223 lines
18 KiB
XML
223 lines
18 KiB
XML
<document ID-DOI="10.25221/fee.393.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7164662" approvalRequired="22" approvalRequired_for_textStreams="22" checkinTime="1665244485009" checkinUser="felipe" docAuthor="Pathomwattananurak, W., Jeenthong, T. & Tasen, W." docDate="2019" docId="03F687F7FFF43B29FF33E0A6C281FA27" docLanguage="en" docName="FarEastEntomol.393.1-10.pdf" docOrigin="Far Eastern Entomologist 393" docSource="http://dx.doi.org/10.25221/fee.393.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Clyster scaurus Prell 1911" docType="treatment" docVersion="2" lastPageNumber="4" masterDocId="FFCFFF8FFFF53B2AFFD0E458C154FFD5" masterDocTitle="To the knowledge of the subfamily Dynastinae (Coleoptera: Scarabaeidae) from Thailand" masterLastPageNumber="10" masterPageNumber="1" pageNumber="2" updateTime="1665257173838" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>To the knowledge of the subfamily Dynastinae (Coleoptera: Scarabaeidae) from Thailand</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pathomwattananurak, W.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jeenthong, T.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tasen, W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Far Eastern Entomologist</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2019-10-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>393</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>10</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.25221/fee.393.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.25221/fee.393.1</mods:identifier>
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<mods:identifier type="ISSN">2713-2196</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7164662</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03F687F7FFF43B29FF33E0A6C281FA27" httpUri="http://treatment.plazi.org/id/03F687F7FFF43B29FF33E0A6C281FA27" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection box="[227,525,1278,1303]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph blockId="1.[227,525,1278,1333]" box="[227,525,1278,1303]" pageId="1" pageNumber="2">
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<heading box="[227,525,1278,1303]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="2" reason="2">
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<taxonomicName authority="Prell, 1911" authorityName="Prell" authorityYear="1911" box="[227,525,1278,1303]" class="Insecta" family="Dynastidae" genus="Clyster" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="scaurus">
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<emphasis bold="true" box="[227,525,1278,1303]" pageId="1" pageNumber="2">
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<emphasis bold="true" box="[227,397,1278,1302]" italics="true" pageId="1" pageNumber="2">Clyster scaurus</emphasis>
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Prell, 1911
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[227,320,1309,1333]" pageId="1" pageNumber="2" type="description">
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<paragraph blockId="1.[227,525,1278,1333]" box="[227,320,1309,1333]" pageId="1" pageNumber="2">
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<figureCitation box="[227,320,1309,1333]" captionStart="Figs 1–5" captionStartId="2.[265,306,1202,1224]" captionTargetBox="[246,1103,276,1177]" captionTargetId="figure-0@2.[227,1123,257,1196]" captionTargetPageId="2" captionText="Figs 1–5. Clyster scaurus: 1 – male, lateral view; 2 – male, dorsal view; 3 – male" figureDoi="http://doi.org/10.5281/zenodo.7164664" httpUri="https://zenodo.org/record/7164664/files/figure.png" pageId="1" pageNumber="2">Figs 1–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="reference_group">
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<paragraph blockId="1.[227,1127,1369,1449]" box="[265,1127,1369,1391]" pageId="1" pageNumber="2">
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<taxonomicName authority="Prell, 1911: 208" authorityName="Prell" authorityPageNumber="208" authorityYear="1911" box="[265,587,1369,1391]" class="Insecta" family="Dynastidae" genus="Clyster" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="scaurus">
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<emphasis box="[265,417,1369,1391]" italics="true" pageId="1" pageNumber="2">Clyster scaurus</emphasis>
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<materialsCitation box="[426,587,1369,1391]" country="Vietnam" location="Cochinchine" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Ho Chi Minh City">Prell, 1911: 208</materialsCitation>
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</taxonomicName>
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.
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<typeStatus box="[599,649,1370,1391]" pageId="1" pageNumber="2">Type</typeStatus>
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locality:
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<location LSID="urn:lsid:plazi:treatment:03F687F7FFF43B29FF33E0A6C281FA27:8E80603AFFF43B2BFD38E101C235FABA" box="[744,865,1369,1391]" country="Vietnam" name="Cochinchine" pageId="1" pageNumber="2" stateProvince="Ho Chi Minh City">Cochinchine</location>
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[
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<collectingCountry box="[879,997,1370,1391]" name="Vietnam" pageId="1" pageNumber="2">VIETNAM</collectingCountry>
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],
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<collectingRegion box="[1014,1081,1369,1391]" country="Vietnam" name="Ho Chi Minh City" pageId="1" pageNumber="2">Saigon</collectingRegion>
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[
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<collectingRegion box="[1095,1127,1370,1391]" country="Vietnam" name="Ho Chi Minh City" pageId="1" pageNumber="2">Ho</collectingRegion>
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</paragraph>
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<paragraph blockId="1.[227,1127,1369,1449]" box="[227,403,1397,1419]" pageId="1" pageNumber="2">Chi Minh] (MfN).</paragraph>
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<paragraph blockId="1.[227,1127,1369,1449]" box="[265,683,1426,1449]" pageId="1" pageNumber="2">
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<taxonomicName authority=": Endrodi, 1974: 320" authorityName=": Endrodi" authorityPageNumber="320" authorityYear="1974" box="[265,616,1427,1449]" class="Insecta" family="Dynastidae" genus="Clyster" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="scaurus">
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<emphasis box="[265,415,1427,1449]" italics="true" pageId="1" pageNumber="2">Clyster scaurus</emphasis>
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: Endrodi, 1974: 320
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</taxonomicName>
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, fig. 1.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph blockId="1.[227,1127,1487,1603]" box="[265,1127,1487,1511]" pageId="1" pageNumber="2">
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MATERIAL.
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<emphasis bold="true" box="[416,518,1487,1511]" pageId="1" pageNumber="2">
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<collectingCountry box="[416,518,1487,1511]" name="Thailand" pageId="1" pageNumber="2">Thailand</collectingCountry>
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</emphasis>
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:
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<collectingRegion box="[532,678,1487,1511]" country="Thailand" name="Trat" pageId="1" pageNumber="2">Trat Province</collectingRegion>
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, Ko Kud district, Ban Khlong Chao, 19.IV
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</paragraph>
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<paragraph blockId="1.[227,1127,1487,1603]" pageId="1" pageNumber="2">
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2009,
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<materialsCitation collectingDate="2016-05-29" collectionCode="THNHM, T" collectorName="Jeenthong" location="Muang district" pageId="1" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Trat">
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<specimenCount box="[297,330,1518,1542]" pageId="1" pageNumber="2" type="male">1♂</specimenCount>
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(THNHM-I-10052,
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<collectionCode box="[556,658,1518,1542]" pageId="1" pageNumber="2">THNHM</collectionCode>
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),
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<collectionCode box="[679,698,1518,1541]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="2">T</collectionCode>
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.
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<collectorName box="[712,820,1517,1541]" pageId="1" pageNumber="2">Jeenthong</collectorName>
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leg.;
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<collectingRegion box="[886,1038,1517,1541]" country="Thailand" name="Trat" pageId="1" pageNumber="2">Trat Province</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:03F687F7FFF43B29FF33E0A6C281FA27:8E80603AFFF43B2BFBCBE1B6C064F9F1" name="Muang district" pageId="1" pageNumber="2" stateProvince="Trat">Muang district</location>
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, Laemglad, Lan Sai beach,
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<date box="[601,717,1548,1572]" pageId="1" pageNumber="2" value="2016-05-29">
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<collectingDate box="[601,717,1548,1572]" pageId="1" pageNumber="2" value="2016-05-29">29.V 2016</collectingDate>
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</date>
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</materialsCitation>
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,
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<materialsCitation box="[728,1122,1548,1572]" collectionCode="THNHM, T" pageId="1" pageNumber="2" specimenCount="1" specimenCount-female="1" stateProvince="Trat">
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<specimenCount box="[728,762,1548,1572]" pageId="1" pageNumber="2" type="female">1♀</specimenCount>
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(THNHM-I-10010,
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<collectionCode box="[983,1085,1548,1572]" pageId="1" pageNumber="2">THNHM</collectionCode>
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),
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<collectionCode box="[1103,1122,1549,1572]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="2">T</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="1.[227,1127,1487,1603]" box="[227,382,1579,1603]" pageId="1" pageNumber="2">Jeenthong leg.</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7164664" ID-Zenodo-Dep="7164664" box="[265,1127,1201,1224]" httpUri="https://zenodo.org/record/7164664/files/figure.png" pageId="2" pageNumber="3" startId="2.[265,306,1202,1224]" targetBox="[246,1103,276,1177]" targetPageId="2">
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<paragraph blockId="2.[227,1127,1201,1251]" box="[265,1127,1201,1224]" pageId="2" pageNumber="3">
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Figs 1–5.
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<taxonomicName authorityName=": Endrodi" authorityYear="1974" box="[371,526,1202,1223]" class="Insecta" family="Dynastidae" genus="Clyster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="scaurus">
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<emphasis box="[371,526,1202,1223]" italics="true" pageId="2" pageNumber="3">Clyster scaurus</emphasis>
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</taxonomicName>
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: 1 – male, lateral view; 2 – male, dorsal view; 3 – male
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</paragraph>
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</caption>
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<paragraph blockId="2.[227,1127,1201,1251]" box="[227,1064,1229,1251]" pageId="2" pageNumber="3">genitalia, dorsal view; 4 – male genitalia, ventral view; 5 – male genitalia, lateral view.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph blockId="2.[227,1127,1288,1619]" pageId="2" pageNumber="3">
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DIAGNOSIS.
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<emphasis box="[426,546,1288,1312]" italics="true" pageId="2" pageNumber="3">Adult male</emphasis>
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(
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<figureCitation box="[563,662,1288,1313]" captionStart="Figs 1–5" captionStartId="2.[265,306,1202,1224]" captionTargetBox="[246,1103,276,1177]" captionTargetId="figure-0@2.[227,1123,257,1196]" captionTargetPageId="2" captionText="Figs 1–5. Clyster scaurus: 1 – male, lateral view; 2 – male, dorsal view; 3 – male" figureDoi="http://doi.org/10.5281/zenodo.7164664" httpUri="https://zenodo.org/record/7164664/files/figure.png" pageId="2" pageNumber="3">Figs 1–2</figureCitation>
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). BL
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<quantity box="[732,810,1288,1312]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="3" unit="mm" value="26.0">26 mm</quantity>
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, BW
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<quantity box="[881,959,1289,1313]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="mm" value="12.0">12 mm</quantity>
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. Body entirely dark brown; anterior face of pronotum punctate, while posterior face smooth and shiny; elytra finely punctate. This species is most closely related to
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<taxonomicName box="[1001,1126,1350,1374]" pageId="2" pageNumber="3">
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<emphasis box="[1001,1126,1350,1374]" italics="true" pageId="2" pageNumber="3">Clyster itys</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph blockId="2.[227,1127,1288,1619]" lastBlockId="3.[227,1127,264,472]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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(Olivier, 1789), a Sundaland species (
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<figureCitation box="[651,766,1380,1404]" captionStart="Figs 6–10" captionStartId="3.[265,306,1472,1494]" captionTargetBox="[246,1098,478,1460]" captionTargetId="figure-84@3.[227,1123,472,1467]" captionTargetPageId="3" captionText="Figs 6–10. Clyster itys: 6 – male, lateral view; 7 – male, dorsal view; 8 – male genitalia," figureDoi="http://doi.org/10.5281/zenodo.7164666" httpUri="https://zenodo.org/record/7164666/files/figure.png" pageId="2" pageNumber="3">Figs 6–10</figureCitation>
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), but can be separated from the latter by the following characteristics: 1) anterior face of pronotum flat and steep anteriorly, its posterior margin laterally with two small acute denticles, and ridge connecting the denticles sinuate (with two large truncate denticles, and ridge connecting the denticles with a distinct denticle in the latter), 2) area around lateral denticles shallowly depressed (distinctly deeply depressed in the latter, see figs 6–7).
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<emphasis italics="true" pageId="2" pageNumber="3">Male genitalia</emphasis>
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(
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<figureCitation box="[340,442,1564,1588]" captionStart="Figs 1–5" captionStartId="2.[265,306,1202,1224]" captionTargetBox="[246,1103,276,1177]" captionTargetId="figure-0@2.[227,1123,257,1196]" captionTargetPageId="2" captionText="Figs 1–5. Clyster scaurus: 1 – male, lateral view; 2 – male, dorsal view; 3 – male" figureDoi="http://doi.org/10.5281/zenodo.7164664" httpUri="https://zenodo.org/record/7164664/files/figure.png" pageId="2" pageNumber="3">Figs 3–5</figureCitation>
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). In dorsal view, paramere relatively long, broader anteriorly and gradually narrowed medially then suddenly broaden apically; apex of paramere with strongly convex lateral margin but almost straight inner margin (
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<figureCitation box="[1011,1080,264,288]" captionStart="Figs 1–5" captionStartId="2.[265,306,1202,1224]" captionTargetBox="[246,1103,276,1177]" captionTargetId="figure-0@2.[227,1123,257,1196]" captionTargetPageId="2" captionText="Figs 1–5. Clyster scaurus: 1 – male, lateral view; 2 – male, dorsal view; 3 – male" figureDoi="http://doi.org/10.5281/zenodo.7164664" httpUri="https://zenodo.org/record/7164664/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
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). In lateral view, paramere strongly curved down, its apex relatively sharp (
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<figureCitation box="[991,1068,294,318]" captionStart="Figs" captionStartId="4.[265,309,1610,1632]" captionTargetBox="[242,1111,559,1592]" captionTargetId="figure-79@4.[227,1122,547,1604]" captionTargetPageId="4" captionText="Figs. 11 – 15. Dichodontus crassus: 11– male, lateral view; 12 – male, dorsal view; 13–" figureDoi="http://doi.org/10.5281/zenodo.7164668" httpUri="https://zenodo.org/record/7164668/files/figure.png" pageId="3" pageNumber="4">Fig. 15</figureCitation>
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) (see male genitalia of the closely related species
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<taxonomicName baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[697,765,325,349]" class="Insecta" family="Dynastidae" genus="Clyster" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="itys">
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<emphasis box="[697,765,325,349]" italics="true" pageId="3" pageNumber="4">C. itys</emphasis>
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</taxonomicName>
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,
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<figureCitation box="[779,886,325,349]" captionStart="Figs 6–10" captionStartId="3.[265,306,1472,1494]" captionTargetBox="[246,1098,478,1460]" captionTargetId="figure-84@3.[227,1123,472,1467]" captionTargetPageId="3" captionText="Figs 6–10. Clyster itys: 6 – male, lateral view; 7 – male, dorsal view; 8 – male genitalia," figureDoi="http://doi.org/10.5281/zenodo.7164666" httpUri="https://zenodo.org/record/7164666/files/figure.png" pageId="3" pageNumber="4">Figs 8–10</figureCitation>
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, for comparison).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="distribution">
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<paragraph blockId="3.[227,1127,264,472]" pageId="3" pageNumber="4">
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DISTRIBUTION.
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<collectingCountry box="[472,564,356,380]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
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(Prell, 1911; Endrodi, 1974) and
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<collectingCountry box="[964,1059,356,380]" name="Thailand" pageId="3" pageNumber="4">Thailand</collectingCountry>
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(
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<emphasis bold="true" pageId="3" pageNumber="4">new record</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="discussion">
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<paragraph blockId="3.[227,1127,264,472]" box="[265,1126,417,442]" pageId="3" pageNumber="4">
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REMARKS. To date,
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<taxonomicName authorityName=": Endrodi" authorityYear="1974" box="[512,629,417,441]" class="Insecta" family="Dynastidae" genus="Clyster" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="scaurus">
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<emphasis box="[512,629,417,441]" italics="true" pageId="3" pageNumber="4">C. scaurus</emphasis>
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</taxonomicName>
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has been known only from the
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<typeStatus box="[990,1036,418,442]" pageId="3" pageNumber="4">type</typeStatus>
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locality
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</paragraph>
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<paragraph blockId="3.[227,1127,264,472]" box="[227,1063,448,472]" pageId="3" pageNumber="4">
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(
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<collectingRegion box="[234,380,448,472]" country="Vietnam" name="Ho Chi Minh City" pageId="3" pageNumber="4">Ho Chi Minh</collectingRegion>
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,
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<collectingCountry box="[391,488,448,472]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
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). We record this species in
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<collectingCountry box="[780,875,448,472]" name="Thailand" pageId="3" pageNumber="4">Thailand</collectingCountry>
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for the first time.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7164666" ID-Zenodo-Dep="7164666" box="[265,1127,1472,1494]" httpUri="https://zenodo.org/record/7164666/files/figure.png" pageId="3" pageNumber="4" startId="3.[265,306,1472,1494]" targetBox="[246,1098,478,1460]" targetPageId="3">
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<paragraph blockId="3.[227,1127,1472,1522]" box="[265,1127,1472,1494]" pageId="3" pageNumber="4">
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Figs 6–10.
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<taxonomicName baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[374,484,1472,1494]" class="Insecta" family="Dynastidae" genus="Clyster" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="itys">
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<emphasis box="[374,484,1472,1494]" italics="true" pageId="3" pageNumber="4">Clyster itys</emphasis>
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</taxonomicName>
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: 6 – male, lateral view; 7 – male, dorsal view; 8 – male genitalia,
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="3.[227,1127,1472,1522]" box="[227,981,1500,1522]" pageId="3" pageNumber="4">dorsal view; 9 – male genitalia, ventral view; 10 – male genitalia, lateral view.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |