treatments-xml/data/03/F6/87/03F687CED76A3F3129FA5CAEFB16E67F.xml

<document id="9182C416A620C0C0BAA589FA41D8B31E" ID-DOI="10.5281/zenodo.176686" ID-GBIF-Dataset="34350dcf-319e-4531-87df-93777a969625" ID-ISSN="1175-5326" ID-Zenodo-Dep="176686" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459838008040" checkinUser="plazi" docAuthor="Campos-Paiva, Rafael M. &amp; Costa, Wilson J. E. M." docDate="2007" docId="03F687CED76A3F3129FA5CAEFB16E67F" docLanguage="en" docName="zt01471p059.pdf" docOrigin="Zootaxa 1471" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ituglanis paraguassuensis Campos-Paiva &amp; Costa, 2007, new species" docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FFCFFFB6D7683F34296D5C39FF96E77D" masterDocTitle="Ituglanis paraguassuensis sp. n. (Teleostei: Siluriformes: Trichomycteridae): a new catfish from the rio Paraguaçu, northeastern Brazil" masterLastPageNumber="59" masterPageNumber="53" pageNumber="55" updateTime="1698223930602" updateUser="plazi">
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<mods:title id="3042FAB7C9336D74F30CCDB5DCD2BDC5">Ituglanis paraguassuensis sp. n. (Teleostei: Siluriformes: Trichomycteridae): a new catfish from the rio Paraguaçu, northeastern Brazil</mods:title>
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<mods:namePart id="DA447F8566FBDBC985BEA3C6672EEF74">Campos-Paiva, Rafael M.</mods:namePart>
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<mods:namePart id="D0E80841A79E25FB99921421C4FED6F0">Costa, Wilson J. E. M.</mods:namePart>
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<mods:date id="07809A4F0C540C85F110D78ABA9B55DE">2007</mods:date>
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<subSubSection id="C3456553D76A3F3629FA5CAEFF79E7A9" pageId="2" pageNumber="55" type="nomenclature">
<paragraph id="8BE036D8D76A3F3629FA5CAEFDFDE7CC" blockId="2.[151,619,151,212]" box="[151,619,151,177]" pageId="2" pageNumber="55">
<heading id="D0A881B4D76A3F3629FA5CAEFDFDE7CC" bold="true" box="[151,619,151,177]" fontSize="11" level="1" pageId="2" pageNumber="55" reason="1">
<emphasis id="B92BEACAD76A3F3629FA5CAEFDFDE7CC" bold="true" box="[151,619,151,177]" pageId="2" pageNumber="55">
<taxonomicName id="4C5F4D5BD76A3F3629FA5CAEFE45E7CC" ID-CoL="6NCR7" box="[151,467,151,177]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="paraguassuensis" status="sp. nov.">
<emphasis id="B92BEACAD76A3F3629FA5CAEFE45E7CC" bold="true" box="[151,467,151,177]" italics="true" pageId="2" pageNumber="55">Ituglanis paraguassuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21857B1D76A3F3628B65CAEFDFDE7CC" box="[475,619,151,177]" pageId="2" pageNumber="55" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE036D8D76A3F3629FA5C83FF79E7A9" blockId="2.[151,619,151,212]" box="[151,239,186,212]" pageId="2" pageNumber="55">
(
<figureCitation id="13642A5DD76A3F3629F25C83FF75E7A9" box="[159,227,186,212]" captionStart="FIGURE 1" captionStartId="2.[151,255,658,682]" captionTargetBox="[275,1311,407,645]" captionTargetId="figure@2.[275,1312,405,646]" captionTargetPageId="2" captionText="FIGURE 1. Ituglanis paraguassuensis, holotype, 41.2 mm SL, LIRP 5780." httpUri="https://zenodo.org/record/176687/files/figure.png" pageId="2" pageNumber="55">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3456553D76A3F3629FA5D3CFE93E43A" pageId="2" pageNumber="55" type="materials_examined">
<paragraph id="8BE036D8D76A3F3629FA5D3CFE87E612" blockId="2.[151,1436,261,367]" pageId="2" pageNumber="55">
<typeStatus id="54E4887AD76A3F3629FA5D3CFE9FE662" box="[151,265,261,287]" pageId="2" pageNumber="55" type="holotype">
<emphasis id="B92BEACAD76A3F3629FA5D3CFE9FE662" bold="true" box="[151,265,261,287]" pageId="2" pageNumber="55">Holotype</emphasis>
</typeStatus>
. 
<collectionCode id="ED4EAE1DD76A3F3628755D3CFECEE662" box="[280,344,261,287]" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="2" pageNumber="55">LIRP</collectionCode>
5780, 
<quantity id="4CA79B3DD76A3F3628C45D3CFD84E662" box="[425,530,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="4.12" pageId="2" pageNumber="55" unit="mm" value="41.2">41.2 mm</quantity>
SL; 
<collectingCountry id="F3487648D76A3F362B215D3CFD0EE662" box="[588,664,261,287]" name="Brazil" pageId="2" pageNumber="55">Brazil</collectingCountry>
: Estado da Bahia, rio Paraguaçu near Iassu, side channel far from main river channel, forming pool with current (
<geoCoordinate id="EE6B501FD76A3F362BA05D14FCD6E63A" box="[717,832,301,327]" direction="south" orientation="latitude" pageId="2" pageNumber="55" precision="925" value="-12.75">12º 45’ S</geoCoordinate>
, 
<geoCoordinate id="EE6B501FD76A3F362A235D14FC58E63A" box="[846,974,301,327]" direction="west" orientation="longitude" pageId="2" pageNumber="55" precision="925" value="-40.2">40º 12’ W</geoCoordinate>
); S. L. Jewett and R. M. C. Castro, 
<date id="FFE11018D76A3F362C125D14FE9BE612" pageId="2" pageNumber="55" value="1988-07-25">25 July 1988</date>
.
</paragraph>
<caption id="DF206650D76A3F3629FA5EABFC28E5D4" box="[151,958,658,682]" httpUri="https://zenodo.org/record/176687/files/figure.png" pageId="2" pageNumber="55" targetBox="[275,1311,407,645]" targetPageId="2">
<paragraph id="8BE036D8D76A3F3629FA5EABFC28E5D4" blockId="2.[151,958,658,682]" box="[151,958,658,682]" pageId="2" pageNumber="55">
<emphasis id="B92BEACAD76A3F3629FA5EABFE84E5D7" bold="true" box="[151,274,658,682]" pageId="2" pageNumber="55">FIGURE 1</emphasis>
. 
<taxonomicName id="4C5F4D5BD76A3F36284D5EABFDA1E5D4" box="[288,567,658,681]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="paraguassuensis">
<emphasis id="B92BEACAD76A3F36284D5EABFDA1E5D4" box="[288,567,658,681]" italics="true" pageId="2" pageNumber="55">Ituglanis paraguassuensis</emphasis>
</taxonomicName>
, holotype, 41.2 mm SL, LIRP 5780.
</paragraph>
</caption>
<paragraph id="8BE036D8D76A3F3629AB5EE4FE93E43A" blockId="2.[151,1436,733,839]" pageId="2" pageNumber="55">
<typeStatus id="54E4887AD76A3F3629AB5EE4FED5E58A" box="[198,323,733,759]" pageId="2" pageNumber="55" type="paratype">
<emphasis id="B92BEACAD76A3F3629AB5EE4FED5E58A" bold="true" box="[198,323,733,759]" pageId="2" pageNumber="55">Paratypes</emphasis>
</typeStatus>
. 
<collectionCode id="ED4EAE1DD76A3F36283F5EE4FE07E58A" box="[338,401,733,759]" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="2" pageNumber="55">LIRP</collectionCode>
5834, 5 ex., 
<quantity id="4CA79B3DD76A3F362B475EE4FD43E58A" box="[554,725,733,759]" metricMagnitude="-2" metricUnit="m" metricValue="3.8049999999999997" metricValueMax="4.13" metricValueMin="3.4799999999999995" pageId="2" pageNumber="55" unit="mm" value="38.05" valueMax="41.3" valueMin="34.8">34.8–41.3 mm</quantity>
SL; 
<collectionCode id="ED4EAE1DD76A3F362A605EE4FCC7E58A" box="[781,849,733,759]" pageId="2" pageNumber="55">UFRJ</collectionCode>
7209, 1 ex., 
<quantity id="4CA79B3DD76A3F362A875EE4FBC5E58A" box="[1002,1107,733,759]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="2" pageNumber="55" unit="mm" value="36.5">36.5 mm</quantity>
SL; 
<collectionCode id="ED4EAE1DD76A3F362DE65EE4FB59E58A" box="[1163,1231,733,759]" pageId="2" pageNumber="55">UFRJ</collectionCode>
7282, 1 ex., 
<quantity id="4CA79B3DD76A3F362C055EE4FF53E462" metricMagnitude="-2" metricUnit="m" metricValue="3.96" pageId="2" pageNumber="55" unit="mm" value="39.6">39.6 mm</quantity>
SL (c&amp;s); 
<collectionCode id="ED4EAE1DD76A3F36283D5F3CFE33E462" LSID="urn:lsid:biocol.org:col:34871" box="[336,421,773,799]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="55">USNM</collectionCode>
<date id="FFE11018D76A3F3628C35F3CFD9FE462" box="[430,521,773,799]" pageId="2" pageNumber="55" value="16-10-30">301016</date>
, 5 ex., 
<quantity id="4CA79B3DD76A3F362B085F3CFC84E462" box="[613,786,773,799]" metricMagnitude="-2" metricUnit="m" metricValue="3.765" metricValueMax="4.16" metricValueMin="3.37" pageId="2" pageNumber="55" unit="mm" value="37.65" valueMax="41.6" valueMin="33.7">33.7–41.6 mm</quantity>
SL (2 c&amp;s), 
<quantity id="4CA79B3DD76A3F362AD85F3CFBF4E462" box="[949,1122,773,799]" metricMagnitude="-2" metricUnit="m" metricValue="3.405" metricValueMax="3.78" metricValueMin="3.03" pageId="2" pageNumber="55" unit="mm" value="34.05" valueMax="37.8" valueMin="30.3">30.3–37.8 mm</quantity>
SL, all collected with the 
<typeStatus id="54E4887AD76A3F3629FA5F14FE94E43A" box="[151,258,813,839]" pageId="2" pageNumber="55" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
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<paragraph id="8BE036D8D76A3F3629FA5B03FE5DE08C" blockId="2.[151,1436,1850,2033]" pageId="2" pageNumber="55">
<emphasis id="B92BEACAD76A3F3629FA5B03FE84E02F" bold="true" box="[151,274,1850,1874]" pageId="2" pageNumber="55">FIGURE 2</emphasis>
. Dorsal view of neurocranium and upper jaw and associated structures of 
<taxonomicName id="4C5F4D5BD76A3F362D415B03FB63E02C" box="[1068,1269,1850,1873]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="paraguassuensis">
<emphasis id="B92BEACAD76A3F362D415B03FB63E02C" box="[1068,1269,1850,1873]" italics="true" pageId="2" pageNumber="55">I. paraguassuensis</emphasis>
</taxonomicName>
. AOR – antorbital; APA – autopalatine; EPE - epioccipital; FPR – parietal fontanel; FSO – postparieto-supraoccipital fontanel; GBC – swim bladder; i1-i3, i10-i11 – infraorbital pores; ll1 – lateral line pore; MAX – maxilla; MET - mesethmoid; PAR – parietal; PMX - premaxilla; po1 – preopercular pores; PP-SO – postparieto-supraoccipital; PTE - pterotic; PTT-SCL – postemporo- supracleithrum; s1-s6 – supraorbital pores; SPH - sphenotic; STB – supraorbital tendon bone; VCU – vertebral centrum; VOM - vomer.
</paragraph>
</caption>
<subSubSection id="C3456553D76B3F3729AB5CAEFB20E517" pageId="3" pageNumber="56" type="diagnosis">
<paragraph id="8BE036D8D76B3F3729AB5CAEFB20E517" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="56">
<emphasis id="B92BEACAD76B3F3729AB5CAEFEDEE7CC" bold="true" box="[198,328,151,177]" pageId="3" pageNumber="56">Diagnosis:</emphasis>
Distinguished from all other species of the genus by the following combination of characteristics: color pale yellow on lateral and dorsal regions of head and body, whiter on ventral. Skin covered by several irregular pale brown blotches aligned along the body (vs. distinct color patterns in all other 
<taxonomicName id="4C5F4D5BD76B3F372C4F5CD1FA1CE67C" box="[1314,1418,232,257]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="B92BEACAD76B3F372C4F5CD1FA1CE67C" box="[1314,1418,232,257]" italics="true" pageId="3" pageNumber="56">Ituglanis</emphasis>
</taxonomicName>
); pectoral fin-rays i,6 (vs. i, 
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<emphasis id="B92BEACAD76B3F37289A5D29FCA6E654" box="[503,816,272,297]" italics="true" pageId="3" pageNumber="56">
<taxonomicName id="4C5F4D5BD76B3F37289A5D29FD0DE654" box="[503,667,272,297]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="macuniama">I. macuniama</taxonomicName>
, I, 
<taxonomicName id="4C5F4D5BD76B3F372BD35D29FCA6E654" box="[702,816,272,297]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="cahyensis">cahyensis</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C5F4D5BD76B3F372A055D29FC6AE654" box="[872,1020,272,297]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="parahybae">
<emphasis id="B92BEACAD76B3F372A055D29FC6AE654" box="[872,1020,272,297]" italics="true" pageId="3" pageNumber="56">I. parahybae</emphasis>
</taxonomicName>
; i, 
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<taxonomicName id="4C5F4D5BD76B3F372D235D29FB66E654" box="[1102,1264,272,297]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="amazonicus">
<emphasis id="B92BEACAD76B3F372D235D29FB66E654" box="[1102,1264,272,297]" italics="true" pageId="3" pageNumber="56">I. amazonicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372D935D29FF55E62C" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="eichorniarium">
<emphasis id="B92BEACAD76B3F372D935D29FF55E62C" italics="true" pageId="3" pageNumber="56">I. eichorniarium</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F3729B85D01FEA1E62C" box="[213,311,312,337]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="metae">
<emphasis id="B92BEACAD76B3F3729B85D01FEA1E62C" box="[213,311,312,337]" italics="true" pageId="3" pageNumber="56">I. metae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D5BD76B3F37281A5D01FD9FE62C" box="[375,521,312,337]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="B92BEACAD76B3F37281A5D01FD9FE62C" box="[375,521,312,337]" italics="true" pageId="3" pageNumber="56">I. nebulosus</emphasis>
</taxonomicName>
; i, 
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<taxonomicName id="4C5F4D5BD76B3F372B0A5D01FD4DE62C" box="[615,731,312,337]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="bambui">
<emphasis id="B92BEACAD76B3F372B0A5D01FD4DE62C" box="[615,731,312,337]" italics="true" pageId="3" pageNumber="56">I. bambui</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372B805D01FCEEE62C" box="[749,888,312,337]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="passensis">
<emphasis id="B92BEACAD76B3F372B805D01FCEEE62C" box="[749,888,312,337]" italics="true" pageId="3" pageNumber="56">I. passensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372AE75D01FBA5E62C" box="[906,1075,312,337]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="epikarsticus">
<emphasis id="B92BEACAD76B3F372AE75D01FBA5E62C" box="[906,1075,312,337]" italics="true" pageId="3" pageNumber="56">I. epikarsticus</emphasis>
</taxonomicName>
; i, 
<quantity id="4CA79B3DD76B3F372D3B5D01FB10E62F" box="[1110,1158,312,338]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="3" pageNumber="56" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="4C5F4D5BD76B3F372DFF5D01FA91E62C" box="[1170,1287,312,337]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="ramiroi">
<emphasis id="B92BEACAD76B3F372DFF5D01FA91E62C" box="[1170,1287,312,337]" italics="true" pageId="3" pageNumber="56">I. ramiroi</emphasis>
</taxonomicName>
except in 
<taxonomicName id="4C5F4D5BD76B3F372CE65D01FF75E604" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="parkoi">
<emphasis id="B92BEACAD76B3F372CE65D01FF75E604" italics="true" pageId="3" pageNumber="56">I. parkoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F37299E5D59FECBE604" box="[243,349,352,377]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="proops">
<emphasis id="B92BEACAD76B3F37299E5D59FECBE604" box="[243,349,352,377]" italics="true" pageId="3" pageNumber="56">I. proops</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D5BD76B3F3728FA5D59FD9AE604" box="[407,524,352,377]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B92BEACAD76B3F3728FA5D59FD9AE604" box="[407,524,352,377]" italics="true" pageId="3" pageNumber="56">I. laticeps</emphasis>
</taxonomicName>
); pelvic-fin-rays i,4 (vs. i, 
<quantity id="4CA79B3DD76B3F372A2E5D59FCE4E607" box="[835,882,352,378]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="56" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C5F4D5BD76B3F372A175D59FBADE604" box="[890,1083,352,377]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="eichorniarium">
<emphasis id="B92BEACAD76B3F372A175D59FBADE604" box="[890,1083,352,377]" italics="true" pageId="3" pageNumber="56">I. eichorniarium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D5BD76B3F372D1B5D59FA97E604" box="[1142,1281,352,377]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="cahyensis">
<emphasis id="B92BEACAD76B3F372D1B5D59FA97E604" box="[1142,1281,352,377]" italics="true" pageId="3" pageNumber="56">I. cahyensis</emphasis>
</taxonomicName>
or pelvic-fin absent in 
<taxonomicName id="4C5F4D5BD76B3F3728645DB1FE09E6DC" box="[265,415,392,417]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="parahybae">
<emphasis id="B92BEACAD76B3F3728645DB1FE09E6DC" box="[265,415,392,417]" italics="true" pageId="3" pageNumber="56">I. parahybae</emphasis>
</taxonomicName>
; i, 
<quantity id="4CA79B3DD76B3F3728D35DB1FE7AE6DF" box="[446,492,392,418]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="56" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4C5F4D5BD76B3F3728985DB1FD0EE6DC" box="[501,664,392,417]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="amazonicus">
<emphasis id="B92BEACAD76B3F3728985DB1FD0EE6DC" box="[501,664,392,417]" italics="true" pageId="3" pageNumber="56">I. amazonicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372BC55DB1FCA6E6DC" box="[680,816,392,417]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="passensis">
<emphasis id="B92BEACAD76B3F372BC55DB1FCA6E6DC" box="[680,816,392,417]" italics="true" pageId="3" pageNumber="56">I. passensis</emphasis>
</taxonomicName>
except in 
<taxonomicName id="4C5F4D5BD76B3F372AC15DB1FBBAE6DC" box="[940,1068,392,417]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="gracilior">
<emphasis id="B92BEACAD76B3F372AC15DB1FBBAE6DC" box="[940,1068,392,417]" italics="true" pageId="3" pageNumber="56">I. gracilior</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372D515DB1FB5CE6DC" box="[1084,1226,392,417]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="B92BEACAD76B3F372D515DB1FB5CE6DC" box="[1084,1226,392,417]" italics="true" pageId="3" pageNumber="56">I. nebulosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372DB45DB1FAEAE6DC" box="[1241,1404,392,417]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="macunaima">
<emphasis id="B92BEACAD76B3F372DB45DB1FAEAE6DC" box="[1241,1404,392,417]" italics="true" pageId="3" pageNumber="56">I. macunaima</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372CE65DB1FF79E6B4" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="bambui">
<emphasis id="B92BEACAD76B3F372CE65DB1FF79E6B4" italics="true" pageId="3" pageNumber="56">I. bambui</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F3728675D89FEEBE6B4" box="[266,381,432,457]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="ramiroi">
<emphasis id="B92BEACAD76B3F3728675D89FEEBE6B4" box="[266,381,432,457]" italics="true" pageId="3" pageNumber="56">I. ramiroi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F3728E25D89FDBAE6B4" box="[399,556,432,457]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="epikasticus">
<emphasis id="B92BEACAD76B3F3728E25D89FDBAE6B4" box="[399,556,432,457]" italics="true" pageId="3" pageNumber="56">I. epikasticus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372B505D89FD32E6B4" box="[573,676,432,457]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="parkoi">
<emphasis id="B92BEACAD76B3F372B505D89FD32E6B4" box="[573,676,432,457]" italics="true" pageId="3" pageNumber="56">I. parkoi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C5F4D5BD76B3F372B875D89FC3BE6B4" box="[746,941,432,457]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="guayaberensis">
<emphasis id="B92BEACAD76B3F372B875D89FC3BE6B4" box="[746,941,432,457]" italics="true" pageId="3" pageNumber="56">I. guayaberensis</emphasis>
</taxonomicName>
); parietal fontanel extending to posterior edge of medial parietal border (
<figureCitation id="13642A5DD76B3F372B685DE1FDDCE68F" box="[517,586,472,498]" captionStart="FIGURE 2" captionStartId="2.[151,255,1850,1874]" captionTargetBox="[472,1107,893,1824]" captionTargetId="figure@2.[467,1120,885,1825]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal view of neurocranium and upper jaw and associated structures of I. paraguassuensis. AOR – antorbital; APA – autopalatine; EPE - epioccipital; FPR – parietal fontanel; FSO – postparieto-supraoccipital fontanel; GBC – swim bladder; i 1 - i 3, i 10 - i 11 – infraorbital pores; ll 1 – lateral line pore; MAX – maxilla; MET - mesethmoid; PAR – parietal; PMX - premaxilla; po 1 – preopercular pores; PP-SO – postparieto-supraoccipital; PTE - pterotic; PTT-SCL – postemporo- supracleithrum; s 1 - s 6 – supraorbital pores; SPH - sphenotic; STB – supraorbital tendon bone; VCU – vertebral centrum; VOM - vomer." httpUri="https://zenodo.org/record/176688/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
) (vs. parietal fontanel reduced or vestigial, restricted to middle of medial parietal border in all other 
<taxonomicName id="4C5F4D5BD76B3F3728A25E39FDA1E564" box="[463,567,512,537]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="B92BEACAD76B3F3728A25E39FDA1E564" box="[463,567,512,537]" italics="true" pageId="3" pageNumber="56">Ituglanis</emphasis>
</taxonomicName>
except in some 
<taxonomicName id="4C5F4D5BD76B3F372B995E39FC01E564" box="[756,919,512,537]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="amazonicus">
<emphasis id="B92BEACAD76B3F372B995E39FC01E564" box="[756,919,512,537]" italics="true" pageId="3" pageNumber="56">I. amazonicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D5BD76B3F372ABD5E39FBAEE564" box="[976,1080,512,537]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="proops">
<emphasis id="B92BEACAD76B3F372ABD5E39FBAEE564" box="[976,1080,512,537]" italics="true" pageId="3" pageNumber="56">I. proops</emphasis>
</taxonomicName>
); the unusual reduced number of vertebrae 34–36, where vertebral counts normally range 39 or more (in 
<taxonomicName id="4C5F4D5BD76B3F372D5D5E11FB40E53C" box="[1072,1238,552,577]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="amazonicus">
<emphasis id="B92BEACAD76B3F372D5D5E11FB40E53C" box="[1072,1238,552,577]" italics="true" pageId="3" pageNumber="56">I. amazonicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372D875E11FAEEE53C" box="[1258,1400,552,577]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="cahyensis">
<emphasis id="B92BEACAD76B3F372D875E11FAEEE53C" box="[1258,1400,552,577]" italics="true" pageId="3" pageNumber="56">I. cahyensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372CE65E11FEA9E514" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="eichorniarium">
<emphasis id="B92BEACAD76B3F372CE65E11FEA9E514" italics="true" pageId="3" pageNumber="56">I. eichorniarium</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F3728395E69FE7CE514" box="[340,490,592,617]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="parahybae">
<emphasis id="B92BEACAD76B3F3728395E69FE7CE514" box="[340,490,592,617]" italics="true" pageId="3" pageNumber="56">I. parahybae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F3728955E69FDF7E514" box="[504,609,592,617]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="proops">
<emphasis id="B92BEACAD76B3F3728955E69FDF7E514" box="[504,609,592,617]" italics="true" pageId="3" pageNumber="56">I. proops</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372B1D5E69FD79E514" box="[624,751,592,617]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="gracilior">
<emphasis id="B92BEACAD76B3F372B1D5E69FD79E514" box="[624,751,592,617]" italics="true" pageId="3" pageNumber="56">I. gracilior</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372B935E69FCE3E514" box="[766,885,592,617]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="herberti">
<emphasis id="B92BEACAD76B3F372B935E69FCE3E514" box="[766,885,592,617]" italics="true" pageId="3" pageNumber="56">I. herberti</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372AE65E69FC7FE514" box="[907,1001,592,617]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="metae">
<emphasis id="B92BEACAD76B3F372AE65E69FC7FE514" box="[907,1001,592,617]" italics="true" pageId="3" pageNumber="56">I. metae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D5BD76B3F372A955E69FBF6E514" box="[1016,1120,592,617]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="B92BEACAD76B3F372A955E69FBF6E514" box="[1016,1120,592,617]" italics="true" pageId="3" pageNumber="56">Ituglanis</emphasis>
</taxonomicName>
sp. A).
</paragraph>
</subSubSection>
<subSubSection id="C3456553D76B3F3029AB5E4EFA1FE67F" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="description">
<paragraph id="8BE036D8D76B3F3729AB5E4EFC69E477" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="56">
<emphasis id="B92BEACAD76B3F3729AB5E4EFEF7E5EC" bold="true" box="[198,353,631,657]" pageId="3" pageNumber="56">Description:</emphasis>
Morphometric data for 
<typeStatus id="54E4887AD76B3F372B135E41FD72E5EF" box="[638,740,632,658]" pageId="3" pageNumber="56" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54E4887AD76B3F372A715E41FC1AE5EF" box="[796,908,632,658]" pageId="3" pageNumber="56" type="paratype">paratypes</typeStatus>
are given in 
<tableCitation id="C6DD0363D76B3F372D4E5E41FBECE5EF" box="[1059,1146,632,658]" captionStart="TABLE 1" captionStartId="4.[167,255,1046,1070]" captionTargetBox="[167,1334,1092,1987]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data of Ituglanis paraguassuensis n. sp. H: holotype (LIRP 5780)" httpUri="http://table.plazi.org/id/DF206650D76C3F3029CA582FFBEDE350" pageId="3" pageNumber="56" tableUuid="DF206650D76C3F3029CA582FFBEDE350">Table 1</tableCitation>
. Body elongated, subcylindrical, compressed in caudal peduncle. Dorsal profile straight on head, convex between nape and dorsal-fin origin, approximately straight on caudal peduncle. Ventral profile straight to slightly convex between lower jaw and end of anal-fin base. Lips and barbels covered by minute papillae.
</paragraph>
<paragraph id="8BE036D8D76B3F3729AB5F21FDDDE35F" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="56">Dorsal-fin origin on vertical between centrum of 19th and 21st vertebrae. Anal-fin origin on vertical through base of 6th and 8th segmented dorsal-fin rays and through the centrum of 22nd and 24th vertebrae. First pectoral-fin ray filamentous and 20 to 30 % longer than other pectoral-fin rays. Tip of pelvic-fin reaching vertical between base of 4th and 6th segmented dorsal-fin rays. Caudal-fin subtruncate. Dorsal-fin rays 12–13 (5,vii or 5,viii); anal-fin rays 9–10 (4,v or 5,v); pectoral-fin rays 7 (i,6); pelvic-fin rays 5 (i,3,i); caudal-fin principal rays 13, dorsal procurrent rays 14–15, ventral procurrent rays 11–12. Total vertebrae 34–36; epipleural ribs 6; upper hypural plate united.</paragraph>
<paragraph id="8BE036D8D76B3F3729AB5809FABAE0EF" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="56">
Head trapezoidal in dorsal view. Eye located on anterior half of head. Mouth subventral. Maxilla longer than premaxilla. Premaxilla trapezoidal with about 20 conical teeth, arranged in two irregular rows. Dentary slender, with about 25 conical teeth, arranged in two or three irregular rows. Tip of nasal barbel reaching posterior edge of opercular patch of odontodes; tip of maxillary barbel reaching pectoral-fin base; tip of rictal barbel reaching anterior edge of opercular patch of odontodes. Branchiostegal rays 8. Hypobranchials ossified; two anterior basibranchials ossified, third basibranchial consisting of rounded cartilage. Interopercular odontodes 15; interopercular patch of odontodes about equal to preopercle in length; opercular odontodes 14–17. Odontodes conical. Cephalic laterosensory canal with simple tubes ending in simple pores, continuous and connected to each other on pterotic, sphenotic and lateral border of parietal (
<figureCitation id="13642A5DD76B3F372D7E5949FBCDE2F7" box="[1043,1115,1392,1418]" captionStart="FIGURE 2" captionStartId="2.[151,255,1850,1874]" captionTargetBox="[472,1107,893,1824]" captionTargetId="figure@2.[467,1120,885,1825]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal view of neurocranium and upper jaw and associated structures of I. paraguassuensis. AOR – antorbital; APA – autopalatine; EPE - epioccipital; FPR – parietal fontanel; FSO – postparieto-supraoccipital fontanel; GBC – swim bladder; i 1 - i 3, i 10 - i 11 – infraorbital pores; ll 1 – lateral line pore; MAX – maxilla; MET - mesethmoid; PAR – parietal; PMX - premaxilla; po 1 – preopercular pores; PP-SO – postparieto-supraoccipital; PTE - pterotic; PTT-SCL – postemporo- supracleithrum; s 1 - s 6 – supraorbital pores; SPH - sphenotic; STB – supraorbital tendon bone; VCU – vertebral centrum; VOM - vomer." httpUri="https://zenodo.org/record/176688/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
). All cephalic pores paired: supraorbital 4; s1 and s2 connected by single canal; s1 between palatine and mesethmoid, s2 on vomer. s3 on vomer, s6 on parietal; connected to one another. Infraorbital 4; 
<date id="FFE11018D76B3F372A1359F9FC73E2A7" box="[894,997,1472,1498]" pageId="3" pageNumber="56">i1 and i3</date>
connected by single canal; both situated next to antorbital; 
<date id="FFE11018D76B3F3728C959D1FDB0E17F" box="[420,550,1512,1538]" pageId="3" pageNumber="56">i10 and i11</date>
connected, on sphenotic. Preopercular 1; on pterotic. Lateral line 3, all connected; ll1 on postemporo- supracleithrum, ll2 and ll3 on lateral side of body near opercle. Parietal fontanel extending to posterior border of parietal; postparieto-supraoccipital fontanel restricted to middle region; sphenotic narrow, anterior tip anteriorly directed (
<figureCitation id="13642A5DD76B3F372BC55A59FD79E107" box="[680,751,1632,1658]" captionStart="FIGURE 2" captionStartId="2.[151,255,1850,1874]" captionTargetBox="[472,1107,893,1824]" captionTargetId="figure@2.[467,1120,885,1825]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal view of neurocranium and upper jaw and associated structures of I. paraguassuensis. AOR – antorbital; APA – autopalatine; EPE - epioccipital; FPR – parietal fontanel; FSO – postparieto-supraoccipital fontanel; GBC – swim bladder; i 1 - i 3, i 10 - i 11 – infraorbital pores; ll 1 – lateral line pore; MAX – maxilla; MET - mesethmoid; PAR – parietal; PMX - premaxilla; po 1 – preopercular pores; PP-SO – postparieto-supraoccipital; PTE - pterotic; PTT-SCL – postemporo- supracleithrum; s 1 - s 6 – supraorbital pores; SPH - sphenotic; STB – supraorbital tendon bone; VCU – vertebral centrum; VOM - vomer." httpUri="https://zenodo.org/record/176688/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
). Mesethmoid somewhat widened, about equal in width to anterior portion of palatine; autopalatine with deep concavity along medial border; posterior process of palatine moderate, length about 50% of length of autopalatine without such process. Supraorbital tendon bone elongate and robust, becoming slender and curved outwards, approximately equal to autopalatine in length (
<figureCitation id="13642A5DD76B3F3729F25B39FF7EE067" box="[159,232,1792,1818]" captionStart="FIGURE 2" captionStartId="2.[151,255,1850,1874]" captionTargetBox="[472,1107,893,1824]" captionTargetId="figure@2.[467,1120,885,1825]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal view of neurocranium and upper jaw and associated structures of I. paraguassuensis. AOR – antorbital; APA – autopalatine; EPE - epioccipital; FPR – parietal fontanel; FSO – postparieto-supraoccipital fontanel; GBC – swim bladder; i 1 - i 3, i 10 - i 11 – infraorbital pores; ll 1 – lateral line pore; MAX – maxilla; MET - mesethmoid; PAR – parietal; PMX - premaxilla; po 1 – preopercular pores; PP-SO – postparieto-supraoccipital; PTE - pterotic; PTT-SCL – postemporo- supracleithrum; s 1 - s 6 – supraorbital pores; SPH - sphenotic; STB – supraorbital tendon bone; VCU – vertebral centrum; VOM - vomer." httpUri="https://zenodo.org/record/176688/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
); no process on supraorbital tendon bone (
<figureCitation id="13642A5DD76B3F372BB25B39FCBEE067" box="[735,808,1792,1818]" captionStart="FIGURE 2" captionStartId="2.[151,255,1850,1874]" captionTargetBox="[472,1107,893,1824]" captionTargetId="figure@2.[467,1120,885,1825]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal view of neurocranium and upper jaw and associated structures of I. paraguassuensis. AOR – antorbital; APA – autopalatine; EPE - epioccipital; FPR – parietal fontanel; FSO – postparieto-supraoccipital fontanel; GBC – swim bladder; i 1 - i 3, i 10 - i 11 – infraorbital pores; ll 1 – lateral line pore; MAX – maxilla; MET - mesethmoid; PAR – parietal; PMX - premaxilla; po 1 – preopercular pores; PP-SO – postparieto-supraoccipital; PTE - pterotic; PTT-SCL – postemporo- supracleithrum; s 1 - s 6 – supraorbital pores; SPH - sphenotic; STB – supraorbital tendon bone; VCU – vertebral centrum; VOM - vomer." httpUri="https://zenodo.org/record/176688/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
). Metapterygoid moderate, longest length about 60% of horizontal length of quadrate; dorsal portion of quadrate anteriorly expanded (
<figureCitation id="13642A5DD76B3F372D275B11FB04E03F" box="[1098,1170,1832,1858]" captionStart="FIGURE 3" captionStartId="4.[151,255,887,911]" captionTargetBox="[405,1165,337,853]" captionTargetId="figure@4.[401,1186,314,863]" captionTargetPageId="4" captionText="FIGURE 3. Lateral view of the left suspensorium and opercular apparatus of I. paraguassuensis. HYO - hyomandibula; INT - interopercle; MTG - metapterygoid; OPC - opercle; PPC - preopercle; QDT - quadrate." httpUri="https://zenodo.org/record/176689/files/figure.png" pageId="3" pageNumber="56">Fig. 3</figureCitation>
). Urohyal lateral wing somewhat elongate, about equal to anterior ceratohyal in length. Two pectoral radials, first cartilaginous, second ossified, length of latter about twice first. Neural prezygapophyses conspicuous in all vertebrae.
</paragraph>
<paragraph id="8BE036D8D76B3F3029AB5BA6FA1FE67F" blockId="3.[151,1437,151,2018]" lastBlockId="4.[151,1436,152,258]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
<emphasis id="B92BEACAD76B3F3729AB5BA6FE74E0C4" bold="true" box="[198,482,1951,1977]" pageId="3" pageNumber="56">Coloration in alcohol:</emphasis>
Lateral and dorsal regions of head and body pale yellow, with pale brown rounded blotches. Dark brown pigmentation concentrated on head, in front of eyes, and in front of opercular and interopercular odontodes patches. Nasal and maxillary barbels whitish with dark brown marginal pigment. Rictal barbel whitish. Dorsal and caudal fins hyaline with small concentrations of dark brown chromatophores scattered on fin rays. Pectoral, pelvic and anal fins whitish. The blotches may vanish in alcohol.
</paragraph>
</subSubSection>
<caption id="DF206650D76C3F3029FA5F4EFBEDE4CC" httpUri="https://zenodo.org/record/176689/files/figure.png" pageId="4" pageNumber="57" targetBox="[405,1165,337,853]" targetPageId="4">
<paragraph id="8BE036D8D76C3F3029FA5F4EFBEDE4CC" blockId="4.[151,1436,887,945]" pageId="4" pageNumber="57">
<emphasis id="B92BEACAD76C3F3029FA5F4EFE84E4F2" bold="true" box="[151,274,887,911]" pageId="4" pageNumber="57">FIGURE 3</emphasis>
. Lateral view of the left suspensorium and opercular apparatus of 
<taxonomicName id="4C5F4D5BD76C3F302ABF5F41FB0CE4F2" box="[978,1178,888,911]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="paraguassuensis">
<emphasis id="B92BEACAD76C3F302ABF5F41FB0CE4F2" box="[978,1178,888,911]" italics="true" pageId="4" pageNumber="57">I. paraguassuensis</emphasis>
</taxonomicName>
. HYO - hyomandibula; INT - interopercle; MTG - metapterygoid; OPC - opercle; PPC - preopercle; QDT - quadrate.
</paragraph>
</caption>
<caption id="DF206650D76C3F3029CA582FFBEDE350" ID-Table-UUID="DF206650D76C3F3029CA582FFBEDE350" box="[167,1147,1046,1070]" httpUri="http://table.plazi.org/id/DF206650D76C3F3029CA582FFBEDE350" pageId="4" pageNumber="57" targetBox="[167,1334,1092,1987]" targetIsTable="true" targetPageId="4">
<paragraph id="8BE036D8D76C3F3029CA582FFBEDE350" blockId="4.[167,1147,1046,1070]" box="[167,1147,1046,1070]" pageId="4" pageNumber="57">
<emphasis id="B92BEACAD76C3F3029CA582FFE8CE353" bold="true" box="[167,282,1046,1070]" pageId="4" pageNumber="57">TABLE 1.</emphasis>
Morphometric data of 
<taxonomicName id="4C5F4D5BD76C3F302B7C582FFCBEE350" box="[529,808,1046,1069]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="paraguassuensis" status="sp. nov.">
<emphasis id="B92BEACAD76C3F302B7C582FFCBEE350" box="[529,808,1046,1069]" italics="true" pageId="4" pageNumber="57">Ituglanis paraguassuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21857B1D76C3F302A42582FFCFEE350" box="[815,872,1046,1069]" pageId="4" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
H: holotype (LIRP 5780)
</paragraph>
</caption>
<paragraph id="8BE036D8D76C3F302BE4587DFA96E0BE" pageId="4" pageNumber="57">
<table id="F95FC478D76CC0CB29CA587DFAA0E0BE" box="[167,1334,1092,1987]" gridcols="3" gridrows="20" pageId="4" pageNumber="57">
<tr id="356F349AD76CC0CB29CA587DFAA0E326" box="[167,1334,1092,1115]" gridrow="0" pageId="4" pageNumber="57" rowspan-0="1">
<th id="76BE5DE6D76CC0CB2BE4587DFB98E326" box="[649,1038,1092,1115]" gridcol="1" gridrow="0" pageId="4" pageNumber="57">H Paratypes range (n= 14)</th>
<th id="76BE5DE6D76CC0CB2DB3587DFAA0E326" box="[1246,1334,1092,1115]" gridcol="2" gridrow="0" pageId="4" pageNumber="57">Average</th>
</tr>
<tr id="356F349AD76CC0CB29CA584DFAA0E3F6" box="[167,1334,1140,1163]" gridrow="1" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA584DFE5FE3F6" box="[167,457,1140,1163]" gridcol="0" gridrow="1" pageId="4" pageNumber="57">Standard length (mm)</th>
<td id="76BE5DE6D76CC0CB2BE4584DFB98E3F6" box="[649,1038,1140,1163]" gridcol="1" gridrow="1" pageId="4" pageNumber="57">41.2 30.3 – 41.6</td>
<td id="76BE5DE6D76CC0CB2DB3584DFAA0E3F6" box="[1246,1334,1140,1163]" gridcol="2" gridrow="1" pageId="4" pageNumber="57">37.2</td>
</tr>
<tr id="356F349AD76CC0CB29CA5898FAA0E3C4" box="[167,1334,1185,1209]" gridrow="2" pageId="4" pageNumber="57" rowspan-1="1" rowspan-2="1">
<th id="76BE5DE6D76CC0CB29CA5898FE5FE3C4" box="[167,457,1185,1209]" gridcol="0" gridrow="2" pageId="4" pageNumber="57">
<emphasis id="B92BEACAD76C3F3029CA5898FE5FE3C4" bold="true" box="[167,457,1185,1209]" pageId="4" pageNumber="57">Percents, standard length</emphasis>
</th>
</tr>
<tr id="356F349AD76CC0CB29CA58E8FAA0E395" box="[167,1334,1233,1256]" gridrow="3" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA58E8FE5FE395" box="[167,457,1233,1256]" gridcol="0" gridrow="3" pageId="4" pageNumber="57">Body depth</th>
<td id="76BE5DE6D76CC0CB2BE458E8FB98E395" box="[649,1038,1233,1256]" gridcol="1" gridrow="3" pageId="4" pageNumber="57">13.3 11.0 – 15.6</td>
<td id="76BE5DE6D76CC0CB2DB358E8FAA0E395" box="[1246,1334,1233,1256]" gridcol="2" gridrow="3" pageId="4" pageNumber="57">13.4</td>
</tr>
<tr id="356F349AD76CC0CB29CA58C7FAA0E268" box="[167,1334,1278,1301]" gridrow="4" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA58C7FE5FE268" box="[167,457,1278,1301]" gridcol="0" gridrow="4" pageId="4" pageNumber="57">Caudal-peduncle depth</th>
<td id="76BE5DE6D76CC0CB2BE458C7FB98E268" box="[649,1038,1278,1301]" gridcol="1" gridrow="4" pageId="4" pageNumber="57">10.5 8.3 – 11.5</td>
<td id="76BE5DE6D76CC0CB2DB358C7FAA0E268" box="[1246,1334,1278,1301]" gridcol="2" gridrow="4" pageId="4" pageNumber="57">9.7</td>
</tr>
<tr id="356F349AD76CC0CB29CA5915FAA0E23E" box="[167,1334,1324,1347]" gridrow="5" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5915FE5FE23E" box="[167,457,1324,1347]" gridcol="0" gridrow="5" pageId="4" pageNumber="57">Body width</th>
<td id="76BE5DE6D76CC0CB2BE45915FB98E23E" box="[649,1038,1324,1347]" gridcol="1" gridrow="5" pageId="4" pageNumber="57">7.1 5.4 – 8.4</td>
<td id="76BE5DE6D76CC0CB2DB35915FAA0E23E" box="[1246,1334,1324,1347]" gridcol="2" gridrow="5" pageId="4" pageNumber="57">6.6</td>
</tr>
<tr id="356F349AD76CC0CB29CA5963FAA0E20C" box="[167,1334,1370,1393]" gridrow="6" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5963FE5FE20C" box="[167,457,1370,1393]" gridcol="0" gridrow="6" pageId="4" pageNumber="57">Caudal peduncle width</th>
<td id="76BE5DE6D76CC0CB2BE45963FB98E20C" box="[649,1038,1370,1393]" gridcol="1" gridrow="6" pageId="4" pageNumber="57">3.0 2.4 – 5.1</td>
<td id="76BE5DE6D76CC0CB2DB35963FAA0E20C" box="[1246,1334,1370,1393]" gridcol="2" gridrow="6" pageId="4" pageNumber="57">3.4</td>
</tr>
<tr id="356F349AD76CC0CB29CA59BEFAA0E2E3" box="[167,1334,1415,1438]" gridrow="7" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA59BEFE5FE2E3" box="[167,457,1415,1438]" gridcol="0" gridrow="7" pageId="4" pageNumber="57">Dorsal-fin base length</th>
<td id="76BE5DE6D76CC0CB2BE459BEFB98E2E3" box="[649,1038,1415,1438]" gridcol="1" gridrow="7" pageId="4" pageNumber="57">13.4 10.7 – 15.3</td>
<td id="76BE5DE6D76CC0CB2DB359BEFAA0E2E3" box="[1246,1334,1415,1438]" gridcol="2" gridrow="7" pageId="4" pageNumber="57">12.7</td>
</tr>
<tr id="356F349AD76CC0CB29CA598DFAA0E2B6" box="[167,1334,1460,1483]" gridrow="8" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA598DFE5FE2B6" box="[167,457,1460,1483]" gridcol="0" gridrow="8" pageId="4" pageNumber="57">Anal-fin base length</th>
<td id="76BE5DE6D76CC0CB2BE4598DFB98E2B6" box="[649,1038,1460,1483]" gridcol="1" gridrow="8" pageId="4" pageNumber="57">10.6 8.5 – 11.4</td>
<td id="76BE5DE6D76CC0CB2DB3598DFAA0E2B6" box="[1246,1334,1460,1483]" gridcol="2" gridrow="8" pageId="4" pageNumber="57">10.3</td>
</tr>
<tr id="356F349AD76CC0CB29CA59DBFAA0E284" box="[167,1334,1506,1529]" gridrow="9" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA59DBFE5FE284" box="[167,457,1506,1529]" gridcol="0" gridrow="9" pageId="4" pageNumber="57">Pelvic-fin length</th>
<td id="76BE5DE6D76CC0CB2BE459DBFB98E284" box="[649,1038,1506,1529]" gridcol="1" gridrow="9" pageId="4" pageNumber="57">7.7 7.5 – 10.6</td>
<td id="76BE5DE6D76CC0CB2DB359DBFAA0E284" box="[1246,1334,1506,1529]" gridcol="2" gridrow="9" pageId="4" pageNumber="57">9.0</td>
</tr>
<tr id="356F349AD76CC0CB29CA5A29FAA0E15A" box="[167,1334,1552,1575]" gridrow="10" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5A29FE5FE15A" box="[167,457,1552,1575]" gridcol="0" gridrow="10" pageId="4" pageNumber="57">Pectoral-fin length</th>
<td id="76BE5DE6D76CC0CB2BE45A29FB98E15A" box="[649,1038,1552,1575]" gridcol="1" gridrow="10" pageId="4" pageNumber="57">11.2 10.5 – 14.0</td>
<td id="76BE5DE6D76CC0CB2DB35A29FAA0E15A" box="[1246,1334,1552,1575]" gridcol="2" gridrow="10" pageId="4" pageNumber="57">12.2</td>
</tr>
<tr id="356F349AD76CC0CB29CA5A04FAA0E129" box="[167,1334,1597,1620]" gridrow="11" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5A04FE5FE129" box="[167,457,1597,1620]" gridcol="0" gridrow="11" pageId="4" pageNumber="57">Predorsal length</th>
<td id="76BE5DE6D76CC0CB2BE45A04FB98E129" box="[649,1038,1597,1620]" gridcol="1" gridrow="11" pageId="4" pageNumber="57">66.3 61.3 – 69.1</td>
<td id="76BE5DE6D76CC0CB2DB35A04FAA0E129" box="[1246,1334,1597,1620]" gridcol="2" gridrow="11" pageId="4" pageNumber="57">65.5</td>
</tr>
<tr id="356F349AD76CC0CB29CA5A52FAA0E1FF" box="[167,1334,1643,1666]" gridrow="12" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5A52FE5FE1FF" box="[167,457,1643,1666]" gridcol="0" gridrow="12" pageId="4" pageNumber="57">Prepelvic length</th>
<td id="76BE5DE6D76CC0CB2BE45A52FB98E1FF" box="[649,1038,1643,1666]" gridcol="1" gridrow="12" pageId="4" pageNumber="57">59.5 54.8 – 61.2</td>
<td id="76BE5DE6D76CC0CB2DB35A52FAA0E1FF" box="[1246,1334,1643,1666]" gridcol="2" gridrow="12" pageId="4" pageNumber="57">58.2</td>
</tr>
<tr id="356F349AD76CC0CB29CA5AA1FAA0E1D2" box="[167,1334,1688,1711]" gridrow="13" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5AA1FE5FE1D2" box="[167,457,1688,1711]" gridcol="0" gridrow="13" pageId="4" pageNumber="57">Head length</th>
<td id="76BE5DE6D76CC0CB2BE45AA1FB98E1D2" box="[649,1038,1688,1711]" gridcol="1" gridrow="13" pageId="4" pageNumber="57">18.6 17.5 – 19.4</td>
<td id="76BE5DE6D76CC0CB2DB35AA1FAA0E1D2" box="[1246,1334,1688,1711]" gridcol="2" gridrow="13" pageId="4" pageNumber="57">18.5</td>
</tr>
<tr id="356F349AD76CC0CB29CA5AFCFAA0E1A0" box="[167,1334,1733,1757]" gridrow="14" pageId="4" pageNumber="57" rowspan-1="1" rowspan-2="1">
<th id="76BE5DE6D76CC0CB29CA5AFCFE5FE1A0" box="[167,457,1733,1757]" gridcol="0" gridrow="14" pageId="4" pageNumber="57">
<emphasis id="B92BEACAD76C3F3029CA5AFCFE0CE1A0" bold="true" box="[167,410,1733,1757]" pageId="4" pageNumber="57">Percents, head length</emphasis>
</th>
</tr>
<tr id="356F349AD76CC0CB29CA5ACCFAA0E071" box="[167,1334,1781,1804]" gridrow="15" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5ACCFE5FE071" box="[167,457,1781,1804]" gridcol="0" gridrow="15" pageId="4" pageNumber="57">Head depth</th>
<td id="76BE5DE6D76CC0CB2BE45ACCFB98E071" box="[649,1038,1781,1804]" gridcol="1" gridrow="15" pageId="4" pageNumber="57">13.9 12.0 – 15.6</td>
<td id="76BE5DE6D76CC0CB2DB35ACCFAA0E071" box="[1246,1334,1781,1804]" gridcol="2" gridrow="15" pageId="4" pageNumber="57">13.8</td>
</tr>
<tr id="356F349AD76CC0CB29CA5B1AFAA0E047" box="[167,1334,1827,1850]" gridrow="16" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5B1AFE5FE047" box="[167,457,1827,1850]" gridcol="0" gridrow="16" pageId="4" pageNumber="57">Head width</th>
<td id="76BE5DE6D76CC0CB2BE45B1AFB98E047" box="[649,1038,1827,1850]" gridcol="1" gridrow="16" pageId="4" pageNumber="57">24.2 21.7 – 25.5</td>
<td id="76BE5DE6D76CC0CB2DB35B1AFAA0E047" box="[1246,1334,1827,1850]" gridcol="2" gridrow="16" pageId="4" pageNumber="57">23.9</td>
</tr>
<tr id="356F349AD76CC0CB29CA5B68FAA0E015" box="[167,1334,1873,1896]" gridrow="17" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5B68FE5FE015" box="[167,457,1873,1896]" gridcol="0" gridrow="17" pageId="4" pageNumber="57">Interorbital width</th>
<td id="76BE5DE6D76CC0CB2BE45B68FB98E015" box="[649,1038,1873,1896]" gridcol="1" gridrow="17" pageId="4" pageNumber="57">7.4 6.8 – 8.2</td>
<td id="76BE5DE6D76CC0CB2DB35B68FAA0E015" box="[1246,1334,1873,1896]" gridcol="2" gridrow="17" pageId="4" pageNumber="57">7.6</td>
</tr>
<tr id="356F349AD76CC0CB29CA5B47FAA0E0E8" box="[167,1334,1918,1941]" gridrow="18" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5B47FE5FE0E8" box="[167,457,1918,1941]" gridcol="0" gridrow="18" pageId="4" pageNumber="57">Preorbital length</th>
<td id="76BE5DE6D76CC0CB2BE45B47FB98E0E8" box="[649,1038,1918,1941]" gridcol="1" gridrow="18" pageId="4" pageNumber="57">9.5 9.4 – 10.2</td>
<td id="76BE5DE6D76CC0CB2DB35B47FAA0E0E8" box="[1246,1334,1918,1941]" gridcol="2" gridrow="18" pageId="4" pageNumber="57">9.8</td>
</tr>
<tr id="356F349AD76CC0CB29CA5B95FAA0E0BE" box="[167,1334,1964,1987]" gridrow="19" pageId="4" pageNumber="57">
<th id="76BE5DE6D76CC0CB29CA5B95FE5FE0BE" box="[167,457,1964,1987]" gridcol="0" gridrow="19" pageId="4" pageNumber="57">Eye diameter</th>
<td id="76BE5DE6D76CC0CB2BE45B95FB98E0BE" box="[649,1038,1964,1987]" gridcol="1" gridrow="19" pageId="4" pageNumber="57">2.4 2.3 – 2.7</td>
<td id="76BE5DE6D76CC0CB2DB35B95FAA0E0BE" box="[1246,1334,1964,1987]" gridcol="2" gridrow="19" pageId="4" pageNumber="57">2.5</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3456553D76D3F3129AB5CAEFF70E7A7" pageId="5" pageNumber="58" type="distribution">
<paragraph id="8BE036D8D76D3F3129AB5CAEFF70E7A7" blockId="5.[151,1436,151,258]" pageId="5" pageNumber="58">
<emphasis id="B92BEACAD76D3F3129AB5CAEFEFFE7CC" bold="true" box="[198,361,151,177]" pageId="5" pageNumber="58">Distribution:</emphasis>
Known only from the 
<typeStatus id="54E4887AD76D3F312B195CA1FD31E7CF" box="[628,679,152,178]" pageId="5" pageNumber="58">type</typeStatus>
locality, rio Paraguaçu near Iassu, Estado da Bahia, northeastern 
<collectingCountry id="F3487648D76D3F3129FA5CF9FF75E7A7" box="[151,227,192,218]" name="Brazil" pageId="5" pageNumber="58">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3456553D76D3F3129AB5CDEFB16E67F" box="[198,1152,231,258]" pageId="5" pageNumber="58" type="etymology">
<paragraph id="8BE036D8D76D3F3129AB5CDEFB16E67F" blockId="5.[151,1436,151,258]" box="[198,1152,231,258]" pageId="5" pageNumber="58">
<emphasis id="B92BEACAD76D3F3129AB5CDEFEC1E67C" bold="true" box="[198,343,231,257]" pageId="5" pageNumber="58">Etymology:</emphasis>
The name 
<taxonomicName id="4C5F4D5BD76D3F3128B75CD1FD08E67C" box="[474,670,232,257]" class="Actinopterygii" family="Trichomycteridae" genus="Ituglanis" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="paraguassuensis">
<emphasis id="B92BEACAD76D3F3128B75CD1FD08E67C" box="[474,670,232,257]" italics="true" pageId="5" pageNumber="58">paraguassuensis</emphasis>
</taxonomicName>
refers to the 
<typeStatus id="54E4887AD76D3F312A555CD1FCFDE67F" box="[824,875,232,258]" pageId="5" pageNumber="58">type</typeStatus>
locality, rio Paraguaçu.
</paragraph>
</subSubSection>
</treatment>
</document>