151,280,1935,1958]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="ansorgei">
<emphasis id="A0DFEA89524D5745FF7D0EB39B458AA7" box="[151,280,1935,1958]" italics="true" pageId="1" pageNumber="126">N. ansorgei</emphasis>
</taxonomicName>
in that Sc terminates on the wing margin before fork of Rs into R2+3 and R4+5, from
<taxonomicName id="55AB4D18524D5745FB440EB39F108AA7" box="[1198,1357,1934,1958]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="krzeminskii">
<emphasis id="A0DFEA89524D5745FB440EB39F108AA7" box="[1198,1357,1934,1958]" italics="true" pageId="1" pageNumber="126">N. krzeminskii</emphasis>
</taxonomicName>
and
<taxonomicName id="55AB4D18524D5745FA680EB39B768ACC" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="kubekovensis">
<emphasis id="A0DFEA89524D5745FA680EB39B768ACC" italics="true" pageId="1" pageNumber="126">N. kubekovensis</emphasis>
</taxonomicName>
in that m-cu is more distal to the fork of M3+4 into M3 and M4, and from
<emphasis id="A0DFEA89524D5745FB960E8A9F5B8ACC" box="[1148,1286,1974,1997]" italics="true" pageId="1" pageNumber="126">
?
<taxonomicName id="55AB4D18524D5745FB6D0E8A9F5B8ACC" box="[1159,1286,1974,1997]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="incertus">N. incertus</taxonomicName>
</emphasis>
in that Rs is shorter. The anal lobe is also more developed than in
<taxonomicName id="55AB4D18524D5745FD070EE199D18AF5" box="[749,908,2012,2036]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="krzeminskii">
<emphasis id="A0DFEA89524D5745FD070EE199D18AF5" box="[749,908,2012,2036]" italics="true" pageId="1" pageNumber="126">N. krzeminskii</emphasis>
</taxonomicName>
and
<taxonomicName id="55AB4D18524D5745FC290EE19E188AF5" box="[963,1093,2013,2036]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="ansorgei">
<emphasis id="A0DFEA89524D5745FC290EE19E188AF5" box="[963,1093,2013,2036]" italics="true" pageId="1" pageNumber="126">N. ansorgei</emphasis>
</taxonomicName>
(not known in other species).
</paragraph>
</subSubSection>
<subSubSection id="DAB16510524E5746FF2D09AB9FDE8C23" pageId="2" pageNumber="127" type="description">
<paragraph id="9214369B524E5746FF2D09AB9FDE8C23" blockId="2.[151,1436,151,401]" pageId="2" pageNumber="127">
<emphasis id="A0DFEA89524E5746FF2D09AB9B048DB1" bold="true" box="[199,345,151,176]" pageId="2" pageNumber="127">Description.</emphasis>
<typeStatus id="4D108839524E5746FE8B09AB9B908DB1" box="[353,461,151,176]" pageId="2" pageNumber="127" type="holotype">Holotype</typeStatus>
. Wing length
<quantity id="55539B7E524E5746FD8709A498958DAE" box="[621,712,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="127" unit="mm" value="2.8">2.8 mm</quantity>
. Sc short, extending half wing length, ending before fork of Rs into R2+3 and R4+5; R2+3 1.6 times longer than Rs and 2.6 times longer than R2; Rs as long as R3 and forming straight line with vein R4; R5 originates directly from R4 a short distance distal to its fork from Rs; vein M1 about 1.3 times longer than the upper edge of d cell; m-cu leaves M4 distal to the fork of M3+4 into M3 and M4. Explicit anal lobe.
</paragraph>
</subSubSection>
<subSubSection id="DAB16510524E5746FF2D080C9BDD8C90" pageId="2" pageNumber="127" type="discussion">
<paragraph id="9214369B524E5746FF2D080C9BDD8C90" blockId="2.[151,1436,151,401]" pageId="2" pageNumber="127">
<emphasis id="A0DFEA89524E5746FF2D080C9B668C48" bold="true" box="[199,315,304,329]" pageId="2" pageNumber="127">Remarks.</emphasis>
Although similar to other
<taxonomicName id="55AB4D18524E5746FD80080C997E8C49" box="[618,803,304,328]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="A0DFEA89524E5746FD80080C997E8C49" box="[618,803,304,328]" italics="true" pageId="2" pageNumber="127">Nannotanyderus</emphasis>
</taxonomicName>
species, in particular
<taxonomicName id="55AB4D18524E5746FBF6080D9EE38C49" box="[1052,1214,304,328]" class="Insecta" family="Tanyderidae" genus="Nannotanyderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="krzeminskii">
<emphasis id="A0DFEA89524E5746FBF6080D9EE38C49" box="[1052,1214,304,328]" italics="true" pageId="2" pageNumber="127">N. krzeminskii</emphasis>
</taxonomicName>
, the observed wing venation differences and significantly older age (in excess of 12 Ma), justify the description of a new species for the English material.
</paragraph>
</subSubSection>
</treatment>
</document>