316 lines
63 KiB
XML
316 lines
63 KiB
XML
<document id="58630A197B696EBD5E23A08FD5B82D04" ID-CLB-Dataset="36382" ID-DOI="10.11646/zootaxa.3828.1.1" ID-GBIF-Dataset="8c1cda9b-0d82-41f3-a1c1-d8eb574a1ba7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286513" ID-ZooBank="B2FEAE60-7E51-45FA-A38F-930A084A5AAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452080076" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="03F68791FFD1180F8FFE3876025EFFD1" docLanguage="en" docName="zt03828p076.pdf" docOrigin="Zootaxa 3828 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Horaeomorphus Schaufuss" docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="FFCFFFE9FFD718088F693A1C034CFC3C" masterDocTitle="' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="7" updateTime="1698338417320" updateUser="ExternalLinkService">
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<mods:title id="FAD4DB010EC35E4A40D1DD984EA54F1A">' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="801AAF406624677B6E24C0EC92328D00">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="FE7FD6BAEF9BF88477BF572C14E97210">Zootaxa</mods:title>
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<mods:date id="871B36134423AB8CC6EFCE8C984ABCD8">2014</mods:date>
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<mods:number id="644C707347E912E887C6F16EF81F08B1">3828</mods:number>
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<treatment id="03F68791FFD1180F8FFE3876025EFFD1" ID-DOI="http://doi.org/10.5281/zenodo.6141050" ID-GBIF-Taxon="127711769" ID-Zenodo-Dep="6141050" LSID="urn:lsid:plazi:treatment:03F68791FFD1180F8FFE3876025EFFD1" httpUri="http://treatment.plazi.org/id/03F68791FFD1180F8FFE3876025EFFD1" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<subSubSection id="C345650CFFD1180E8FFE387602AEFEB8" box="[151,482,618,644]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="8BE03687FFD1180E8FFE387602AEFEB8" blockId="6.[151,482,618,644]" box="[151,482,618,644]" pageId="6" pageNumber="7">
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<heading id="D0A881EBFFD1180E8FFE387602AEFEB8" bold="true" box="[151,482,618,644]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
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<taxonomicName id="4C5F4D04FFD1180E8FFE387602AEFEB8" ID-CoL="8XLVM" authority="Schaufuss" authorityName="Schaufuss" box="[151,482,618,644]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="B92BEA95FFD1180E8FFE387602AEFEB8" bold="true" box="[151,482,618,644]" pageId="6" pageNumber="7">
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<emphasis id="B92BEA95FFD1180E8FFE38760211FEB8" bold="true" box="[151,349,618,644]" italics="true" pageId="6" pageNumber="7">Horaeomorphus</emphasis>
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Schaufuss
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</emphasis>
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||
</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C345650CFFD1180E8FFE38AD0015FAE1" pageId="6" pageNumber="7" type="reference_group">
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<paragraph id="8BE03687FFD1180E8FFE38AD0761FF7E" blockId="6.[151,1436,689,895]" pageId="6" pageNumber="7">
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<treatmentCitationGroup id="AB4F11A9FFD1180E8FFE38AD0761FF7E" infered="true" pageId="6" pageNumber="7">
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<taxonomicName id="4C5F4D04FFD1180E8FFE38AD015EFEFB" ID-CoL="8XLVM" authority="Schaufuss, 1889: 21" authorityName="Schaufuss" authorityPageNumber="21" authorityYear="1889" box="[151,530,689,711]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="B92BEA95FFD1180E8FFE38AD0271FEFB" box="[151,317,689,711]" italics="true" pageId="6" pageNumber="7">Horaeomorphus</emphasis>
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<treatmentCitation id="0AFE1096FFD1180E8E2A38AD015EFEFB" author="Schaufuss" box="[323,530,689,711]" page="21" pageId="6" pageNumber="7" year="1889">
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<bibRefCitation id="EFCE4B76FFD1180E8E2A38AD02A0FEFB" author="Schaufuss" box="[323,492,689,711]" pageId="6" pageNumber="7" refString="Schaufuss, L. W. (1889) Neue Scydmaeniden im Museum Ludwig Salvator. Berliner Entomologische Zeitschrift, 33, 1 - 42. http: // dx. doi. org / 10.1002 / mmnd. 18890330103" type="journal article" year="1889">Schaufuss, 1889</bibRefCitation>
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: 21
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</treatmentCitation>
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||
</taxonomicName>
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.
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<typeStatus id="54E48825FFD1180E8D7538AE0102FEFB" box="[540,590,690,711]" pageId="6" pageNumber="7">Type</typeStatus>
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species:
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<taxonomicName id="4C5F4D04FFD1180E8DC238AD0733FEFB" ID-CoL="3MNJX" authority="Schaufuss, 1889" authorityName="Schaufuss" authorityYear="1889" box="[683,1151,689,711]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="eumicroides">
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<emphasis id="B92BEA95FFD1180E8DC238AD009FFEFB" box="[683,979,689,711]" italics="true" pageId="6" pageNumber="7">Horaeomorphus eumicroides</emphasis>
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<bibRefCitation id="EFCE4B76FFD1180E8CB038AD0733FEFB" author="Schaufuss" box="[985,1151,689,711]" pageId="6" pageNumber="7" refString="Schaufuss, L. W. (1889) Neue Scydmaeniden im Museum Ludwig Salvator. Berliner Entomologische Zeitschrift, 33, 1 - 42. http: // dx. doi. org / 10.1002 / mmnd. 18890330103" type="journal article" year="1889">Schaufuss, 1889</bibRefCitation>
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</taxonomicName>
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(orig. des.).
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<taxonomicName id="4C5F4D04FFD1180E8FFE38CC02BDFEDA" authority="Franz, 1975: 175" authorityName="Franz" authorityPageNumber="175" authorityYear="1975" box="[151,497,720,742]" class="Insecta" family="Scydmaenidae" genus="Anthicimorphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="B92BEA95FFD1180E8FFE38CC0275FEDA" box="[151,313,720,742]" italics="true" pageId="6" pageNumber="7">Anthicimorphus</emphasis>
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<treatmentCitation id="0AFE1096FFD1180E8E2838CD02BDFEDA" author="Franz" box="[321,497,720,742]" page="175" pageId="6" pageNumber="7" year="1975">
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||
<bibRefCitation id="EFCE4B76FFD1180E8E2838CD02F3FEDA" author="Franz" box="[321,447,720,742]" pageId="6" pageNumber="7" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz, 1975</bibRefCitation>
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: 175
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</treatmentCitation>
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</taxonomicName>
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<emphasis id="B92BEA95FFD1180E8E9338CC010EFED9" bold="true" box="[506,578,720,741]" pageId="6" pageNumber="7">syn. n.</emphasis>
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(as subgenus of
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<taxonomicName id="4C5F4D04FFD1180E8D9C38CD0015FEDA" box="[757,857,721,742]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="B92BEA95FFD1180E8D9C38CD0015FEDA" box="[757,857,721,742]" italics="true" pageId="6" pageNumber="7">Euconnus</emphasis>
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</taxonomicName>
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||
; not as new).
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<typeStatus id="54E48825FFD1180E8C9838CD076FFEDA" box="[1009,1059,721,742]" pageId="6" pageNumber="7">Type</typeStatus>
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species:
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<taxonomicName id="4C5F4D04FFD1180E8BED38CC0260FF38" ID-CoL="4FQ9W" authority="Lea, 1915" authorityName="Lea" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="anthicoides">
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<emphasis id="B92BEA95FFD1180E8BED38CC06D0FEDA" box="[1156,1436,720,742]" italics="true" pageId="6" pageNumber="7">Phagonophana anthicoides</emphasis>
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Lea, 1915
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</taxonomicName>
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(monotypy). Also spelled
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<emphasis id="B92BEA95FFD1180E8D5038F20182FF38" box="[569,718,750,772]" italics="true" pageId="6" pageNumber="7">Anticimorphus</emphasis>
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(
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<treatmentCitation id="0AFE1096FFD1180E8DB338F200C8FF38" author="Franz" box="[730,900,750,772]" page="303" pageId="6" pageNumber="7" year="1975">
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||
<bibRefCitation id="EFCE4B76FFD1180E8DB338F200C8FF38" author="Franz" box="[730,900,750,772]" pageId="6" pageNumber="7" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz 1975: 303</bibRefCitation>
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||
</treatmentCitation>
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), selected as incorrect original spelling by Newton & Franz (1998).
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<bibRefCitation id="EFCE4B76FFD1180E8E3F39120295FF1F" author="Franz" box="[342,473,781,803]" pageId="6" pageNumber="7" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz (1975)</bibRefCitation>
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referred to Madagascan
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<taxonomicName id="4C5F4D04FFD1180E8DBA39110039FF1F" box="[723,885,781,803]" class="Insecta" family="Scydmaenidae" genus="Anthicimorphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="B92BEA95FFD1180E8DBA39110039FF1F" box="[723,885,781,803]" italics="true" pageId="6" pageNumber="7">Anthicimorphus</emphasis>
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</taxonomicName>
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, evidently having in mind a paper that has appeared in print much later (
|
||
<bibRefCitation id="EFCE4B76FFD1180E8E1F393002B5FF7E" author="Franz" box="[374,505,812,834]" pageId="6" pageNumber="7" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986" yearSuffix="b">Franz 1986b</bibRefCitation>
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), but presumably was supposed to be published as first.
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BE03687FFD1180E8FFE39560132FF43" blockId="6.[151,1436,689,895]" pageId="6" pageNumber="7">
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||
<treatmentCitationGroup id="AB4F11A9FFD1180E8FFE39560132FF43" pageId="6" pageNumber="7">
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||
<taxonomicName id="4C5F4D04FFD1180E8FFE3956014EFF5C" authority="Franz, 1986b: 234" authorityName="Franz" authorityPageNumber="234" authorityYear="1986" box="[151,514,842,864]" class="Insecta" family="Scydmaenidae" genus="Anthicimorphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="B92BEA95FFD1180E8FFE39560275FF5C" box="[151,313,842,864]" italics="true" pageId="6" pageNumber="7">Anthicimorphus</emphasis>
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||
<treatmentCitation id="0AFE1096FFD1180E8E2B3957014EFF5C" author="Franz" box="[322,514,842,864]" page="234" pageId="6" pageNumber="7" year="1986">
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||
<bibRefCitation id="EFCE4B76FFD1180E8E2B39570282FF5C" author="Franz" box="[322,462,842,864]" pageId="6" pageNumber="7" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986" yearSuffix="b">Franz, 1986b</bibRefCitation>
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: 234
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</treatmentCitation>
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||
</taxonomicName>
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(as a new subgenus of
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<taxonomicName id="4C5F4D04FFD1180E8C6A3957002BFF5C" box="[771,871,843,864]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92BEA95FFD1180E8C6A3957002BFF5C" box="[771,871,843,864]" italics="true" pageId="6" pageNumber="7">Euconnus</emphasis>
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||
</taxonomicName>
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||
).
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<typeStatus id="54E48825FFD1180E8C16395700FDFF5C" box="[895,945,843,864]" pageId="6" pageNumber="7">Type</typeStatus>
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||
species:
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<taxonomicName id="4C5F4D04FFD1180E8B7D395706D0FF5C" authority="Franz, 1986" authorityName="Franz" authorityYear="1986" box="[1044,1436,842,864]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="anthiciformis">
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||
<emphasis id="B92BEA95FFD1180E8B7D39570645FF5C" box="[1044,1289,842,864]" italics="true" pageId="6" pageNumber="7">Euconnus anthiciformis</emphasis>
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||
<bibRefCitation id="EFCE4B76FFD1180E8A7A395706D0FF5C" author="Franz" box="[1299,1436,842,864]" pageId="6" pageNumber="7" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986" yearSuffix="b">Franz, 1986b</bibRefCitation>
|
||
</taxonomicName>
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||
(preoccupied, nec
|
||
<bibRefCitation id="EFCE4B76FFD1180E8EE8397602B7FF43" author="Franz" box="[385,507,873,895]" pageId="6" pageNumber="7" refString="Franz, H. (1967 b) Zur Kenntnis der Scydmaenidenfauna von Lateinamerika. I Teil: Scydmaeniden aus Chile und dem sudlichen Argentinien. II Teil: Scydmaeniden aus dem subtropischen und tropischen Amerika. Biologie de l'Amerique Australe, 3, 611 - 724." type="journal article" year="1967">Franz, 1967</bibRefCitation>
|
||
) (des. orig.).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFD1180E8FFE39B00015FAE1" blockId="6.[151,1437,940,2009]" pageId="6" pageNumber="7">
|
||
<emphasis id="B92BEA95FFD1180E8FFE39B0023EFFF9" bold="true" box="[151,370,940,965]" pageId="6" pageNumber="7">Revised diagnosis.</emphasis>
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Body (
|
||
<figureCitation id="13642A02FFD1180E8EA139B00179FFF9" box="[456,565,940,965]" captionStart="FIGURES 1 – 5" captionStartId="7.[151,264,1767,1790]" captionTargetBox="[157,1436,1065,1740]" captionTargetId="figure@7.[151,1436,1060,1747]" captionTargetPageId="7" captionText="FIGURES 1 – 5. Examples of body form in Horaeomorphus, males. Horaeomorphus valdepunctatus Franz from Malay Peninsula (1); H. antennatus Jałoszyński from Malay Peninsula (2); H. ishiianus Jałoszyński from Borneo (3); H. punctatissimus Franz from Borneo (4), H. calcarifer (Franz) from Fiji (photograph from Jałoszyński 2012 a) (5)." httpUri="https://zenodo.org/record/286514/files/figure.png" pageId="6" pageNumber="7">Figs. 1–5</figureCitation>
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||
,
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||
<figureCitation id="13642A02FFD1180E8D2A39B001ADFFF9" box="[579,737,940,965]" captionStart="FIGURES 22 – 28" captionStartId="14.[151,264,1902,1925]" captionTargetBox="[216,1351,198,1842]" captionTargetId="figure@14.[196,1359,193,1852]" captionTargetPageId="14" captionText="FIGURES 22 – 28. Horaeomorphus spp. Dorsal habitus (22 – 23, 26 – 28), original set of labels of lectotype (24) and paralectotype (25). Horaeomorphus anthicoides (Lea) comb. n., lectotype male (22), paralectotype female (23) and their labels, respectively (24 – 25); Horaeomorphus verus sp. n., holotype male (26); female of Horaeomorphus sp. from Royal National Park (27); and female of Horaeomorphus sp. from Blue Mountains (28)." httpUri="https://zenodo.org/record/286519/files/figure.png" pageId="6" pageNumber="7">22–23, 26–28</figureCitation>
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||
) strongly elongate; head (
|
||
<figureCitation id="13642A02FFD1180E8B6239B00734FFF9" box="[1035,1144,940,965]" captionStart="FIGURES 1 – 5" captionStartId="7.[151,264,1767,1790]" captionTargetBox="[157,1436,1065,1740]" captionTargetId="figure@7.[151,1436,1060,1747]" captionTargetPageId="7" captionText="FIGURES 1 – 5. Examples of body form in Horaeomorphus, males. Horaeomorphus valdepunctatus Franz from Malay Peninsula (1); H. antennatus Jałoszyński from Malay Peninsula (2); H. ishiianus Jałoszyński from Borneo (3); H. punctatissimus Franz from Borneo (4), H. calcarifer (Franz) from Fiji (photograph from Jałoszyński 2012 a) (5)." httpUri="https://zenodo.org/record/286514/files/figure.png" pageId="6" pageNumber="7">Figs. 1–5</figureCitation>
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,
|
||
<figureCitation id="13642A02FFD1180E8BEF39B0066FFFF9" box="[1158,1315,940,965]" captionStart="FIGURES 22 – 28" captionStartId="14.[151,264,1902,1925]" captionTargetBox="[216,1351,198,1842]" captionTargetId="figure@14.[196,1359,193,1852]" captionTargetPageId="14" captionText="FIGURES 22 – 28. Horaeomorphus spp. Dorsal habitus (22 – 23, 26 – 28), original set of labels of lectotype (24) and paralectotype (25). Horaeomorphus anthicoides (Lea) comb. n., lectotype male (22), paralectotype female (23) and their labels, respectively (24 – 25); Horaeomorphus verus sp. n., holotype male (26); female of Horaeomorphus sp. from Royal National Park (27); and female of Horaeomorphus sp. from Blue Mountains (28)." httpUri="https://zenodo.org/record/286519/files/figure.png" pageId="6" pageNumber="7">22–23, 26–28</figureCitation>
|
||
) distinctly flattened, with vertex not projecting dorsocaudad, its posterior margin arcuate and distinctly concave; supraantennal tubercles prominent and in most cases accompanied posteriorly by a pair of shallow but distinct pits; frontoclypeal suture or groove absent; eyes located anteriorly and therefore tempora long and strongly curved toward occipital constriction; tempora, genae and postgenae with sparse and short thick bristles mostly located lateroventrally; 'neck region' (
|
||
<figureCitation id="13642A02FFD1180E8E803E430161F844" box="[489,557,1119,1144]" captionStart="FIGURES 6 – 9" captionStartId="8.[151,264,1678,1701]" captionTargetBox="[151,1426,193,1642]" captionTargetId="figure@8.[151,1436,193,1657]" captionTargetPageId="8" captionText="FIGURES 6 – 9. Morphology of Horaeomorphus sakishimanus Jałoszyński. Head capsule in ventral view (6), labrum in dorsal view (7), right mandible in dorsal view (8), and left maxilla in ventral view (9). Abbreviations: bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lp, labial palpus; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286515/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
|
||
) distinctly demarcated from anterior part of head capsule by constriction and much narrower than vertex; hypostomal ridges (
|
||
<figureCitation id="13642A02FFD1180E8DDC3E9801B5F8A1" box="[693,761,1156,1181]" captionStart="FIGURES 6 – 9" captionStartId="8.[151,264,1678,1701]" captionTargetBox="[151,1426,193,1642]" captionTargetId="figure@8.[151,1436,193,1657]" captionTargetPageId="8" captionText="FIGURES 6 – 9. Morphology of Horaeomorphus sakishimanus Jałoszyński. Head capsule in ventral view (6), labrum in dorsal view (7), right mandible in dorsal view (8), and left maxilla in ventral view (9). Abbreviations: bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lp, labial palpus; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286515/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8C6E3E98006CF8A1" box="[775,800,1156,1181]" italics="true" pageId="6" pageNumber="7">hr</emphasis>
|
||
) extending from cardines to posterior tentorial pits (
|
||
<figureCitation id="13642A02FFD1180E8A073E9803EBF8FC" captionStart="FIGURES 6 – 9" captionStartId="8.[151,264,1678,1701]" captionTargetBox="[151,1426,193,1642]" captionTargetId="figure@8.[151,1436,193,1657]" captionTargetPageId="8" captionText="FIGURES 6 – 9. Morphology of Horaeomorphus sakishimanus Jałoszyński. Head capsule in ventral view (6), labrum in dorsal view (7), right mandible in dorsal view (8), and left maxilla in ventral view (9). Abbreviations: bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lp, labial palpus; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286515/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8FD13EB50390F8FC" box="[184,220,1193,1216]" italics="true" pageId="6" pageNumber="7">ptp</emphasis>
|
||
), which are elongate and located on anterior margin of transverse groove demarcating 'neck' ventrally; submentum (
|
||
<figureCitation id="13642A02FFD1180E8E433ED00222F8D9" box="[298,366,1228,1253]" captionStart="FIGURES 6 – 9" captionStartId="8.[151,264,1678,1701]" captionTargetBox="[151,1426,193,1642]" captionTargetId="figure@8.[151,1436,193,1657]" captionTargetPageId="8" captionText="FIGURES 6 – 9. Morphology of Horaeomorphus sakishimanus Jałoszyński. Head capsule in ventral view (6), labrum in dorsal view (7), right mandible in dorsal view (8), and left maxilla in ventral view (9). Abbreviations: bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lp, labial palpus; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286515/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8E153ED102E5F8D8" box="[380,425,1229,1252]" italics="true" pageId="6" pageNumber="7">smn</emphasis>
|
||
) without lateral sutures; each mandible with two subapical teeth (
|
||
<figureCitation id="13642A02FFD1180E8BFB3ED0079BF8D8" box="[1170,1239,1228,1253]" captionStart="FIGURES 6 – 9" captionStartId="8.[151,264,1678,1701]" captionTargetBox="[151,1426,193,1642]" captionTargetId="figure@8.[151,1436,193,1657]" captionTargetPageId="8" captionText="FIGURES 6 – 9. Morphology of Horaeomorphus sakishimanus Jałoszyński. Head capsule in ventral view (6), labrum in dorsal view (7), right mandible in dorsal view (8), and left maxilla in ventral view (9). Abbreviations: bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lp, labial palpus; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286515/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8B8D3ED10649F8D8" box="[1252,1285,1229,1252]" italics="true" pageId="6" pageNumber="7">sat</emphasis>
|
||
) and densely setose submedian mesal prostheca (
|
||
<figureCitation id="13642A02FFD1180E8D4C3EF30126F934" box="[549,618,1263,1288]" captionStart="FIGURES 6 – 9" captionStartId="8.[151,264,1678,1701]" captionTargetBox="[151,1426,193,1642]" captionTargetId="figure@8.[151,1436,193,1657]" captionTargetPageId="8" captionText="FIGURES 6 – 9. Morphology of Horaeomorphus sakishimanus Jałoszyński. Head capsule in ventral view (6), labrum in dorsal view (7), right mandible in dorsal view (8), and left maxilla in ventral view (9). Abbreviations: bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lp, labial palpus; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286515/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8D1F3EED01DAF934" box="[630,662,1265,1288]" italics="true" pageId="6" pageNumber="7">pst</emphasis>
|
||
); maxillary palpomeres III and IV (
|
||
<figureCitation id="13642A02FFD1180E8B413EF30721F934" box="[1064,1133,1263,1288]" captionStart="FIGURES 6 – 9" captionStartId="8.[151,264,1678,1701]" captionTargetBox="[151,1426,193,1642]" captionTargetId="figure@8.[151,1436,193,1657]" captionTargetPageId="8" captionText="FIGURES 6 – 9. Morphology of Horaeomorphus sakishimanus Jałoszyński. Head capsule in ventral view (6), labrum in dorsal view (7), right mandible in dorsal view (8), and left maxilla in ventral view (9). Abbreviations: bst, basistipes; cd, cardo; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lp, labial palpus; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286515/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8B133EEC079EF934" box="[1146,1234,1264,1288]" italics="true" pageId="6" pageNumber="7">mxp3–4</emphasis>
|
||
) slender, strongly elongate; antennae (
|
||
<figureCitation id="13642A02FFD1180E8E153F0802A5F911" box="[380,489,1300,1325]" captionStart="FIGURES 1 – 5" captionStartId="7.[151,264,1767,1790]" captionTargetBox="[157,1436,1065,1740]" captionTargetId="figure@7.[151,1436,1060,1747]" captionTargetPageId="7" captionText="FIGURES 1 – 5. Examples of body form in Horaeomorphus, males. Horaeomorphus valdepunctatus Franz from Malay Peninsula (1); H. antennatus Jałoszyński from Malay Peninsula (2); H. ishiianus Jałoszyński from Borneo (3); H. punctatissimus Franz from Borneo (4), H. calcarifer (Franz) from Fiji (photograph from Jałoszyński 2012 a) (5)." httpUri="https://zenodo.org/record/286514/files/figure.png" pageId="6" pageNumber="7">Figs. 1–5</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFD1180E8E913F0801DAF911" box="[504,662,1300,1325]" captionStart="FIGURES 22 – 28" captionStartId="14.[151,264,1902,1925]" captionTargetBox="[216,1351,198,1842]" captionTargetId="figure@14.[196,1359,193,1852]" captionTargetPageId="14" captionText="FIGURES 22 – 28. Horaeomorphus spp. Dorsal habitus (22 – 23, 26 – 28), original set of labels of lectotype (24) and paralectotype (25). Horaeomorphus anthicoides (Lea) comb. n., lectotype male (22), paralectotype female (23) and their labels, respectively (24 – 25); Horaeomorphus verus sp. n., holotype male (26); female of Horaeomorphus sp. from Royal National Park (27); and female of Horaeomorphus sp. from Blue Mountains (28)." httpUri="https://zenodo.org/record/286519/files/figure.png" pageId="6" pageNumber="7">22–23, 26–28</figureCitation>
|
||
) gradually thickening distally; pronotum (
|
||
<figureCitation id="13642A02FFD1180E8B123F0807A4F911" box="[1147,1256,1300,1325]" captionStart="FIGURES 1 – 5" captionStartId="7.[151,264,1767,1790]" captionTargetBox="[157,1436,1065,1740]" captionTargetId="figure@7.[151,1436,1060,1747]" captionTargetPageId="7" captionText="FIGURES 1 – 5. Examples of body form in Horaeomorphus, males. Horaeomorphus valdepunctatus Franz from Malay Peninsula (1); H. antennatus Jałoszyński from Malay Peninsula (2); H. ishiianus Jałoszyński from Borneo (3); H. punctatissimus Franz from Borneo (4), H. calcarifer (Franz) from Fiji (photograph from Jałoszyński 2012 a) (5)." httpUri="https://zenodo.org/record/286514/files/figure.png" pageId="6" pageNumber="7">Figs. 1–5</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFD1180E8B9E3F0806D9F911" box="[1271,1429,1300,1325]" captionStart="FIGURES 22 – 28" captionStartId="14.[151,264,1902,1925]" captionTargetBox="[216,1351,198,1842]" captionTargetId="figure@14.[196,1359,193,1852]" captionTargetPageId="14" captionText="FIGURES 22 – 28. Horaeomorphus spp. Dorsal habitus (22 – 23, 26 – 28), original set of labels of lectotype (24) and paralectotype (25). Horaeomorphus anthicoides (Lea) comb. n., lectotype male (22), paralectotype female (23) and their labels, respectively (24 – 25); Horaeomorphus verus sp. n., holotype male (26); female of Horaeomorphus sp. from Royal National Park (27); and female of Horaeomorphus sp. from Blue Mountains (28)." httpUri="https://zenodo.org/record/286519/files/figure.png" pageId="6" pageNumber="7">22–23, 26–28</figureCitation>
|
||
) with odd (three or five) number of antebasal pits (
|
||
<figureCitation id="13642A02FFD1180E8DB93F2B0069F96C" box="[720,805,1335,1360]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8C5B3F2B0004F96C" box="[818,840,1335,1360]" italics="true" pageId="6" pageNumber="7">lp</emphasis>
|
||
,
|
||
<emphasis id="B92BEA95FFD1180E8C3E3F250035F96C" box="[855,889,1337,1360]" italics="true" pageId="6" pageNumber="7">mp</emphasis>
|
||
) often connected by transverse groove (
|
||
<figureCitation id="13642A02FFD1180E8A2A3F2B06DBF96C" box="[1347,1431,1335,1360]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8FFE3F4103E1F948" box="[151,173,1373,1396]" italics="true" pageId="6" pageNumber="7">tg</emphasis>
|
||
); prosternum laterally demarcated from hypomera by pronotosternal sutures (
|
||
<figureCitation id="13642A02FFD1180E8B303F4007FEF948" box="[1113,1202,1372,1397]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8BAF3F4107A6F948" box="[1222,1258,1373,1396]" italics="true" pageId="6" pageNumber="7">nss</emphasis>
|
||
), with distinct prosternal process (
|
||
<figureCitation id="13642A02FFD1180E8E103F630282F9A4" box="[377,462,1407,1432]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8EB63F9D0149F9A4" box="[479,517,1409,1432]" italics="true" pageId="6" pageNumber="7">psp</emphasis>
|
||
) developed as low and narrow carina; procoxal sockets (
|
||
<figureCitation id="13642A02FFD1180E8BCE3F6307B1F9A4" box="[1191,1277,1407,1432]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8A643F9D067EF9A4" box="[1293,1330,1409,1432]" italics="true" pageId="6" pageNumber="7">pcs</emphasis>
|
||
) broadly closed; prothoracic hypomera (
|
||
<figureCitation id="13642A02FFD1180E8E953FB8011CF980" box="[508,592,1444,1469]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8D093FB80136F981" box="[608,634,1444,1469]" italics="true" pageId="6" pageNumber="7">hy</emphasis>
|
||
) with large internal (adcoxal) parts demarcated laterally by complete hypomeral ridges (
|
||
<figureCitation id="13642A02FFD1180E8E063FDB0288F9DC" box="[367,452,1479,1504]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8EBB3FDB02BBF9DC" box="[466,503,1479,1504]" italics="true" pageId="6" pageNumber="7">hyr</emphasis>
|
||
); sides of pronotum (
|
||
<figureCitation id="13642A02FFD1180E8D9D3FDB000BF9DC" box="[756,839,1479,1504]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
|
||
) with short and sparse thick bristles mostly located lateroventrally; mesoventrite with strikingly long asetose and reticulated anterior ridge (
|
||
<figureCitation id="13642A02FFD1180E8B1C3FF00785FA39" box="[1141,1225,1516,1541]" captionStart="FIGURES 15 – 17" captionStartId="10.[151,264,1238,1261]" captionTargetBox="[151,1436,193,1217]" captionTargetId="figure@10.[151,1436,193,1218]" captionTargetPageId="10" captionText="FIGURES 15 – 17. Morphology of Horaeomorphus sakishimanus Jałoszyński. Mesothorax in ventral view (15), pterothorax in left lateral view (16), base of elytron in dorsal view, dotted line showing portion of inversely U-shaped groove (17). Abbreviations: aest 2 – 3, meso- and metanepisternum; ar, anterior ridge; bef, basal elytral fovea; cx 3, metacoxa; eal, elytral articulating lobe; epm 2 – 3, mes- and metepimeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msvp, mesoventral process; pre, prepectus; v 3, metaventrite." httpUri="https://zenodo.org/record/286517/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8BBC3FF107A1FA38" box="[1237,1261,1517,1540]" italics="true" pageId="6" pageNumber="7">ar</emphasis>
|
||
) with posterior median projection; mesoventral process (
|
||
<figureCitation id="13642A02FFD1180E8D043C13018EFA14" box="[621,706,1551,1576]" captionStart="FIGURES 15 – 17" captionStartId="10.[151,264,1238,1261]" captionTargetBox="[151,1436,193,1217]" captionTargetId="figure@10.[151,1436,193,1218]" captionTargetPageId="10" captionText="FIGURES 15 – 17. Morphology of Horaeomorphus sakishimanus Jałoszyński. Mesothorax in ventral view (15), pterothorax in left lateral view (16), base of elytron in dorsal view, dotted line showing portion of inversely U-shaped groove (17). Abbreviations: aest 2 – 3, meso- and metanepisternum; ar, anterior ridge; bef, basal elytral fovea; cx 3, metacoxa; eal, elytral articulating lobe; epm 2 – 3, mes- and metepimeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msvp, mesoventral process; pre, prepectus; v 3, metaventrite." httpUri="https://zenodo.org/record/286517/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8DB93C0D0046FA14" box="[720,778,1553,1576]" italics="true" pageId="6" pageNumber="7">msvp</emphasis>
|
||
) narrow and weakly expanding ventrally but distinct and clearly separating mesocoxae; ventro- and dorsolateral foveae absent; anterior metaventral process absent; metaventral intercoxal process (
|
||
<figureCitation id="13642A02FFD1180E8E943C4B01CAFA4C" box="[509,646,1623,1648]" captionStart="FIGURES 18 – 21" captionStartId="12.[151,264,1461,1484]" captionTargetBox="[151,1436,193,1440]" captionTargetId="figure@12.[151,1436,193,1441]" captionTargetPageId="12" captionText="FIGURES 18 – 21. Morphology of Horaeomorphus sakishimanus Jałoszyński. Pterotorax in ventral view (18 – 19), distal part of right metacoxa, metatrochanter and proximal portion of metafemur in ventral view (20), aedeagus in ventral view (21). Abbreviations: aml, apex of median lobe; ar, anterior ridge; cx 3, metacoxa; fm 3, metafemur; lmfa, lateral metafurcal arm; ma, membranous area; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pm, paramere; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vw, ventral wall." httpUri="https://zenodo.org/record/286518/files/figure.png" pageId="6" pageNumber="7">Figs. 18–19</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8DFB3C450184FA4C" box="[658,712,1625,1648]" italics="true" pageId="6" pageNumber="7">mtvp</emphasis>
|
||
) narrowly separating metacoxae, with two long spines and deep median notch; each elytron with two distinct and asetose basal foveae (
|
||
<figureCitation id="13642A02FFD1180E8CBA3C600765FAA8" box="[979,1065,1660,1685]" captionStart="FIGURES 15 – 17" captionStartId="10.[151,264,1238,1261]" captionTargetBox="[151,1436,193,1217]" captionTargetId="figure@10.[151,1436,193,1218]" captionTargetPageId="10" captionText="FIGURES 15 – 17. Morphology of Horaeomorphus sakishimanus Jałoszyński. Mesothorax in ventral view (15), pterothorax in left lateral view (16), base of elytron in dorsal view, dotted line showing portion of inversely U-shaped groove (17). Abbreviations: aest 2 – 3, meso- and metanepisternum; ar, anterior ridge; bef, basal elytral fovea; cx 3, metacoxa; eal, elytral articulating lobe; epm 2 – 3, mes- and metepimeron; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msvp, mesoventral process; pre, prepectus; v 3, metaventrite." httpUri="https://zenodo.org/record/286517/files/figure.png" pageId="6" pageNumber="7">Fig. 17</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8B5E3C600715FAA9" box="[1079,1113,1660,1685]" italics="true" pageId="6" pageNumber="7">bef</emphasis>
|
||
) connected by inversely Ushaped groove located on basal articulating lobe of elytron; aedeagus (
|
||
<figureCitation id="13642A02FFD1180E8CBE3C830775FA84" box="[983,1081,1695,1720]" captionStart="FIGURES 18 – 21" captionStartId="12.[151,264,1461,1484]" captionTargetBox="[151,1436,193,1440]" captionTargetId="figure@12.[151,1436,193,1441]" captionTargetPageId="12" captionText="FIGURES 18 – 21. Morphology of Horaeomorphus sakishimanus Jałoszyński. Pterotorax in ventral view (18 – 19), distal part of right metacoxa, metatrochanter and proximal portion of metafemur in ventral view (20), aedeagus in ventral view (21). Abbreviations: aml, apex of median lobe; ar, anterior ridge; cx 3, metacoxa; fm 3, metafemur; lmfa, lateral metafurcal arm; ma, membranous area; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pm, paramere; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vw, ventral wall." httpUri="https://zenodo.org/record/286518/files/figure.png" pageId="6" pageNumber="7">Figs. 21</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFD1180E8B203C8307DCFA84" box="[1097,1168,1695,1720]" captionStart-0="FIGURES 29 – 32" captionStart-1="FIGURES 33 – 36" captionStartId-0="15.[151,264,793,816]" captionStartId-1="15.[151,264,1487,1510]" captionTargetBox-0="[161,1432,199,770]" captionTargetBox-1="[160,1428,910,1456]" captionTargetId-0="figure@15.[151,1436,193,773]" captionTargetId-1="figure@15.[151,1436,896,1466]" captionTargetPageId-0="15" captionTargetPageId-1="15" captionText-0="FIGURES 29 – 32. Aedeagi of Horaeomorphus spp. in ventral (29, 31) and lateral (30, 32) views. Horaeomorphus anthicoides (Lea) comb. n. (29 – 30), Horaeomorphus verus sp. n. (31 – 32). Abbreviations: cc, central complex; ts, tubular structure." captionText-1="FIGURES 33 – 36. Aedeagi of Horaeomorphus spp. in ventral (33, 35) and lateral (34, 36) views. Horaeomorphus anthicoides (Lea) comb. n. (33 – 34), Horaeomorphus verus sp. n. (35 – 36)." httpUri-0="https://zenodo.org/record/286520/files/figure.png" httpUri-1="https://zenodo.org/record/286521/files/figure.png" pageId="6" pageNumber="7">29–36</figureCitation>
|
||
) with free and slender parameres (
|
||
<figureCitation id="13642A02FFD1180E8E703CD80220FAE0" box="[281,364,1732,1757]" captionStart="FIGURES 18 – 21" captionStartId="12.[151,264,1461,1484]" captionTargetBox="[151,1436,193,1440]" captionTargetId="figure@12.[151,1436,193,1441]" captionTargetPageId="12" captionText="FIGURES 18 – 21. Morphology of Horaeomorphus sakishimanus Jałoszyński. Pterotorax in ventral view (18 – 19), distal part of right metacoxa, metatrochanter and proximal portion of metafemur in ventral view (20), aedeagus in ventral view (21). Abbreviations: aml, apex of median lobe; ar, anterior ridge; cx 3, metacoxa; fm 3, metafemur; lmfa, lateral metafurcal arm; ma, membranous area; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pm, paramere; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vw, ventral wall." httpUri="https://zenodo.org/record/286518/files/figure.png" pageId="6" pageNumber="7">Fig. 21</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFD1180E8E113CD902D6FAE0" box="[376,410,1733,1756]" italics="true" pageId="6" pageNumber="7">pm</emphasis>
|
||
); spermatheca sclerotized and globular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345650CFFD1180F8FAE3CFB025EFFD1" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph id="8BE03687FFD1180F8FAE3CFB0001FDC9" blockId="6.[151,1437,940,2009]" lastBlockId="7.[151,1437,151,1005]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<emphasis id="B92BEA95FFD1180E8FAE3CFB0277FB3C" bold="true" box="[199,315,1767,1792]" pageId="6" pageNumber="7">Remarks.</emphasis>
|
||
Although
|
||
<taxonomicName id="4C5F4D04FFD1180E8ED23CFB013FFB3C" box="[443,627,1767,1792]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFD1180E8ED23CFB013FFB3C" box="[443,627,1767,1792]" italics="true" pageId="6" pageNumber="7">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
is highly diverse and includes species small and large, flattened and strongly convex, with the pronotum strongly elongate or nearly circular, with or without antebasal transverse groove, yet with some experience it is easy to identify the genus even without examining the ventral characters. Independently of the variable body shape, the general shape of the head is nearly identical in all species of
|
||
<taxonomicName id="4C5F4D04FFD1180E8FFE3D6B0203FBAC" box="[151,335,1911,1936]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFD1180E8FFE3D6B0203FBAC" box="[151,335,1911,1936]" italics="true" pageId="6" pageNumber="7">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
: always flattened, with the vertex never projecting dorsocaudad and its posterior margin concave; the bristles on tempora, genae and postgenae are sparse and inconspicuous in dry-mounted specimens, visible only in transparent mounts under high magnifications of a compound microscope; the supraantenal tubercles always prominent and in nearly all species accompanied posteromesally by a pair of distinct pits (absent only in
|
||
<taxonomicName id="4C5F4D04FFD0180F8AE83A85022DFCE8" authority="Franz" authorityName="Franz" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="B92BEA95FFD0180F8AE83A850249FCE9" italics="true" pageId="7" pageNumber="8">H. calcarifer</emphasis>
|
||
(Franz)
|
||
</taxonomicName>
|
||
and barely discernible in some specimens of
|
||
<taxonomicName id="4C5F4D04FFD0180F8C193AA10738FCE8" authority="Franz" authorityName="Franz" box="[880,1140,188,213]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="B92BEA95FFD0180F8C193AA10767FCE9" box="[880,1067,188,213]" italics="true" pageId="7" pageNumber="8">H. sarawakensis</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFCE4B76FFD0180F8B143AA00627FCE8" author="Jaloszynski" box="[1149,1387,188,213]" pageId="7" pageNumber="8" refString="Jaloszynski, P. (2012 a) Taxonomic status of Allohoraeomorphus Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3313, 62 - 68." type="journal article" year="2012" yearSuffix="a">(Jałoszyński 2012a))</bibRefCitation>
|
||
. All Australian specimens seen by the author have these pits. Another unique character of
|
||
<taxonomicName id="4C5F4D04FFD0180F8BFE3AC30603FCC4" box="[1175,1359,223,248]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFD0180F8BFE3AC30603FCC4" box="[1175,1359,223,248]" italics="true" pageId="7" pageNumber="8">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
is the inversely U-shaped groove connecting a pair of basal foveae on each elytron; the groove extends from each fovea anteriorly onto the elytral articulating lobe overlapped by the pronotal base, but is usually partly visible in intact specimens. Many species of
|
||
<taxonomicName id="4C5F4D04FFD0180F8EBF3B5001C2FD59" box="[470,654,332,357]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFD0180F8EBF3B5001C2FD59" box="[470,654,332,357]" italics="true" pageId="7" pageNumber="8">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
show a distinct sexual dimorphism, frequently expressed in elongated metatrochanters of males, sometimes to such extent that the distal part of trochanter forms a long rod-like projection exceeding half length of femur (examples are shown in
|
||
<figureCitation id="13642A02FFD0180F8CC83B880762FD91" box="[929,1070,404,429]" captionStart="FIGURES 1 – 5" captionStartId="7.[151,264,1767,1790]" captionTargetBox="[157,1436,1065,1740]" captionTargetId="figure@7.[151,1436,1060,1747]" captionTargetPageId="7" captionText="FIGURES 1 – 5. Examples of body form in Horaeomorphus, males. Horaeomorphus valdepunctatus Franz from Malay Peninsula (1); H. antennatus Jałoszyński from Malay Peninsula (2); H. ishiianus Jałoszyński from Borneo (3); H. punctatissimus Franz from Borneo (4), H. calcarifer (Franz) from Fiji (photograph from Jałoszyński 2012 a) (5)." httpUri="https://zenodo.org/record/286514/files/figure.png" pageId="7" pageNumber="8">Figs. 2, 4–5</figureCitation>
|
||
, and details in
|
||
<figureCitation id="13642A02FFD0180F8B8A3B880676FD90" box="[1251,1338,404,429]" captionStart="FIGURES 18 – 21" captionStartId="12.[151,264,1461,1484]" captionTargetBox="[151,1436,193,1440]" captionTargetId="figure@12.[151,1436,193,1441]" captionTargetPageId="12" captionText="FIGURES 18 – 21. Morphology of Horaeomorphus sakishimanus Jałoszyński. Pterotorax in ventral view (18 – 19), distal part of right metacoxa, metatrochanter and proximal portion of metafemur in ventral view (20), aedeagus in ventral view (21). Abbreviations: aml, apex of median lobe; ar, anterior ridge; cx 3, metacoxa; fm 3, metafemur; lmfa, lateral metafurcal arm; ma, membranous area; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pm, paramere; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vw, ventral wall." httpUri="https://zenodo.org/record/286518/files/figure.png" pageId="7" pageNumber="8">Fig. 20</figureCitation>
|
||
). In two species males differ from females in metafemora with a small ventral tooth. Australian species, at least those known from males, do not have any modifications of the legs.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFD0180F8FAE381C01C8FECC" blockId="7.[151,1437,151,1005]" pageId="7" pageNumber="8">
|
||
A character previously unknown was discovered during the present study.
|
||
<taxonomicName id="4C5F4D04FFD0180F8B7E3BE30622FE24" box="[1047,1390,511,536]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sakishimanus">
|
||
<emphasis id="B92BEA95FFD0180F8B7E3BE30622FE24" box="[1047,1390,511,536]" italics="true" pageId="7" pageNumber="8">Horaeomorphus sakishimanus</emphasis>
|
||
</taxonomicName>
|
||
has the mesocoxae with dense and relatively long setae on the dorsal surface (
|
||
<figureCitation id="13642A02FFD0180F8C8D38380722FE00" box="[996,1134,548,573]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="7" pageNumber="8">Figs. 12–13</figureCitation>
|
||
), and bunches of two-nine setae are inserted on small papillae (
|
||
<figureCitation id="13642A02FFD0180F8D42385B0132FE5C" box="[555,638,583,608]" captionStart="FIGURES 10 – 14" captionStartId="9.[151,264,1584,1607]" captionTargetBox="[151,1436,193,1562]" captionTargetId="figure@9.[151,1436,193,1563]" captionTargetPageId="9" captionText="FIGURES 10 – 14. Morphology of Horaeomorphus sakishimanus Jałoszyński. Base of pronotum in dorsal view (10), prothorax in ventral view (11), left fore leg in dorsal view (12), left procoxa, protrochanter and proximal part of profemur in dorsal view (13), coxal papilla with setae (14). Abbreviations: bst, basisternal part of prosternum; cx 2, mesocoxa; fm 2, mesofemur; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; psp, prosternal intercoxal process; tg, transverse antebasal groove; tr 2, mesotrochanter; mp, median antebasal pit." httpUri="https://zenodo.org/record/286516/files/figure.png" pageId="7" pageNumber="8">Fig. 14</figureCitation>
|
||
). Examination of 15 species of Oriental
|
||
<taxonomicName id="4C5F4D04FFD0180F8B56385B07BAFE5C" box="[1087,1270,583,608]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFD0180F8B56385B07BAFE5C" box="[1087,1270,583,608]" italics="true" pageId="7" pageNumber="8">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
(cPJ) revealed that all of them have densely setose dorsal surface of mesocoxae. Among Australian
|
||
<taxonomicName id="4C5F4D04FFD0180F8B193870065AFEB9" box="[1136,1302,620,645]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
|
||
bunches of setae on the dorsal surface of mesocoxae were found only in
|
||
<taxonomicName id="4C5F4D04FFD0180F8C3538930748FE94" box="[860,1028,655,680]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFD0180F8C3538930748FE94" box="[860,1028,655,680]" italics="true" pageId="7" pageNumber="8">Scydmaenozila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92BEA95FFD0180F8B6438930713FE94" bold="true" box="[1037,1119,655,680]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="A21857EEFFD0180F8B6438930713FE94" box="[1037,1119,655,680]" pageId="7" pageNumber="8" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13642A02FFD0180F8B193893078AFE94" box="[1136,1222,655,680]" captionStart="FIGURES 62 – 66" captionStartId="30.[151,264,1734,1757]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@30.[151,1436,193,1713]" captionTargetPageId="30" captionText="FIGURES 62 – 66. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Prothorax in ventral and slightly lateral view (62 – 63), base of pronotum in dorsal view (64), mesonotum and elytral base in dorsal view (65), and mesocoxa in dorsal view (66). Abbreviations: bef, basal elytral foveae; bst, basisternal part of prosternum; cx 2, mesocoxa; eal, elytral articulating lobe; fm 2, mesofemur; hy, hypomeron; ip, internal antebasal pit; lp, lateral antebasal pit; nss, pronosternal suture; pcc. procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; sss, mesoscutoscutellar suture; tr 2, mesotrochanter." httpUri="https://zenodo.org/record/286528/files/figure.png" pageId="7" pageNumber="8">Fig. 66</figureCitation>
|
||
), but in this genus there are two groups of setae, each seta inserted separately, the surface of coxa is not papillate, and there are numerous pores visible on the coxal cuticle.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFD0180F8FAE38E0025EFFD1" blockId="7.[151,1437,151,1005]" pageId="7" pageNumber="8">
|
||
<emphasis id="B92BEA95FFD0180F8FAE38E0017DFF29" bold="true" box="[199,561,764,789]" pageId="7" pageNumber="8">Composition and distribution.</emphasis>
|
||
Thirty-nine species (including those redescribed and described in the present paper) can be unambiguously placed in
|
||
<taxonomicName id="4C5F4D04FFD0180F8D1139030063FF04" box="[632,815,799,824]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFD0180F8D1139030063FF04" box="[632,815,799,824]" italics="true" pageId="7" pageNumber="8">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
. Thirty six are Asian (Oriental, Himalayan and Far Eastern), one Melanesian (from
|
||
<collectingCountry id="F3487617FFD0180F8D6E3958017CFF61" box="[519,560,836,861]" name="Fiji" pageId="7" pageNumber="8">Fiji</collectingCountry>
|
||
) and two Australian. Numerous additional undescribed species were seen by the author, mostly represented by unidentifiable females, from SE Asia (including the Indonesian New
|
||
<collectingCountry id="F3487617FFD0180F8A52397B06DCFFBC" box="[1339,1424,871,896]" name="Guinea-Bissau" pageId="7" pageNumber="8">Guinea</collectingCountry>
|
||
), and two more undescribed species (known from females) occur in
|
||
<collectingCountry id="F3487617FFD0180F8CC939900746FF99" box="[928,1034,908,933]" name="Australia" pageId="7" pageNumber="8">Australia</collectingCountry>
|
||
. Australian findings (
|
||
<figureCitation id="13642A02FFD0180F8A6C3990062BFF98" box="[1285,1383,908,933]" captionStart="FIGURES 193 – 198" captionStartId="72.[151,264,1762,1785]" captionTargetBox="[195,1376,193,1707]" captionTargetId="figure@72.[191,1378,193,1739]" captionTargetPageId="72" captionText="FIGURES 193 – 198. Distribution of revised Australian species of Horaeomorphus Schaufuss (193), Leascydmus gen. n. (194), Scydmaenozila gen. n. (195), Spinosciacharis gen. n. (196), Sciacharoides gen. n. (197), and Sciacharis Broun. Horaeomorphus anthicoides (Lea) comb. n. (193 a), Horaeomorphus verus sp. n. (193 b), Leascydmus simplicicornis (Lea) comb. n. (194 a), Leascydmus armstrongi sp. n. (194 b), Scydmaenozila macrosticta (Lea) comb. n. (195 a), Scydmaenozila latipennis (Lea) comb. n. (195 b), Scydmaenozila mallacootana sp. n. (195 c), Scydmaenozila inexpectata sp. n. (195 d), Spinosciacharis puncticeps (Franz) comb. n. (196 a), Spinosciacharis sassy sp. n. (196 b), Spinosciacharis clydensis sp. n. (196 c), Spinosciacharis acheron sp. n. (196 d), Sciacharoides thompsoni (Franz) comb. n. (197 a), Sciacharoides eucalypti (Franz) comb. n. (197 b), Sciacharoides maipotonensis (Franz) comb. n. (197 c), Sciacharoides mantistamborinensis (Franz) comb. n. (197 d), and Sciacharis (Maorinus) tasmaniensis (Franz) comb. n. (198)." httpUri="https://zenodo.org/record/286555/files/figure.png" pageId="7" pageNumber="8">Fig. 193</figureCitation>
|
||
) are located near the eastern coastline and fill a distributional gap between SE Asia / New
|
||
<collectingCountry id="F3487617FFD0180F8BD039B30646FFF4" box="[1209,1290,943,968]" name="Guinea-Bissau" pageId="7" pageNumber="8">Guinea</collectingCountry>
|
||
and eastern Melanesia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |