210 lines
21 KiB
XML
210 lines
21 KiB
XML
<document id="689C8A325DFC8583FA7ADDDB943DE0EA" ID-DOI="10.5281/zenodo.274671" ID-GBIF-Dataset="de46898d-f662-46aa-ab26-7e75546f425d" ID-ISSN="1175-5326" ID-Zenodo-Dep="274671" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459995356039" checkinUser="plazi" docAuthor="Schmied, Heiko, Wappler, Torsten & Č, Ji Ř Í Kolibá" docDate="2009" docId="0394BD6DD367354C4390FA03FBF0ED22" docLanguage="en" docName="zt01993p026.pdf" docOrigin="Zootaxa 1993" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ancyrona eocenica Schmied, Wappler & Č, 2009, sp. nov." docType="treatment" docVersion="7" lastPageNumber="20" masterDocId="FFADC515D366354F4307FFAAFF8AEE48" masterDocTitle="A new bark-gnawing beetle (Coleoptera, Trogossitidae) from the middle Eocene of Europe, with a checklist of fossil Trogossitidae" masterLastPageNumber="26" masterPageNumber="17" pageNumber="18" updateTime="1698232808219" updateUser="plazi">
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<mods:title id="D5B73B7396C7932E0E3D37EDF0F588B7">A new bark-gnawing beetle (Coleoptera, Trogossitidae) from the middle Eocene of Europe, with a checklist of fossil Trogossitidae</mods:title>
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<mods:namePart id="322B492B300F0CA28DC18CA5FD627E6D">Schmied, Heiko</mods:namePart>
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<mods:namePart id="1CA713A734B102802DC566EEC758F0A8">Wappler, Torsten</mods:namePart>
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<mods:namePart id="E4BD316038DA4B191661BC8D5BA57949">Č, Ji Ř Í Kolibá</mods:namePart>
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<mods:date id="6F654025581C3D24ADAA48195C3E4CB4">2009</mods:date>
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<mods:number id="AE495FDD2EE49D23C623CB009820BA33">1993</mods:number>
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<mods:identifier id="E5673C524DCC3180345E5354EC6192C1" type="DOI">10.5281/zenodo.274671</mods:identifier>
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<treatment id="0394BD6DD367354C4390FA03FBF0ED22" ID-DOI="http://doi.org/10.5281/zenodo.6218170" ID-GBIF-Taxon="119380274" ID-Zenodo-Dep="6218170" LSID="urn:lsid:plazi:treatment:0394BD6DD367354C4390FA03FBF0ED22" httpUri="http://treatment.plazi.org/id/0394BD6DD367354C4390FA03FBF0ED22" lastPageId="3" lastPageNumber="20" pageId="1" pageNumber="18">
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<subSubSection id="C3275FF0D367354E4390FA03FEDEEBAE" pageId="1" pageNumber="18" type="nomenclature">
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<paragraph id="8B820C7BD367354E4390FA03FE69EB8B" blockId="1.[151,483,1449,1510]" box="[151,483,1449,1475]" pageId="1" pageNumber="18">
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<heading id="D0CABB17D367354E4390FA03FE69EB8B" bold="true" box="[151,483,1449,1475]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
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<emphasis id="B949D069D367354E4390FA03FE69EB8B" bold="true" box="[151,483,1449,1475]" pageId="1" pageNumber="18">
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<taxonomicName id="4C3D77F8D367354E4390FA03FEF5EB8B" box="[151,383,1449,1475]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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<emphasis id="B949D069D367354E4390FA03FEF5EB8B" bold="true" box="[151,383,1449,1475]" italics="true" pageId="1" pageNumber="18">Ancyrona eocenica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A6D12D367354E4281FA03FE69EB8B" box="[390,483,1449,1475]" pageId="1" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B820C7BD367354E4390FA66FEDEEBAE" blockId="1.[151,483,1449,1510]" box="[151,340,1484,1510]" pageId="1" pageNumber="18">(Figs. 1A, B, C)</paragraph>
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</subSubSection>
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<subSubSection id="C3275FF0D367354E4390F9B3FC2AE878" box="[151,928,1561,1584]" pageId="1" pageNumber="18" type="description">
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||
<paragraph id="8B820C7BD367354E4390F9B3FC2AE878" blockId="1.[151,928,1561,1584]" box="[151,928,1561,1584]" pageId="1" pageNumber="18">
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<taxonomicName id="4C3D77F8D367354E4390F9B3FE81E878" box="[151,267,1561,1584]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Cassidinae">Cassidinae</taxonomicName>
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gen. et sp. indet.
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<bibRefCitation id="EFAC718AD367354E42C0F9B3FDFAE878" author="Wappler" box="[455,624,1561,1584]" pageId="1" pageNumber="22" refString="Wappler, T. (2003 a) Die Insekten aus dem Mittel-Eozan des Eckfelder Maares, Vulkaneifel. Mainzer Naturwissenschaftliches Archiv, 27, 1 - 234." type="journal article" year="2003" yearSuffix="a">Wappler, 2003a</bibRefCitation>
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, pl. 13, fig. i, p. 110, fig. 75.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3275FF0D367354E4390F9CAFCAAE8A6" pageId="1" pageNumber="18" type="materials_examined">
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<paragraph id="8B820C7BD367354E4390F9CAFD93E8E9" blockId="1.[151,1437,1632,2006]" pageId="1" pageNumber="18">
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<emphasis id="B949D069D367354E4390F9CAFE92E832" bold="true" box="[151,280,1632,1658]" pageId="1" pageNumber="18">
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<typeStatus id="5486B2D9D367354E4390F9CAFE99E832" box="[151,275,1632,1658]" pageId="1" pageNumber="18" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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PE_2000/475, LS, completely preserved, dorsal side visible. Deposited in the Eckfeld insect collection at the
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<collectionCode id="ED2C94BED367354E425EF92DFE31E8E9" box="[345,443,1671,1697]" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="1" pageNumber="18">NHMM</collectionCode>
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, Mainz.
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</paragraph>
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<paragraph id="8B820C7BD367354E43C2F907FCAAE8A6" blockId="1.[151,1437,1632,2006]" pageId="1" pageNumber="18">
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<emphasis id="B949D069D367354E43C2F907FD98E88F" bold="true" box="[197,530,1709,1735]" pageId="1" pageNumber="18">
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<typeStatus id="5486B2D9D367354E43C2F907FE89E88F" box="[197,259,1709,1735]" pageId="1" pageNumber="18">Type</typeStatus>
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locality and horizon:
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</emphasis>
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Eckfeld Maar near Manderscheid; middle Eocene, middle Lutetian,
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<collectionCode id="ED2C94BED367354E464FF907FA16E88F" box="[1352,1436,1709,1735]" pageId="1" pageNumber="18">ELMA</collectionCode>
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Geiseltalian, MP 13, 44.3 ± 0.4 Ma (
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<bibRefCitation id="EFAC718AD367354E4144F97EFC9EE8A6" author="Mertz" box="[579,788,1748,1774]" pageId="1" pageNumber="22" refString="Mertz, D. F., Swisher, C. C., Franzen, J. L., Neuffer, F. O. & Lutz, H. (2000) Numerical dating of the Eckfeld maar fossil site, Eifel, Germany: A calibration mark for the Eocene time scale. Naturwissenschaften, 87, 270 - 274." type="journal article" year="2000">
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Mertz
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<emphasis id="B949D069D367354E4195F97EFD47E8A5" box="[658,717,1748,1773]" italics="true" pageId="1" pageNumber="18">et al.</emphasis>
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2000
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3275FF0D367354C43C2F950FBDAEFD2" lastPageId="3" lastPageNumber="20" pageId="1" pageNumber="18" type="description">
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<paragraph id="8B820C7BD367354E43C2F950FD81E9C1" blockId="1.[151,1437,1632,2006]" pageId="1" pageNumber="18">
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<emphasis id="B949D069D367354E43C2F950FED5E95C" bold="true" box="[197,351,1786,1812]" pageId="1" pageNumber="18">Description:</emphasis>
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Body length 2.66 mm, width 1.63 mm, distance between eyes 0.34 mm, pronotal length 0.55 mm, pronotal width 1.41 mm (fragmentary length), elytral length 1.83 mm, width of compressed part of elytra 0.16 mm. Dorsal surface black-brown; head, pronotum, and elytra conspicuously and regularly punctate. Pubescence of body not visible.
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</paragraph>
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<paragraph id="8B820C7BD367354D43C2F83FFBCAE9A6" blockId="1.[151,1437,1632,2006]" lastBlockId="2.[151,1436,1786,2031]" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
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Two small apical teeth oriented in horizontal axis, with one tooth visible on right mandible; frontoclypeal suture present; compound eyes large, exopthalmic, not emarginate (Fig. 1C). Pronotum transverse, twice as
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<emphasis id="B949D069D364354D4390F950FE95E95C" bold="true" box="[151,287,1786,1812]" pageId="2" pageNumber="19">FIGURE 1.</emphasis>
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A, B, C:
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<taxonomicName id="4C3D77F8D364354D428EF957FDDCE95C" box="[393,598,1789,1812]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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<emphasis id="B949D069D364354D428EF957FDDCE95C" box="[393,598,1789,1812]" italics="true" pageId="2" pageNumber="19">Ancyrona eocenica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A6D12D364354D4159F956FD3FE95C" box="[606,693,1788,1812]" pageId="2" pageNumber="19" rank="species">
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<emphasis id="B949D069D364354D4159F956FD3FE95C" bold="true" box="[606,693,1788,1812]" pageId="2" pageNumber="19">sp. nov.</emphasis>
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</taxonomicNameLabel>
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(PE_2000/475, LS): (A)
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<typeStatus id="5486B2D9D364354D40CEF956FBA7E95B" box="[969,1069,1788,1811]" pageId="2" pageNumber="19" type="holotype">Holotype</typeStatus>
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in dorsal view, (B) camera lucida drawing of the
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<typeStatus id="5486B2D9D364354D423DF888FE10E971" box="[314,410,1826,1849]" pageId="2" pageNumber="19" type="holotype">holotype</typeStatus>
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, (C) detail of the head, showing details of the mandible (M) and fragmentary preserved parts of the first antennal segments (Ant). D:
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<taxonomicName id="4C3D77F8D364354D4131F8ECFC1EE914" authority="Leveille, 1899" authorityName="Leveille" authorityYear="1899" box="[566,916,1861,1885]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="vicina">
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<emphasis id="B949D069D364354D4131F8ECFD66E915" box="[566,748,1862,1885]" italics="true" pageId="2" pageNumber="19">Ancyrona vicina</emphasis>
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Léveillé, 1899
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</taxonomicName>
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“
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<collectingCountry id="F32A4CEBD364354D40A2F8EFFB9AE914" box="[933,1040,1861,1884]" name="Cameroon" pageId="2" pageNumber="19">Kamerun</collectingCountry>
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, Col. Kraatz, Grouvelle det.” (DEI, ZALF); general habitus. E:
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<taxonomicName id="4C3D77F8D364354D42CEF8C0FC86E9C9" authority="Pic, 1921" authorityName="Pic" authorityYear="1921" box="[457,780,1898,1921]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="diversa">
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<emphasis id="B949D069D364354D42CEF8C0FD00E9C9" box="[457,650,1898,1921]" italics="true" pageId="2" pageNumber="19">Ancyrona diversa</emphasis>
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(Pic, 1921)
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</taxonomicName>
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“
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<collectingCountry id="F32A4CEBD364354D401AF8C0FCE8E9C9" box="[797,866,1898,1921]" name="South Korea" pageId="2" pageNumber="19">Korea</collectingCountry>
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Prov. Gang-von, district On-dzong, Kum-gang san, near hotel Gon-song,
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<quantity id="4CC5A19ED364354D4286F827FE42E9EC" box="[385,456,1933,1956]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="19" unit="m" value="250.0">250 m</quantity>
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;
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<date id="FF832ABBD364354D42D3F827FDE4E9EC" box="[468,622,1933,1956]" pageId="2" pageNumber="19" value="1975-08-04">4 August1975</date>
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, leg. J. Papp et A. Vojnits”; general habitus. F:
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<taxonomicName id="4C3D77F8D364354D4769F824FF5FE981" authority="Linneaus, 1758" authorityName="Linneaus" authorityYear="1758" class="Insecta" family="Trogossitidae" genus="Peltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="ferruginea">
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<emphasis id="B949D069D364354D4769F824FAA9E9ED" box="[1134,1315,1934,1957]" italics="true" pageId="2" pageNumber="19">Peltis ferruginea</emphasis>
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(Linneaus, 1758)
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</taxonomicName>
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(ZFMK, Bonn); detail of the head; black arrows indicating the distinctly extended corners of the pronotum. G:
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<taxonomicName id="4C3D77F8D364354D4682F818FEEFE9A6" authority="Pic, 1921" authorityName="Pic" authorityYear="1921" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="diversa">
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<emphasis id="B949D069D364354D4682F818FF6CE9A7" italics="true" pageId="2" pageNumber="19">A. diversa</emphasis>
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(Pic, 1921)
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</taxonomicName>
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; detail of head and pronotum. Scale bars: A–C,
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<quantity id="4CC5A19ED364354D4062F87DFC2CE9A6" box="[869,934,2007,2030]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="19" unit="mm" value="1.0">1 mm</quantity>
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. E–G 0.5 mm.
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</paragraph>
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<paragraph id="8B820C7BD365354C4390FF32FCFBEF6E" blockId="3.[151,1437,152,874]" pageId="3" pageNumber="20">wide as head, anterior margin weakly concave, anterior corners not distinctly projecting, posterolateral corners rounded, posterior margin straight and as wide as elytra. Scutellum small, semicircular. Elytral humeri evenly rounded. Elytra regularly punctate, with approximately ten rows of punctures and ten carinae. Dorsal lateral margins of elytra flattened along entire length of elytra.</paragraph>
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<paragraph id="8B820C7BD365354C43C2FE98FBDAEFD2" blockId="3.[151,1437,152,874]" pageId="3" pageNumber="20">
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<emphasis id="B949D069D365354C43C2FE98FE7AEF04" bold="true" box="[197,496,306,332]" pageId="3" pageNumber="20">Differential diagnosis:</emphasis>
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The new species differs from recent species of
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<taxonomicName id="4C3D77F8D365354C4757FE98FB42EF03" box="[1104,1224,306,331]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="B949D069D365354C4757FE98FB42EF03" box="[1104,1224,306,331]" italics="true" pageId="3" pageNumber="20">Ancyrona</emphasis>
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</taxonomicName>
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in the following combination of characters: (1) minute body (2.66 mm), (2) anterior margin of pronotum weakly concave, (3) anterior corners of pronotum not distinctly developed, and (4) body broadly oval.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3275FF0D365354C43C2FE0CFAFEEF88" box="[197,1396,422,448]" pageId="3" pageNumber="20" type="etymology">
|
||
<paragraph id="8B820C7BD365354C43C2FE0CFAFEEF88" blockId="3.[151,1437,152,874]" box="[197,1396,422,448]" pageId="3" pageNumber="20">
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<emphasis id="B949D069D365354C43C2FE0CFEDFEF88" bold="true" box="[197,341,422,448]" pageId="3" pageNumber="20">Etymology:</emphasis>
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The specific epithet is taken from the Greek
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<emphasis id="B949D069D365354C4060FE0DFC1CEF88" box="[871,918,423,448]" italics="true" pageId="3" pageNumber="20">eos,</emphasis>
|
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referring to the Eocene age of the fossil.
|
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3275FF0D365354C43C2FE67FBF0ED22" pageId="3" pageNumber="20" type="discussion">
|
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<paragraph id="8B820C7BD365354C43C2FE67FE6FECBE" blockId="3.[151,1437,152,874]" pageId="3" pageNumber="20">
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<emphasis id="B949D069D365354C43C2FE67FEDCEFAF" bold="true" box="[197,342,461,487]" pageId="3" pageNumber="20">Comments:</emphasis>
|
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The diverse genus
|
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<taxonomicName id="4C3D77F8D365354C4139FE67FD3AEFAE" box="[574,688,461,486]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="B949D069D365354C4139FE67FD3AEFAE" box="[574,688,461,486]" italics="true" pageId="3" pageNumber="20">Ancyrona</emphasis>
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</taxonomicName>
|
||
(~ 50 described and many undescribed species) is widespread, extending from Africa south of the Sahara across to
|
||
<collectingCountry id="F32A4CEBD365354C4007FE5EFC05EC46" box="[768,911,500,526]" name="Madagascar" pageId="3" pageNumber="20">Madagascar</collectingCountry>
|
||
, and throughout southeast Asia to
|
||
<collectingCountry id="F32A4CEBD365354C4620FE5EFAE0EC46" box="[1319,1386,500,526]" name="Japan" pageId="3" pageNumber="20">Japan</collectingCountry>
|
||
and east Siberia, New
|
||
<collectingCountry id="F32A4CEBD365354C4270FDB0FE58EC7C" box="[375,466,538,564]" name="Guinea-Bissau" pageId="3" pageNumber="20">Guinea</collectingCountry>
|
||
, and northern
|
||
<collectingCountry id="F32A4CEBD365354C4183FDB0FD72EC7C" box="[644,760,538,564]" name="Australia" pageId="3" pageNumber="20">Australia</collectingCountry>
|
||
. One species (
|
||
<taxonomicName id="4C3D77F8D365354C40ADFDB0FB6BEC7C" authority="Reitter, 1889" authorityName="Reitter" authorityYear="1889" box="[938,1249,538,564]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="B949D069D365354C40ADFDB0FBB2EC7B" box="[938,1080,538,563]" italics="true" pageId="3" pageNumber="20">A. japonica</emphasis>
|
||
Reitter, 1889
|
||
</taxonomicName>
|
||
) also occurs in Europe (
|
||
<collectingCountry id="F32A4CEBD365354C43FBFDEBFEECEC13" box="[252,358,577,603]" name="Bulgaria" pageId="3" pageNumber="20">Bulgaria</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A4CEBD365354C4274FDEBFE51EC13" box="[371,475,577,603]" name="Hungary" pageId="3" pageNumber="20">Hungary</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F32A4CEBD365354C411DFDEBFD0CEC13" box="[538,646,577,603]" name="Slovakia" pageId="3" pageNumber="20">Slovakia</collectingCountry>
|
||
). Distribution records in South
|
||
<collectingCountry id="F32A4CEBD365354C40FCFDEBFBEAEC13" box="[1019,1120,577,603]" name="United States of America" pageId="3" pageNumber="20">America</collectingCountry>
|
||
(two species reported) are dubious and should be double-checked. Body size varies from
|
||
<quantity id="4CC5A19ED365354C408FFDC2FB8DECCA" box="[904,1031,616,642]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="3" pageNumber="20" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">3 to 8 mm</quantity>
|
||
; dorsal surface is covered by wax scales, thick setae, fine hairs, or is glabrous; anterior margin of pronotum is emarginate (concave), anterior corners are more or less projecting; eyes exopthalmic; joints of antennal club rounded to serrate; body rounded or weakly elongate.
|
||
</paragraph>
|
||
<paragraph id="8B820C7BD365354C43C1FCA8FBF0ED22" blockId="3.[151,1437,152,874]" pageId="3" pageNumber="20">
|
||
<bibRefCitation id="EFAC718AD365354C43C1FCA8FE08ED54" author="Kolibac" box="[198,386,770,796]" pageId="3" pageNumber="21" refString="Kolibac, J. (2007) Further observations on the tribe Ancyronini Kolibac, 2006 (Coleoptera, Trogossitidae, Peltinae). Entomologica Basiliensia et Collectionis Frey, 29, 53 - 76." type="journal article" year="2007">Kolibáč (2007)</bibRefCitation>
|
||
divided
|
||
<taxonomicName id="4C3D77F8D365354C42FAFCA8FDE4ED53" box="[509,622,770,795]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949D069D365354C42FAFCA8FDE4ED53" box="[509,622,770,795]" italics="true" pageId="3" pageNumber="20">Ancyrona</emphasis>
|
||
</taxonomicName>
|
||
into five species groups based on common body habitus and distribution rather than phylogenetic affinities. Thus, there is inherent difficulty in classifying
|
||
<taxonomicName id="4C3D77F8D365354C47EDFC83FAF9ED0A" box="[1258,1395,809,834]" class="Insecta" family="Trogossitidae" genus="Ancyrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
|
||
<emphasis id="B949D069D365354C47EDFC83FAF9ED0A" box="[1258,1395,809,834]" italics="true" pageId="3" pageNumber="20">A. eocenica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A6D12D365354C467CFC83FF42ED22" pageId="3" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in any of these groups due to a lack of sculpture and antennae in the fossil form.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |