treatments-xml/data/03/F6/87/03F687FE435ABC760CD7FBC0FB807669.xml

<document id="53EA6362CD7B265662979CBF6033F1D8" ID-CLB-Dataset="2609" ID-DOI="10.11646/zootaxa.5091.2.3" ID-GBIF-Dataset="36ace73e-babc-4c20-a78f-fa772ce5f539" ID-ISSN="1175-5326" ID-Zenodo-Dep="5843837" ID-ZooBank="C8081B60-C872-4A19-9291-22A42DC8B763" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1642061981252" checkinUser="plazi" docAuthor="Mai, Zuqi, Hu, Jian, Minoshima, Yûsuke N., Jia, Fenglong &amp; Fikáček, Martin" docDate="2022" docId="03F687FE435ABC760CD7FBC0FB807669" docLanguage="en" docName="zootaxa.5091.2.3.pdf" docOrigin="Zootaxa 5091 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dactylosternum pseudolatum Mai et Jia 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="292" masterDocId="FFCFFF86434EBC610C40FFFEFF88770C" masterDocTitle="Review of Dactylosternum Wollaston, 1854 from China and Japan (Coleoptera, Hydrophilidae, Sphaeridiinae)" masterLastPageNumber="300" masterPageNumber="269" pageNumber="289" updateTime="1699302224171" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="92E680BD704DADB0B88AA93B477E0C50">Review of Dactylosternum Wollaston, 1854 from China and Japan (Coleoptera, Hydrophilidae, Sphaeridiinae)</mods:title>
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<mods:namePart id="BF81F5D47DF5FA435365969F632F6F66">Mai, Zuqi</mods:namePart>
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<mods:affiliation id="89428F2DAC9F2DDD4C3339B25D61220E">School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, Guangdong, China. &amp; maizq 5 @ mail 2. sysu. edu. cn; http: // orcid. org / 0000 - 0003 - 3124 - 2021</mods:affiliation>
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<mods:namePart id="7652A0924C3E233C10AEACA096C1E4A0">Hu, Jian</mods:namePart>
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<mods:namePart id="69DBBC409B90999B2084A37D57A2071E">Minoshima, Yûsuke N.</mods:namePart>
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<mods:affiliation id="481FB53CADD301281E59AF04F6C96B4D">Natural History Division, Kitakyushu Museum of Natural History and Human History, 2 - 4 - 1 Higashida, Yahatahigashi-ku, Kitakyushushi, Fukuoka, 805 - 0071 Japan. minoshima @ kmnh. jp; http: // orcid. org / 0000 - 0002 - 2575 - 4082</mods:affiliation>
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<mods:namePart id="B27460BED198AF6D5AFC86FE7D787B5E">Jia, Fenglong</mods:namePart>
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<mods:affiliation id="A91D41FDF8BB16A6A2E86A3AAEA7909B">School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, Guangdong, China. &amp; Institute of Entomology, Life Sciences School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China. lssjfl @ mail. sysu. edu. cn; fenglongjia @ alilyun. com; http: // orcid. org / 0000 - 0003 - 2391 - 5038</mods:affiliation>
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<mods:namePart id="7A606B76BEF5BAC411E8E06D46E91E4F">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="2BD6DBD3ABCD3FDA215D1C66F348FE4F">Department of Biological Sciences, National Sun Yat-sen University, No. 70 Lien-hai Rd., Kaohsiung City 80424. mfikacek @ gmail. com; http: // orcid. org / 0000 - 0002 - 2078 - 6798 &amp; National Museum, Department of Entomology, Cirkusová 1740, CZ- 19300 Praha 9, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 12843 Praha 2, Czech Republic Corresponding author</mods:affiliation>
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<taxonomicName id="4C5F4D6B435ABC750CD7FBC0FD087354" ID-CoL="9GCNV" authority="Mai et Jia" authorityName="Mai et Jia" authorityYear="2022" box="[151,640,1086,1113]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="pseudolatum" status="sp. nov.">
<emphasis id="B92BEAFA435ABC750CD7FBC0FE727355" bold="true" box="[151,506,1086,1113]" italics="true" pageId="20" pageNumber="289">Dactylosternum pseudolatum</emphasis>
Mai et Jia
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(
<figureCitation id="13642A6D435ABC750CDFFB9DFF677371" box="[159,239,1123,1149]" captionStart="FIGURES 5–10" captionStartId="8.[151,264,1933,1957]" captionTargetBox="[260,1303,188,1900]" captionTargetId="figure-30@8.[214,1373,181,1908]" captionTargetPageId="8" captionText="FIGURES 5–10. Habitus of Dactylosternum spp. from China and Japan. 5: D. songxiaobini sp. nov., 6: D. hydrophiloides (MacLeay, 1825), 7: D. pseudolatum sp. nov., 8: D. salvazai Orchymont, 1925, 9: D. latum (Sharp, 1873), 10: D. frater sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5843841" httpUri="https://zenodo.org/record/5843841/files/figure.png" pageId="20" pageNumber="289">Figs. 7</figureCitation>
, 
<figureCitation id="13642A6D435ABC750CBBFB9DFE917371" box="[251,281,1123,1149]" captionStart="FIGURES 17-19" captionStartId="11.[151,264,1231,1255]" captionTargetBox="[151,1435,190,1207]" captionTargetId="figure-19@11.[151,1435,181,1207]" captionTargetPageId="11" captionText="FIGURES 17-19. Morphology of Dactylosternum spp. 17: hind wing of D. nanlingensis sp. nov., 18–19: Anterior margin of clypeus of D. hydrophiloides and D. pseudolatum sp. nov. (18: D. hydrophiloides (arrow indicates the marginal rim on anterior margin of clypeus), 19: D. pseudolatum sp. nov. (without marginal rim on anterior margin of clypeus))" figureDoi="http://doi.org/10.5281/zenodo.5843853" httpUri="https://zenodo.org/record/5843853/files/figure.png" pageId="20" pageNumber="289">19</figureCitation>
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<figureCitation id="13642A6D435ABC750D65FB9DFEE67371" box="[293,366,1123,1149]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="20" pageNumber="289">27–29</figureCitation>
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<figureCitation id="13642A6D435ABC750D39FB9CFE627372" box="[377,490,1122,1150]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="20" pageNumber="289">52, 56, 60</figureCitation>
)
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<emphasis id="B92BEAFA435ABC750CD7FB55FE4073C8" bold="true" box="[151,456,1194,1221]" pageId="20" pageNumber="289">Type material examined.</emphasis>
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male (
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&amp; 
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leg.
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<materialsCitation id="3B373CB5435ABC75087EFB30FD8B723C" ID-GBIF-Occurrence="3436469327" collectingDate="1918-03-28" collectionCode="IRSNB" collectorName="Fujian" country="Laos" determinerName="A. d'Orchymont" location="Laos" pageId="20" pageNumber="289" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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(
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/ Vitalis // male symbol // co- / type // Para- / type // 
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det. / 
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”; 
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, 
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&amp; 
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leg.
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, 
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; 
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(
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, 
<collectingMunicipality id="6B84AC92435ABC750FD3FA7DFBDB7291" box="[915,1107,1411,1437]" pageId="20" pageNumber="289">Dongwu District</collectingMunicipality>
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<collectingDate id="EFA5E9C0435ABC75081FFA7DFB4D7291" box="[1119,1221,1411,1437]" pageId="20" pageNumber="289" value="2011-05-08">8.V.2011</collectingDate>
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, 
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leg.
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; 
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, 
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, 
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leg.
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; 
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, 
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, 
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leg.
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; 
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, 
<collectingMunicipality id="6B84AC92435ABC750FA6FA35FBEC72E9" box="[998,1124,1483,1509]" pageId="20" pageNumber="289">Xiangzhou</collectingMunicipality>
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leg.
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; 
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(
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, 
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<collectingDate id="EFA5E9C0435ABC750EEFFA11FCA47105" box="[687,812,1519,1545]" pageId="20" pageNumber="289" value="2011-04-26">26.IV.2011</collectingDate>
</date>
, 
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leg.
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; 
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,
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(
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, 
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, 
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<collectingDate id="EFA5E9C0435ABC750E0AF9ECFD417121" box="[586,713,1554,1581]" pageId="20" pageNumber="289" value="2015-04-18">18.IV.2015</collectingDate>
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, 
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&amp; 
<collectorName id="26AA533E435ABC750F5BF9EDFCDB7121" box="[795,851,1555,1581]" pageId="20" pageNumber="289">Tang</collectorName>
leg.
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; 
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, 
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<collectingDate id="EFA5E9C0435ABC750CD7F9C9FE9C715D" box="[151,276,1591,1617]" pageId="20" pageNumber="289" value="2016-05-23">23. V.2016</collectingDate>
</date>
, 
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leg.
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; 
<materialsCitation id="3B373CB5435ABC750DF5F9C9FB90715D" ID-GBIF-Occurrence="3436469352" box="[437,1048,1590,1617]" collectingDate="1957-04-22" collectionCode="SYSU" collectorName="Hong" country="Laos" elevation="600" location="Cheli" pageId="20" pageNumber="289" specimenCount="2" typeStatus="paratype">
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(
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), 
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, 
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<elevation id="0072D1DB435ABC750E82F9C9FC85715D" box="[706,781,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="289" unit="m" value="600.0">600 m</elevation>
</quantity>
, 
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<collectingDate id="EFA5E9C0435ABC750F5AF9C8FC11715D" box="[794,921,1590,1617]" pageId="20" pageNumber="289" value="1957-04-22">22.IV.1957</collectingDate>
</date>
, 
<collectorName id="26AA533E435ABC750FE6F9C9FC6C715D" box="[934,996,1591,1617]" pageId="20" pageNumber="289">Hong</collectorName>
leg.
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; 
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<specimenCount id="9D59FD61435ABC750866F9C9FBFD715D" box="[1062,1141,1591,1617]" pageId="20" pageNumber="289" type="generic">2 spec.</specimenCount>
(
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), 
<location id="8E806033435ABC7508A7F9C9FA10715D" LSID="urn:lsid:plazi:treatment:03F687FE435ABC760CD7FBC0FB807669:8E806033435ABC7508A7F9C9FA10715D" box="[1255,1432,1591,1617]" country="Laos" name="Xishuangbanna" pageId="20" pageNumber="289">Xishuangbanna</location>
, 
<quantity id="4CA79B0D435ABC750CD7F9A4FF577179" box="[151,223,1626,1653]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="20" pageNumber="289" unit="m" value="580.0">
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, 
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<collectingDate id="EFA5E9C0435ABC750CAAF9A4FEE07179" box="[234,360,1626,1653]" pageId="20" pageNumber="289" value="1957-04-22">22.IV.1957</collectingDate>
</date>
, 
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leg.
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; 
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(
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), 
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, 
<collectingMunicipality id="6B84AC92435ABC750F55F9A5FC4B7179" box="[789,963,1627,1653]" pageId="20" pageNumber="289">Xinzhai village</collectingMunicipality>
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<geoCoordinate id="EE6B502F435ABC75087FF9A5FB317179" box="[1087,1209,1627,1653]" degrees="101.132" direction="east" orientation="longitude" pageId="20" pageNumber="289" precision="55" value="101.132">101.132°E</geoCoordinate>
, 
<quantity id="4CA79B0D435ABC750883F9A5FA917179" box="[1219,1305,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.4869999999999999" pageId="20" pageNumber="289" unit="m" value="1487.0">
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, 
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<collectingDate id="EFA5E9C0435ABC750964F9A5FA107179" box="[1316,1432,1627,1653]" pageId="20" pageNumber="289" value="2011-05-21">21.V.2011</collectingDate>
</date>
, 
<collectorName id="26AA533E435ABC750CD7F981FF597195" box="[151,209,1663,1689]" pageId="20" pageNumber="289">Song</collectorName>
leg.
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</paragraph>
</subSubSection>
<subSubSection id="C3456563435ABC750C87F95CFDB370D1" pageId="20" pageNumber="289" type="diagnosis">
<paragraph id="8BE036E8435ABC750C87F95CFDB370D1" blockId="20.[151,1437,1194,2013]" pageId="20" pageNumber="289">
<emphasis id="B92BEAFA435ABC750C87F95CFECA71B0" bold="true" box="[199,322,1698,1724]" pageId="20" pageNumber="289">Diagnosis.</emphasis>
Length 
<quantity id="4CA79B0D435ABC750DE3F95CFDA771B1" box="[419,559,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.2" metricValueMin="5.9" pageId="20" pageNumber="289" unit="mm" value="6.55" valueMax="7.2" valueMin="5.9">5.9–7.2 mm</quantity>
. Body oval, anterior half of elytra slightly parallel-sided (fig. 7), moderately convex. Dorsal black, surface with dense and fine punctures. Anterior margin of clypeus without marginal rim (
<figureCitation id="13642A6D435ABC75092FF939FF3E7009" captionStart="FIGURES 17-19" captionStartId="11.[151,264,1231,1255]" captionTargetBox="[151,1435,190,1207]" captionTargetId="figure-19@11.[151,1435,181,1207]" captionTargetPageId="11" captionText="FIGURES 17-19. Morphology of Dactylosternum spp. 17: hind wing of D. nanlingensis sp. nov., 18–19: Anterior margin of clypeus of D. hydrophiloides and D. pseudolatum sp. nov. (18: D. hydrophiloides (arrow indicates the marginal rim on anterior margin of clypeus), 19: D. pseudolatum sp. nov. (without marginal rim on anterior margin of clypeus))" figureDoi="http://doi.org/10.5281/zenodo.5843853" httpUri="https://zenodo.org/record/5843853/files/figure.png" pageId="20" pageNumber="289">Fig. 19</figureCitation>
). Antenna yellowish brown, club compact. Elytron with sutural stria and 10 distinct rows of serial punctures, all nearly reaching elytral base, ground punctures attached with one/two short transverse microsculptures.Anteromedial portion of metaventral process with a short transverse groove (
<figureCitation id="13642A6D435ABC750F16F8CDFC237041" box="[854,939,1842,1869]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="20" pageNumber="289">Fig. 56</figureCitation>
). Posterior margin of ventrite 5 with distinct marginal bead and a stria behind the marginal bead (
<figureCitation id="13642A6D435ABC750EBFF8A9FCDF707E" box="[767,855,1879,1906]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="20" pageNumber="289">Fig. 60</figureCitation>
). Aedeagus (
<figureCitation id="13642A6D435ABC750FADF8A9FBF3707D" box="[1005,1147,1879,1905]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="20" pageNumber="289">Figs. 27–29</figureCitation>
): Phallobase shorter than parameres. Parameres distinctly sinuate on outer margin, and strongly curved subapically. Median lobe widest in the middle, a little shorter than parameres; ventral plate triangular, strongly pointed at apex, gonopore situated at apex of ventral plate (
<figureCitation id="13642A6D435ABC750DCCF83DFDA670D1" box="[396,558,1987,2013]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="20" pageNumber="289">Figs. 27c, 28c</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF206660435BBC740CD7FA29FE5371CB" ID-DOI="http://doi.org/10.5281/zenodo.5843883" ID-Zenodo-Dep="5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="21" pageNumber="290" startId="21.[151,264,1495,1519]" targetBox="[153,1430,183,1463]" targetPageId="21">
<paragraph id="8BE036E8435BBC740CD7FA29FE5371CB" blockId="21.[151,1437,1495,1735]" pageId="21" pageNumber="290">
<emphasis id="B92BEAFA435BBC740CD7FA29FEDF72E3" bold="true" box="[151,343,1495,1519]" pageId="21" pageNumber="290">FIGURES 48–63.</emphasis>
Morphology characters of 
<taxonomicName id="4C5F4D6B435BBC740E33FA29FCC072E3" box="[627,840,1495,1519]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92BEAFA435BBC740E33FA29FC9D72E3" box="[627,789,1495,1519]" italics="true" pageId="21" pageNumber="290">Dactylosternum</emphasis>
spp.
</taxonomicName>
SEM micrographs. 48–50: mediobasal portion of elytron (48: 
<taxonomicName id="4C5F4D6B435BBC740C88FA02FEE3711F" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[200,363,1531,1555]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="songxiaobini" status="sp. nov.">
<emphasis id="B92BEAFA435BBC740C88FA02FEE3711F" box="[200,363,1531,1555]" italics="true" pageId="21" pageNumber="290">D. songxiaobini</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435BBC740D32FA05FE4C711F" bold="true" box="[370,452,1531,1555]" pageId="21" pageNumber="290">
<taxonomicNameLabel id="A2185781435BBC740D32FA05FE4C711F" box="[370,452,1531,1555]" pageId="21" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: 
<taxonomicName id="4C5F4D6B435BBC740CD7F9DEFEBF713B" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[151,311,1567,1591]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B92BEAFA435BBC740CD7F9DEFEBF713B" box="[151,311,1567,1591]" italics="true" pageId="21" pageNumber="290">D. nanlingensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435BBC740D7EF9E1FE18713B" bold="true" box="[318,400,1567,1591]" pageId="21" pageNumber="290">
<taxonomicNameLabel id="A2185781435BBC740D7EF9E1FE18713B" box="[318,400,1567,1591]" pageId="21" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 50: 
<taxonomicName id="4C5F4D6B435BBC740D85F9DEFDBC713B" authorityName="Balfour-Browne" authorityYear="1942" box="[453,564,1567,1591]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="corbetti">
<emphasis id="B92BEAFA435BBC740D85F9DEFDBC713B" box="[453,564,1567,1591]" italics="true" pageId="21" pageNumber="290">D. corbetti</emphasis>
</taxonomicName>
), 51–55: punctures on posterior half of elytron (51: 
<taxonomicName id="4C5F4D6B435BBC74080DF9DEFA8C713B" baseAuthorityName="MacLeay" baseAuthorityYear="1825" box="[1101,1284,1567,1591]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Fungi" order="Coleoptera" pageId="21" pageNumber="290" phylum="Ascomycota" rank="species" species="hydrophiloides">
<emphasis id="B92BEAFA435BBC74080DF9DEFA8C713B" box="[1101,1284,1567,1591]" italics="true" pageId="21" pageNumber="290">D. hydrophiloides</emphasis>
</taxonomicName>
(the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: 
<taxonomicName id="4C5F4D6B435BBC740FF8F9BAFBDF7157" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[952,1111,1603,1627]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="pseudolatum" status="sp. nov.">
<emphasis id="B92BEAFA435BBC740FF8F9BAFBDF7157" box="[952,1111,1603,1627]" italics="true" pageId="21" pageNumber="290">D. pseudolatum</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435BBC74081DF9BDFB257157" bold="true" box="[1117,1197,1603,1627]" pageId="21" pageNumber="290">
<taxonomicNameLabel id="A2185781435BBC74081DF9BDFB257157" box="[1117,1197,1603,1627]" pageId="21" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 53: 
<taxonomicName id="4C5F4D6B435BBC7408A0F9BAFAD97157" authorityName="Orchymont" authorityYear="1925" box="[1248,1361,1603,1627]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA435BBC7408A0F9BAFAD97157" box="[1248,1361,1603,1627]" italics="true" pageId="21" pageNumber="290">D. salvazai</emphasis>
</taxonomicName>
, 54: 
<taxonomicName id="4C5F4D6B435BBC7409C4F9BAFF597173" authorityName="Sharp" authorityYear="1873" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435BBC7409C4F9BAFF597173" italics="true" pageId="21" pageNumber="290">D. latum</emphasis>
</taxonomicName>
, 55: 
<taxonomicName id="4C5F4D6B435BBC740D46F996FEE87173" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[262,352,1639,1663]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="frater" status="sp. nov.">
<emphasis id="B92BEAFA435BBC740D46F996FEE87173" box="[262,352,1639,1663]" italics="true" pageId="21" pageNumber="290">D. frater</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435BBC740D27F999FE317173" bold="true" box="[359,441,1639,1663]" pageId="21" pageNumber="290">
<taxonomicNameLabel id="A2185781435BBC740D27F999FE317173" box="[359,441,1639,1663]" pageId="21" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), 56–59: anteromedial portion of metaventrite (56: 
<taxonomicName id="4C5F4D6B435BBC740F8BF996FBE47173" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[971,1132,1639,1663]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="pseudolatum" status="sp. nov.">
<emphasis id="B92BEAFA435BBC740F8BF996FBE47173" box="[971,1132,1639,1663]" italics="true" pageId="21" pageNumber="290">D. pseudolatum</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435BBC740834F999FB4D7173" bold="true" box="[1140,1221,1639,1663]" pageId="21" pageNumber="290">
<taxonomicNameLabel id="A2185781435BBC740834F999FB4D7173" box="[1140,1221,1639,1663]" pageId="21" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 57: 
<taxonomicName id="4C5F4D6B435BBC7408BBF996FAE67173" authorityName="Orchymont" authorityYear="1925" box="[1275,1390,1639,1663]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA435BBC7408BBF996FAE67173" box="[1275,1390,1639,1663]" italics="true" pageId="21" pageNumber="290">D. salvazai</emphasis>
</taxonomicName>
, 58: 
<taxonomicName id="4C5F4D6B435BBC740CD7F972FF7871AF" authorityName="Sharp" authorityYear="1873" box="[151,240,1675,1699]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435BBC740CD7F972FF7871AF" box="[151,240,1675,1699]" italics="true" pageId="21" pageNumber="290">D. latum</emphasis>
</taxonomicName>
, 59: 
<taxonomicName id="4C5F4D6B435BBC740D64F972FEF571AF" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[292,381,1675,1699]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="frater" status="sp. nov.">
<emphasis id="B92BEAFA435BBC740D64F972FEF571AF" box="[292,381,1675,1699]" italics="true" pageId="21" pageNumber="290">D. frater</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435BBC740DC4F975FE5D71AF" bold="true" box="[388,469,1675,1699]" pageId="21" pageNumber="290">
<taxonomicNameLabel id="A2185781435BBC740DC4F975FE5D71AF" box="[388,469,1675,1699]" pageId="21" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), 60–63: posteromedial portion of ventrite 5 (60: 
<taxonomicName id="4C5F4D6B435BBC740F90F972FBF871AF" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[976,1136,1675,1699]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="pseudolatum" status="sp. nov.">
<emphasis id="B92BEAFA435BBC740F90F972FBF871AF" box="[976,1136,1675,1699]" italics="true" pageId="21" pageNumber="290">D. pseudolatum</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435BBC740837F975FB4071AF" bold="true" box="[1143,1224,1675,1699]" pageId="21" pageNumber="290">
<taxonomicNameLabel id="A2185781435BBC740837F975FB4071AF" box="[1143,1224,1675,1699]" pageId="21" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 61: 
<taxonomicName id="4C5F4D6B435BBC7408BDF972FAE771AF" authorityName="Orchymont" authorityYear="1925" box="[1277,1391,1675,1699]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA435BBC7408BDF972FAE771AF" box="[1277,1391,1675,1699]" italics="true" pageId="21" pageNumber="290">D. salvazai</emphasis>
</taxonomicName>
, 62: 
<taxonomicName id="4C5F4D6B435BBC740CD7F94EFF7871CB" authorityName="Sharp" authorityYear="1873" box="[151,240,1711,1735]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435BBC740CD7F94EFF7871CB" box="[151,240,1711,1735]" italics="true" pageId="21" pageNumber="290">D. latum</emphasis>
</taxonomicName>
, 63: 
<taxonomicName id="4C5F4D6B435BBC740D63F94EFEF471CB" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[291,380,1711,1735]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="290" phylum="Arthropoda" rank="species" species="frater" status="sp. nov.">
<emphasis id="B92BEAFA435BBC740D63F94EFEF471CB" box="[291,380,1711,1735]" italics="true" pageId="21" pageNumber="290">D. frater</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435BBC740DC2F951FE5B71CB" bold="true" box="[386,467,1711,1735]" pageId="21" pageNumber="290">
<taxonomicNameLabel id="A2185781435BBC740DC2F951FE5B71CB" box="[386,467,1711,1735]" pageId="21" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
)
</paragraph>
</caption>
<subSubSection id="C3456563435BBC770C87F90BFAD67371" lastPageId="22" lastPageNumber="291" pageId="21" pageNumber="290" type="description">
<paragraph id="8BE036E8435BBC740C87F90BFAB9701C" blockId="21.[151,1437,1781,2024]" box="[199,1329,1781,1808]" pageId="21" pageNumber="290">
<emphasis id="B92BEAFA435BBC740C87F90BFED17003" bold="true" box="[199,345,1781,1807]" pageId="21" pageNumber="290">Description.</emphasis>
Total length 
<quantity id="4CA79B0D435BBC740DADF90BFDFF701C" box="[493,631,1781,1808]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.2" metricValueMin="5.9" pageId="21" pageNumber="290" unit="mm" value="6.55" valueMax="7.2" valueMin="5.9">5.9–7.2 mm</quantity>
; maximum width 
<quantity id="4CA79B0D435BBC740F01F90BFC43701C" box="[833,971,1781,1808]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.2" metricValueMin="3.6" pageId="21" pageNumber="290" unit="mm" value="3.9" valueMax="4.2" valueMin="3.6">3.6–4.2 mm</quantity>
; body oval, moderately convex.
</paragraph>
<paragraph id="8BE036E8435BBC740C87F8E7FC2C705B" blockId="21.[151,1437,1781,2024]" pageId="21" pageNumber="290">
<emphasis id="B92BEAFA435BBC740C87F8E7FEC3703F" box="[199,331,1817,1844]" italics="true" pageId="21" pageNumber="290">
<emphasis id="B92BEAFA435BBC740C87F8E7FECB7038" bold="true" box="[199,323,1817,1844]" italics="true" pageId="21" pageNumber="290">Coloration</emphasis>
.
</emphasis>
Dorsal black, with lateral margin of elytron reddish brown (
<figureCitation id="13642A6D435BBC740FBDF8E7FBCB7038" box="[1021,1091,1817,1844]" captionStart="FIGURES 5–10" captionStartId="8.[151,264,1933,1957]" captionTargetBox="[260,1303,188,1900]" captionTargetId="figure-30@8.[214,1373,181,1908]" captionTargetPageId="8" captionText="FIGURES 5–10. Habitus of Dactylosternum spp. from China and Japan. 5: D. songxiaobini sp. nov., 6: D. hydrophiloides (MacLeay, 1825), 7: D. pseudolatum sp. nov., 8: D. salvazai Orchymont, 1925, 9: D. latum (Sharp, 1873), 10: D. frater sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5843841" httpUri="https://zenodo.org/record/5843841/files/figure.png" pageId="21" pageNumber="290">Fig. 7</figureCitation>
). Maxillary palps and antenna yellowish brown. Ventral black to reddish brown, legs reddish brown.
</paragraph>
<paragraph id="8BE036E8435BBC770C87F89FFF7C7669" blockId="21.[151,1437,1781,2024]" lastBlockId="22.[151,1437,151,2014]" lastPageId="22" lastPageNumber="291" pageId="21" pageNumber="290">
<emphasis id="B92BEAFA435BBC740C87F89FFE847070" bold="true" box="[199,268,1889,1916]" italics="true" pageId="21" pageNumber="290">Head.</emphasis>
Clypeus weakly convex, surface with dense and fine punctures, anterior portion with denser punctures. Interstices between punctures smooth. Anterior margin of clypeus without marginal rim (
<figureCitation id="13642A6D435BBC7408EEF87BFA8E70AC" box="[1198,1286,1925,1952]" captionStart="FIGURES 17-19" captionStartId="11.[151,264,1231,1255]" captionTargetBox="[151,1435,190,1207]" captionTargetId="figure-19@11.[151,1435,181,1207]" captionTargetPageId="11" captionText="FIGURES 17-19. Morphology of Dactylosternum spp. 17: hind wing of D. nanlingensis sp. nov., 18–19: Anterior margin of clypeus of D. hydrophiloides and D. pseudolatum sp. nov. (18: D. hydrophiloides (arrow indicates the marginal rim on anterior margin of clypeus), 19: D. pseudolatum sp. nov. (without marginal rim on anterior margin of clypeus))" figureDoi="http://doi.org/10.5281/zenodo.5843853" httpUri="https://zenodo.org/record/5843853/files/figure.png" pageId="21" pageNumber="290">Fig. 19</figureCitation>
). Frons with punctures sparser than those on clypeus. Eyes of moderate size, separated by ca. 6× as wide as one eye, distinctly emarginate anteriorly in lateral view, smooth medially. Maxilla with glabrous basistipes. Mentum transverse, glabrous, strongly impressed anteromedially, lateral sides with series of fine setae. Submentum reddish brown, pubescent. Labial palps rather stout, palpomere 1 minute, palpomere 2 strongly widened distally, with dense brush of long yellow setae on its inner face and distal margin, palpomere 3 longest. Antennal scapus longest, rather thick, ca. 7.5× as long as pedicellus, antennomere 3 thin and a little shorter than pedicellus, antennomeres 4 and 5 almost equal in length, slightly widening distally, club densely pubescent, with few long setae, compact, ca. 2.5× as long as wide.
</paragraph>
<paragraph id="8BE036E84358BC770C87FE90FCFC76DD" blockId="22.[151,1437,151,2014]" pageId="22" pageNumber="291">
<emphasis id="B92BEAFA4358BC770C87FE90FECC7685" bold="true" box="[199,324,366,393]" italics="true" pageId="22" pageNumber="291">Prothorax.</emphasis>
Pronotum moderately convex, widest basally, arcuately narrowed in front, with punctures sparser than those on vertex, interstices between punctures smooth; posterior margin slightly arcuate. Prosternum strongly tectiform, with median carina and a distinct tooth anteromedially.
</paragraph>
<paragraph id="8BE036E84358BC770C87FE25FD8575C1" blockId="22.[151,1437,151,2014]" pageId="22" pageNumber="291">
<emphasis id="B92BEAFA4358BC770C87FE25FE5076F9" bold="true" box="[199,472,474,501]" italics="true" pageId="22" pageNumber="291">Meso- and metathorax.</emphasis>
Scutellum distinctly longer than wide, apex rounded, in shape of isosceles triangle, with punctation similar to those on pronotum. Elytron moderately convex, sutural stria deeply impressed in posterior half; each elytron with 10 longitudinal serial punctures, deeply impressed in posterior half, intervals with fine ground punctures, each puncture with one/two short transverse lines especially in posterior half (
<figureCitation id="13642A6D4358BC7708AFFDB9FACE756D" box="[1263,1350,582,609]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 52</figureCitation>
); series 3+4 and 6+7 linked together subposteriorly, others serial punctures reaching end of elytron; lateral four series with punctures larger and deeper than those in the inner six series. Anteromedial portion of metaventral process with a short transverse groove (
<figureCitation id="13642A6D4358BC770DECFD4DFD8875C1" box="[428,512,690,717]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 56</figureCitation>
).
</paragraph>
<paragraph id="8BE036E84358BC770C87FD29FF6D7451" blockId="22.[151,1437,151,2014]" pageId="22" pageNumber="291">
<emphasis id="B92BEAFA4358BC770C87FD29FE8D75FD" bold="true" box="[199,261,727,753]" italics="true" pageId="22" pageNumber="291">Legs.</emphasis>
Reddish brown, rather short. Procoxae globular, pubescent; profemora without pubescence ventrally; protarsomeres pubescent ventrally. Ventral surface of meso- and metafemora without pubescence, with sparse punctures and microsculptures; meso- and metatarsomeres with sparse pubescence ventrally, almost equal in length.
</paragraph>
<paragraph id="8BE036E84358BC770C87FC98FC2F74C6" blockId="22.[151,1437,151,2014]" pageId="22" pageNumber="291">
<emphasis id="B92BEAFA4358BC770C87FC98FEB4748D" bold="true" box="[199,316,870,897]" italics="true" pageId="22" pageNumber="291">Abdomen.</emphasis>
Pubescent, ventrite 1 with complete median carina, other ventrites uniformly convex, with complete marginal bead. Posteromedial portion of ventrite 5 without pubescence, with fine punctures, posterior margin of ventrite 5 rounded, with a stria in front of the marginal bead (
<figureCitation id="13642A6D4358BC770F06FC51FC1274C6" box="[838,922,943,970]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 60</figureCitation>
).
</paragraph>
<paragraph id="8BE036E84358BC770C87FC2CFAD67371" blockId="22.[151,1437,151,2014]" pageId="22" pageNumber="291">
<emphasis id="B92BEAFA4358BC770C87FC2CFEE774E1" bold="true" box="[199,367,978,1005]" italics="true" pageId="22" pageNumber="291">Male genitalia</emphasis>
(
<figureCitation id="13642A6D4358BC770D3FFC2DFD8274E1" box="[383,522,979,1005]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="22" pageNumber="291">Figs. 27–29</figureCitation>
). Aedeagus about 
<quantity id="4CA79B0D4358BC770E98FC2DFCA774E1" box="[728,815,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="291" unit="mm" value="1.1">1.1 mm</quantity>
long, ca. 2.2× as long as wide. Phallobase shorter than parameres, with short manubrium and rounded posteriorly. Parameres strongly curved inwards apically, pointed at apex, sinuate on outer margin. Median lobe ca. 0.94× as long as parameres, widest in the middle, gradually narrowing towards apex; ventral plate triangular, widest basally, tapering towards apex, strongly pointed at apex (
<figureCitation id="13642A6D4358BC770CDFFB9DFEC97371" box="[159,321,1123,1149]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="22" pageNumber="291">Figs. 27c, 28c</figureCitation>
). Gonopore triangular, situated at apex of ventral plate, reaching ca. distal 0.8 of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C34565634358BC770C87FB79FE9873E5" pageId="22" pageNumber="291" type="etymology">
<paragraph id="8BE036E84358BC770C87FB79FE9873E5" blockId="22.[151,1437,151,2014]" pageId="22" pageNumber="291">
<emphasis id="B92BEAFA4358BC770C87FB79FEC673AD" bold="true" box="[199,334,1159,1185]" pageId="22" pageNumber="291">Etymology.</emphasis>
This species name is a combination of the Latin 
<emphasis id="B92BEAFA4358BC770FFCFB79FB8373AD" box="[956,1035,1159,1185]" italics="true" pageId="22" pageNumber="291">pseudo</emphasis>
-, false, and the species name of 
<taxonomicName id="4C5F4D6B4358BC770CD7FB55FE1C73C9" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[151,404,1195,1221]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA4358BC770CD7FB55FE1C73C9" box="[151,404,1195,1221]" italics="true" pageId="22" pageNumber="291">Dactylosternum latum</emphasis>
</taxonomicName>
. The name refers to the fact that this species is quite similar to 
<taxonomicName id="4C5F4D6B4358BC7708C4FB55FB6173C9" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[1156,1257,1195,1221]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA4358BC7708C4FB55FB6173C9" box="[1156,1257,1195,1221]" italics="true" pageId="22" pageNumber="291">D. latum</emphasis>
</taxonomicName>
in morphology characters.
</paragraph>
</subSubSection>
<subSubSection id="C34565634358BC770C87FB0CFADC7201" box="[199,1364,1266,1293]" pageId="22" pageNumber="291" type="biology_ecology">
<paragraph id="8BE036E84358BC770C87FB0CFADC7201" blockId="22.[151,1437,151,2014]" box="[199,1364,1266,1293]" pageId="22" pageNumber="291">
<emphasis id="B92BEAFA4358BC770C87FB0CFEAF7200" bold="true" box="[199,295,1266,1292]" pageId="22" pageNumber="291">Biology.</emphasis>
Living in moist plant material, such as decaying pseudostem of banana trunks and in leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="C34565634358BC760C87FAE9FEF3764C" lastPageId="23" lastPageNumber="292" pageId="22" pageNumber="291" type="discussion">
<paragraph id="8BE036E84358BC770C87FAE9FD6E70D2" blockId="22.[151,1437,151,2014]" pageId="22" pageNumber="291">
<emphasis id="B92BEAFA4358BC770C87FAE9FEB3723D" bold="true" box="[199,315,1303,1329]" pageId="22" pageNumber="291">Remarks.</emphasis>
We examined 
<specimenCount id="9D59FD614358BC770D9BFAE9FDF7723D" box="[475,639,1303,1329]" pageId="22" pageNumber="291" type="generic">two specimens</specimenCount>
of the type series of 
<taxonomicName id="4C5F4D6B4358BC770F1DFAE9FBE9723D" authorityName="d' Orchymont" authorityYear="1925" box="[861,1121,1303,1329]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA4358BC770F1DFAE9FBE9723D" box="[861,1121,1303,1329]" italics="true" pageId="22" pageNumber="291">Dactylosternum vitalisi</emphasis>
</taxonomicName>
(the 
<typeStatus id="54E4884A4358BC7708D6FAE9FB70723D" box="[1174,1272,1303,1329]" pageId="22" pageNumber="291" type="holotype">holotype</typeStatus>
and a 
<typeStatus id="54E4884A4358BC77097CFAE9FA14723D" box="[1340,1436,1303,1329]" pageId="22" pageNumber="291" type="paratype">paratype</typeStatus>
from 
<collectingCountry id="F34876784358BC770C97FAC5FE997259" box="[215,273,1339,1365]" name="Laos" pageId="22" pageNumber="291">Laos</collectingCountry>
), in which we found several morphological differences. The 
<typeStatus id="54E4884A4358BC770F8BFAC5FBA57259" box="[971,1069,1339,1365]" pageId="22" pageNumber="291" type="holotype">holotype</typeStatus>
differs from the 
<typeStatus id="54E4884A4358BC7708B2FAC5FADA7259" box="[1266,1362,1339,1365]" pageId="22" pageNumber="291" type="paratype">paratype</typeStatus>
in the morphology of the anterior part of the metaventral elevation, the mesoventral elevation, especially in the shapes of the aedeagus (compare 
<figureCitation id="13642A6D4358BC770DE1FA7DFE7D7291" box="[417,501,1411,1437]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="22" pageNumber="291">Figs 29</figureCitation>
and 
<figureCitation id="13642A6D4358BC770E6DFA7DFDC57292" box="[557,589,1411,1438]" captionStart="FIGURES 33–38" captionStartId="18.[151,264,1947,1971]" captionTargetBox="[225,1404,347,1884]" captionTargetId="figure-85@18.[151,1436,309,1924]" captionTargetPageId="18" captionText="FIGURES 33–38. Aedeagus of D. latum (dorsal view). 33: from Hunan, Shimen County, 34: syntype, 35: from Guangxi, 36: holotype of D. vitalisi, 37: from Hunan, Shimen County, 38: from Yunnan" figureDoi="http://doi.org/10.5281/zenodo.5843873" httpUri="https://zenodo.org/record/5843873/files/figure.png" pageId="22" pageNumber="291">36</figureCitation>
). This indicates that the 
<typeStatus id="54E4884A4358BC770F22FA7DFC4C7291" box="[866,964,1411,1437]" pageId="22" pageNumber="291" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54E4884A4358BC770FBCFA7DFBD47291" box="[1020,1116,1411,1437]" pageId="22" pageNumber="291" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C5F4D6B4358BC7708C3FA7DFB7A7291" authorityName="d' Orchymont" authorityYear="1925" box="[1155,1266,1411,1437]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="vitalisi">D. vitalisi</taxonomicName>
each represent a different species. The 
<typeStatus id="54E4884A4358BC770DEDFA59FD8572CD" box="[429,525,1447,1473]" pageId="22" pageNumber="291" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C5F4D6B4358BC770E78FA59FD2072CD" authorityName="d' Orchymont" authorityYear="1925" box="[568,680,1447,1473]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="vitalisi">D. vitalisi</taxonomicName>
is possibly represent an undescribed species. This new species is similar to 
<taxonomicName id="4C5F4D6B4358BC770D4FFA35FD9572E9" authority="Orchymont" authorityName="Orchymont" authorityYear="1925" box="[271,541,1483,1509]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA4358BC770D4FFA35FE1972E9" box="[271,401,1483,1509]" italics="true" pageId="22" pageNumber="291">D. salvazai</emphasis>
Orchymont
</taxonomicName>
, 
<taxonomicName id="4C5F4D6B4358BC770E6CFA35FCB072E9" authority="(Sharp, 1873)" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[556,824,1483,1509]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA4358BC770E6CFA35FD1972E9" box="[556,657,1483,1509]" italics="true" pageId="22" pageNumber="291">D. latum</emphasis>
(
<bibRefCitation id="EFCE4B194358BC770EE3FA35FCB872E9" author="Sharp, D." box="[675,816,1483,1509]" pageId="22" pageNumber="291" pagination="64 - 65" refId="ref23391" refString="Sharp, D. (1873) The water beetles of Japan. Transactions of the Entomological Society of London, 1874, 64 - 65." type="journal article" year="1873">Sharp, 1873</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C5F4D6B4358BC770F35FA35FC5372E9" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[885,987,1483,1509]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="frater" status="sp. nov.">
<emphasis id="B92BEAFA4358BC770F35FA35FC5372E9" box="[885,987,1483,1509]" italics="true" pageId="22" pageNumber="291">D. frater</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA4358BC770FA5FA35FBCA72E9" bold="true" box="[997,1090,1483,1509]" pageId="22" pageNumber="291">
<taxonomicNameLabel id="A21857814358BC770FA5FA35FBCA72E9" box="[997,1090,1483,1509]" pageId="22" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in both habitus and aedeagus morphology. It can be distinguished from the latter three species by dorsal surface with rather densely arranged fine punctures, less shiny (with sparser punctures on the dorsal surface and shiny in the latter three species). It can be distinguished from 
<taxonomicName id="4C5F4D6B4358BC770DD8F9C9FD90715D" authorityName="Orchymont" authorityYear="1925" box="[408,536,1591,1617]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA4358BC770DD8F9C9FD90715D" box="[408,536,1591,1617]" italics="true" pageId="22" pageNumber="291">D. salvazai</emphasis>
</taxonomicName>
by anteromedial portion of metaventral process with a short transverse groove (
<figureCitation id="13642A6D4358BC770CE0F9A5FF7A7179" box="[160,242,1626,1653]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 56</figureCitation>
) (anteromedial portion of metaventrite, behind metaventral process with a Λ-shape divergent ridge in 
<taxonomicName id="4C5F4D6B4358BC7709C1F9A5FF7B7195" authorityName="Orchymont" authorityYear="1925" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA4358BC7709C1F9A5FF7B7195" italics="true" pageId="22" pageNumber="291">D. salvazai</emphasis>
</taxonomicName>
, 
<figureCitation id="13642A6D4358BC770D42F981FEDE7195" box="[258,342,1662,1689]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 57</figureCitation>
), each elytral ground puncture with one/two short transverse line especially in the posterior half of elytra (
<figureCitation id="13642A6D4358BC770D46F95DFED071B1" box="[262,344,1698,1725]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 52</figureCitation>
) (each puncture with a K/H-shape microsculpture in 
<taxonomicName id="4C5F4D6B4358BC770FF0F95DFBA771B1" authorityName="Orchymont" authorityYear="1925" box="[944,1071,1699,1725]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA4358BC770FF0F95DFBA771B1" box="[944,1071,1699,1725]" italics="true" pageId="22" pageNumber="291">D. salvazai</emphasis>
</taxonomicName>
, 
<figureCitation id="13642A6D4358BC77087DF95DFB1871B1" box="[1085,1168,1698,1725]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 53</figureCitation>
), posteromedial portion of ventrite 5 with fine punctures (
<figureCitation id="13642A6D4358BC770E55F939FDE071EE" box="[533,616,1735,1762]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 60</figureCitation>
) (with mesh-like microsculpture instead of puncture in 
<taxonomicName id="4C5F4D6B4358BC7708A2F939FAE971ED" authorityName="Orchymont" authorityYear="1925" box="[1250,1377,1735,1761]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA4358BC7708A2F939FAE971ED" box="[1250,1377,1735,1761]" italics="true" pageId="22" pageNumber="291">D. salvazai</emphasis>
</taxonomicName>
, 
<figureCitation id="13642A6D4358BC770930F939FF3D700A" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 61</figureCitation>
), parameres distinctly sinuate on outer margin and more curved on lateral margin subapically (
<figureCitation id="13642A6D4358BC77089AF915FAE87009" box="[1242,1376,1771,1797]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="22" pageNumber="291">Figs. 27–29</figureCitation>
) (not sinuate and less curved on lateral margin subapically in 
<taxonomicName id="4C5F4D6B4358BC770F59F8F1FC117025" authorityName="Orchymont" authorityYear="1925" box="[793,921,1807,1833]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA4358BC770F59F8F1FC117025" box="[793,921,1807,1833]" italics="true" pageId="22" pageNumber="291">D. salvazai</emphasis>
</taxonomicName>
, 
<figureCitation id="13642A6D4358BC770FE9F8F1FBAF7025" box="[937,1063,1806,1833]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="22" pageNumber="291">Figs. 25a,b</figureCitation>
) and ventral plate of the median lobe strongly pointed apically (
<figureCitation id="13642A6D4358BC770DB9F8CDFD1D7041" box="[505,661,1843,1869]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="22" pageNumber="291">Figs 27c, 28c</figureCitation>
) (rounded at apex in 
<taxonomicName id="4C5F4D6B4358BC770FCBF8CDFB837041" authorityName="Orchymont" authorityYear="1925" box="[907,1035,1843,1869]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B92BEAFA4358BC770FCBF8CDFB837041" box="[907,1035,1843,1869]" italics="true" pageId="22" pageNumber="291">D. salvazai</emphasis>
</taxonomicName>
, 
<figureCitation id="13642A6D4358BC77085AF8CDFBF37041" box="[1050,1147,1842,1869]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="22" pageNumber="291">Fig. 25c</figureCitation>
). The new species differs from 
<taxonomicName id="4C5F4D6B4358BC770C94F8A9FEBD707D" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[212,309,1879,1905]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA4358BC770C94F8A9FEBD707D" box="[212,309,1879,1905]" italics="true" pageId="22" pageNumber="291">D. latum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D6B4358BC770D2AF8A9FE44707D" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[362,460,1879,1905]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BEAFA4358BC770D2AF8A9FE44707D" box="[362,460,1879,1905]" italics="true" pageId="22" pageNumber="291">D. frater</emphasis>
</taxonomicName>
by anteromedial portion of metaventral process with a short transverse groove (
<figureCitation id="13642A6D4358BC770903F8A9FA1C707D" box="[1347,1428,1878,1905]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 56</figureCitation>
) (with a strongly impressed pit in 
<taxonomicName id="4C5F4D6B4358BC770E45F885FDEF7099" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[517,615,1915,1941]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA4358BC770E45F885FDEF7099" box="[517,615,1915,1941]" italics="true" pageId="22" pageNumber="291">D. latum</emphasis>
</taxonomicName>
(
<figureCitation id="13642A6D4358BC770E35F885FD4E7099" box="[629,710,1914,1941]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 58</figureCitation>
) or slightly impressed pit in 
<taxonomicName id="4C5F4D6B4358BC770844F885FBEE7099" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[1028,1126,1915,1941]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BEAFA4358BC770844F885FBEE7099" box="[1028,1126,1915,1941]" italics="true" pageId="22" pageNumber="291">D. frater</emphasis>
</taxonomicName>
(
<figureCitation id="13642A6D4358BC770834F885FB4E7099" box="[1140,1222,1914,1941]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 59</figureCitation>
)), posterior margin of ventrite 5 with distinct marginal bead and a stria behind the marginal bead (
<figureCitation id="13642A6D4358BC770FBCF861FBC570B6" box="[1020,1101,1951,1978]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Fig. 60</figureCitation>
) (with fine marginal bead and without stria in 
<taxonomicName id="4C5F4D6B4358BC770D08F83DFE2270D1" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[328,426,1987,2013]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA4358BC770D08F83DFE2270D1" box="[328,426,1987,2013]" italics="true" pageId="22" pageNumber="291">D. latum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D6B4358BC770DA0F83DFDCA70D1" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[480,578,1987,2013]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="291" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="B92BEAFA4358BC770DA0F83DFDCA70D1" box="[480,578,1987,2013]" italics="true" pageId="22" pageNumber="291">D. frater</emphasis>
</taxonomicName>
; 
<figureCitation id="13642A6D4358BC770E11F83DFD5270D2" box="[593,730,1987,2014]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="22" pageNumber="291">Figs. 62–63</figureCitation>
).
</paragraph>
<paragraph id="8BE036E84359BC760C87FF68FEF3764C" blockId="23.[151,1437,150,357]" pageId="23" pageNumber="292">
<emphasis id="B92BEAFA4359BC760C87FF68FEC577BC" bold="true" box="[199,333,150,176]" pageId="23" pageNumber="292">Variability.</emphasis>
Outer margin of parameres of the specimen from Hainan Island not so sinuated in apical twothirds as in specimens from other areas (
<figureCitation id="13642A6D4359BC760E19FF44FD5C77D9" box="[601,724,186,213]" captionStart="FIGURES 25–29" captionStartId="14.[151,264,1908,1932]" captionTargetBox="[286,1384,199,1845]" captionTargetId="figure-30@14.[182,1405,181,1883]" captionTargetPageId="14" captionText="FIGURES 25–29. Aedeagus of Dactylosternum spp. and type specimen. 25: D. salvazai (25a: ventral view, 25b: dorsal view, 25c: ventral plate), 26: D. frater sp. nov., holotype (26a: ventral view, 26b: dorsal view, 26c: ventral plate), 27–29: D. pseudolatum sp. nov. (27: from Guangdong, 28: from Hainan, 27a, 28a: ventral view, 27b, 28b: dorsal view, 27c, 28c: ventral plate of the median lobe), 29: aedeagus of paratype of D. vitalisi (dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.5843861" httpUri="https://zenodo.org/record/5843861/files/figure.png" pageId="23" pageNumber="292">Figs 28a,b</figureCitation>
). In this aspect, the aedeagus of the Hainan specimens is almost the same as that of the 
<typeStatus id="54E4884A4359BC760DE0FF21FD8877F5" box="[416,512,223,249]" pageId="23" pageNumber="292" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C5F4D6B4359BC760E69FF21FD1077F4" authorityName="d' Orchymont" authorityYear="1925" box="[553,664,222,248]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="292" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA4359BC760E69FF21FD1077F4" box="[553,664,222,248]" italics="true" pageId="23" pageNumber="292">D. vitalisi</emphasis>
</taxonomicName>
from 
<collectingCountry id="F34876784359BC760EA0FF21FC9277F5" box="[736,794,223,249]" name="Laos" pageId="23" pageNumber="292">Laos</collectingCountry>
. The latter specimens also corresponds with the Chinese specimens of 
<taxonomicName id="4C5F4D6B4359BC760D71FEFDFE6A7610" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[305,482,258,284]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="292" phylum="Arthropoda" rank="species" species="pseudolatum">
<emphasis id="B92BEAFA4359BC760D71FEFDFE6A7610" box="[305,482,258,284]" italics="true" pageId="23" pageNumber="292">D. pseudolatum</emphasis>
</taxonomicName>
in external morphology. We hence treat the 
<typeStatus id="54E4884A4359BC760F90FEFDFBB87611" box="[976,1072,259,285]" pageId="23" pageNumber="292" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C5F4D6B4359BC760815FEFDFB4B7610" authorityName="d' Orchymont" authorityYear="1925" box="[1109,1219,258,284]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="292" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA4359BC760815FEFDFB4B7610" box="[1109,1219,258,284]" italics="true" pageId="23" pageNumber="292">D. vitalisi</emphasis>
</taxonomicName>
as a 
<typeStatus id="54E4884A4359BC7608BCFEFDFAD47611" box="[1276,1372,259,285]" pageId="23" pageNumber="292" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C5F4D6B4359BC7609C1FEFDFEAE764C" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="292" phylum="Arthropoda" rank="species" species="pseudolatum">
<emphasis id="B92BEAFA4359BC7609C1FEFDFEAE764C" italics="true" pageId="23" pageNumber="292">D. pseudolatum</emphasis>
</taxonomicName>
herein.
</paragraph>
</subSubSection>
<subSubSection id="C34565634359BC760C87FEB4FB807669" box="[199,1032,330,357]" pageId="23" pageNumber="292" type="distribution">
<paragraph id="8BE036E84359BC760C87FEB4FB807669" blockId="23.[151,1437,150,357]" box="[199,1032,330,357]" pageId="23" pageNumber="292">
<emphasis id="B92BEAFA4359BC760C87FEB4FEE97668" bold="true" box="[199,353,330,356]" pageId="23" pageNumber="292">Distribution.</emphasis>
<collectingCountry id="F34876784359BC760D27FEB4FE227668" box="[359,426,330,356]" name="China" pageId="23" pageNumber="292">China</collectingCountry>
(
<collectingRegion id="499BF80A4359BC760DF9FEB4FDC87668" box="[441,576,330,356]" country="China" name="Guangdong" pageId="23" pageNumber="292">Guangdong</collectingRegion>
, 
<collectingRegion id="499BF80A4359BC760E0BFEB4FD177668" box="[587,671,330,356]" country="China" name="Jiangxi" pageId="23" pageNumber="292">Jiangxi</collectingRegion>
, 
<collectingRegion id="499BF80A4359BC760EEAFEB4FD7D7668" box="[682,757,330,356]" country="China" name="Fujian" pageId="23" pageNumber="292">Fujian</collectingRegion>
, 
<collectingRegion id="499BF80A4359BC760F40FEB4FCDC7668" box="[768,852,330,356]" country="China" name="Hainan" pageId="23" pageNumber="292">Hainan</collectingRegion>
, 
<collectingRegion id="499BF80A4359BC760F1EFEB5FC307669" box="[862,952,331,357]" country="China" name="Yunnan" pageId="23" pageNumber="292">Yunnan</collectingRegion>
), 
<collectingCountry id="F34876784359BC760F8AFEB5FB8C7669" box="[970,1028,331,357]" name="Laos" pageId="23" pageNumber="292">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>