301 lines
36 KiB
XML
301 lines
36 KiB
XML
<document id="11F7A286A46736FD4F0853E42926D7E6" ID-DOI="10.11646/zootaxa.5497.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13618112" ID-ZooBank="F29D526F-79AC-469B-BA88-3EF640EEA1E1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725025687576" checkinUser="plazi" docAuthor="Dell’Angelo, Bruno, Sosso, Maurizio & Taviani, Marco" docDate="2024" docId="039487A1FF8EFFD8FF0155C4EB69A2F4" docLanguage="en" docName="zootaxa.5497.2.2.pdf" docOrigin="Zootaxa 5497 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hanleya schwabei Dell’Angelo, Sosso & Taviani, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="199" masterDocId="FFADFFD9FF8DFFDDFF965251E803A325" masterDocTitle="A critical update of Mediterranean chitons (Mollusca: Polyplacophora) with the description of new taxa" masterLastPageNumber="208" masterPageNumber="194" pageNumber="197" updateTime="1725026850595" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AD336F7017475FC8013C7315B212F7BF">A critical update of Mediterranean chitons (Mollusca: Polyplacophora) with the description of new taxa</mods:title>
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<mods:name id="841C0A42C60033BB12F846F087499D30" type="personal">
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<mods:namePart id="A9DF63AB637843EADAFC2377EF4D040A">Dell’Angelo, Bruno</mods:namePart>
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<mods:namePart id="CF824F2EA15670B26EF45EC3C4CD8D2B">Sosso, Maurizio</mods:namePart>
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<mods:affiliation id="D809F00D285285B1B0AFB41EEC4F8099">Independent Researcher, via Bengasi 4, 16153 Genova, Italy</mods:affiliation>
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<mods:namePart id="EFD19E3854E0036E075AFFD0A4729D8B">Taviani, Marco</mods:namePart>
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<mods:affiliation id="C5F14B45F3BE5ED8BB4A04CB0A989E37">Istituto di Scienze Marine (ISMAR-CNR), via Gobetti 101, 40129 Bologna, Italy & Stazione Zoologica Anton Dohrn, Villa Comunale, 80121 Napoli, Italy & Independent Researcher, via Briscata</mods:affiliation>
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<mods:title id="A99925B4FEDFD189E31D6D775B07E8C6">Zootaxa</mods:title>
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<mods:part id="DFE62C6BC910C88FF3EBFBA4BAE048CE">
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<mods:date id="E4130ED5F495894C3C49534BD0443582">2024</mods:date>
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<mods:number id="8AB2A4219C8C4EA64F5B627EC1838861">2024-08-23</mods:number>
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<treatment id="039487A1FF8EFFD8FF0155C4EB69A2F4" LSID="urn:lsid:plazi:treatment:039487A1FF8EFFD8FF0155C4EB69A2F4" httpUri="http://treatment.plazi.org/id/039487A1FF8EFFD8FF0155C4EB69A2F4" lastPageId="5" lastPageNumber="199" pageId="3" pageNumber="197">
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<subSubSection id="C327653CFF8EFFDEFF0155C4EA08A495" box="[151,523,1941,1968]" pageId="3" pageNumber="197" type="nomenclature">
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<paragraph id="8B8236B7FF8EFFDEFF0155C4EA08A495" blockId="3.[151,523,1941,2004]" box="[151,523,1941,1968]" pageId="3" pageNumber="197">
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<heading id="D0CA81DBFF8EFFDEFF0155C4EA08A495" bold="true" box="[151,523,1941,1968]" fontSize="11" level="1" pageId="3" pageNumber="197" reason="1">
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<emphasis id="B949EAA5FF8EFFDEFF0155C4EA08A495" bold="true" box="[151,523,1941,1968]" pageId="3" pageNumber="197">
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<taxonomicName id="4C3D4D34FF8EFFDEFF0155C4E977A495" authorityName="Dell’Angelo & Sosso & Taviani" authorityYear="2024" box="[151,372,1941,1968]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="197" phylum="Mollusca" rank="species" species="schwabei" status="sp. nov.">
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<emphasis id="B949EAA5FF8EFFDEFF0155C4E977A495" bold="true" box="[151,372,1941,1968]" italics="true" pageId="3" pageNumber="197">Hanleya schwabei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57DEFF8EFFDEFEED55C7EA08A495" box="[379,523,1942,1968]" pageId="3" pageNumber="197" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C327653CFF8EFFDEFF0155EBE90BA4F1" box="[151,264,1978,2004]" pageId="3" pageNumber="197" type="description">
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<paragraph id="8B8236B7FF8EFFDEFF0155EBE90BA4F1" blockId="3.[151,523,1941,2004]" box="[151,264,1978,2004]" pageId="3" pageNumber="197">
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(
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<figureCitation id="13062A32FF8EFFDEFF0955EBE8FCA4F1" box="[159,255,1978,2004]" captionStart="FIGURE 3" captionStartId="4.[151,250,1821,1845]" captionTargetBox="[151,1436,1200,1797]" captionTargetId="figure-399@4.[151,1436,1200,1797]" captionTargetPageId="4" captionText="FIGURE 3. Hanleya schwabei sp. nov., Tyrrhenian Sea: A. offshore Capraia island-Capo Corso, depth ca. 350 m, (possibly last glacial Pleistocene), Paratype 1 (MNHN-IM-2022-2410), intermediate valve, width 3.4 mm, dorsal view. B–E. Off Scilla, Holotype (MZUB 60338), intermediate valve, width 3.8 mm, dorsal view (B), close-up of surface ornamentation of pleural (C) and lateral (D) areas, and frontal view (E). F–H. Off Scilla, Paratype 4 (ZISP 2423), tail valve, width 2.7 mm, dorsal view (F), close-up of surface ornamentation of antemucronal area (G) and lateral view (H). Scale bars = 400 μm (G), 100 μm (D), 40 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.13618120" httpUri="https://zenodo.org/record/13618120/files/figure.png" pageId="3" pageNumber="197">Figure 3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C327653CFF89FFD9FF0152C6EC4EA38A" box="[151,1101,150,175]" pageId="4" pageNumber="198" type="reference_group">
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<paragraph id="8B8236B7FF89FFD9FF0152C6EC4EA38A" blockId="4.[151,1101,150,175]" box="[151,1101,150,175]" pageId="4" pageNumber="198">
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<treatmentCitationGroup id="AB2D1199FF89FFD9FF0152C6EC4EA38A" box="[151,1101,150,175]" pageId="4" pageNumber="198">
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<taxonomicName id="4C3D4D34FF89FFD9FF0152C6E9A7A38A" ID-CoL="6LC6L" authorityName="Sirenko" authorityYear="2014" box="[151,420,151,175]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" isUncertain="true" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="198" phylum="Mollusca" rank="species" species="mediterranea">
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<emphasis id="B949EAA5FF89FFD9FF0152C6E8EEA38A" box="[151,237,151,175]" italics="true" pageId="4" pageNumber="198">Hanleya</emphasis>
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aff.
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<emphasis id="B949EAA5FF89FFD9FE8A52C6E9A7A38A" box="[284,420,151,175]" italics="true" pageId="4" pageNumber="198">mediterranea</emphasis>
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</taxonomicName>
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(
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<emphasis id="B949EAA5FF89FFD9FE2552C6E9DAA38B" box="[435,473,151,174]" italics="true" pageId="4" pageNumber="198">non</emphasis>
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<bibRefCitation id="EFAC4B46FF89FFD9FE4952C6EA71A38A" author="Sirenko, B. I." box="[479,626,151,175]" pageId="4" pageNumber="198" pagination="2913 - 2945" refId="ref10023" refString="Sirenko, B. I. (2014) Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History, 48, 2913 - 2945. https: // doi. org / 10.1080 / 00222933.2014.963722" type="journal article" year="2014">Sirenko, 2014</bibRefCitation>
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);
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<treatmentCitation id="0A9C10A6FF89FFD9FD1552C6EBC1A38A" author="Dell'Angelo, B. & Renda, W. & Sirenko, B. I. & Sosso, M. & Giacobbe, S." box="[643,962,151,175]" page="131" pageId="4" pageNumber="198" year="2021">
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<bibRefCitation id="EFAC4B46FF89FFD9FD1552C6EB75A38A" author="Dell'Angelo, B. & Renda, W. & Sirenko, B. I. & Sosso, M. & Giacobbe, S." box="[643,886,151,175]" pageId="4" pageNumber="198" pagination="124 - 135" refId="ref7731" refString="Dell'Angelo, B., Renda, W., Sirenko, B. I., Sosso, M. & Giacobbe, S. (2021) The Mediterranean distribution of Hanleya hanleyi (Bean in Thorpe, 1844) and H. mediterranea Sirenko, 2014 (Polyplacophora). Bollettino Malacologico, 57 (1), 124 - 135." type="journal article" year="2021">
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Dell’Angelo
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||
<emphasis id="B949EAA5FF89FFD9FC9E52C6EB36A38A" box="[776,821,151,175]" italics="true" pageId="4" pageNumber="198">et al</emphasis>
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. 2021
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</bibRefCitation>
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, p. 131
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</treatmentCitation>
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, fig. 5, tab. 3.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C327653CFF89FFD9FF015287EADAA281" pageId="4" pageNumber="198" type="materials_examined">
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<paragraph id="8B8236B7FF89FFD9FF015287E8EBA2A5" blockId="4.[151,1437,214,1176]" pageId="4" pageNumber="198">
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<emphasis id="B949EAA5FF89FFD9FF015287E94BA3D5" bold="true" box="[151,328,214,240]" pageId="4" pageNumber="198">Type material.</emphasis>
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<typeStatus id="54868815FF89FFD9FEC55287E9C3A3D5" box="[339,448,214,240]" pageId="4" pageNumber="198" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DB9B9ECCFF89FFD9FE5B5286EA72A3D5" box="[461,625,214,241]" collectionCode="MZUB" country="Italy" httpUri="http://grbio.org/cool/xa0v-1feb" name="Museo di Zoologia" pageId="4" pageNumber="198">MZUB 60338</specimenCode>
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, intermediate valve, width
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<quantity id="4CC59B52FF89FFD9FC385287EC0BA3D4" box="[942,1032,214,241]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="198" unit="mm" value="3.8">3.8 mm</quantity>
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(
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<figureCitation id="13062A32FF89FFD9FB8F5287EC9AA3D5" box="[1049,1177,214,240]" captionStart="FIGURE 3" captionStartId="4.[151,250,1821,1845]" captionTargetBox="[151,1436,1200,1797]" captionTargetId="figure-399@4.[151,1436,1200,1797]" captionTargetPageId="4" captionText="FIGURE 3. Hanleya schwabei sp. nov., Tyrrhenian Sea: A. offshore Capraia island-Capo Corso, depth ca. 350 m, (possibly last glacial Pleistocene), Paratype 1 (MNHN-IM-2022-2410), intermediate valve, width 3.4 mm, dorsal view. B–E. Off Scilla, Holotype (MZUB 60338), intermediate valve, width 3.8 mm, dorsal view (B), close-up of surface ornamentation of pleural (C) and lateral (D) areas, and frontal view (E). F–H. Off Scilla, Paratype 4 (ZISP 2423), tail valve, width 2.7 mm, dorsal view (F), close-up of surface ornamentation of antemucronal area (G) and lateral view (H). Scale bars = 400 μm (G), 100 μm (D), 40 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.13618120" httpUri="https://zenodo.org/record/13618120/files/figure.png" pageId="4" pageNumber="198">Figs 3B–E</figureCitation>
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)
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;
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<materialsCitation id="3B553CEAFF89FFD9FB275286E8B3A21C" collectionCode="MNHN-" elevation="500" pageId="4" pageNumber="198" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54868815FF89FFD9FB275286ED10A3D4" box="[1201,1299,215,241]" pageId="4" pageNumber="198" type="paratype">Paratype</typeStatus>
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1:
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<collectionCode id="ED2CAE72FF89FFD9FAAB5286ED9FA3D4" box="[1341,1436,215,241]" pageId="4" pageNumber="198">MNHN-</collectionCode>
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IM-2022-2410, intermediate valve, width
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<quantity id="4CC59B52FF89FFD9FDEC52ABEAD0A230" box="[634,723,250,277]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="198" unit="mm" value="3.4">3.4 mm</quantity>
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(
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<figureCitation id="13062A32FF89FFD9FD7352ABEB3CA231" box="[741,831,250,276]" captionStart="FIGURE 3" captionStartId="4.[151,250,1821,1845]" captionTargetBox="[151,1436,1200,1797]" captionTargetId="figure-399@4.[151,1436,1200,1797]" captionTargetPageId="4" captionText="FIGURE 3. Hanleya schwabei sp. nov., Tyrrhenian Sea: A. offshore Capraia island-Capo Corso, depth ca. 350 m, (possibly last glacial Pleistocene), Paratype 1 (MNHN-IM-2022-2410), intermediate valve, width 3.4 mm, dorsal view. B–E. Off Scilla, Holotype (MZUB 60338), intermediate valve, width 3.8 mm, dorsal view (B), close-up of surface ornamentation of pleural (C) and lateral (D) areas, and frontal view (E). F–H. Off Scilla, Paratype 4 (ZISP 2423), tail valve, width 2.7 mm, dorsal view (F), close-up of surface ornamentation of antemucronal area (G) and lateral view (H). Scale bars = 400 μm (G), 100 μm (D), 40 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.13618120" httpUri="https://zenodo.org/record/13618120/files/figure.png" pageId="4" pageNumber="198">Fig. 3A</figureCitation>
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) from offshore Capraia island-Capo Corso -350/
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<quantity id="4CC59B52FF89FFD9FAE452ABE8B3A21C" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="198" unit="m" value="500.0">
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<elevation id="0010D184FF89FFD9FAE452ABE8B3A21C" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="198" unit="m" value="500.0">500 m</elevation>
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</quantity>
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;
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<materialsCitation id="3B553CEAFF89FFD9FF28534EEC75A21C" box="[190,1142,286,313]" collectionCode="MZUB" country="Algeria" location="Tipaza" pageId="4" pageNumber="198" specimenCode="MZUB 60339" specimenCount="1" stateProvince="Tipaza" typeStatus="paratype">
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<typeStatus id="54868815FF89FFD9FF28534EE923A21C" box="[190,288,287,313]" pageId="4" pageNumber="198" type="paratype">Paratype</typeStatus>
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2:
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<specimenCode id="DB9B9ECCFF89FFD9FEDE534EE9EFA21D" box="[328,492,286,313]" collectionCode="MZUB" country="Italy" httpUri="http://grbio.org/cool/xa0v-1feb" name="Museo di Zoologia" pageId="4" pageNumber="198">MZUB 60339</specimenCode>
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, intermediate valve from
|
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<collectingRegion id="49F9F855FF89FFD9FC85534FEB62A21D" box="[787,865,286,312]" country="Algeria" name="Tipaza" pageId="4" pageNumber="198">Tipaza</collectingRegion>
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,
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<collectingCountry id="F32A7627FF89FFD9FCFB534FEBC7A21D" box="[877,964,286,312]" name="Algeria" pageId="4" pageNumber="198">Algeria</collectingCountry>
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, width
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<quantity id="4CC59B52FF89FFD9FB8D534FEC75A21C" box="[1051,1142,286,313]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="198" unit="mm" value="3.5">3.5 mm</quantity>
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</materialsCitation>
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;
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<materialsCitation id="3B553CEAFF89FFD9FB12534EEB1BA278" collectionCode="SMF" country="Algeria" location="Gorgona island" pageId="4" pageNumber="198" specimenCode="SMF 66407" specimenCount="1" stateProvince="Tipaza" typeStatus="paratype">
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<typeStatus id="54868815FF89FFD9FB12534EECE5A21C" box="[1156,1254,287,313]" pageId="4" pageNumber="198" type="paratype">Paratype</typeStatus>
|
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3:
|
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<specimenCode id="DB9B9ECCFF89FFD9FA98534FED9BA21D" box="[1294,1432,286,312]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="198">SMF 66407</specimenCode>
|
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, intermediate valve from
|
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<location id="8EE2606CFF89FFD9FE255313EA66A279" LSID="urn:lsid:plazi:treatment:039487A1FF8EFFD8FF0155C4EB69A2F4:8EE2606CFF89FFD9FE255313EA66A279" box="[435,613,322,348]" country="Algeria" name="Gorgona island" pageId="4" pageNumber="198" stateProvince="Tipaza">Gorgona island</location>
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, width
|
||
<quantity id="4CC59B52FF89FFD9FD2A5313EB1BA278" box="[700,792,322,349]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="198" unit="mm" value="3.5">3.5 mm</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B553CEAFF89FFD9FCB05312E8E0A2A5" collectionCode="ZISP" country="Algeria" pageId="4" pageNumber="198" specimenCode="ZISP 2423" specimenCount="1" stateProvince="Tipaza" typeStatus="paratype">
|
||
<typeStatus id="54868815FF89FFD9FCB05312EB8BA278" box="[806,904,323,349]" pageId="4" pageNumber="198" type="paratype">Paratype</typeStatus>
|
||
4:
|
||
<specimenCode id="DB9B9ECCFF89FFD9FC275313EC2DA279" box="[945,1070,322,348]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="198">ZISP 2423</specimenCode>
|
||
, tail valve, width
|
||
<quantity id="4CC59B52FF89FFD9FA975313ED5AA278" box="[1281,1369,322,349]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="198" unit="mm" value="2.7">2.7 mm</quantity>
|
||
(
|
||
<figureCitation id="13062A32FF89FFD9FAFC5313E8D8A2A5" captionStart="FIGURE 3" captionStartId="4.[151,250,1821,1845]" captionTargetBox="[151,1436,1200,1797]" captionTargetId="figure-399@4.[151,1436,1200,1797]" captionTargetPageId="4" captionText="FIGURE 3. Hanleya schwabei sp. nov., Tyrrhenian Sea: A. offshore Capraia island-Capo Corso, depth ca. 350 m, (possibly last glacial Pleistocene), Paratype 1 (MNHN-IM-2022-2410), intermediate valve, width 3.4 mm, dorsal view. B–E. Off Scilla, Holotype (MZUB 60338), intermediate valve, width 3.8 mm, dorsal view (B), close-up of surface ornamentation of pleural (C) and lateral (D) areas, and frontal view (E). F–H. Off Scilla, Paratype 4 (ZISP 2423), tail valve, width 2.7 mm, dorsal view (F), close-up of surface ornamentation of antemucronal area (G) and lateral view (H). Scale bars = 400 μm (G), 100 μm (D), 40 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.13618120" httpUri="https://zenodo.org/record/13618120/files/figure.png" pageId="4" pageNumber="198">Figs 3F–H</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236B7FF89FFD9FF5153DBEADAA281" blockId="4.[151,1437,214,1176]" box="[199,729,394,420]" pageId="4" pageNumber="198">
|
||
<materialsCitation id="3B553CEAFF89FFD9FF5153DBEAD6A281" box="[199,725,394,420]" location="Scilla" municipality="Tyrrhenian Sea" pageId="4" pageNumber="198" specimenCount="1" stateProvince="Calabria" typeStatus="holotype">
|
||
<emphasis id="B949EAA5FF89FFD9FF5153DBE965A281" bold="true" box="[199,358,394,420]" pageId="4" pageNumber="198">
|
||
<typeStatus id="54868815FF89FFD9FF5153DBE901A281" box="[199,258,394,420]" pageId="4" pageNumber="198">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingMunicipality id="6BE6ACCDFF89FFD9FEFA53DBEA1AA281" box="[364,537,394,420]" pageId="4" pageNumber="198">Tyrrhenian Sea</collectingMunicipality>
|
||
,
|
||
<location id="8EE2606CFF89FFD9FDB253DBEA64A281" LSID="urn:lsid:plazi:treatment:039487A1FF8EFFD8FF0155C4EB69A2F4:8EE2606CFF89FFD9FDB253DBEA64A281" box="[548,615,394,420]" municipality="Tyrrhenian Sea" name="Scilla" pageId="4" pageNumber="198" stateProvince="Calabria">Scilla</location>
|
||
,
|
||
<collectingRegion id="49F9F855FF89FFD9FDE453DBEAD6A281" box="[626,725,394,420]" country="Italy" name="Calabria" pageId="4" pageNumber="198">Calabria</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327653CFF89FFD9FF5153FFE9B3A2C9" pageId="4" pageNumber="198" type="etymology">
|
||
<paragraph id="8B8236B7FF89FFD9FF5153FFE9B3A2C9" blockId="4.[151,1437,214,1176]" pageId="4" pageNumber="198">
|
||
<emphasis id="B949EAA5FF89FFD9FF5153FFE94DA2ED" bold="true" box="[199,334,430,456]" pageId="4" pageNumber="198">Etymology.</emphasis>
|
||
This species is named after Enrico Schwabe in recognition of his important contribution to the study of
|
||
<taxonomicName id="4C3D4D34FF89FFD9FF6F5383E9AAA2C9" authorityName="Gray" authorityYear="1821" box="[249,425,466,492]" class="Polyplacophora" kingdom="Animalia" pageId="4" pageNumber="198" phylum="Mollusca" rank="class">Polyplacophora</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327653CFF89FFD9FF5153A7EB92A110" pageId="4" pageNumber="198" type="materials_examined">
|
||
<paragraph id="8B8236B7FF89FFD9FF5153A7EB92A110" blockId="4.[151,1437,214,1176]" pageId="4" pageNumber="198">
|
||
<emphasis id="B949EAA5FF89FFD9FF5153A7E9B1A135" bold="true" box="[199,434,502,528]" pageId="4" pageNumber="198">Material examined.</emphasis>
|
||
Western Mediterranean, Tyrrhenian Sea, Capraia island: 2 valves (BD 249,
|
||
<collectionCode id="ED2CAE72FF89FFD9FA8553A7ED4FA135" box="[1299,1356,502,528]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="198">ZISP</collectionCode>
|
||
2424), Giglio island: 1 valve (
|
||
<collectionCode id="ED2CAE72FF89FFD9FE0E504BE9C3A111" box="[408,448,538,564]" pageId="4" pageNumber="198">BD</collectionCode>
|
||
250). Maximum width: -- / 3.8 /
|
||
<quantity id="4CC59B52FF89FFD9FCA2504BEB8EA110" box="[820,909,538,565]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="198" unit="mm" value="2.7">2.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327653CFF89FFD9FF51506FEC56A1E1" pageId="4" pageNumber="198" type="diagnosis">
|
||
<paragraph id="8B8236B7FF89FFD9FF51506FEC56A1E1" blockId="4.[151,1437,214,1176]" pageId="4" pageNumber="198">
|
||
<emphasis id="B949EAA5FF89FFD9FF51506FE941A17D" bold="true" box="[199,322,574,600]" pageId="4" pageNumber="198">Diagnosis.</emphasis>
|
||
Intermediate valves trapezoidal, semicarinate, elevated, apex well developed, tail valve circular, mucro subcentral. Tegmentum slightly rough, sculptured with granules irregularly arranged, oval (up to 127 μm) in PA, AMA, smaller and more irregular in JA, each granule with a central aesthete and 12–15 others, all more or less of same size. Articulamentum strongly developed, apophyses wide, oriented laterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327653CFF89FFD9FF51509FE9DDA7BD" pageId="4" pageNumber="198" type="description">
|
||
<paragraph id="8B8236B7FF89FFD9FF51509FEA7DA0B9" blockId="4.[151,1437,214,1176]" pageId="4" pageNumber="198">
|
||
<emphasis id="B949EAA5FF89FFD9FF51509FE95AA1CD" bold="true" box="[199,345,718,744]" pageId="4" pageNumber="198">Description.</emphasis>
|
||
Head valve unknown. Intermediate valves trapezoidal (W/L = 1.50–1.60), elevated (H/W = 0.39–0.44), semicarinate in anterior profile, anterior margin greatly splayed laterally, almost straight to slightly concave in jugal area, posterior margin with protruding, well developed apex in middle, lateral areas not raised, only marked by a difference in sculpture. Tail valve circular (W/L = 1.16), anterior margin almost straight between apophyses, mucro subcentral, antemucronal slope convex, postmucronal slope decidedly concave directly behind mucro, straightening near posterior margin.
|
||
</paragraph>
|
||
<paragraph id="8B8236B7FF89FFD9FF5151F6EC30A775" blockId="4.[151,1437,214,1176]" pageId="4" pageNumber="198">Tegmentum surface slightly rough. LA and PMA uniformly sculptured with roundish randomly arranged granules (diameter up to 100 μm). PA and AMA sculptured with oval granules (up to 127 μm long) arranged without pattern, JA sculptured with smaller granules, more irregular and closer to each other, without interspaces. Each granule contains one aesthete in central position surrounded by up to 15 others in CA, AMA, a little less (up to 12) in LA, PMA; all aesthetes more or less of same size, no aesthetes among granules.</paragraph>
|
||
<paragraph id="8B8236B7FF89FFD9FF51560BE9DDA7BD" blockId="4.[151,1437,214,1176]" pageId="4" pageNumber="198">Articulamentum strongly developed, apophyses wide, oriented laterally, connected to primordium of insertion plates in intermediate valves.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF42663FFF89FFD9FF01554CE8C3A4CC" ID-DOI="http://doi.org/10.5281/zenodo.13618120" ID-Zenodo-Dep="13618120" httpUri="https://zenodo.org/record/13618120/files/figure.png" pageId="4" pageNumber="198" startId="4.[151,250,1821,1845]" targetBox="[151,1436,1200,1797]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B8236B7FF89FFD9FF01554CE8C3A4CC" blockId="4.[151,1436,1821,2025]" pageId="4" pageNumber="198">
|
||
<emphasis id="B949EAA5FF89FFD9FF01554CE916A410" bold="true" box="[151,277,1821,1845]" pageId="4" pageNumber="198">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C3D4D34FF89FFD9FE8B554CE9DBA410" authorityName="Dell’Angelo & Sosso & Taviani" authorityYear="2024" box="[285,472,1821,1845]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="198" phylum="Mollusca" rank="species" species="schwabei" status="sp. nov.">
|
||
<emphasis id="B949EAA5FF89FFD9FE8B554CE9DBA410" box="[285,472,1821,1845]" italics="true" pageId="4" pageNumber="198">Hanleya schwabei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949EAA5FF89FFD9FE77554CEA30A410" bold="true" box="[481,563,1821,1845]" pageId="4" pageNumber="198">
|
||
<taxonomicNameLabel id="A27A57DEFF89FFD9FE77554CEA30A410" box="[481,563,1821,1845]" pageId="4" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Tyrrhenian Sea: A. offshore Capraia island-Capo Corso, depth ca. 350 m, (possibly last glacial Pleistocene), Paratype 1 (MNHN-IM-2022-2410), intermediate valve, width 3.4 mm, dorsal view. B–E. Off Scilla, Holotype (MZUB 60338), intermediate valve, width 3.8 mm, dorsal view (B), close-up of surface ornamentation of pleural (C) and lateral (D) areas, and frontal view (E). F–H. Off Scilla, Paratype 4 (ZISP 2423), tail valve, width 2.7 mm, dorsal view (F), close-up of surface ornamentation of antemucronal area (G) and lateral view (H). Scale bars = 400 μm (G), 100 μm (D), 40 μm (C).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C327653CFF88FFD8FF5152C7EBFCA2AC" pageId="5" pageNumber="199" type="discussion">
|
||
<paragraph id="8B8236B7FF88FFD8FF5152C7E9B2A239" blockId="5.[151,1437,150,465]" pageId="5" pageNumber="199">
|
||
<emphasis id="B949EAA5FF88FFD8FF5152C7E938A395" bold="true" box="[199,315,150,176]" pageId="5" pageNumber="199">Remarks.</emphasis>
|
||
<taxonomicName id="4C3D4D34FF88FFD8FED352C7EA16A395" authorityName="Dell’Angelo & Sosso & Taviani" authorityYear="2024" box="[325,533,150,176]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="199" phylum="Mollusca" rank="species" species="schwabei" status="sp. nov.">
|
||
<emphasis id="B949EAA5FF88FFD8FED352C7EA16A395" box="[325,533,150,176]" italics="true" pageId="5" pageNumber="199">Hanleya schwabei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949EAA5FF88FFD8FD8952C7EA78A395" bold="true" box="[543,635,150,176]" pageId="5" pageNumber="199">
|
||
<taxonomicNameLabel id="A27A57DEFF88FFD8FD8952C7EA78A395" box="[543,635,150,176]" pageId="5" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is recorded from various Mediterranean localities, previously labeled as
|
||
<taxonomicName id="4C3D4D34FF88FFD8FF2E52EAE9A1A3F1" authorityName="Sirenko" authorityYear="2014" box="[184,418,186,212]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" isUncertain="true" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="199" phylum="Mollusca" rank="species" species="mediterranea">
|
||
<emphasis id="B949EAA5FF88FFD8FF2E52EAE8D0A3F1" box="[184,211,187,212]" italics="true" pageId="5" pageNumber="199">H.</emphasis>
|
||
aff.
|
||
<emphasis id="B949EAA5FF88FFD8FE9D52EBE9A1A3F1" box="[267,418,186,212]" italics="true" pageId="5" pageNumber="199">mediterranea</emphasis>
|
||
</taxonomicName>
|
||
in collections. Some such valves were illustrated by
|
||
<bibRefCitation id="EFAC4B46FF88FFD8FB9152EBED26A3F1" author="Dell'Angelo, B. & Renda, W. & Sirenko, B. I. & Sosso, M. & Giacobbe, S." box="[1031,1317,186,213]" pageId="5" pageNumber="199" pagination="124 - 135" refId="ref7731" refString="Dell'Angelo, B., Renda, W., Sirenko, B. I., Sosso, M. & Giacobbe, S. (2021) The Mediterranean distribution of Hanleya hanleyi (Bean in Thorpe, 1844) and H. mediterranea Sirenko, 2014 (Polyplacophora). Bollettino Malacologico, 57 (1), 124 - 135." type="journal article" year="2021">
|
||
Dell’Angelo
|
||
<emphasis id="B949EAA5FF88FFD8FB0A52EAECD3A3F1" box="[1180,1232,186,212]" italics="true" pageId="5" pageNumber="199">et al</emphasis>
|
||
. (2021
|
||
</bibRefCitation>
|
||
: figs 5.A– F). A unique intermediate valve offshore Capraia/Capo Corso -350/
|
||
<quantity id="4CC59B52FF88FFD8FC0B528FEBE4A3DC" box="[925,999,222,249]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="199" unit="m" value="500.0">500 m</quantity>
|
||
(
|
||
<figureCitation id="13062A32FF88FFD8FC6F528FEC50A3DD" box="[1017,1107,222,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1821,1845]" captionTargetBox="[151,1436,1200,1797]" captionTargetId="figure-399@4.[151,1436,1200,1797]" captionTargetPageId="4" captionText="FIGURE 3. Hanleya schwabei sp. nov., Tyrrhenian Sea: A. offshore Capraia island-Capo Corso, depth ca. 350 m, (possibly last glacial Pleistocene), Paratype 1 (MNHN-IM-2022-2410), intermediate valve, width 3.4 mm, dorsal view. B–E. Off Scilla, Holotype (MZUB 60338), intermediate valve, width 3.8 mm, dorsal view (B), close-up of surface ornamentation of pleural (C) and lateral (D) areas, and frontal view (E). F–H. Off Scilla, Paratype 4 (ZISP 2423), tail valve, width 2.7 mm, dorsal view (F), close-up of surface ornamentation of antemucronal area (G) and lateral view (H). Scale bars = 400 μm (G), 100 μm (D), 40 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.13618120" httpUri="https://zenodo.org/record/13618120/files/figure.png" pageId="5" pageNumber="199">Fig. 3A</figureCitation>
|
||
) could belong to last glacial Pleistocene assemblages.
|
||
</paragraph>
|
||
<paragraph id="8B8236B7FF88FFD8FF515377EBFCA2AC" blockId="5.[151,1437,150,465]" pageId="5" pageNumber="199">
|
||
<emphasis id="B949EAA5FF88FFD8FF515377E96EA265" bold="true" box="[199,365,294,320]" pageId="5" pageNumber="199">Comparisons.</emphasis>
|
||
The most similar species is
|
||
<taxonomicName id="4C3D4D34FF88FFD8FD285376EB78A265" authorityName="Sirenko" authorityYear="2014" box="[702,891,294,320]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="199" phylum="Mollusca" rank="species" species="mediterranea">
|
||
<emphasis id="B949EAA5FF88FFD8FD285376EB78A265" box="[702,891,294,320]" italics="true" pageId="5" pageNumber="199">H. mediterranea</emphasis>
|
||
</taxonomicName>
|
||
, that differs from
|
||
<taxonomicName id="4C3D4D34FF88FFD8FBC25376ECE1A265" authorityName="Dell’Angelo & Sosso & Taviani" authorityYear="2024" box="[1108,1250,294,320]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="199" phylum="Mollusca" rank="species" species="schwabei" status="sp. nov.">
|
||
<emphasis id="B949EAA5FF88FFD8FBC25376ECE1A265" box="[1108,1250,294,320]" italics="true" pageId="5" pageNumber="199">H. schwabei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949EAA5FF88FFD8FB7A5377ED4AA265" bold="true" box="[1260,1353,294,320]" pageId="5" pageNumber="199">
|
||
<taxonomicNameLabel id="A27A57DEFF88FFD8FB7A5377ED4AA265" box="[1260,1353,294,320]" pageId="5" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the characteristic large granules, not present in
|
||
<taxonomicName id="4C3D4D34FF88FFD8FDEA531AEB06A241" authorityName="Dell’Angelo & Sosso & Taviani" authorityYear="2024" box="[636,773,330,356]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="199" phylum="Mollusca" rank="species" species="schwabei" status="sp. nov.">
|
||
<emphasis id="B949EAA5FF88FFD8FDEA531AEB06A241" box="[636,773,330,356]" italics="true" pageId="5" pageNumber="199">H. schwabei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949EAA5FF88FFD8FC9A531BEB66A241" bold="true" box="[780,869,330,356]" pageId="5" pageNumber="199">
|
||
<taxonomicNameLabel id="A27A57DEFF88FFD8FC9A531BEB66A241" box="[780,869,330,356]" pageId="5" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Moreover, the shape of the tail valves is different, almost circular in
|
||
<taxonomicName id="4C3D4D34FF88FFD8FEF4533EE9EFA2AD" authorityName="Dell’Angelo & Sosso & Taviani" authorityYear="2024" box="[354,492,366,392]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="199" phylum="Mollusca" rank="species" species="schwabei" status="sp. nov.">
|
||
<emphasis id="B949EAA5FF88FFD8FEF4533EE9EFA2AD" box="[354,492,366,392]" italics="true" pageId="5" pageNumber="199">H. schwabei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949EAA5FF88FFD8FE62533FEA4EA2AD" bold="true" box="[500,589,366,392]" pageId="5" pageNumber="199">
|
||
<taxonomicNameLabel id="A27A57DEFF88FFD8FE62533FEA4EA2AD" box="[500,589,366,392]" pageId="5" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, more or less oval in
|
||
<taxonomicName id="4C3D4D34FF88FFD8FCA9533EEBFBA2AD" authorityName="Sirenko" authorityYear="2014" box="[831,1016,366,392]" class="Polyplacophora" family="Hanleyidae" genus="Hanleya" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="199" phylum="Mollusca" rank="species" species="mediterranea">
|
||
<emphasis id="B949EAA5FF88FFD8FCA9533EEBFBA2AD" box="[831,1016,366,392]" italics="true" pageId="5" pageNumber="199">H. mediterranea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327653CFF88FFD8FF5153C3EB69A2F4" pageId="5" pageNumber="199" type="distribution">
|
||
<paragraph id="8B8236B7FF88FFD8FF5153C3EB69A2F4" blockId="5.[151,1437,150,465]" pageId="5" pageNumber="199">
|
||
<emphasis id="B949EAA5FF88FFD8FF5153C3E962A289" bold="true" box="[199,353,402,428]" pageId="5" pageNumber="199">Distribution.</emphasis>
|
||
Recent:
|
||
<collectingCountry id="F32A7627FF88FFD8FE5753C3E9F7A289" box="[449,500,402,428]" name="Italy" pageId="5" pageNumber="199">Italy</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32A7627FF88FFD8FDB053C3EA79A289" box="[550,634,402,428]" name="Algeria" pageId="5" pageNumber="199">Algeria</collectingCountry>
|
||
(this study); possibly from last glacial Pleistocene assemblages, Western Mediterranean, offshore Capraia island/Capo Corso -350/
|
||
<specimenCount id="9D3BFD3EFF88FFD8FC8D53E7EB69A2F4" box="[795,874,438,465]" count="500" pageId="5" pageNumber="199" type="generic">
|
||
<quantity id="4CC59B52FF88FFD8FC8D53E7EB65A2F4" box="[795,870,438,465]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="199" unit="m" value="500.0">500 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |