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<document id="A7C8F01E6D71027A9DD28300F8798987" ID-DOI="10.5281/zenodo.12584512" ID-ISSN="2064-2474" ID-Zenodo-Dep="12584512" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719592314640" checkinUser="felipe" docAuthor="Csuzdi, C. S." docDate="2010" docId="03F687CF9968FFBEFDC119F0FD49604C" docLanguage="en" docName="ActaZoolAcadSciH.56.2.103-118.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 56 (2)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Pickfordiella eudrilina Csuzdi, 2010, sp. n." docType="treatment" docVersion="3" lastPageNumber="113" masterDocId="FFCFFFB79961FFB5FFA81D0DFFA5626C" masterDocTitle="Review Of The Paleotropical Neogastrini Earthworms (Oligochaeta, Acanthodrilidae: Benhamiinae) With Description Of Two New Genera" masterLastPageNumber="118" masterPageNumber="103" pageNumber="112" updateTime="1719597088924" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="4E5D62105F88A1265429512B38BBD3C4">Review Of The Paleotropical Neogastrini Earthworms (Oligochaeta, Acanthodrilidae: Benhamiinae) With Description Of Two New Genera</mods:title>
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<mods:namePart id="21BE7B2579BA6738F4A0C7FB5646B877">Csuzdi, C. S.</mods:namePart>
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<mods:title id="C922DF9852DF42D97BC61982BE7D8FD6">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date id="81D362C5D16BD6203B277177E8CC56F0">2010</mods:date>
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<mods:number id="0454EEFE0D808FB852AEF85C36308C3C">2010-05-30</mods:number>
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<mods:number id="11B253013ABF5A31CC711089EFB4FD8E">56</mods:number>
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<mods:identifier id="8DABF860AEF20A00B2AFEE005EF191CB" type="ISSN">2064-2474</mods:identifier>
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<treatment id="03F687CF9968FFBEFDC119F0FD49604C" ID-DOI="http://doi.org/10.5281/zenodo.12585370" ID-Zenodo-Dep="12585370" LSID="urn:lsid:plazi:treatment:03F687CF9968FFBEFDC119F0FD49604C" httpUri="http://treatment.plazi.org/id/03F687CF9968FFBEFDC119F0FD49604C" lastPageId="11" lastPageNumber="113" pageId="9" pageNumber="112">
<subSubSection id="C34565529968FFBCFDC119F0FC6F6774" box="[617,970,1277,1304]" pageId="9" pageNumber="112" type="nomenclature">
<paragraph id="8BE036D99968FFBCFDC119F0FC6F6774" blockId="9.[617,970,1277,1304]" box="[617,970,1277,1304]" pageId="9" pageNumber="112">
<heading id="D0A881B59968FFBCFDC119F0FC6F6774" box="[617,970,1277,1304]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="112" reason="2">
<taxonomicName id="4C5F4D5A9968FFBCFDC119F0FC21677B" authorityName="Csuzdi" authorityYear="2010" box="[617,900,1277,1303]" class="Clitellata" family="Acanthodrilidae" genus="Pickfordiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="112" phylum="Annelida" rank="species" species="eudrilina" status="sp. nov.">
<emphasis id="B92BEACB9968FFBCFDC119F0FC21677B" bold="true" box="[617,900,1277,1303]" pageId="9" pageNumber="112">Pickfordiella eudrilina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21857B09968FFBCFC2319F3FC6F6774" box="[907,970,1278,1304]" pageId="9" pageNumber="112" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34565529968FFBEFD66182CFD49604C" lastPageId="11" lastPageNumber="114" pageId="9" pageNumber="112" type="description">
<paragraph id="8BE036D99968FFBCFD66182CFCC06757" blockId="9.[718,869,1313,1339]" box="[718,869,1313,1339]" pageId="9" pageNumber="112">
<heading id="D0A881B59968FFBCFD66182CFCC06757" box="[718,869,1313,1339]" centered="true" fontSize="11" level="3" pageId="9" pageNumber="112" reason="3">
(
<figureCitation id="13642A5C9968FFBCFD7F182CFCFD6757" box="[727,856,1313,1339]" captionStart="Fig" captionStartId="10.[317,352,1752,1773]" captionTargetBox="[338,1256,985,1724]" captionTargetId="figure-230@10.[330,1258,984,1724]" captionTargetPageId="10" captionText="Fig. 2. Pickfordiella eudrilina gen et sp. n. A: ventral side of the clitellum. fp = female pore, pp = prostatic pore., B: prostate. p = glandular part of the prostate, cs = copulatory chamber., C: Spermatheca. d = diverticulum, s = septum" figureDoi="http://doi.org/10.5281/zenodo.12584516" httpUri="https://zenodo.org/record/12584516/files/figure.png" pageId="9" pageNumber="112">Fig. 2A–C</figureCitation>
.)
</heading>
</paragraph>
<paragraph id="8BE036D99968FFBCFED61869FED967BC" blockId="9.[317,1270,1380,1488]" pageId="9" pageNumber="112">
<materialsCitation id="3B373C849968FFBCFED61869FB706715" box="[382,1237,1380,1401]" collectingDate="1968-04-10" collectionCode="NILES" collectorName="Campus. Leg. J. J." country="Ghana" location="Ghana" pageId="9" pageNumber="112" specimenCode="BMHN 1997.1270" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E4887B9968FFBCFED61869FE7E6715" box="[382,475,1380,1401]" pageId="9" pageNumber="112" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBF99EA29968FFBCFE4A1869FD336715" box="[482,662,1380,1401]" collectionCode="BMHN" pageId="9" pageNumber="112">BMHN 1997.1270</specimenCode>
,
<collectingCountry id="F34876499968FFBCFD371869FD456715" box="[671,736,1380,1401]" name="Ghana" pageId="9" pageNumber="112">Ghana</collectingCountry>
, Kumasi, UST
<collectorName id="26AA530F9968FFBCFCC61869FBB06715" box="[878,1045,1380,1401]" pageId="9" pageNumber="112">Campus. Leg. J.J.</collectorName>
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<collectionCode id="ED4EAE1C9968FFBCFBB11869FBF76714" box="[1049,1106,1380,1401]" pageId="9" pageNumber="112">NILES</collectionCode>
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,
<date id="FFE110199968FFBCFBF41869FB706715" box="[1116,1237,1380,1401]" pageId="9" pageNumber="112" value="1968-04-10">
<collectingDate id="EFA5E9F19968FFBCFBF41869FB706715" box="[1116,1237,1380,1401]" pageId="9" pageNumber="112" value="1968-04-10">10. IV. 1968</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B373C849968FFBCFED6188CFB9967FA" box="[382,1084,1409,1430]" collectionCode="HNHM" country="Ghana" pageId="9" pageNumber="112" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E4887B9968FFBCFED6188CFE7367FA" box="[382,470,1409,1430]" pageId="9" pageNumber="112" type="paratype">Paratype</typeStatus>
:
<collectionCode id="ED4EAE1C9968FFBCFE48188CFD8C67FA" box="[480,553,1409,1430]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="112">HNHM</collectionCode>
AF/ 5128,
<specimenCount id="9D59FD509968FFBCFD33188CFD6967FA" box="[667,716,1409,1430]" count="1" pageId="9" pageNumber="112" type="generic">1 ex.</specimenCount>
Locality and date same as that of the
</materialsCitation>
<typeStatus id="54E4887B9968FFBCFBEC188CFB0567FA" box="[1092,1184,1409,1430]" pageId="9" pageNumber="112" type="holotype">Holotype</typeStatus>
. Etymology. The specific epithet refers to shape of the new species which resembles an eudrilid worm.
</paragraph>
<paragraph id="8BE036D99968FFBCFED618FAFB3B6417" blockId="9.[317,1270,1527,1765]" pageId="9" pageNumber="112">
Diagnosis.
<emphasis id="B92BEACB9968FFBCFDA918F5FDB4647E" box="[513,529,1528,1554]" italics="true" pageId="9" pageNumber="112">L</emphasis>
.
<quantity id="4CA79B3C9968FFBCFD8818FAFD04647D" box="[544,673,1527,1553]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="4.0" metricValueMin="3.2" pageId="9" pageNumber="112" unit="mm" value="36.0" valueMax="40.0" valueMin="32.0">32–40 mm</quantity>
,
<emphasis id="B92BEACB9968FFBCFD0418F5FD64647E" box="[684,705,1528,1554]" italics="true" pageId="9" pageNumber="112">D</emphasis>
. 1.5–1.8 mm.
<emphasis id="B92BEACB9968FFBCFCC118F5FC2E647E" box="[873,907,1528,1554]" italics="true" pageId="9" pageNumber="112">No</emphasis>
. segments 88–92. Colour pale. Head epilobous, first dorsal pore 6/7. Spermathecal pores 7/8
<emphasis id="B92BEACB9968FFBCFBB81B16FB886459" box="[1040,1069,1563,1589]" italics="true" pageId="9" pageNumber="112">ab</emphasis>
. Clitellum circular on 14–½18. f 14, single ventralmedian. m 19. Genital field lacking, prostatic pores 19 open together with the male pores through copulatory chambers.
</paragraph>
<paragraph id="8BE036D99968FFBCFED61B89FB8B6489" blockId="9.[317,1270,1527,1765]" pageId="9" pageNumber="112">Septa 6/7–10/11 somewhat thickened. Muscular gizzard 6 well developed. Calciferous glands 14–15 open in the oesophagus through a common duct. Last pair of hearts 13. Typhlosole vestigial. Excretory system holoic.</paragraph>
<paragraph id="8BE036D9996BFFBFFED61CBAFDD66057" blockId="10.[317,1270,439,571]" pageId="10" pageNumber="113">
Testes
<quantity id="4CA79B3C996BFFBFFE661CBAFD9763BD" box="[462,562,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="2.5679399999999997" pageId="10" pageNumber="113" unit="in" value="10.11">10, 11 in</quantity>
perioesophageal testis sacs. Vesicles 9, 12. Prostates 19 large, open in hemispherical copulatory chambers. Penial setae lacking. Spermathecae sac-shaped with an elongated ampoule and a shorter duct which bears entally a hose-shaped diverticulum.
</paragraph>
<paragraph id="8BE036D9996BFFBFFED61F53FB21616F" blockId="10.[317,1271,606,944]" pageId="10" pageNumber="113">
<emphasis id="B92BEACB996BFFBFFED61F53FE53601F" box="[382,502,606,627]" italics="true" pageId="10" pageNumber="113">Description.</emphasis>
<typeStatus id="54E4887B996BFFBFFE521F53FDF3601F" box="[506,598,606,627]" pageId="10" pageNumber="113" type="holotype">Holotype</typeStatus>
. Length
<quantity id="4CA79B3C996BFFBFFD0E1F53FD4E601F" box="[678,747,606,627]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="113" unit="mm" value="32.0">32 mm</quantity>
, diameter 1.8 mm, number of segments 88.
<typeStatus id="54E4887B996BFFBFFB2D1F53FB7C601F" box="[1157,1241,606,627]" pageId="10" pageNumber="113" type="paratype">Paratype</typeStatus>
<quantity id="4CA79B3C996BFFBFFB761F53FEC760FC" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="113" unit="mm" value="40.0">40 mm</quantity>
in length, 1.5 mm in diameter, number of segments 92. Colour pale, pigmentation lacking. Prostomium epilobous Y-shaped. First dorsal pore in 6/7. Setae all ventral, setal arrangement after the clitellum aa:ab:bc:cd:dd = 4:1:4:1:25. Clitellum circular, extends over segments 14–½18. Male pores and prostatic pores open through a large orifice on 19, genital field lacking. Female pore single ventralmedian on 14 (
<figureCitation id="13642A5C996BFFBFFDBC1FE3FDC6616F" box="[532,611,750,771]" captionStart="Fig" captionStartId="10.[317,352,1752,1773]" captionTargetBox="[338,1256,985,1724]" captionTargetId="figure-230@10.[330,1258,984,1724]" captionTargetPageId="10" captionText="Fig. 2. Pickfordiella eudrilina gen et sp. n. A: ventral side of the clitellum. fp = female pore, pp = prostatic pore., B: prostate. p = glandular part of the prostate, cs = copulatory chamber., C: Spermatheca. d = diverticulum, s = septum" figureDoi="http://doi.org/10.5281/zenodo.12584516" httpUri="https://zenodo.org/record/12584516/files/figure.png" pageId="10" pageNumber="113">Fig. 2A</figureCitation>
). Spermathecal pores paired in 7/
<quantity id="4CA79B3C996BFFBFFC041FE3FC77616F" box="[940,978,750,771]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="10" pageNumber="113" unit="in" value="8.0">8 in</quantity>
the setal lines
<emphasis id="B92BEACB996BFFBFFBCE1FE3FBDB616F" box="[1126,1150,750,771]" italics="true" pageId="10" pageNumber="113">ab</emphasis>
.
</paragraph>
<paragraph id="8BE036D9996BFFBFFED61E06FBAB611A" blockId="10.[317,1271,606,944]" pageId="10" pageNumber="113">
<emphasis id="B92BEACB996BFFBFFED61E06FDE5614C" box="[382,576,779,800]" italics="true" pageId="10" pageNumber="113">Internal characters.</emphasis>
The first visible septum 5/6, septa 6/7–10/11 somewhat thickened. Oesophageal gizzard large, in segment 6. Two pairs of calciferous glands located in segment 14–15, they are of equal size and open in the oesophagus through a common duct at 14/15. Excretory system holoic. Paired hearts are present in segments 10–13. Typhlosole vestigial.
</paragraph>
<paragraph id="8BE036D9996BFFBEFED61E73FD49604C" blockId="10.[317,1271,606,944]" lastBlockId="11.[317,1270,436,544]" lastPageId="11" lastPageNumber="114" pageId="10" pageNumber="113">
Testes in 10 and 11 enclosed in perioesophageal testis sacs. Seminal vesicles large in 9, 12. Ovary in segment 13. Seminal ducts apparent, discharging in copulatory chambers in segment 19. A pair of prostatic glands present in 19, they open in the copulatory chambers as well (
<figureCitation id="13642A5C996AFFBEFB331CB9FB4F63A5" box="[1179,1258,436,457]" captionStart="Fig" captionStartId="10.[317,352,1752,1773]" captionTargetBox="[338,1256,985,1724]" captionTargetId="figure-230@10.[330,1258,984,1724]" captionTargetPageId="10" captionText="Fig. 2. Pickfordiella eudrilina gen et sp. n. A: ventral side of the clitellum. fp = female pore, pp = prostatic pore., B: prostate. p = glandular part of the prostate, cs = copulatory chamber., C: Spermatheca. d = diverticulum, s = septum" figureDoi="http://doi.org/10.5281/zenodo.12584516" httpUri="https://zenodo.org/record/12584516/files/figure.png" pageId="11" pageNumber="114">Fig. 2B</figureCitation>
). Spermatheca one pair in segment 8. Ampoule elongated sac-shaped, the duct somewhat shorter than the ampoule. The ental part of the duct bears a hose-shaped diverticulum which pass through the septum 7/8 and located in segment 7 (
<figureCitation id="13642A5C996AFFBEFD3B1F06FD44604C" box="[659,737,523,544]" captionStart="Fig" captionStartId="10.[317,352,1752,1773]" captionTargetBox="[338,1256,985,1724]" captionTargetId="figure-230@10.[330,1258,984,1724]" captionTargetPageId="10" captionText="Fig. 2. Pickfordiella eudrilina gen et sp. n. A: ventral side of the clitellum. fp = female pore, pp = prostatic pore., B: prostate. p = glandular part of the prostate, cs = copulatory chamber., C: Spermatheca. d = diverticulum, s = septum" figureDoi="http://doi.org/10.5281/zenodo.12584516" httpUri="https://zenodo.org/record/12584516/files/figure.png" pageId="11" pageNumber="114">Fig. 2C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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