606 lines
101 KiB
XML
606 lines
101 KiB
XML
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<mods:title id="DFE299DAAD78C898CF780D7108CEFC07">New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships</mods:title>
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<mods:title id="ABC07677EB8FBEAD417BCC1653A8ECC6">Fossil Record</mods:title>
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<mods:date id="C9C21E89AA1E777F8071DA2464FD5400">2016</mods:date>
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<treatment id="03F68791BA332F11FFFFFD43FF55D291" ID-DOI="http://doi.org/10.5281/zenodo.11045888" ID-Zenodo-Dep="11045888" LSID="urn:lsid:plazi:treatment:03F68791BA332F11FFFFFD43FF55D291" httpUri="http://treatment.plazi.org/id/03F68791BA332F11FFFFFD43FF55D291" lastPageId="22" lastPageNumber="53" pageId="18" pageNumber="49">
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<subSubSection id="C345650CBA332F15FFFFFD43FDD6D49F" box="[124,587,717,741]" pageId="18" pageNumber="49" type="nomenclature">
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<paragraph id="8BE03687BA332F15FFFFFD43FDD6D49F" blockId="18.[124,587,717,741]" box="[124,587,717,741]" pageId="18" pageNumber="49">
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<heading id="D0A881EBBA332F15FFFFFD43FDD6D49F" box="[124,587,717,741]" fontSize="10" level="2" pageId="18" pageNumber="49" reason="4">
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<emphasis id="B92BEA95BA332F15FFFFFD43FDD6D49F" bold="true" box="[124,587,717,741]" pageId="18" pageNumber="49">
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<taxonomicName id="4C5F4D04BA332F15FFFFFD43FEE9D49F" authority="Graf" authorityName="Graf" box="[124,372,717,741]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithii">
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<emphasis id="B92BEA95BA332F15FFFFFD43FEABD49F" bold="true" box="[124,310,717,741]" italics="true" pageId="18" pageNumber="49">Ascalabos voithii</emphasis>
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Graf
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</taxonomicName>
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zu
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<bibRefCitation id="EFCE4B76BA332F15FE18FD43FDD6D49F" author="Munster, G." box="[411,587,717,741]" pageId="18" pageNumber="49" pagination="112 - 114" refId="ref18498" refString="Graf zu Munster, G.: Ascalabos voithii, Beitrage zur Petrefacten- Kunde, 2, 112 - 114, 1839 b." type="journal article" year="1839">Münster, 1839b</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C345650CBA332F11FFFFFC80FF55D291" lastPageId="22" lastPageNumber="53" pageId="18" pageNumber="49" type="description">
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<paragraph id="8BE03687BA332F15FFFFFC80FE88D55F" blockId="18.[124,767,782,1889]" box="[124,277,782,805]" pageId="18" pageNumber="49">
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<figureCitation id="13642A02BA332F15FFFFFC80FE88D55F" box="[124,277,782,805]" captionStart-0="Figure 14" captionStart-1="Figure 15" captionStart-2="Figure 16" captionStart-3="Figure 17" captionStart-4="Figure 18" captionStart-5="Figure 19" captionStartId-0="19.[124,191,1569,1591]" captionStartId-1="20.[124,191,1546,1568]" captionStartId-2="21.[124,191,647,669]" captionStartId-3="22.[124,191,738,760]" captionStartId-4="23.[124,191,1346,1368]" captionStartId-5="24.[124,191,1198,1220]" captionTargetBox-0="[169,1416,177,1526]" captionTargetBox-1="[169,1416,177,1515]" captionTargetBox-2="[169,1416,177,615]" captionTargetBox-3="[369,1215,187,694]" captionTargetBox-4="[146,1448,197,1298]" captionTargetBox-5="[169,1407,185,1146]" captionTargetId-0="figure-11@19.[169,1416,177,1538]" captionTargetId-1="figure-11@20.[169,1416,177,1515]" captionTargetId-2="figure-11@21.[169,1416,177,615]" captionTargetId-3="figure-11@22.[358,1227,177,706]" captionTargetId-4="figure-11@23.[132,1455,177,1315]" captionTargetId-5="figure-11@24.[169,1416,177,1167]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionTargetPageId-4="23" captionTargetPageId-5="24" captionText-0="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." captionText-1="Figure 15. Ascalabos voithii from Schernfeld in the Eichstätt Basin. Uncatalogued specimen from U. Eller (Dümpelfeld). (a) Photograph of specimen in lateral view. (b) Enlargement of the head. Note the peculiar shape and distribution of the sensory tubules of the preopercular canal. Photographs courtesy of U. Eller." captionText-2="Figure 16. Ascalabos voithii. Restoration in lateral view, updated from Arratia (1997)." captionText-3="Figure 17. Ascalabos voithii (neotype, JME SOS 537). Midcaudal vertebrae illustrating the presence of a lateral crest and grooves on the lateral side of the vertebral centra. Arrows point to a lateral longitudinal crest. Abbreviations: na, neural arch; ns, neural spine; ha, haemal arch; hs, haemal spine; vc, vertebral centrum." captionText-4="Figure 18. Ascalabos voithii. Caudal skeleton in lateral view slightly modified from Arratia (1997). Abbreviations: Ant, anterior; dpr, dorsal or epaxial procurrent rays; d.scu, dorsal caudal scute; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–11, hypurals 1–11; hsPU2, haemal spine of preural centrum 2; mo, membranous outgrowth on anterodorsal margin of first uroneural; mo’, membranous outgrowth on anterodorsal margin of second uroneural; nsPU2–4, neural spine of preural centra 2–4; PR1,19, principal caudal rays 1, 19; rer, rudimentary fin ray; U1+2+H1–2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–7, uroneurals 1–7; v.scu., ventral caudal scute." captionText-5="Figure 19. Ascalabos voithii. Caudal skeleton in lateral view with only two epaxial basal fulcra preserved (neotype, JME SOS 537). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; dpr, dorsal or epaxial procurrent rays; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–4, hypurals 1–4; hbfu, hypaxial basal fulcra; hsPU4, haemal spine of preural centrum 4; naPU1, neural arch of preural centrum 1; nsPU2–4, neural spine of preural centra 2–4; PH, parhypural; PU1,4, preural centrum 1, 4; rer, rudimentary fin ray; sc, dorsal and ventral caudal scutes; U1+2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–8, uroneurals 1–8; PR1–19, principal rays 1–19; vpr, ventral procurrent rays; v.scu, ventral caudal scute." figureDoi-0="http://doi.org/10.5281/zenodo.11045902" figureDoi-1="http://doi.org/10.5281/zenodo.11045904" figureDoi-2="http://doi.org/10.5281/zenodo.11045906" figureDoi-3="http://doi.org/10.5281/zenodo.11045908" figureDoi-4="http://doi.org/10.5281/zenodo.11045910" figureDoi-5="http://doi.org/10.5281/zenodo.11045912" httpUri-0="https://zenodo.org/record/11045902/files/figure.png" httpUri-1="https://zenodo.org/record/11045904/files/figure.png" httpUri-2="https://zenodo.org/record/11045906/files/figure.png" httpUri-3="https://zenodo.org/record/11045908/files/figure.png" httpUri-4="https://zenodo.org/record/11045910/files/figure.png" httpUri-5="https://zenodo.org/record/11045912/files/figure.png" pageId="18" pageNumber="49">Figures 14–19</figureCitation>
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFCA0FF24D51F" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
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1839b
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<taxonomicName id="4C5F4D04BA332F15FF45FCA3FE1DD53F" box="[198,384,813,837]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithii">
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<emphasis id="B92BEA95BA332F15FF45FCA3FE1DD53F" box="[198,384,813,837]" italics="true" pageId="18" pageNumber="49">Ascalabos Voithii</emphasis>
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</taxonomicName>
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– Graf zu Münster, p. 112, pl. XII, fig. 5.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFCE0FF42D5DF" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
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1843
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<taxonomicName id="4C5F4D04BA332F15FF3BFCE3FEEFD5FF" box="[184,370,877,901]" class="Liliopsida" family="Cyperaceae" genus="Leptolepis" kingdom="Plantae" order="Poales" pageId="18" pageNumber="49" phylum="Tracheophyta" rank="species" species="voithii">
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<emphasis id="B92BEA95BA332F15FF3BFCE3FEEFD5FF" box="[184,370,877,901]" italics="true" pageId="18" pageNumber="49">Leptolepis Voithii</emphasis>
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</taxonomicName>
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– Agassiz, p. 131, 165, 295, pl. LXI, figs. 2–3.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFC23FF42D59E" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
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1843
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<taxonomicName id="4C5F4D04BA332F15FF38FC23FE56D5BF" authorityName="Agassiz" authorityYear="1844" box="[187,459,941,965]" class="Liliopsida" family="Cyperaceae" genus="Leptolepis" kingdom="Plantae" order="Poales" pageId="18" pageNumber="49" phylum="Tracheophyta" rank="species" species="polyspondylus">
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<emphasis id="B92BEA95BA332F15FF38FC23FE56D5BF" box="[187,459,941,965]" italics="true" pageId="18" pageNumber="49">Leptolepis polyspondylus</emphasis>
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</taxonomicName>
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– Agassiz, p. 133, pl. LXI, figs. 7–8.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFC63FF5BD25E" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
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1843
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<taxonomicName id="4C5F4D04BA332F15FF34FC63FE43D27F" authorityName="Agassiz" authorityYear="1844" box="[183,478,1005,1029]" class="Liliopsida" family="Cyperaceae" genus="Leptolepis" kingdom="Plantae" order="Poales" pageId="18" pageNumber="49" phylum="Tracheophyta" rank="species" species="paucispondylous">
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<emphasis id="B92BEA95BA332F15FF34FC63FE43D27F" box="[183,478,1005,1029]" italics="true" pageId="18" pageNumber="49">Leptolepis paucispondylous</emphasis>
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</taxonomicName>
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(nomen nudum) – Agassiz, p. 134.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFBA3FDB6D23E" blockId="18.[124,767,782,1889]" box="[124,555,1069,1093]" pageId="18" pageNumber="49">
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1848
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<taxonomicName id="4C5F4D04BA332F15FF3BFBA3FEEFD23F" box="[184,370,1069,1093]" class="Liliopsida" family="Cyperaceae" genus="Leptolepis" kingdom="Plantae" order="Poales" pageId="18" pageNumber="49" phylum="Tracheophyta" rank="species" species="voithii">
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<emphasis id="B92BEA95BA332F15FF3BFBA3FEEFD23F" box="[184,370,1069,1093]" italics="true" pageId="18" pageNumber="49">Leptolepis Voithii</emphasis>
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</taxonomicName>
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– Giebel, p. 143.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFBC3FD83D21E" blockId="18.[124,767,782,1889]" box="[124,542,1100,1124]" pageId="18" pageNumber="49">
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1848
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<taxonomicName id="4C5F4D04BA332F15FF3BFBC2FEFBD21E" authorityName="Giebel" authorityYear="1848" box="[184,358,1100,1124]" class="Gastropoda" family="Skeneidae" genus="Tharsis" kingdom="Animalia" order="Trochida" pageId="18" pageNumber="49" phylum="Mollusca" rank="species" species="radiatus">
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<emphasis id="B92BEA95BA332F15FF3BFBC2FEFBD21E" box="[184,358,1100,1124]" italics="true" pageId="18" pageNumber="49">Tharsis radiatus</emphasis>
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</taxonomicName>
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– Giebel, p. 146.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFBE3FDC0D2FE" blockId="18.[124,767,782,1889]" box="[124,605,1132,1156]" pageId="18" pageNumber="49">
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1863
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<taxonomicName id="4C5F4D04BA332F15FF3BFBE2FE49D2FE" authority=", Wagner" authorityName="Wagner" box="[184,468,1132,1156]" class="Liliopsida" family="Cyperaceae" genus="Leptolepis" kingdom="Plantae" order="Poales" pageId="18" pageNumber="49" phylum="Tracheophyta" rank="species" species="voithii">
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<emphasis id="B92BEA95BA332F15FF3BFBE2FEEFD2FE" box="[184,370,1132,1156]" italics="true" pageId="18" pageNumber="49">Leptolepis Voithii</emphasis>
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, Wagner
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</taxonomicName>
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, p. 131, 134.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFB03FDCBD2DE" blockId="18.[124,767,782,1889]" box="[124,598,1164,1188]" pageId="18" pageNumber="49">
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1895
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<taxonomicName id="4C5F4D04BA332F15FF3BFB02FEECD2DE" box="[184,369,1164,1188]" class="Liliopsida" family="Cyperaceae" genus="Leptolepis" kingdom="Plantae" order="Poales" pageId="18" pageNumber="49" phylum="Tracheophyta" rank="species" species="voithii">
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<emphasis id="B92BEA95BA332F15FF3BFB02FEECD2DE" box="[184,369,1164,1188]" italics="true" pageId="18" pageNumber="49">Leptolepis voithii</emphasis>
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</taxonomicName>
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– Woodward, p. 512.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFB23FE24D299" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
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1974
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<taxonomicName id="4C5F4D04BA332F15FF3BFB22FEF4D2BE" box="[184,361,1196,1220]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithi">
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<emphasis id="B92BEA95BA332F15FF3BFB22FEF4D2BE" box="[184,361,1196,1220]" italics="true" pageId="18" pageNumber="49">Ascalabos voithi</emphasis>
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</taxonomicName>
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– Nybelin, 173–180, text-figs. 35, 36, pl. XXIX, figs. 1–5, pl. XXX.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFB62FF43D359" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
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1975a
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<taxonomicName id="4C5F4D04BA332F15FF49FB62FD90D37E" authorityName="Agassiz" authorityYear="1834" box="[202,525,1260,1284]" class="Liliopsida" family="Cyperaceae" genus="Leptolepis" kingdom="Plantae" order="Poales" pageId="18" pageNumber="49" phylum="Tracheophyta" rank="species" species="voithi" subGenus="Ascalabos">
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<emphasis id="B92BEA95BA332F15FF49FB62FEA4D37E" box="[202,313,1260,1284]" italics="true" pageId="18" pageNumber="49">Leptolepis</emphasis>
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(
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<emphasis id="B92BEA95BA332F15FECDFB62FE26D37E" box="[334,443,1260,1284]" italics="true" pageId="18" pageNumber="49">Ascalabos</emphasis>
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)
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<emphasis id="B92BEA95BA332F15FE53FB62FD90D37E" box="[464,525,1260,1284]" italics="true" pageId="18" pageNumber="49">voithi</emphasis>
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</taxonomicName>
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– Taverne, 233–243, figs. 1–6.
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</paragraph>
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<paragraph id="8BE03687BA332F15FFFFFAA2FF64D319" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
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1977
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<taxonomicName id="4C5F4D04BA332F15FF3DFAA2FEE1D33E" box="[190,380,1324,1348]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithii">
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<emphasis id="B92BEA95BA332F15FF3DFAA2FEE1D33E" box="[190,380,1324,1348]" italics="true" pageId="18" pageNumber="49">Ascalabos voithii</emphasis>
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</taxonomicName>
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– Patterson and Rosen, 151–152, figs. 52–53.
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</paragraph>
|
||
<paragraph id="8BE03687BA332F15FFFFFAE2FD1ED3F9" blockId="18.[124,767,782,1889]" box="[124,643,1387,1411]" pageId="18" pageNumber="49">
|
||
1991
|
||
<taxonomicName id="4C5F4D04BA332F15FF3BFAE5FEF4D3F9" box="[184,361,1387,1411]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithi">
|
||
<emphasis id="B92BEA95BA332F15FF3BFAE5FEF4D3F9" box="[184,361,1387,1411]" italics="true" pageId="18" pageNumber="49">Ascalabos voithi</emphasis>
|
||
</taxonomicName>
|
||
– Arratia, 268–271, fig. 9.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F15FFFFFA02FD2DD3D9" blockId="18.[124,767,782,1889]" box="[124,688,1419,1443]" pageId="18" pageNumber="49">
|
||
1997
|
||
<taxonomicName id="4C5F4D04BA332F15FF3BFA05FEEDD3D9" box="[184,368,1419,1443]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA332F15FF3BFA05FEEDD3D9" box="[184,368,1419,1443]" italics="true" pageId="18" pageNumber="49">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
– Arratia, 31–39, figs. 14–21.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F15FFFFFA22FF76D078" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
|
||
<materialsCitation id="3B373CDABA332F15FFFFFA22FF76D078" collectionCode="SOS" collectorName="The & Eichstatt I" pageId="18" pageNumber="49" specimenCode="SOS 537" specimenCount="3" typeStatus="neotype">
|
||
<typeStatus id="54E48825BA332F15FFFFFA22FF47D3B9" box="[124,218,1452,1475]" pageId="18" pageNumber="49" type="neotype">Neotype</typeStatus>
|
||
:
|
||
<collectorName id="26AA5351BA332F15FF67FA22FE90D3B9" box="[228,269,1452,1475]" pageId="18" pageNumber="49">The</collectorName>
|
||
holotype is lost.
|
||
<bibRefCitation id="EFCE4B76BA332F15FE46FA22FDF4D3B9" author="Nybelin, O." box="[453,617,1452,1475]" pageId="18" pageNumber="49" pagination="1 - 202" refId="ref18872" refString="Nybelin, O.: A revision of the leptolepid fishes, Acta Regiae Societatis Scientiarum et Litterarum Gothoburgensis, Zoologica, 9, 1 - 202, 1974." type="journal article" year="1974">Nybelin (1974)</bibRefCitation>
|
||
designated as neotype a specimen identified as
|
||
<collectorName id="26AA5351BA332F15FE66FA45FDCED398" box="[485,595,1483,1506]" pageId="18" pageNumber="49">Eichstätt I</collectorName>
|
||
(currently JME
|
||
<specimenCode id="DBF99EFCBA332F15FFFFFA65FF42D078" box="[124,223,1515,1538]" collectionCode="SOS" pageId="18" pageNumber="49">SOS 537</specimenCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F15FFFFF985FE76D11B" blockId="18.[124,767,782,1889]" pageId="18" pageNumber="49">
|
||
Additional specimens:
|
||
<collectionCode id="ED4EAE42BA332F15FE09F985FE2ED058" box="[394,435,1547,1570]" country="China" name="Chongqing Museum" pageId="18" pageNumber="49">CM</collectionCode>
|
||
9491.
|
||
<collectionCode id="ED4EAE42BA332F15FD95F985FDF7D058" box="[534,618,1547,1570]" pageId="18" pageNumber="49">CMMH</collectionCode>
|
||
9491. JME
|
||
<collectionCode id="ED4EAE42BA332F15FFFFF9A5FF30D038" box="[124,173,1579,1602]" pageId="18" pageNumber="49">SOS</collectionCode>
|
||
2362, JME
|
||
<collectionCode id="ED4EAE42BA332F15FEAFF9A5FEC0D038" box="[300,349,1579,1602]" pageId="18" pageNumber="49">SOS</collectionCode>
|
||
2364, JME
|
||
<collectionCode id="ED4EAE42BA332F15FE5FF9A5FD90D038" box="[476,525,1579,1602]" pageId="18" pageNumber="49">SOS</collectionCode>
|
||
2365, JME
|
||
<collectionCode id="ED4EAE42BA332F15FD0FF9A5FD20D038" box="[652,701,1579,1602]" pageId="18" pageNumber="49">SOS</collectionCode>
|
||
2458, JME
|
||
<collectionCode id="ED4EAE42BA332F15FF38F9C5FF71D018" box="[187,236,1611,1634]" pageId="18" pageNumber="49">SOS</collectionCode>
|
||
2483, JME
|
||
<collectionCode id="ED4EAE42BA332F15FE02F9C5FE2FD018" box="[385,434,1611,1634]" pageId="18" pageNumber="49">SOS</collectionCode>
|
||
2496, JME
|
||
<collectionCode id="ED4EAE42BA332F15FDC5F9C5FDEAD018" box="[582,631,1611,1634]" pageId="18" pageNumber="49">SOS</collectionCode>
|
||
2497, JME
|
||
<collectionCode id="ED4EAE42BA332F15FFFFF9E5FF30D0F8" box="[124,173,1643,1666]" pageId="18" pageNumber="49">SOS</collectionCode>
|
||
2886 (peels of scales), and many other specimens deposited at the JME. NHM 3672, NHM 3673a, NHM 37062, and NHM 37080. SHL, collection of H. Leich (Bochum,
|
||
<collectingCountry id="F3487617BA332F15FFFFF944FF7DD09B" box="[124,224,1738,1761]" name="Germany" pageId="18" pageNumber="49">Germany</collectingCountry>
|
||
) currently deposited in Tierpark und Fossilium Bochum (Bochum).
|
||
<collectionCode id="ED4EAE42BA332F15FEDDF964FE36D17B" box="[350,427,1770,1793]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="18" pageNumber="49">SMNH</collectionCode>
|
||
P5683.
|
||
<collectionCode id="ED4EAE42BA332F15FD89F964FDAFD17B" box="[522,562,1770,1793]" country="Germany" name="Teylers Museum, Paleontologische" pageId="18" pageNumber="49">TM</collectionCode>
|
||
6651,
|
||
<collectionCode id="ED4EAE42BA332F15FD00F964FD36D17B" box="[643,683,1770,1793]" country="Germany" name="Teylers Museum, Paleontologische" pageId="18" pageNumber="49">TM</collectionCode>
|
||
10307, and
|
||
<collectionCode id="ED4EAE42BA332F15FF2BF884FF4DD15B" box="[168,208,1802,1825]" country="Germany" name="Teylers Museum, Paleontologische" pageId="18" pageNumber="49">TM</collectionCode>
|
||
10325. Additionally, one uncatalogued specimen belonging to U. Eller (Dümpelfeld,
|
||
<collectingCountry id="F3487617BA332F15FE58F8A4FDA2D13B" box="[475,575,1834,1857]" name="Germany" pageId="18" pageNumber="49">Germany</collectingCountry>
|
||
) was included because of its excellent preservation.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F15FCB0FF3EFA38D7FD" blockId="18.[819,1463,176,1921]" pageId="18" pageNumber="49">
|
||
Locality and geological time: Blumenberg, Eichstätt, Hofstetten, Kelheim, and Wintershof (
|
||
<figureCitation id="13642A02BA332F15FB53FF5EFA87D69D" box="[1232,1306,208,231]" captionStart="Figure 1" captionStartId="2.[124,191,1081,1103]" captionTargetBox="[150,1435,177,1050]" captionTargetId="figure-11@2.[150,1435,177,1050]" captionTargetPageId="2" captionText="Figure 1. Distribution of Plattenkalk basins and reef areas in the southern Franconian Alb during the Early Tithonian (slightly modified from Viohl, 1996). The new fish described herein was recovered in Ettling." figureDoi="http://doi.org/10.5281/zenodo.11045857" httpUri="https://zenodo.org/record/11045857/files/figure.png" pageId="18" pageNumber="49">Fig. 1</figureCitation>
|
||
) in
|
||
<collectingRegion id="499BF865BA332F15FADDFF5EFA2FD69D" box="[1374,1458,208,231]" country="Germany" name="Bayern" pageId="18" pageNumber="49">Bavaria</collectingRegion>
|
||
, southern
|
||
<collectingCountry id="F3487617BA332F15FC18FF7EFC62D77D" box="[923,1023,240,263]" name="Germany" pageId="18" pageNumber="49">Germany</collectingCountry>
|
||
. Upper Jurassic, Tithonian (
|
||
<bibRefCitation id="EFCE4B76BA332F15FABEFF7EFB84D75D" author="Meyer, R. & Schmidt-Kaler, H." pageId="18" pageNumber="49" pagination="3 - 77" refId="ref18708" refString="Meyer, R. and Schmidt-Kaler, H.: Palaogeographischer Atlas des suddeutschen Oberjura (Malm), Geol. Jahrbucher A, 225, 3 - 77, 1989." type="journal article" year="1989">Meyer and Schmidt-Kaler, 1989</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCE4B76BA332F15FBABFE9EFBFDD75D" author="Meyer, R. & Schmidt-Kaler, H." box="[1064,1120,272,295]" pageId="18" pageNumber="49" refId="ref18740" refString="Meyer, R. and Schmidt-Kaler, H.: Wanderungen in die Erdgeschichte, (1) Treuchtlingen - Solnhofen - Mornsheim, Dollnstein, Verlag Dr. F. Pfeil, Munchen, 1990." type="book" year="1990">1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCE4B76BA332F15FBECFE9EFAAED75D" author="Schweigert, G." box="[1135,1331,272,295]" pageId="18" pageNumber="49" pagination="63 - 66" refId="ref19429" refString="Schweigert, G.: Biostratigraphie der Plattenkalke der Sudlichen Frankenalb, in: Solnhofen, ein Frenster in die Jurazeit, edited by: Arratia, G., Schultze, H. - P., Tischlinger, H., and Viohl, G., Verlag Dr. F. Pfeil, Munchen, 63 - 66, 2015." type="book chapter" year="2015">Schweigert, 2015</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C5F4D04BA332F15FACAFE9EFCEAD73D" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA332F15FACAFE9EFCEAD73D" italics="true" pageId="18" pageNumber="49">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
has also been cited from Cerin (
|
||
<collectingCountry id="F3487617BA332F15FB77FEBEFADCD73D" box="[1268,1345,304,327]" name="France" pageId="18" pageNumber="49">France</collectingCountry>
|
||
;
|
||
<bibRefCitation id="EFCE4B76BA332F15FAD2FEBEFC2ED71D" author="de Saint-Seine, P." pageId="18" pageNumber="49" pagination="1 - 357" refId="ref18161" refString="de Saint-Seine, P.: Les poissons des calcaires lithographiques de Cerin (Ain), Nouvelle Archive du Museum d'Histoire naturelle, Lyon, 2, 1 - 357, 1949." type="journal article" year="1949">de Saint-Seine, 1949</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCE4B76BA332F15FC3DFEDEFBEAD71D" author="Wenz, S. & Bernier, P. & Barale, G. & Bourseau, J. P. & Buffetaut, E. & Gaillard, C. & Gall, J. C." box="[958,1143,336,359]" pageId="18" pageNumber="49" pagination="61 - 70" refId="ref19912" refString="Wenz, S., Bernier, P., Barale, G., Bourseau, J. P., Buffetaut, E., Gaillard, C., and Gall, J. C.: L' ichthyofaune des calcaires lithographiques du Kimmeridgien superieur de Cerin (Ain, France), Geobios, M. S., 16, 61 - 70, 1993." type="journal article" year="1993">Wenz et al., 1993</bibRefCitation>
|
||
), which is interpreted as Kimmeridgian in age. This material is currently under revision.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F15FCB0FE1EFC32D7DD" blockId="18.[819,1463,176,1921]" box="[819,943,400,423]" pageId="18" pageNumber="49">Description</paragraph>
|
||
<paragraph id="8BE03687BA332F15FCCDFE21FA18D41C" blockId="18.[819,1463,176,1921]" pageId="18" pageNumber="49">
|
||
A description of
|
||
<taxonomicName id="4C5F4D04BA332F15FB84FE21FB5CD7BD" box="[1031,1217,431,455]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA332F15FB84FE21FB5CD7BD" box="[1031,1217,431,455]" italics="true" pageId="18" pageNumber="49">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
can be found in
|
||
<bibRefCitation id="EFCE4B76BA332F15FAFFFE21FC02D79C" author="Arratia, G." pageId="18" pageNumber="49" pagination="1 - 168" refId="ref17245" refString="Arratia, G.: Basal teleosts and teleostean phylogeny, Palaeo Ichthyol., 7, 1 - 168, 1997." type="journal article" year="1997">Arratia (1997)</bibRefCitation>
|
||
. Here, I only describe and illustrate additional information after of a re-study of the
|
||
<typeStatus id="54E48825BA332F15FB33FE61FA99D47C" box="[1200,1284,495,518]" pageId="18" pageNumber="49" type="neotype">neotype</typeStatus>
|
||
and other specimens. I describe some characters that add new interpretations or a more complete understanding of certain structures. Some are compared with the structures in
|
||
<taxonomicName id="4C5F4D04BA332F15FB33FDC0FAABD41C" authority="Arratia, 2016" authorityName="Arratia" authorityYear="2016" box="[1200,1334,590,614]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95BA332F15FB33FDC0FAABD41C" box="[1200,1334,590,614]" italics="true" pageId="18" pageNumber="49">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEBA332F15FABEFDC1FA18D41C" box="[1341,1413,591,614]" pageId="18" pageNumber="49" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F15FCB0FDE1FADFD5DF" blockId="18.[819,1463,176,1921]" pageId="18" pageNumber="49">
|
||
General description:
|
||
<taxonomicName id="4C5F4D04BA332F15FB93FDE0FB5BD4FC" box="[1040,1222,622,646]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA332F15FB93FDE0FB5BD4FC" box="[1040,1222,622,646]" italics="true" pageId="18" pageNumber="49">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
has a head proportionally large, about 30 % of standard length; however, the head in
|
||
<taxonomicName id="4C5F4D04BA332F15FCCCFD20FC48D4BC" authorityName="Arratia" authorityYear="2016" box="[847,981,686,710]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA332F15FCCCFD20FC48D4BC" box="[847,981,686,710]" italics="true" pageId="18" pageNumber="49">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
is comparative larger, about 34 % of SL. The diameter of the eye is large, 22 to 25 % of the head length (
|
||
<figureCitation id="13642A02BA332F15FCB8FD60FC3CD57F" box="[827,929,750,773]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="18" pageNumber="49">Figs. 14b</figureCitation>
|
||
and
|
||
<figureCitation id="13642A02BA332F15FC54FD60FB9FD57F" box="[983,1026,750,773]" captionStart="Figure 15" captionStartId="20.[124,191,1546,1568]" captionTargetBox="[169,1416,177,1515]" captionTargetId="figure-11@20.[169,1416,177,1515]" captionTargetPageId="20" captionText="Figure 15. Ascalabos voithii from Schernfeld in the Eichstätt Basin. Uncatalogued specimen from U. Eller (Dümpelfeld). (a) Photograph of specimen in lateral view. (b) Enlargement of the head. Note the peculiar shape and distribution of the sensory tubules of the preopercular canal. Photographs courtesy of U. Eller." figureDoi="http://doi.org/10.5281/zenodo.11045904" httpUri="https://zenodo.org/record/11045904/files/figure.png" pageId="18" pageNumber="49">15b</figureCitation>
|
||
), whereas the eye in
|
||
<taxonomicName id="4C5F4D04BA332F15FB6BFD60FCE8D55C" authorityName="Arratia" authorityYear="2016" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="ettlingensis">
|
||
<emphasis id="B92BEA95BA332F15FB6BFD60FCE8D55C" italics="true" pageId="18" pageNumber="49">Ebertichthys ettlingensis</emphasis>
|
||
</taxonomicName>
|
||
is even larger, 34 to 36 % of the head length. The insertion of the dorsal fin (
|
||
<figureCitation id="13642A02BA332F15FBB9FCA0FB3DD53F" box="[1082,1184,814,837]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="18" pageNumber="49">Figs. 14a</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02BA332F15FB28FCA0FB4ED53F" box="[1195,1235,814,837]" captionStart="Figure 15" captionStartId="20.[124,191,1546,1568]" captionTargetBox="[169,1416,177,1515]" captionTargetId="figure-11@20.[169,1416,177,1515]" captionTargetPageId="20" captionText="Figure 15. Ascalabos voithii from Schernfeld in the Eichstätt Basin. Uncatalogued specimen from U. Eller (Dümpelfeld). (a) Photograph of specimen in lateral view. (b) Enlargement of the head. Note the peculiar shape and distribution of the sensory tubules of the preopercular canal. Photographs courtesy of U. Eller." figureDoi="http://doi.org/10.5281/zenodo.11045904" httpUri="https://zenodo.org/record/11045904/files/figure.png" pageId="18" pageNumber="49">15a</figureCitation>
|
||
, and
|
||
<figureCitation id="13642A02BA332F15FA8FFCA0FABAD53F" box="[1292,1319,814,837]" captionStart="Figure 16" captionStartId="21.[124,191,647,669]" captionTargetBox="[169,1416,177,615]" captionTargetId="figure-11@21.[169,1416,177,615]" captionTargetPageId="21" captionText="Figure 16. Ascalabos voithii. Restoration in lateral view, updated from Arratia (1997)." figureDoi="http://doi.org/10.5281/zenodo.11045906" httpUri="https://zenodo.org/record/11045906/files/figure.png" pageId="18" pageNumber="49">16</figureCitation>
|
||
) is above the level of the insertion of the pelvic fins, about 52 to 58 % of SL. In contrast, the insertion of the dorsal fin is posterior to that of the pelvic fins in
|
||
<taxonomicName id="4C5F4D04BA332F15FBB5FC03FAA6D5DF" authorityName="Arratia" authorityYear="2016" box="[1078,1339,909,933]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="ettlingensis">
|
||
<emphasis id="B92BEA95BA332F15FBB5FC03FAA6D5DF" box="[1078,1339,909,933]" italics="true" pageId="18" pageNumber="49">Ebertichthys ettlingensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F15FCB0FC23FAC3D23E" blockId="18.[819,1463,176,1921]" pageId="18" pageNumber="49">
|
||
Cranial bones: The cranial bones are unornamented with the exception of the premaxilla, which bears some small, tubercle-like ornaments (
|
||
<figureCitation id="13642A02BA332F15FBC3FC63FB07D27E" box="[1088,1178,1005,1028]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="18" pageNumber="49">Fig. 14c</figureCitation>
|
||
) covered with a thin layer of ganoine in the
|
||
<typeStatus id="54E48825BA332F15FC7BFB83FBCDD25E" box="[1016,1104,1037,1060]" pageId="18" pageNumber="49" type="neotype">neotype</typeStatus>
|
||
. The braincase of the
|
||
<typeStatus id="54E48825BA332F15FAC4FB83FA06D25E" box="[1351,1435,1037,1060]" pageId="18" pageNumber="49" type="neotype">neotype</typeStatus>
|
||
is poorly preserved and does not add new information.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F15FCB0FBC3FB50D398" blockId="18.[819,1463,176,1921]" pageId="18" pageNumber="49">
|
||
Circumorbital series: The supraorbital is incompletely preserved in most specimens or not preserved at all. It has an expanded anteroventral region, similar to that found in
|
||
<taxonomicName id="4C5F4D04BA332F15FAF9FB02FB95D2BE" authorityName="Arratia" authorityYear="2016" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="ettlingensis">
|
||
<emphasis id="B92BEA95BA332F15FAF9FB02FB95D2BE" italics="true" pageId="18" pageNumber="49">Ebertichthys ettlingensis</emphasis>
|
||
</taxonomicName>
|
||
. Infraorbital 1 is an elongate bone that joins with an elongate infraorbital 2 that is not as long as that of
|
||
<taxonomicName id="4C5F4D04BA332F15FCD2FB62FC4AD37E" authorityName="Arratia" authorityYear="2016" box="[849,983,1260,1284]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA332F15FCD2FB62FC4AD37E" box="[849,983,1260,1284]" italics="true" pageId="18" pageNumber="49">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="13642A02BA332F15FBC9FB62FB3FD379" box="[1098,1186,1260,1283]" captionStart="Figure 3" captionStartId="6.[124,191,1734,1756]" captionTargetBox="[264,1322,177,1693]" captionTargetId="figure-11@6.[264,1322,177,1703]" captionTargetPageId="6" captionText="Figure 3. Ebertichthys ettlingensis n. gen. et n. sp. in lateral view (holotype JME ETT 108). (a) Photograph of cranium and anterior part of body. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Drawing of cranium and anterior part of body. Abbreviations: ant, antorbital; br, branchiostegal rays; cl, cleithrum; cor, coracoid; dpa, dermopalatine; ent, entopterygoid; exc, extrascapula; hy, hyomandibula; io1-5, infraorbitals 1–5; iop, interopercle; l.de, left dentary; leth, lateral ethmoid; l.pmx, left premaxilla; met, mesethmoid; mx, maxilla; op, opercle; orb, orbitosphenoid; pa[= fr], parietal [= frontal bone of traditional terminology]; par, parasphenoid; pcl1-3, postcleithra 1–3; pop, preopercle; ppa[= pa], postparietal [= parietal bone of traditional terminology]; pt, pterotic; ptsp, pterosphenoid; ptt, posttemporal; qu, quadrate; sc, scales; sca, scapula; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sn, supraneurals; sorb.b, broken supraorbital bone; sop, subopercle; sy, symplectic; r.de, right dentary; r.pmx, right premaxilla; vo, vomer." figureDoi="http://doi.org/10.5281/zenodo.11045865" httpUri="https://zenodo.org/record/11045865/files/figure.png" pageId="18" pageNumber="49">Figs. 3b</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02BA332F15FB2DFB62FB54D379" box="[1198,1225,1260,1283]" captionStart="Figure 4" captionStartId="7.[124,191,1748,1770]" captionTargetBox="[265,1294,178,1710]" captionTargetId="figure-11@7.[264,1322,177,1716]" captionTargetPageId="7" captionText="Figure 4. Ebertichthys ettlingensis n. gen. et n. sp. Crania in lateral views illustrating specific morphological features. (a) JME ETT 60. (b) JME ETT 24. Arrow points to a small toothed patch. Abbreviations: ang, angular; aup, autopalatine; b.ant, broken antorbital; a.cer, anterior ceratohyal; br, branchiostegal rays; cl, cleithrum; de, dentary; ect, ectopterygoid; ent, entopterygoid; ghy, glossohyal; d.hy, dorsal hypohyal; iop, interopercle; hy, hyomandibula; io1–5, infraorbitals 1–5; l.ang, left angular; l.de, left dentary; l.pmx, left premaxilla; met, mesethmoid; mtg, metapterygoid; mx, maxilla; op, opercle; par, parasphenoid; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; ra, remnant of a branchial arch; r.de, right dentary; r.pmx, right premaxilla; scl, supracleithrum; scl.b, broken sclerotic bones; smx1–2, supramaxillae 1–2; sorb, supraorbital; sop, subopercle; ur, urohyal; v.hy, ventral hypohyal; vo, vomer." figureDoi="http://doi.org/10.5281/zenodo.11045870" httpUri="https://zenodo.org/record/11045870/files/figure.png" pageId="18" pageNumber="49">4a</figureCitation>
|
||
, and
|
||
<figureCitation id="13642A02BA332F15FA81FB62FA92D379" box="[1282,1295,1260,1283]" captionStart="Figure 5" captionStartId="8.[124,191,942,964]" captionTargetBox="[367,1240,193,899]" captionTargetId="figure-11@8.[339,1246,177,911]" captionTargetPageId="8" captionText="Figure 5. Ebertichthys ettlingensis n. gen. et n. sp. Infraorbital bones, preopercle, suspensorium, and part of upper jaw (JME ETT 148). Abbreviations: hy, hyomandibula; io1–4, infraorbitals 1–4; io5?, broken infraorbital 5?; mtg, metapterygoid; mx, maxilla; pop, preopercle; qu, quadrate; smx1–2, supramaxillae 1–2." figureDoi="http://doi.org/10.5281/zenodo.11045872" httpUri="https://zenodo.org/record/11045872/files/figure.png" pageId="18" pageNumber="49">5</figureCitation>
|
||
with
|
||
<figureCitation id="13642A02BA332F15FACEFB62FA37D379" box="[1357,1450,1260,1283]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="18" pageNumber="49">Fig. 14b</figureCitation>
|
||
). Infraorbital 3 is a small bone at the posteroventral corner of the orbit (
|
||
<figureCitation id="13642A02BA332F15FC18FAA2FC6AD339" box="[923,1015,1324,1347]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="18" pageNumber="49">Fig. 14b</figureCitation>
|
||
). Because of its reduced size, a significant part of the quadrate is exposed. Although dorsoposterior infraorbitals 4 and 5 as the dorsal part of the preopercle are not preserved in the
|
||
<typeStatus id="54E48825BA332F15FC65FA02FBA0D3D9" box="[998,1085,1420,1443]" pageId="18" pageNumber="49" type="neotype">neotype</typeStatus>
|
||
, space left indicates that the region between the inner margin of infraorbitals 3–5 and the anterior margin of the preopercule was narrow.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA332F14FCB0FA65FF20D173" blockId="18.[819,1463,176,1921]" lastBlockId="19.[124,767,1746,1897]" lastPageId="19" lastPageNumber="50" pageId="18" pageNumber="49">
|
||
Upper jaw: It contains the premaxilla, maxilla, and two supramaxillae (
|
||
<figureCitation id="13642A02BA332F15FC54F985FBAED058" box="[983,1075,1547,1570]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="18" pageNumber="49">Fig. 14b</figureCitation>
|
||
), which are all preserved in the
|
||
<typeStatus id="54E48825BA332F15FA04F985FCF9D038" pageId="18" pageNumber="49" type="neotype">neotype</typeStatus>
|
||
. The re-study of the
|
||
<typeStatus id="54E48825BA332F15FBC6F9A5FB04D038" box="[1093,1177,1579,1602]" pageId="18" pageNumber="49" type="neotype">neotype</typeStatus>
|
||
confirms that the premaxilla has a short and small ascendent process as in the premaxilla in
|
||
<taxonomicName id="4C5F4D04BA332F15FCF4F9E4FC60D0F8" authority="Arratia, 2016" authorityName="Arratia" authorityYear="2016" box="[887,1021,1642,1666]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95BA332F15FCF4F9E4FC60D0F8" box="[887,1021,1642,1666]" italics="true" pageId="18" pageNumber="49">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEBA332F15FB80F9E5FBD6D0F8" box="[1027,1099,1643,1666]" pageId="18" pageNumber="49" rank="genus">n. gen.</taxonomicNameLabel>
|
||
However, the articulatory anterior process of the maxilla is not as long as that in
|
||
<taxonomicName id="4C5F4D04BA332F15FA8EF904FA0ED0D8" authorityName="Arratia" authorityYear="2016" box="[1293,1427,1674,1698]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA332F15FA8EF904FA0ED0D8" box="[1293,1427,1674,1698]" italics="true" pageId="18" pageNumber="49">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
, so that the posterior blade of the premaxilla is shorter. The maxilla is long and partially covers the anterior part of quadrate. The two supramaxillae of the
|
||
<typeStatus id="54E48825BA332F15FB0BF964FB41D17B" box="[1160,1244,1770,1793]" pageId="18" pageNumber="49" type="neotype">neotype</typeStatus>
|
||
are very similar to that of
|
||
<taxonomicName id="4C5F4D04BA332F15FCFCF884FB19D158" authorityName="Arratia" authorityYear="2016" box="[895,1156,1802,1826]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="49" phylum="Chordata" rank="species" species="ettlingensis">
|
||
<emphasis id="B92BEA95BA332F15FCFCF884FB19D158" box="[895,1156,1802,1826]" italics="true" pageId="18" pageNumber="49">Ebertichthys ettlingensis</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="13642A02BA332F15FB76F884FAD7D15B" box="[1269,1354,1802,1825]" captionStart="Figure 3" captionStartId="6.[124,191,1734,1756]" captionTargetBox="[264,1322,177,1693]" captionTargetId="figure-11@6.[264,1322,177,1703]" captionTargetPageId="6" captionText="Figure 3. Ebertichthys ettlingensis n. gen. et n. sp. in lateral view (holotype JME ETT 108). (a) Photograph of cranium and anterior part of body. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Drawing of cranium and anterior part of body. Abbreviations: ant, antorbital; br, branchiostegal rays; cl, cleithrum; cor, coracoid; dpa, dermopalatine; ent, entopterygoid; exc, extrascapula; hy, hyomandibula; io1-5, infraorbitals 1–5; iop, interopercle; l.de, left dentary; leth, lateral ethmoid; l.pmx, left premaxilla; met, mesethmoid; mx, maxilla; op, opercle; orb, orbitosphenoid; pa[= fr], parietal [= frontal bone of traditional terminology]; par, parasphenoid; pcl1-3, postcleithra 1–3; pop, preopercle; ppa[= pa], postparietal [= parietal bone of traditional terminology]; pt, pterotic; ptsp, pterosphenoid; ptt, posttemporal; qu, quadrate; sc, scales; sca, scapula; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sn, supraneurals; sorb.b, broken supraorbital bone; sop, subopercle; sy, symplectic; r.de, right dentary; r.pmx, right premaxilla; vo, vomer." figureDoi="http://doi.org/10.5281/zenodo.11045865" httpUri="https://zenodo.org/record/11045865/files/figure.png" pageId="18" pageNumber="49">Figs. 3b</figureCitation>
|
||
and
|
||
<figureCitation id="13642A02BA332F15FAFCF884FA37D15B" box="[1407,1450,1802,1825]" captionStart="Figure 12" captionStartId="16.[124,191,1206,1228]" captionTargetBox="[180,1415,194,1164]" captionTargetId="figure-11@16.[169,1416,177,1175]" captionTargetPageId="16" captionText="Figure 12. Ebertichthys ettlingensis n. gen. et n. sp. Caudal fin and endoskeleton in lateral view (holotype JME ETT 108). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; ebfu, epaxial basal fulcra; ffu, fringing fulcra; H1–7, hypurals 1–7; hsPU4, haemal spine of preural centrum 4; nsPU2–4, neural spines of preural centra 2–4; naU1+2, neural arches of ural centrum 1 + 2; PH, parhypural; prp, procurrent rays; PU1, 4, preural centra 1, 4; U1+2, 3, ural centrum 1 + 2 and 3 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermal”; UN1–4, uroneurals 1–4; PR1–19, principal rays 1–19; v.csu, ventral caudal scute." figureDoi="http://doi.org/10.5281/zenodo.11045894" httpUri="https://zenodo.org/record/11045894/files/figure.png" pageId="18" pageNumber="49">12b</figureCitation>
|
||
). Supramaxilla 2 is broadly expanded posteriorly, forming the main body of the bone. The anterodorsal process is very long and narrow. Supramaxilla 1 is elongate and ovoid-shaped. Both supramaxillae cover the dorsal margin of the maxillary blade.
|
||
</paragraph>
|
||
<caption id="DF20660FBA322F14FFFFF9AFFAE7D0F5" ID-DOI="http://doi.org/10.5281/zenodo.11045902" ID-Zenodo-Dep="11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="19" pageNumber="50" startId="19.[124,191,1569,1591]" targetBox="[169,1416,177,1526]" targetPageId="19" targetType="figure">
|
||
<paragraph id="8BE03687BA322F14FFFFF9AFFAE7D0F5" blockId="19.[124,1462,1569,1679]" pageId="19" pageNumber="50">
|
||
<emphasis id="B92BEA95BA322F14FFFFF9AFFF78D04D" bold="true" box="[124,229,1569,1591]" pageId="19" pageNumber="50">Figure 14.</emphasis>
|
||
<taxonomicName id="4C5F4D04BA322F14FF68F9ACFE0FD04D" box="[235,402,1570,1591]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="50" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA322F14FF68F9ACFE0FD04D" box="[235,402,1570,1591]" italics="true" pageId="19" pageNumber="50">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
(neotype, JME SOS 537).
|
||
<emphasis id="B92BEA95BA322F14FD19F9AFFD2BD04D" bold="true" box="[666,694,1569,1591]" pageId="19" pageNumber="50">(a)</emphasis>
|
||
Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm.
|
||
<emphasis id="B92BEA95BA322F14FF7EF9B0FE87D02E" bold="true" box="[253,282,1598,1620]" pageId="19" pageNumber="50">(b)</emphasis>
|
||
Cranium in lateral view and enlargement of premaxilla
|
||
<emphasis id="B92BEA95BA322F14FCC4F9B0FCFCD02E" bold="true" box="[839,865,1598,1620]" pageId="19" pageNumber="50">(c)</emphasis>
|
||
. Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE03687BA322F13FFFFF89CFD11D16F" blockId="19.[124,767,1746,1897]" lastBlockId="20.[124,767,1694,1908]" lastPageId="20" lastPageNumber="51" pageId="19" pageNumber="50">
|
||
Lower jaw: The mandible (
|
||
<figureCitation id="13642A02BA322F14FE1FF89CFE6BD153" box="[412,502,1810,1833]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="19" pageNumber="50">Fig. 14b</figureCitation>
|
||
) is strongly ossified, and although it is largely hidden by the maxilla and supramaxillae, the large and high coronoid process is visible in the
|
||
<typeStatus id="54E48825BA322F14FCB0F95CFC16D093" box="[819,907,1746,1769]" pageId="19" pageNumber="50" type="neotype">neotype</typeStatus>
|
||
. The articulation of the mandible with the quadrate is placed at the posterior margin of the orbit in the
|
||
<typeStatus id="54E48825BA322F14FAE1F97CFA2BD173" box="[1378,1462,1778,1801]" pageId="19" pageNumber="50" type="neotype">neotype</typeStatus>
|
||
(specimen with almost closed mouth), which is in contrast to an anterior position in specimens with the mouth open. A “leptolepid” notch is present in the ascending dorsal margin of the dentary in specimen JME
|
||
<specimenCode id="DBF99EFCBA352F13FE57F910FDDED0CF" box="[468,579,1694,1717]" collectionCode="SOS" pageId="20" pageNumber="51">SOS 3228</specimenCode>
|
||
. This notch is not observed in the
|
||
<typeStatus id="54E48825BA352F13FEABF930FEE2D0AF" box="[296,383,1726,1749]" pageId="20" pageNumber="51" type="neotype">neotype</typeStatus>
|
||
, because the maxilla covers this region. The “leptolepid” notch is narrow and partially forms the anterior wall of the massive coronoid process.
|
||
</paragraph>
|
||
<caption id="DF20660FBA352F13FFFFF984FE67D021" ID-DOI="http://doi.org/10.5281/zenodo.11045904" ID-Zenodo-Dep="11045904" httpUri="https://zenodo.org/record/11045904/files/figure.png" pageId="20" pageNumber="51" startId="20.[124,191,1546,1568]" targetBox="[169,1416,177,1515]" targetPageId="20" targetType="figure">
|
||
<paragraph id="8BE03687BA352F13FFFFF984FE67D021" blockId="20.[124,1462,1546,1627]" pageId="20" pageNumber="51">
|
||
<emphasis id="B92BEA95BA352F13FFFFF984FF7ED05A" bold="true" box="[124,227,1546,1568]" pageId="20" pageNumber="51">Figure 15.</emphasis>
|
||
<taxonomicName id="4C5F4D04BA352F13FF6AF985FE13D05A" box="[233,398,1547,1568]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="51" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA352F13FF6AF985FE13D05A" box="[233,398,1547,1568]" italics="true" pageId="20" pageNumber="51">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
from Schernfeld in the Eichstätt Basin. Uncatalogued specimen from U. Eller (Dümpelfeld).
|
||
<emphasis id="B92BEA95BA352F13FA8FF984FAB5D05A" bold="true" box="[1292,1320,1546,1568]" pageId="20" pageNumber="51">(a)</emphasis>
|
||
Photograph of specimen in lateral view.
|
||
<emphasis id="B92BEA95BA352F13FEF6F9A6FE0CD044" bold="true" box="[373,401,1576,1598]" pageId="20" pageNumber="51">(b)</emphasis>
|
||
Enlargement of the head. Note the peculiar shape and distribution of the sensory tubules of the preopercular canal. Photographs courtesy of U. Eller.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE03687BA352F12FFFFF890FE4FD54D" blockId="20.[124,767,1694,1908]" lastBlockId="21.[124,767,736,1907]" lastPageId="21" lastPageNumber="52" pageId="20" pageNumber="51">
|
||
Opercular bones: The preopercle (
|
||
<figureCitation id="13642A02BA352F13FE73F890FDC8D14F" box="[496,597,1822,1845]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="20" pageNumber="51">Figs. 14b</figureCitation>
|
||
and
|
||
<figureCitation id="13642A02BA352F13FD0EF890FD2BD14F" box="[653,694,1822,1845]" captionStart="Figure 15" captionStartId="20.[124,191,1546,1568]" captionTargetBox="[169,1416,177,1515]" captionTargetId="figure-11@20.[169,1416,177,1515]" captionTargetPageId="20" captionText="Figure 15. Ascalabos voithii from Schernfeld in the Eichstätt Basin. Uncatalogued specimen from U. Eller (Dümpelfeld). (a) Photograph of specimen in lateral view. (b) Enlargement of the head. Note the peculiar shape and distribution of the sensory tubules of the preopercular canal. Photographs courtesy of U. Eller." figureDoi="http://doi.org/10.5281/zenodo.11045904" httpUri="https://zenodo.org/record/11045904/files/figure.png" pageId="20" pageNumber="51">15b</figureCitation>
|
||
) is incompletely preserved in the
|
||
<typeStatus id="54E48825BA352F13FE20F8B0FE67D12F" box="[419,506,1854,1877]" pageId="20" pageNumber="51" type="neotype">neotype</typeStatus>
|
||
, and it is probably somehow displaced. The important aspect to be noted is that the bone presents a slightly rounded bony flange at its anterior margin, similar to that in
|
||
<taxonomicName id="4C5F4D04BA352F13FBF7F930FB67D0AC" authority="Arratia, 2016" authorityName="Arratia" authorityYear="2016" box="[1140,1274,1726,1750]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="51" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95BA352F13FBF7F930FB67D0AC" box="[1140,1274,1726,1750]" italics="true" pageId="20" pageNumber="51">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEBA352F13FA82F930FAD6D0AF" box="[1281,1355,1726,1749]" pageId="20" pageNumber="51" rank="genus">n. gen.</taxonomicNameLabel>
|
||
(compare
|
||
<figureCitation id="13642A02BA352F13FCB0F950FC16D08F" box="[819,907,1758,1781]" captionStart="Figure 3" captionStartId="6.[124,191,1734,1756]" captionTargetBox="[264,1322,177,1693]" captionTargetId="figure-11@6.[264,1322,177,1703]" captionTargetPageId="6" captionText="Figure 3. Ebertichthys ettlingensis n. gen. et n. sp. in lateral view (holotype JME ETT 108). (a) Photograph of cranium and anterior part of body. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Drawing of cranium and anterior part of body. Abbreviations: ant, antorbital; br, branchiostegal rays; cl, cleithrum; cor, coracoid; dpa, dermopalatine; ent, entopterygoid; exc, extrascapula; hy, hyomandibula; io1-5, infraorbitals 1–5; iop, interopercle; l.de, left dentary; leth, lateral ethmoid; l.pmx, left premaxilla; met, mesethmoid; mx, maxilla; op, opercle; orb, orbitosphenoid; pa[= fr], parietal [= frontal bone of traditional terminology]; par, parasphenoid; pcl1-3, postcleithra 1–3; pop, preopercle; ppa[= pa], postparietal [= parietal bone of traditional terminology]; pt, pterotic; ptsp, pterosphenoid; ptt, posttemporal; qu, quadrate; sc, scales; sca, scapula; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sn, supraneurals; sorb.b, broken supraorbital bone; sop, subopercle; sy, symplectic; r.de, right dentary; r.pmx, right premaxilla; vo, vomer." figureDoi="http://doi.org/10.5281/zenodo.11045865" httpUri="https://zenodo.org/record/11045865/files/figure.png" pageId="20" pageNumber="51">Figs. 3b</figureCitation>
|
||
and
|
||
<figureCitation id="13642A02BA352F13FC47F950FC40D08F" box="[964,989,1758,1781]" captionStart="Figure 4" captionStartId="7.[124,191,1748,1770]" captionTargetBox="[265,1294,178,1710]" captionTargetId="figure-11@7.[264,1322,177,1716]" captionTargetPageId="7" captionText="Figure 4. Ebertichthys ettlingensis n. gen. et n. sp. Crania in lateral views illustrating specific morphological features. (a) JME ETT 60. (b) JME ETT 24. Arrow points to a small toothed patch. Abbreviations: ang, angular; aup, autopalatine; b.ant, broken antorbital; a.cer, anterior ceratohyal; br, branchiostegal rays; cl, cleithrum; de, dentary; ect, ectopterygoid; ent, entopterygoid; ghy, glossohyal; d.hy, dorsal hypohyal; iop, interopercle; hy, hyomandibula; io1–5, infraorbitals 1–5; l.ang, left angular; l.de, left dentary; l.pmx, left premaxilla; met, mesethmoid; mtg, metapterygoid; mx, maxilla; op, opercle; par, parasphenoid; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; ra, remnant of a branchial arch; r.de, right dentary; r.pmx, right premaxilla; scl, supracleithrum; scl.b, broken sclerotic bones; smx1–2, supramaxillae 1–2; sorb, supraorbital; sop, subopercle; ur, urohyal; v.hy, ventral hypohyal; vo, vomer." figureDoi="http://doi.org/10.5281/zenodo.11045870" httpUri="https://zenodo.org/record/11045870/files/figure.png" pageId="20" pageNumber="51">4a</figureCitation>
|
||
with
|
||
<figureCitation id="13642A02BA352F13FB9DF950FBE1D08F" box="[1054,1148,1758,1781]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="20" pageNumber="51">Fig. 14b</figureCitation>
|
||
). Since this feature has been observed only in a few specimens due to poor preservation, I have not considered it as a diagnostic character of the family, but this feature should be re-evaluated when more specimens of
|
||
<taxonomicName id="4C5F4D04BA352F13FC0DF8D3FC66D10F" box="[910,1019,1885,1909]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="51" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA352F13FC0DF8D3FC66D10F" box="[910,1019,1885,1909]" italics="true" pageId="20" pageNumber="51">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
become available. The characteristic sensory tubules of the preopercular sensory canal are nicely preserved in the
|
||
<typeStatus id="54E48825BA342F12FE8DFC8EFEFFD56D" box="[270,354,768,791]" pageId="21" pageNumber="52" type="neotype">neotype</typeStatus>
|
||
(
|
||
<figureCitation id="13642A02BA342F12FEF1FC8EFE4DD56D" box="[370,464,768,791]" captionStart="Figure 14" captionStartId="19.[124,191,1569,1591]" captionTargetBox="[169,1416,177,1526]" captionTargetId="figure-11@19.[169,1416,177,1538]" captionTargetPageId="19" captionText="Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic." figureDoi="http://doi.org/10.5281/zenodo.11045902" httpUri="https://zenodo.org/record/11045902/files/figure.png" pageId="21" pageNumber="52">Fig. 14b</figureCitation>
|
||
), as well as in an additional specimen illustrated in
|
||
<figureCitation id="13642A02BA342F12FEF0FCAEFE53D54D" box="[371,462,800,823]" captionStart="Figure 15" captionStartId="20.[124,191,1546,1568]" captionTargetBox="[169,1416,177,1515]" captionTargetId="figure-11@20.[169,1416,177,1515]" captionTargetPageId="20" captionText="Figure 15. Ascalabos voithii from Schernfeld in the Eichstätt Basin. Uncatalogued specimen from U. Eller (Dümpelfeld). (a) Photograph of specimen in lateral view. (b) Enlargement of the head. Note the peculiar shape and distribution of the sensory tubules of the preopercular canal. Photographs courtesy of U. Eller." figureDoi="http://doi.org/10.5281/zenodo.11045904" httpUri="https://zenodo.org/record/11045904/files/figure.png" pageId="21" pageNumber="52">Fig. 15b</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF20660FBA342F12FFFFFD09FC25D4E7" ID-DOI="http://doi.org/10.5281/zenodo.11045906" ID-Zenodo-Dep="11045906" box="[124,952,647,669]" httpUri="https://zenodo.org/record/11045906/files/figure.png" pageId="21" pageNumber="52" startId="21.[124,191,647,669]" targetBox="[169,1416,177,615]" targetPageId="21" targetType="figure">
|
||
<paragraph id="8BE03687BA342F12FFFFFD09FC25D4E7" blockId="21.[124,952,647,669]" box="[124,952,647,669]" pageId="21" pageNumber="52">
|
||
<emphasis id="B92BEA95BA342F12FFFFFD09FF7ED4E7" bold="true" box="[124,227,647,669]" pageId="21" pageNumber="52">Figure 16.</emphasis>
|
||
<taxonomicName id="4C5F4D04BA342F12FF6AFD09FE13D4E6" box="[233,398,647,668]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA342F12FF6AFD09FE13D4E6" box="[233,398,647,668]" italics="true" pageId="21" pageNumber="52">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
. Restoration in lateral view, updated from
|
||
<bibRefCitation id="EFCE4B76BA342F12FCA4FD06FC2ED4E7" author="Arratia, G." box="[807,947,648,669]" pageId="21" pageNumber="52" pagination="1 - 168" refId="ref17245" refString="Arratia, G.: Basal teleosts and teleostean phylogeny, Palaeo Ichthyol., 7, 1 - 168, 1997." type="journal article" year="1997">Arratia (1997)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE03687BA342F12FFFFFCCEFEBED34F" blockId="21.[124,767,736,1907]" pageId="21" pageNumber="52">
|
||
Vertebral column: A long and prominent lateral crest (
|
||
<figureCitation id="13642A02BA342F12FF07FCD1FF4CD50C" box="[132,209,863,886]" captionStart="Figure 17" captionStartId="22.[124,191,738,760]" captionTargetBox="[369,1215,187,694]" captionTargetId="figure-11@22.[358,1227,177,706]" captionTargetPageId="22" captionText="Figure 17. Ascalabos voithii (neotype, JME SOS 537). Midcaudal vertebrae illustrating the presence of a lateral crest and grooves on the lateral side of the vertebral centra. Arrows point to a lateral longitudinal crest. Abbreviations: na, neural arch; ns, neural spine; ha, haemal arch; hs, haemal spine; vc, vertebral centrum." figureDoi="http://doi.org/10.5281/zenodo.11045908" httpUri="https://zenodo.org/record/11045908/files/figure.png" pageId="21" pageNumber="52">Fig. 17</figureCitation>
|
||
) characterizes the centra of the caudal region of the vertebral column in large specimens. The prominent crest is still present on the lateral surface of preural centrum 2 (
|
||
<figureCitation id="13642A02BA342F12FF07FC31FF4AD5AC" box="[132,215,959,982]" captionStart="Figure 18" captionStartId="23.[124,191,1346,1368]" captionTargetBox="[146,1448,197,1298]" captionTargetId="figure-11@23.[132,1455,177,1315]" captionTargetPageId="23" captionText="Figure 18. Ascalabos voithii. Caudal skeleton in lateral view slightly modified from Arratia (1997). Abbreviations: Ant, anterior; dpr, dorsal or epaxial procurrent rays; d.scu, dorsal caudal scute; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–11, hypurals 1–11; hsPU2, haemal spine of preural centrum 2; mo, membranous outgrowth on anterodorsal margin of first uroneural; mo’, membranous outgrowth on anterodorsal margin of second uroneural; nsPU2–4, neural spine of preural centra 2–4; PR1,19, principal caudal rays 1, 19; rer, rudimentary fin ray; U1+2+H1–2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–7, uroneurals 1–7; v.scu., ventral caudal scute." figureDoi="http://doi.org/10.5281/zenodo.11045910" httpUri="https://zenodo.org/record/11045910/files/figure.png" pageId="21" pageNumber="52">Fig. 18</figureCitation>
|
||
). This is very different to the condition in
|
||
<taxonomicName id="4C5F4D04BA342F12FD40FC31FF53D58D" authorityName="Arratia" authorityYear="2016" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FD40FC31FF53D58D" italics="true" pageId="21" pageNumber="52">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
with smooth centra.
|
||
<taxonomicName id="4C5F4D04BA342F12FE2DFC51FD86D58D" box="[430,539,991,1015]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FE2DFC51FD86D58D" box="[430,539,991,1015]" italics="true" pageId="21" pageNumber="52">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
may have from 34 to 39 vertebrae (commonly 36), whereas in
|
||
<taxonomicName id="4C5F4D04BA342F12FDCEFC70FD4ED26C" authorityName="Arratia" authorityYear="2016" box="[589,723,1022,1046]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FDCEFC70FD4ED26C" box="[589,723,1022,1046]" italics="true" pageId="21" pageNumber="52">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
the number is commonly 38 or 39. These represent the lowest counts among teleosts from the Solnhofen limestones (see Arratia and
|
||
<bibRefCitation id="EFCE4B76BA342F12FF7FFBD1FE3DD20C" author="Schultze, H. - P." box="[252,416,1119,1142]" pageId="21" pageNumber="52" pagination="560 - 672" refId="ref19148" refString="Schultze, H. - P.: Liste der Fossilien, in: Solnhofen, ein Frenster in die Jurazeit, edited by: Arratia, G., Schultze, H. - P., Tischlinger, H., and Viohl, G., Verlag Dr. F. Pfeil, Munchen, 560 - 672, 2015." type="book chapter" year="2015">Schultze, 2015</bibRefCitation>
|
||
). There are 16 to 19 pairs of ribs (
|
||
<figureCitation id="13642A02BA342F12FF07FBF0FF4ED2EF" box="[132,211,1150,1173]" captionStart="Figure 16" captionStartId="21.[124,191,647,669]" captionTargetBox="[169,1416,177,615]" captionTargetId="figure-11@21.[169,1416,177,615]" captionTargetPageId="21" captionText="Figure 16. Ascalabos voithii. Restoration in lateral view, updated from Arratia (1997)." figureDoi="http://doi.org/10.5281/zenodo.11045906" httpUri="https://zenodo.org/record/11045906/files/figure.png" pageId="21" pageNumber="52">Fig. 16</figureCitation>
|
||
), whereas 18 or 19 pairs of ribs are present in
|
||
<taxonomicName id="4C5F4D04BA342F12FD40FBF0FECCD2CC" authorityName="Arratia" authorityYear="2016" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="species" species="ettlingensis">
|
||
<emphasis id="B92BEA95BA342F12FD40FBF0FECCD2CC" italics="true" pageId="21" pageNumber="52">Ebertichthys ettlingensis</emphasis>
|
||
</taxonomicName>
|
||
. A series of about 14 supraneurals extend between the occiput and below the first dorsal pterygiophores, whereas 16 supraneurals are commonly found in
|
||
<taxonomicName id="4C5F4D04BA342F12FFFFFB73FE9FD36F" authorityName="Arratia" authorityYear="2016" box="[124,258,1277,1301]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FFFFFB73FE9FD36F" box="[124,258,1277,1301]" italics="true" pageId="21" pageNumber="52">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
, but they do not extend below the first dorsal pterygiophores.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA342F12FF14FAB0FE65D0C9" blockId="21.[124,767,736,1907]" pageId="21" pageNumber="52">
|
||
The early formation of the centra of
|
||
<taxonomicName id="4C5F4D04BA342F12FDACFAB3FF23D30F" authorityName="Arratia" authorityYear="2016" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="species" species="ettlingensis">
|
||
<emphasis id="B92BEA95BA342F12FDACFAB3FF23D30F" italics="true" pageId="21" pageNumber="52">Ebertichthys ettlingensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13642A02BA342F12FF51FAD0FE8AD30F" box="[210,279,1374,1397]" captionStart="Figure 9" captionStartId="13.[124,191,1334,1356]" captionTargetBox="[172,1415,177,1291]" captionTargetId="figure-11@13.[172,1415,177,1303]" captionTargetPageId="13" captionText="Figure 9. Ebertichthys ettlingensis n. gen. et n. sp. (a) Photograph of young specimen (JME ETT 61) in lateral view. Photograph courtesy of H. Tischlinger, Scale equals 1 cm. (b) Detail of last caudal vertebrae illustrating pseudo-diplospondyly and caudal endoskeleton incompletely preserved. Abbreviations: chcPU4, chordacentrum of preural centrum 4; chcU1–3, chordacentra of ural centra 1–3; E, epurals (the first one is missing); H1–7, hypurals 1–7; hsPU2, haemal spine of preural centrum 2; PH, parhypural; ‘UD’, “urodermal”; UN1–6, uroneurals 1–6." figureDoi="http://doi.org/10.5281/zenodo.11045886" httpUri="https://zenodo.org/record/11045886/files/figure.png" pageId="21" pageNumber="52">Fig. 9</figureCitation>
|
||
) is similar to that of
|
||
<taxonomicName id="4C5F4D04BA342F12FD94FAD3FD49D30F" box="[535,724,1373,1397]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA342F12FD94FAD3FD49D30F" box="[535,724,1373,1397]" italics="true" pageId="21" pageNumber="52">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
, so that the description above stands for both species. My observations (here and in Arratia, 1991, p. 257) and interpretations differ from those of
|
||
<bibRefCitation id="EFCE4B76BA342F12FE29FA33FD67D3AE" author="Schaeffer, B. & Patterson, C." box="[426,762,1469,1492]" pageId="21" pageNumber="52" pagination="1 - 86" refId="ref18949" refString="Schaeffer, B. and Patterson, C.: Jurassic fishes from the western United States, with comments on Jurassic fish distribution, Am. Museum Novit., 2796, 1 - 86, 1984." type="journal article" year="1984">Schaeffer and Patterson (1984)</bibRefCitation>
|
||
, who interpreted the centra of young
|
||
<taxonomicName id="4C5F4D04BA342F12FDA4FA53FDEBD38F" box="[551,630,1501,1525]" class="Gastropoda" family="Skeneidae" genus="Tharsis" kingdom="Animalia" order="Trochida" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FDA4FA53FDEBD38F" box="[551,630,1501,1525]" italics="true" pageId="21" pageNumber="52">Tharsis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5F4D04BA342F12FD08FA53FD65D38F" box="[651,760,1501,1525]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FD08FA53FD65D38F" box="[651,760,1501,1525]" italics="true" pageId="21" pageNumber="52">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
, and lycopterid osteoglossomorphs as diplospondylous, each formed by a precentrum and a postcentrum perichondrally ossified, and that the chordacentrum is then reabsorbed or suppressed in the middle of the centrum. These interpretations are not supported by my observations in young
|
||
<taxonomicName id="4C5F4D04BA342F12FD2BF9F2FD6AD0EE" box="[680,759,1660,1684]" class="Gastropoda" family="Skeneidae" genus="Tharsis" kingdom="Animalia" order="Trochida" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FD2BF9F2FD6AD0EE" box="[680,759,1660,1684]" italics="true" pageId="21" pageNumber="52">Tharsis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5F4D04BA342F12FFFFF912FF74D0CE" box="[124,233,1692,1716]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FFFFF912FF74D0CE" box="[124,233,1692,1716]" italics="true" pageId="21" pageNumber="52">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5F4D04BA342F12FEA0F912FE34D0CE" authority="Arratia, 2016" authorityName="Arratia" authorityYear="2016" box="[291,425,1692,1716]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95BA342F12FEA0F912FE34D0CE" box="[291,425,1692,1716]" italics="true" pageId="21" pageNumber="52">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEBA342F12FE33F912FE65D0C9" box="[432,504,1692,1715]" pageId="21" pageNumber="52" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA342F12FFFFF932FD51D109" blockId="21.[124,767,736,1907]" pageId="21" pageNumber="52">
|
||
Pelvic plate or basipterygium: The basipterygium of
|
||
<taxonomicName id="4C5F4D04BA342F12FD38F932FF33D08E" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FD38F932FF33D08E" italics="true" pageId="21" pageNumber="52">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
is characterized by its slightly triangular shape and its posterior expansion that has a short and sharp medial posterior process (
|
||
<bibRefCitation id="EFCE4B76BA342F12FEC7F892FE47D149" author="Arratia, G." box="[324,474,1820,1843]" pageId="21" pageNumber="52" pagination="1 - 168" refId="ref17245" refString="Arratia, G.: Basal teleosts and teleostean phylogeny, Palaeo Ichthyol., 7, 1 - 168, 1997." type="journal article" year="1997">Arratia, 1997</bibRefCitation>
|
||
,
|
||
<figureCitation id="13642A02BA342F12FE64F892FDD9D149" box="[487,580,1820,1843]" captionStart="Figure 19" captionStartId="24.[124,191,1198,1220]" captionTargetBox="[169,1407,185,1146]" captionTargetId="figure-11@24.[169,1416,177,1167]" captionTargetPageId="24" captionText="Figure 19. Ascalabos voithii. Caudal skeleton in lateral view with only two epaxial basal fulcra preserved (neotype, JME SOS 537). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; dpr, dorsal or epaxial procurrent rays; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–4, hypurals 1–4; hbfu, hypaxial basal fulcra; hsPU4, haemal spine of preural centrum 4; naPU1, neural arch of preural centrum 1; nsPU2–4, neural spine of preural centra 2–4; PH, parhypural; PU1,4, preural centrum 1, 4; rer, rudimentary fin ray; sc, dorsal and ventral caudal scutes; U1+2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–8, uroneurals 1–8; PR1–19, principal rays 1–19; vpr, ventral procurrent rays; v.scu, ventral caudal scute." figureDoi="http://doi.org/10.5281/zenodo.11045912" httpUri="https://zenodo.org/record/11045912/files/figure.png" pageId="21" pageNumber="52">Fig. 19c</figureCitation>
|
||
). In contrast, the basipterygium of
|
||
<taxonomicName id="4C5F4D04BA342F12FEB6F8B5FDA4D129" authorityName="Arratia" authorityYear="2016" box="[309,569,1851,1875]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="species" species="ettlingensis">
|
||
<emphasis id="B92BEA95BA342F12FEB6F8B5FDA4D129" box="[309,569,1851,1875]" italics="true" pageId="21" pageNumber="52">Ebertichthys ettlingensis</emphasis>
|
||
</taxonomicName>
|
||
lacks the posterior expansion, and a medial process has not been observed.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA342F12FCB0FD6EFC30D22C" blockId="21.[819,1462,736,1907]" pageId="21" pageNumber="52">
|
||
Dorsal and anal fins and their supports: The first dorsal pterygiophore is characteristically expanded and with a few anteroventral processes in
|
||
<taxonomicName id="4C5F4D04BA342F12FBC5FC91FACFD54D" authorityName="Arratia" authorityYear="2016" box="[1094,1362,799,823]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="species" species="ettlingensis">
|
||
<emphasis id="B92BEA95BA342F12FBC5FC91FACFD54D" box="[1094,1362,799,823]" italics="true" pageId="21" pageNumber="52">Ebertichthys ettlingensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13642A02BA342F12FAE4FCAEFA2BD54D" box="[1383,1462,800,823]" captionStart="Figure 7" captionStartId="11.[124,191,1811,1833]" captionTargetBox="[169,1413,182,1776]" captionTargetId="figure-11@11.[169,1416,177,1780]" captionTargetPageId="11" captionText="Figure 7. Ebertichthys ettlingensis n. gen. et n. sp. Dorsal and anal fins and their relationships to the vertebral column (JME ETT 132a). Abbreviations: Ant, anterior; an.r, principal anal rays; epl.b, epipleural bones; dor.r, dorsal principal rays; epin.p, epineural processes; ha, haemal arch; hs, haemal spine; la.pt, last anal pterygiophore; ld.pt, last dorsal pterygiophore; na, neural arch; ns, neural spine; paph, parapophyses; pr.anr, anal procurrent rays; pr.dr, dorsal procurrent rays; ri, ribs; sn, supraneural bones; V21, vertebra 21; 1st a.pt, first anal pterygiophore; 1st d.pt, first dorsal pterygiophore." figureDoi="http://doi.org/10.5281/zenodo.11045878" httpUri="https://zenodo.org/record/11045878/files/figure.png" pageId="21" pageNumber="52">Figs. 7</figureCitation>
|
||
and
|
||
<figureCitation id="13642A02BA342F12FCE7FCCEFC10D52D" box="[868,909,832,855]" captionStart="Figure 11" captionStartId="15.[124,191,836,858]" captionTargetBox="[169,1408,182,799]" captionTargetId="figure-11@15.[169,1416,177,805]" captionTargetPageId="15" captionText="Figure 11. Ebertichthys ettlingensis n. gen. et n. sp. (JME ETT 64a). (a) Details of first two dorsal pterygiophores and first dorsal-fin rays. (b) Detail of last dorsal pterygiophores. Arrows point to the two articular areas for fin rays. Abbreviations: Ant, anterior; 1st d.pt, first dorsal pterygiophore; 2nd d.pt, second dorsal pterygiophore; ld.pt, last dorsal pterygiophore." figureDoi="http://doi.org/10.5281/zenodo.11045892" httpUri="https://zenodo.org/record/11045892/files/figure.png" pageId="21" pageNumber="52">11a</figureCitation>
|
||
), and a similar pterygiophore (
|
||
<figureCitation id="13642A02BA342F12FB6AFCCEFAA7D52D" box="[1257,1338,832,855]" captionStart="Figure 16" captionStartId="21.[124,191,647,669]" captionTargetBox="[169,1416,177,615]" captionTargetId="figure-11@21.[169,1416,177,615]" captionTargetPageId="21" captionText="Figure 16. Ascalabos voithii. Restoration in lateral view, updated from Arratia (1997)." figureDoi="http://doi.org/10.5281/zenodo.11045906" httpUri="https://zenodo.org/record/11045906/files/figure.png" pageId="21" pageNumber="52">Fig. 16</figureCitation>
|
||
) is present in the
|
||
<typeStatus id="54E48825BA342F12FCFFFCD1FC4DD50C" box="[892,976,863,886]" pageId="21" pageNumber="52" type="neotype">neotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5F4D04BA342F12FC79FCD1FBFAD50D" box="[1018,1127,863,887]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FC79FCD1FBFAD50D" box="[1018,1127,863,887]" italics="true" pageId="21" pageNumber="52">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
. A first pterygiophore bearing only two processes (JME
|
||
<collectionCode id="ED4EAE42BA342F12FBD1FCF1FB1ED5EC" box="[1106,1155,895,918]" pageId="21" pageNumber="52">SOS</collectionCode>
|
||
2886) and another partially broken one (
|
||
<collectionCode id="ED4EAE42BA342F12FC3FFC11FC7CD5CC" box="[956,993,927,950]" country="USA" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" pageId="21" pageNumber="52" type="Herbarium">HL</collectionCode>
|
||
309) were illustrated for
|
||
<taxonomicName id="4C5F4D04BA342F12FB7AFC11FAFBD5CD" box="[1273,1382,927,951]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FB7AFC11FAFBD5CD" box="[1273,1382,927,951]" italics="true" pageId="21" pageNumber="52">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFCE4B76BA342F12FA11FC11FC30D5AC" author="Arratia, G." pageId="21" pageNumber="52" pagination="1 - 168" refId="ref17245" refString="Arratia, G.: Basal teleosts and teleostean phylogeny, Palaeo Ichthyol., 7, 1 - 168, 1997." type="journal article" year="1997">Arratia (1997</bibRefCitation>
|
||
,
|
||
<figureCitation id="13642A02BA342F12FC3FFC31FBA8D5AC" box="[956,1077,959,982]" captionStart="Figure 19" captionStartId="24.[124,191,1198,1220]" captionTargetBox="[169,1407,185,1146]" captionTargetId="figure-11@24.[169,1416,177,1167]" captionTargetPageId="24" captionText="Figure 19. Ascalabos voithii. Caudal skeleton in lateral view with only two epaxial basal fulcra preserved (neotype, JME SOS 537). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; dpr, dorsal or epaxial procurrent rays; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–4, hypurals 1–4; hbfu, hypaxial basal fulcra; hsPU4, haemal spine of preural centrum 4; naPU1, neural arch of preural centrum 1; nsPU2–4, neural spine of preural centra 2–4; PH, parhypural; PU1,4, preural centrum 1, 4; rer, rudimentary fin ray; sc, dorsal and ventral caudal scutes; U1+2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–8, uroneurals 1–8; PR1–19, principal rays 1–19; vpr, ventral procurrent rays; v.scu, ventral caudal scute." figureDoi="http://doi.org/10.5281/zenodo.11045912" httpUri="https://zenodo.org/record/11045912/files/figure.png" pageId="21" pageNumber="52">Fig. 19a–b</figureCitation>
|
||
). Currently, I interpret these bones as partially incomplete. According to the evidence gathered here, the characteristic shape of the first dorsal pterygiophore is now proposed as a diagnostic feature of the new family
|
||
<taxonomicName id="4C5F4D04BA342F12FA13FB91FC35D22C" authorityName="Arratia" authorityYear="2016" family="Ascalaboidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="family">Ascalaboidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA342F12FCCDFBD1FCE4D28F" blockId="21.[819,1462,736,1907]" pageId="21" pageNumber="52">
|
||
The first anal pterygiophore is long, broadly expanded and almost rectangular in outline in
|
||
<taxonomicName id="4C5F4D04BA342F12FB06FBF0FA96D2EC" authorityName="Arratia" authorityYear="2016" box="[1157,1291,1150,1174]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FB06FBF0FA96D2EC" box="[1157,1291,1150,1174]" italics="true" pageId="21" pageNumber="52">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
. In contrast, the pterygiophore is thinner and more rod-like in
|
||
<taxonomicName id="4C5F4D04BA342F12FAA9FB10FA0AD2CC" box="[1322,1431,1182,1206]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA342F12FAA9FB10FA0AD2CC" box="[1322,1431,1182,1206]" italics="true" pageId="21" pageNumber="52">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
as well as in other Jurassic teleosts (see
|
||
<bibRefCitation id="EFCE4B76BA342F12FB5BFB30FAF6D2AF" author="Arratia, G." box="[1240,1387,1214,1237]" pageId="21" pageNumber="52" pagination="107 - 139" refId="ref17050" refString="Arratia, G.: Varasichthys ariasi n. gen. et sp. from the Upper Jurassic from Chile (Pisces, Teleostei, Varasichthyidae n. fam.), Palaeontographica Abt. A, 175, 107 - 139, 1981." type="journal article" year="1981">Arratia, 1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCE4B76BA342F12FAF9FB30FA2FD2AF" author="Arratia, G." box="[1402,1458,1214,1237]" pageId="21" pageNumber="52" pagination="1 - 168" refId="ref17245" refString="Arratia, G.: Basal teleosts and teleostean phylogeny, Palaeo Ichthyol., 7, 1 - 168, 1997." type="journal article" year="1997">1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCE4B76BA342F12FCB0FB50FCF0D28F" author="Arratia, G." box="[819,877,1246,1269]" pageId="21" pageNumber="52" pagination="137 - 179" refId="ref17323" refString="Arratia, G.: Remarkable teleostean fishes from the Late Jurassic of southern Germany and their phylogenetic relationships, Foss. Rec., 3, 137 - 179, doi: 10.5194 / fr- 3 - 137 - 2000, 2000." type="journal article" year="2000">2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA342F12FCB0FB70FB82D0A9" blockId="21.[819,1462,736,1907]" pageId="21" pageNumber="52">
|
||
Caudal skeleton and fin: The caudal skeleton of the
|
||
<typeStatus id="54E48825BA342F12FA04FB70FCFCD34F" pageId="21" pageNumber="52" type="neotype">neotype</typeStatus>
|
||
confirms previous descriptions by Arratia (1991, 1997), but adds some new information about variability. While in some specimens the first two uroneurals extend anteriorly to the lateral surface of preural centrum 3 (
|
||
<figureCitation id="13642A02BA342F12FA9EFAF3FAEDD3EE" box="[1309,1392,1405,1428]" captionStart="Figure 18" captionStartId="23.[124,191,1346,1368]" captionTargetBox="[146,1448,197,1298]" captionTargetId="figure-11@23.[132,1455,177,1315]" captionTargetPageId="23" captionText="Figure 18. Ascalabos voithii. Caudal skeleton in lateral view slightly modified from Arratia (1997). Abbreviations: Ant, anterior; dpr, dorsal or epaxial procurrent rays; d.scu, dorsal caudal scute; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–11, hypurals 1–11; hsPU2, haemal spine of preural centrum 2; mo, membranous outgrowth on anterodorsal margin of first uroneural; mo’, membranous outgrowth on anterodorsal margin of second uroneural; nsPU2–4, neural spine of preural centra 2–4; PR1,19, principal caudal rays 1, 19; rer, rudimentary fin ray; U1+2+H1–2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–7, uroneurals 1–7; v.scu., ventral caudal scute." figureDoi="http://doi.org/10.5281/zenodo.11045910" httpUri="https://zenodo.org/record/11045910/files/figure.png" pageId="21" pageNumber="52">Fig. 18</figureCitation>
|
||
), they reach preural centrum
|
||
<quantity id="4CA79B62BA342F12FB9CFA13FBDAD3CE" box="[1055,1095,1437,1460]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="21" pageNumber="52" unit="in" value="2.0">2 in</quantity>
|
||
the
|
||
<typeStatus id="54E48825BA342F12FBF1FA13FB5BD3CE" box="[1138,1222,1437,1460]" pageId="21" pageNumber="52" type="neotype">neotype</typeStatus>
|
||
(
|
||
<figureCitation id="13642A02BA342F12FB50FA13FABCD3CE" box="[1235,1313,1437,1460]" captionStart="Figure 19" captionStartId="24.[124,191,1198,1220]" captionTargetBox="[169,1407,185,1146]" captionTargetId="figure-11@24.[169,1416,177,1167]" captionTargetPageId="24" captionText="Figure 19. Ascalabos voithii. Caudal skeleton in lateral view with only two epaxial basal fulcra preserved (neotype, JME SOS 537). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; dpr, dorsal or epaxial procurrent rays; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–4, hypurals 1–4; hbfu, hypaxial basal fulcra; hsPU4, haemal spine of preural centrum 4; naPU1, neural arch of preural centrum 1; nsPU2–4, neural spine of preural centra 2–4; PH, parhypural; PU1,4, preural centrum 1, 4; rer, rudimentary fin ray; sc, dorsal and ventral caudal scutes; U1+2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–8, uroneurals 1–8; PR1–19, principal rays 1–19; vpr, ventral procurrent rays; v.scu, ventral caudal scute." figureDoi="http://doi.org/10.5281/zenodo.11045912" httpUri="https://zenodo.org/record/11045912/files/figure.png" pageId="21" pageNumber="52">Fig. 19</figureCitation>
|
||
). The
|
||
<typeStatus id="54E48825BA342F12FAE1FA13FA2BD3CE" box="[1378,1462,1437,1460]" pageId="21" pageNumber="52" type="neotype">neotype</typeStatus>
|
||
presents eight uroneurals, whereas in
|
||
<typeStatus id="54E48825BA342F12FB4CFA33FAE3D3AE" box="[1231,1406,1469,1492]" isOtherTypeStatus="true" pageId="21" pageNumber="52">other specimens</typeStatus>
|
||
only seven have been observed, but this could be a result of poor preservation. The fourth uroneural is broken in the
|
||
<typeStatus id="54E48825BA342F12FAD8FA73FA2FD06E" box="[1371,1458,1533,1556]" pageId="21" pageNumber="52" type="neotype">neotype</typeStatus>
|
||
, so that it is unclear whether it reached ural centrum 1 + 2 or not, as it does in
|
||
<typeStatus id="54E48825BA342F12FB91F9B3FB5ED02E" box="[1042,1219,1597,1620]" isOtherTypeStatus="true" pageId="21" pageNumber="52">other specimens</typeStatus>
|
||
. Uroneurals 5 to 8 reduce in size caudally. The neural spine of preural centrum 2 is slightly shorter than that of preural centrum 3, which is broken distally. The neural arch of preural centrum 1 is incompletely preserved.
|
||
</paragraph>
|
||
<paragraph id="8BE03687BA342F11FCCDF952FE17D5B6" blockId="21.[819,1462,736,1907]" lastBlockId="22.[124,767,885,1259]" lastPageId="22" lastPageNumber="53" pageId="21" pageNumber="52">
|
||
There is an incomplete series of epaxial basal fulcra, two segmented procurrent rays, one epaxial rudimentary ray, 19 principal caudal rays, three hypaxial procurrent rays, and five hypaxial basal fulcra in the
|
||
<typeStatus id="54E48825BA342F12FB12F8B2FB78D129" box="[1169,1253,1852,1875]" pageId="21" pageNumber="52" type="neotype">neotype</typeStatus>
|
||
(
|
||
<figureCitation id="13642A02BA342F12FB76F8B2FAD8D129" box="[1269,1349,1852,1875]" captionStart="Figure 19" captionStartId="24.[124,191,1198,1220]" captionTargetBox="[169,1407,185,1146]" captionTargetId="figure-11@24.[169,1416,177,1167]" captionTargetPageId="24" captionText="Figure 19. Ascalabos voithii. Caudal skeleton in lateral view with only two epaxial basal fulcra preserved (neotype, JME SOS 537). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; dpr, dorsal or epaxial procurrent rays; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–4, hypurals 1–4; hbfu, hypaxial basal fulcra; hsPU4, haemal spine of preural centrum 4; naPU1, neural arch of preural centrum 1; nsPU2–4, neural spine of preural centra 2–4; PH, parhypural; PU1,4, preural centrum 1, 4; rer, rudimentary fin ray; sc, dorsal and ventral caudal scutes; U1+2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–8, uroneurals 1–8; PR1–19, principal rays 1–19; vpr, ventral procurrent rays; v.scu, ventral caudal scute." figureDoi="http://doi.org/10.5281/zenodo.11045912" httpUri="https://zenodo.org/record/11045912/files/figure.png" pageId="21" pageNumber="52">Fig. 19</figureCitation>
|
||
). Fringing fulcra are absent. In contrast, other specimens have five or six epaxial basal fulcra and seven hypaxial basal fulcra (
|
||
<figureCitation id="13642A02BA372F11FD27FCFBFD6CD5F6" box="[676,753,885,908]" captionStart="Figure 18" captionStartId="23.[124,191,1346,1368]" captionTargetBox="[146,1448,197,1298]" captionTargetId="figure-11@23.[132,1455,177,1315]" captionTargetPageId="23" captionText="Figure 18. Ascalabos voithii. Caudal skeleton in lateral view slightly modified from Arratia (1997). Abbreviations: Ant, anterior; dpr, dorsal or epaxial procurrent rays; d.scu, dorsal caudal scute; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–11, hypurals 1–11; hsPU2, haemal spine of preural centrum 2; mo, membranous outgrowth on anterodorsal margin of first uroneural; mo’, membranous outgrowth on anterodorsal margin of second uroneural; nsPU2–4, neural spine of preural centra 2–4; PR1,19, principal caudal rays 1, 19; rer, rudimentary fin ray; U1+2+H1–2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; ‘UD’, “urodermals”; UN1–7, uroneurals 1–7; v.scu., ventral caudal scute." figureDoi="http://doi.org/10.5281/zenodo.11045910" httpUri="https://zenodo.org/record/11045910/files/figure.png" pageId="22" pageNumber="53">Fig. 18</figureCitation>
|
||
). Dorsal and ventral scutes precede the epaxial and hypaxial basal fulcra, respectively.
|
||
</paragraph>
|
||
<caption id="DF20660FBA372F11FFFFFD6CFDADD548" ID-DOI="http://doi.org/10.5281/zenodo.11045908" ID-Zenodo-Dep="11045908" httpUri="https://zenodo.org/record/11045908/files/figure.png" pageId="22" pageNumber="53" startId="22.[124,191,738,760]" targetBox="[369,1215,187,694]" targetPageId="22" targetType="figure">
|
||
<paragraph id="8BE03687BA372F11FFFFFD6CFDADD548" blockId="22.[124,1462,738,818]" pageId="22" pageNumber="53">
|
||
<emphasis id="B92BEA95BA372F11FFFFFD6CFF78D482" bold="true" box="[124,229,738,760]" pageId="22" pageNumber="53">Figure 17.</emphasis>
|
||
<taxonomicName id="4C5F4D04BA372F11FF6FFD6CFE0ED48D" box="[236,403,738,759]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="53" phylum="Chordata" rank="species" species="voithii">
|
||
<emphasis id="B92BEA95BA372F11FF6FFD6CFE0ED48D" box="[236,403,738,759]" italics="true" pageId="22" pageNumber="53">Ascalabos voithii</emphasis>
|
||
</taxonomicName>
|
||
(neotype, JME SOS 537). Midcaudal vertebrae illustrating the presence of a lateral crest and grooves on the lateral side of the vertebral centra. Arrows point to a lateral longitudinal crest. Abbreviations: na, neural arch; ns, neural spine; ha, haemal arch; hs, haemal spine; vc, vertebral centrum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE03687BA372F11FFFFFC5BFF55D291" blockId="22.[124,767,885,1259]" pageId="22" pageNumber="53">
|
||
Scales: The scales of
|
||
<taxonomicName id="4C5F4D04BA372F11FEDDFC5BFE56D597" box="[350,459,981,1005]" family="Ascalaboidae" genus="Ascalabos" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92BEA95BA372F11FEDDFC5BFE56D597" box="[350,459,981,1005]" italics="true" pageId="22" pageNumber="53">Ascalabos</emphasis>
|
||
</taxonomicName>
|
||
are incompletely known, because it is difficult to observe isolated scales. The strongly imbricate scales are usually preserved in situ. They are slightly rectangular, with incomplete circuli in the anterior field and with transverse lines that do not completely cross the middle field of the scale. In contrast,
|
||
<taxonomicName id="4C5F4D04BA372F11FDFAFBFAFD62D2F6" authority="Arratia, 2016" authorityName="Arratia" authorityYear="2016" box="[633,767,1140,1164]" family="Ascalaboidae" genus="Ebertichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="53" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95BA372F11FDFAFBFAFD62D2F6" box="[633,767,1140,1164]" italics="true" pageId="22" pageNumber="53">Ebertichthys</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEBA372F11FFFFFB1AFF58D2D1" box="[124,197,1172,1195]" pageId="22" pageNumber="53" rank="genus">n. gen.</taxonomicNameLabel>
|
||
has large, slightly oval scales (
|
||
<figureCitation id="13642A02BA372F11FD95FB1AFDF9D2D1" box="[534,612,1172,1195]" captionStart="Figure 6" captionStartId="10.[124,191,1734,1756]" captionTargetBox="[265,1313,193,1701]" captionTargetId="figure-11@10.[264,1322,190,1703]" captionTargetPageId="10" captionText="Figure 6. Ebertichthys ettlingensis n. gen. et n. sp. Details of certain abdominal vertebrae and pectoral fin. (a) Anterior abdominal region and pectoral girdle and fin (holotype JME ETT 108). (b) Enlargement of a few abdominal vertebrae illustrating details of the epineural processes (holotype JME ETT 108). (c) Details of pectoral fin and radials (JME ETT 132a). Abbreviations: cl, cleithrum; cor, coracoid; d.ra, distal radials; epin.p, epineural process; na, neural arch; ns, neural spine; paph, parapophyses; pcl1–3, postcleithrum 1–3; pec.f, pectoral fin; pec.ra, proximal radials supporting pectoral rays; pel.p, pelvic plate; ptt, posttemporal; ri, ribs; sc, anterior field of cycloid scales; sca, scapula; scl, supracleithrum; sn, supraneurals; vc, abdominal vertebral centra; 1st. pec, first pectoral ray." figureDoi="http://doi.org/10.5281/zenodo.11045876" httpUri="https://zenodo.org/record/11045876/files/figure.png" pageId="22" pageNumber="53">Fig. 6b</figureCitation>
|
||
) with few circuli in the anterior field. No transverse lines have been observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |