1533 lines
168 KiB
XML
1533 lines
168 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4323816" ID-GBIF-Dataset="7fbb4285-59e7-49e6-a890-df9610e94455" ID-GBIF-Taxon="164845016" ID-Zenodo-Dep="4323816" approvalRequired="7" approvalRequired_for_document="1" approvalRequired_for_textStreams="6" checkinTime="1593590792548" checkinUser="plazi" docAuthor="Ballarin, Francesco" docDate="2020" docId="03BCA0121E63FFC6FF2D624FC584FD33" docLanguage="en" docName="JCaveAndKarstStudies.82.2.82_94.pdf" docOrigin="Journal of Cave and Karst Studies 82 (2)" docSource="http://dx.doi.org/10.4311/2019lsc0103" docStyle="DocumentStyle{}" docTitle="Domitius culsu Ballarin 2020, sp. nov." docType="treatment" docVersion="5" lastPageId="11" lastPageNumber="93" masterDocId="FF85D86A1E62FFCDFFBD642DC75AFFD9" masterDocTitle="Domitius Culsu sp. nov. (Araneae, Nestici-dae), a new troglobiont spider from Italy with notes on Italian nesticids of the genus Domitius Ribera, 2018" masterLastPageNumber="94" masterPageNumber="82" pageId="1" pageNumber="83" updateTime="1635489950706" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Domitius Culsu sp. nov. (Araneae, Nestici-dae), a new troglobiont spider from Italy with notes on Italian nesticids of the genus Domitius Ribera, 2018</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Ballarin, Francesco</mods:namePart>
|
||
<mods:affiliation>Systematic Zoology Laboratory, Department of Biological Sciences, Tokyo Metropolitan University 1 - 1 Minami-Osawa, Hachioji-shi, Tokyo, 192 - 0397, Japan</mods:affiliation>
|
||
<mods:nameIdentifier type="email">ballarin.francesco@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Journal of Cave and Karst Studies</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2020</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2020-06-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>82</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>82</mods:start>
|
||
<mods:end>94</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.4311/2019lsc0103</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.4311/2019lsc0103</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">7fbb4285-59e7-49e6-a890-df9610e94455</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">3925078</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4323816" ID-GBIF-Taxon="164845016" ID-Zenodo-Dep="4323816" LSID="urn:lsid:plazi:treatment:03BCA0121E63FFC6FF2D624FC584FD33" httpUri="http://treatment.plazi.org/id/03BCA0121E63FFC6FF2D624FC584FD33" lastPageId="11" lastPageNumber="93" pageId="1" pageNumber="83">
|
||
<subSubSection box="[144,550,1634,1659]" pageId="1" pageNumber="83" type="nomenclature">
|
||
<paragraph blockId="1.[144,849,1462,1726]" box="[144,550,1634,1659]" pageId="1" pageNumber="83">
|
||
<emphasis bold="true" box="[144,550,1634,1659]" pageId="1" pageNumber="83">
|
||
<taxonomicName authority="Ballarin" authorityName="Ballarin" authorityYear="2020" box="[144,443,1634,1659]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="83" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis bold="true" box="[144,337,1634,1659]" italics="true" pageId="1" pageNumber="83">Domitius culsu</emphasis>
|
||
Ballarin
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[451,550,1634,1659]" pageId="1" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[144,484,1667,1693]" pageId="1" pageNumber="83" type="description">
|
||
<paragraph blockId="1.[144,849,1462,1726]" box="[144,484,1667,1693]" pageId="1" pageNumber="83">
|
||
<figureCitation box="[144,249,1667,1693]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="1" pageNumber="83">Figures 1</figureCitation>
|
||
A‒G, 2A‒E, 3A‒D.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[144,849,1701,1726]" pageId="1" pageNumber="83" type="reference_group">
|
||
<paragraph blockId="1.[144,849,1462,1726]" box="[144,849,1701,1726]" pageId="1" pageNumber="83">
|
||
<taxonomicName authority="Brignoli, 1979: 214" authorityName="Brignoli" authorityPageNumber="214" authorityYear="1979" box="[144,635,1701,1726]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="83" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[144,408,1701,1726]" italics="true" pageId="1" pageNumber="83">Nesticus speluncarum</emphasis>
|
||
<bibRefCitation author="Brignoli, P. M." box="[415,635,1701,1726]" journalOrPublisher="Quaderni del Museo di Speleologia " V. Rivera" pageId="1" pageNumber="83" pagination="1 - 48" part="10" refId="ref7203" refString="Brignoli, P. M., 1979, Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae):. Quaderni del Museo di Speleologia " V. Rivera ", v. 5, no. 10, p. 1 - 48." title="Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae)" type="journal article" year="1979">Brignoli, 1979: 214</bibRefCitation>
|
||
</taxonomicName>
|
||
(misidentification)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="2" lastPageNumber="84" pageId="1" pageNumber="83" type="materials_examined">
|
||
<paragraph blockId="1.[144,1537,1750,1937]" pageId="1" pageNumber="83">
|
||
<emphasis bold="true" box="[144,568,1750,1777]" pageId="1" pageNumber="83">
|
||
Type material.
|
||
<materialsCitation ID-GBIF-Occurrence="2802128304" collectingDate="2018-08-21" collectionCode="MSNV" collectorName="F. Ballarin & D. Avesani" country="Italy" elevation="260" latitude="44.046337" location="Lucca Province" longLatPrecision="1" longitude="10.523525" municipality="Garfagnana" pageId="1" pageNumber="83" specimenCount="1" stateProvince="Toscana" typeStatus="holotype">
|
||
<typeStatus box="[336,452,1751,1776]" pageId="1" pageNumber="83">Holotype</typeStatus>
|
||
<specimenCount box="[462,483,1750,1777]" pageId="1" pageNumber="83" type="male">♂</specimenCount>
|
||
.
|
||
<collectingCountry box="[497,568,1751,1776]" name="Italy" pageId="1" pageNumber="83">ITALY</collectingCountry>
|
||
,
|
||
<emphasis bold="true" box="[585,692,1751,1776]" italics="true" pageId="1" pageNumber="83">
|
||
<collectingRegion box="[585,692,1751,1776]" country="Italy" name="Toscana" pageId="1" pageNumber="83">Toscana</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingMunicipality box="[709,852,1751,1777]" pageId="1" pageNumber="83">Garfagnana</collectingMunicipality>
|
||
area,
|
||
<location LSID="urn:lsid:plazi:treatment:03BCA0121E63FFC6FF2D624FC584FD33:8ECA47DF1E63FFCCFC1E62FAC33BF929" box="[931,1121,1751,1776]" country="Italy" latitude="44.046337" longLatPrecision="1" longitude="10.523525" municipality="Garfagnana" name="Lucca Province" pageId="1" pageNumber="83" stateProvince="Toscana">Lucca Province</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:03BCA0121E63FFC6FF2D624FC584FD33:8ECA47DF1E63FFCCFBD062FAC2A4F929" box="[1133,1534,1751,1777]" country="Italy" latitude="44.046337" longLatPrecision="1" longitude="10.523525" municipality="Garfagnana" name="Coreglia Antelminelli Municipality" pageId="1" pageNumber="83" stateProvince="Toscana">Coreglia Antelminelli Municipality</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:03BCA0121E63FFC6FF2D624FC584FD33:8ECA47DF1E63FFCCFF2D62DAC47DF8C9" box="[144,807,1783,1809]" country="Italy" latitude="44.046337" longLatPrecision="1" longitude="10.523525" municipality="Garfagnana" name="Tana delle Fate di Coreglia Antelminelli cave, 141 / T / LU" pageId="1" pageNumber="83" stateProvince="Toscana">Tana delle Fate di Coreglia Antelminelli cave, 141/T/LU</location>
|
||
,
|
||
<quantity box="[819,887,1783,1808]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="83" unit="m" value="260.0">
|
||
<elevation box="[819,887,1783,1808]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="83" unit="m" value="260.0">260m</elevation>
|
||
</quantity>
|
||
a.s.l.,
|
||
<geoCoordinate box="[968,1126,1783,1808]" degrees="44.046336" direction="north" orientation="latitude" pageId="1" pageNumber="83" precision="1" value="44.046337">44.046336°N</geoCoordinate>
|
||
<geoCoordinate box="[1136,1294,1783,1808]" degrees="10.523525" direction="east" orientation="longitude" pageId="1" pageNumber="83" precision="1" value="10.523525">10.523525°E</geoCoordinate>
|
||
,
|
||
<date box="[1306,1450,1783,1808]" pageId="1" pageNumber="83" value="2018-08-21">
|
||
<collectingDate box="[1306,1450,1783,1808]" pageId="1" pageNumber="83" value="2018-08-21">21.VIII.2018</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName pageId="1" pageNumber="83">F. Ballarin</collectorName>
|
||
and
|
||
<collectorName box="[299,429,1815,1840]" pageId="1" pageNumber="83">D. Avesani</collectorName>
|
||
(
|
||
<collectionCode box="[446,533,1815,1841]" collectionName="Italy, Verona, Museo Civico di Storia Naturale" pageId="1" pageNumber="83">MSNV</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="2802128305" collectingDate="1967-09-04" collectionCode="MSNV" collectorName="A. Vigna Taglianti" country="Italy" elevation="260" latitude="44.046337" location="Lucca Province" longLatPrecision="1" longitude="10.523525" municipality="Garfagnana" pageId="1" pageNumber="83" specimenCount="1" specimenCount-female="1" stateProvince="Toscana" typeStatus="paratype">
|
||
<emphasis bold="true" box="[553,682,1815,1840]" pageId="1" pageNumber="83">
|
||
<typeStatus box="[553,682,1815,1840]" pageId="1" pageNumber="83">Paratypes</typeStatus>
|
||
</emphasis>
|
||
. Same locality as the holotype,
|
||
<specimenCount box="[1078,1113,1813,1840]" pageId="1" pageNumber="83" type="female">1♀</specimenCount>
|
||
,
|
||
<date box="[1127,1256,1815,1840]" pageId="1" pageNumber="83" value="1967-09-04">
|
||
<collectingDate box="[1127,1256,1815,1840]" pageId="1" pageNumber="83" value="1967-09-04">04.IX.1967</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[1322,1536,1815,1840]" pageId="1" pageNumber="83">A. Vigna Taglianti</collectorName>
|
||
(
|
||
<collectionCode box="[150,233,1847,1873]" collectionName="Italy, Verona, Museo Civico di Storia Naturale" pageId="1" pageNumber="83">MSNV</collectionCode>
|
||
) (
|
||
<bibRefCitation author="Brignoli, P. M." box="[258,424,1847,1872]" journalOrPublisher="Quaderni del Museo di Speleologia " V. Rivera" pageId="1" pageNumber="83" pagination="1 - 48" part="10" refId="ref7203" refString="Brignoli, P. M., 1979, Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae):. Quaderni del Museo di Speleologia " V. Rivera ", v. 5, no. 10, p. 1 - 48." title="Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae)" type="journal article" year="1979">Brignoli 1979</bibRefCitation>
|
||
, sub
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[492,760,1847,1872]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="83" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[492,760,1847,1872]" italics="true" pageId="1" pageNumber="83">Nesticus speluncarum</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2802128303" box="[787,1464,1845,1873]" collectingDate="2015-08-15" collectionCode="MSNV" collectorName="F. Ballarin & M. Gaiga" country="Italy" elevation="260" latitude="44.046337" location="Lucca Province" longLatPrecision="1" longitude="10.523525" municipality="Garfagnana" pageId="1" pageNumber="83" specimenCount="1" specimenCount-female="1" stateProvince="Toscana">
|
||
<specimenCount box="[787,821,1845,1872]" pageId="1" pageNumber="83" type="female">1♀</specimenCount>
|
||
,
|
||
<date box="[837,981,1847,1872]" pageId="1" pageNumber="83" value="2015-08-15">
|
||
<collectingDate box="[837,981,1847,1872]" pageId="1" pageNumber="83" value="2015-08-15">15.VIII.2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[1050,1172,1847,1872]" pageId="1" pageNumber="83">F. Ballarin</collectorName>
|
||
and
|
||
<collectorName box="[1239,1353,1847,1873]" pageId="1" pageNumber="83">M. Gaiga</collectorName>
|
||
(
|
||
<collectionCode box="[1370,1456,1847,1873]" collectionName="Italy, Verona, Museo Civico di Storia Naturale" pageId="1" pageNumber="83">MSNV</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2802128302" collectingDate="2017-06-24" collectingDateMax="2018-06-28" collectingDateMin="2017-06-24" collectionCode="MSNV" collectorName="F. Ballarin & R. Ballarin" pageId="1" pageNumber="83" specimenCount="9" specimenCount-adult="1" specimenCount-female="7" specimenCount-juvenile="1">
|
||
<specimenCount box="[1478,1534,1845,1872]" pageId="1" pageNumber="83" type="female">4♀♀</specimenCount>
|
||
,
|
||
<date box="[144,271,1879,1904]" pageId="1" pageNumber="83" value="2017-06-24">
|
||
<collectingDate box="[144,271,1879,1904]" pageId="1" pageNumber="83" value="2017-06-24">24.VI.2017</collectingDate>
|
||
</date>
|
||
, (
|
||
<specimenCount box="[288,343,1877,1904]" pageId="1" pageNumber="83" type="female">3♀♀</specimenCount>
|
||
collected as
|
||
<specimenCount box="[500,604,1879,1904]" pageId="1" pageNumber="83" type="juvenile">juveniles</specimenCount>
|
||
and raised in captivity,
|
||
<specimenCount box="[885,960,1879,1904]" pageId="1" pageNumber="83" type="adult">adults</specimenCount>
|
||
:
|
||
<date box="[971,1113,1879,1904]" pageId="1" pageNumber="83" value="2017-08-20">
|
||
<collectingDate box="[971,1113,1879,1904]" pageId="1" pageNumber="83" value="2017-08-20">20.VIII.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<date box="[1124,1251,1879,1904]" pageId="1" pageNumber="83" value="2017-09-15">
|
||
<collectingDate box="[1124,1251,1879,1904]" pageId="1" pageNumber="83" value="2017-09-15">15.IX.2017</collectingDate>
|
||
</date>
|
||
and
|
||
<date box="[1311,1438,1879,1904]" pageId="1" pageNumber="83" value="2018-06-28">
|
||
<collectingDate box="[1311,1438,1879,1904]" pageId="1" pageNumber="83" value="2018-06-28">28.VI.2018</collectingDate>
|
||
</date>
|
||
respec- tively), leg.
|
||
<collectorName box="[278,396,1911,1936]" pageId="1" pageNumber="83">F. Ballarin</collectorName>
|
||
and
|
||
<collectorName box="[455,580,1911,1936]" pageId="1" pageNumber="83">R. Ballarin</collectorName>
|
||
(
|
||
<collectionCode box="[593,679,1911,1937]" collectionName="Italy, Verona, Museo Civico di Storia Naturale" pageId="1" pageNumber="83">MSNV</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2802128301" box="[697,1470,1909,1937]" collectingDate="2018-08-21" collectionCode="MSNV, MSNB" collectorName="F. Ballarin & D. Avesani" pageId="1" pageNumber="83" specimenCount="4" specimenCount-female="4">
|
||
<specimenCount box="[697,753,1909,1936]" pageId="1" pageNumber="83" type="female">4♀♀</specimenCount>
|
||
,
|
||
<date box="[762,906,1911,1936]" pageId="1" pageNumber="83" value="2018-08-21">
|
||
<collectingDate box="[762,906,1911,1936]" pageId="1" pageNumber="83" value="2018-08-21">21.VIII.2018</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[967,1086,1911,1936]" pageId="1" pageNumber="83">F. Ballarin</collectorName>
|
||
and
|
||
<collectorName box="[1145,1272,1911,1936]" pageId="1" pageNumber="83">D. Avesani</collectorName>
|
||
(
|
||
<collectionCode box="[1286,1369,1911,1937]" collectionName="Italy, Verona, Museo Civico di Storia Naturale" pageId="1" pageNumber="83">MSNV</collectionCode>
|
||
,
|
||
<collectionCode box="[1379,1462,1911,1937]" collectionName="MSNB" pageId="1" pageNumber="83">MSNB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="DF6A418C1E60FFCFFFEE64B6C27FFF6B" box="[83,1317,155,178]" httpUri="http://table.plazi.org/id/DF6A418C1E60FFCFFFEE64B6C27FFF6B" pageId="2" pageNumber="84" targetBox="[83,1429,194,878]" targetIsTable="true" targetPageId="2">
|
||
<paragraph blockId="2.[83,1317,155,178]" box="[83,1317,155,178]" pageId="2" pageNumber="84">Table 1. Species, GenBank accession numbers, and locality of the specimens used in the phylogenetic analysis.</paragraph>
|
||
</caption>
|
||
<paragraph pageId="2" pageNumber="84">
|
||
<table box="[83,1429,194,878]" gridcols="6" gridrows="17" pageId="2" pageNumber="84">
|
||
<tr box="[83,1429,194,217]" gridrow="0" pageId="2" pageNumber="84">
|
||
<th box="[83,355,194,217]" gridcol="0" gridrow="0" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[173,266,194,217]" pageId="2" pageNumber="84">Species</emphasis>
|
||
</th>
|
||
<th box="[378,439,194,217]" gridcol="1" gridrow="0" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[378,439,194,217]" pageId="2" pageNumber="84">Code</emphasis>
|
||
</th>
|
||
<th box="[462,575,194,217]" gridcol="2" gridrow="0" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[497,540,194,217]" pageId="2" pageNumber="84">COI</emphasis>
|
||
</th>
|
||
<th box="[598,711,194,217]" gridcol="3" gridrow="0" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[633,675,194,217]" pageId="2" pageNumber="84">16S</emphasis>
|
||
</th>
|
||
<th box="[734,847,194,217]" gridcol="4" gridrow="0" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[775,806,194,217]" pageId="2" pageNumber="84">H3</emphasis>
|
||
</th>
|
||
<th box="[870,1429,194,217]" gridcol="5" gridrow="0" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[1114,1208,194,217]" pageId="2" pageNumber="84">Locality</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[83,1429,233,253]" gridrow="1" pageId="2" pageNumber="84">
|
||
<th box="[83,355,233,253]" gridcol="0" gridrow="1" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="López-Pancorbo & Ribera" baseAuthorityYear="2011" box="[83,254,233,253]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="baeticus">Domitius baeticus</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,233,253]" gridcol="1" gridrow="1" pageId="2" pageNumber="84">Dbae</td>
|
||
<td box="[462,575,233,253]" gridcol="2" gridrow="1" pageId="2" pageNumber="84">MF693114</td>
|
||
<td box="[598,711,233,253]" gridcol="3" gridrow="1" pageId="2" pageNumber="84">MF693118</td>
|
||
<td box="[734,847,233,253]" gridcol="4" gridrow="1" pageId="2" pageNumber="84">MF693106</td>
|
||
<td box="[870,1429,233,253]" gridcol="5" gridrow="1" pageId="2" pageNumber="84">Cueva del Castillo. Siles, Jaén. Spain</td>
|
||
</tr>
|
||
<tr box="[83,1429,269,313]" gridrow="2" pageId="2" pageNumber="84">
|
||
<th box="[83,355,269,313]" gridcol="0" gridrow="2" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Ribera & Guerao" baseAuthorityYear="1995" box="[83,231,270,290]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="luquei">
|
||
<emphasis box="[83,231,270,290]" italics="true" pageId="2" pageNumber="84">Domitius luquei</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,269,313]" gridcol="1" gridrow="2" pageId="2" pageNumber="84">Dluq</td>
|
||
<td box="[462,575,269,313]" gridcol="2" gridrow="2" pageId="2" pageNumber="84">MF693112</td>
|
||
<td box="[598,711,269,313]" gridcol="3" gridrow="2" pageId="2" pageNumber="84">EU746439</td>
|
||
<td box="[734,847,269,313]" gridcol="4" gridrow="2" pageId="2" pageNumber="84">MF693104</td>
|
||
<td box="[870,1429,269,313]" gridcol="5" gridrow="2" pageId="2" pageNumber="84">Cueva de la Picona, San Pedro de Carmona, Cabuérniga, Cantabria, Spain</td>
|
||
</tr>
|
||
<tr box="[83,1429,330,351]" gridrow="3" pageId="2" pageNumber="84">
|
||
<th box="[83,355,330,351]" gridcol="0" gridrow="3" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Fage" baseAuthorityYear="1931" box="[83,276,331,351]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="lusitanicus">
|
||
<emphasis box="[83,276,331,351]" italics="true" pageId="2" pageNumber="84">Domitius lusitanicus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,330,351]" gridcol="1" gridrow="3" pageId="2" pageNumber="84">Dlus</td>
|
||
<td box="[462,575,330,351]" gridcol="2" gridrow="3" pageId="2" pageNumber="84">MF693113</td>
|
||
<td box="[598,711,330,351]" gridcol="3" gridrow="3" pageId="2" pageNumber="84">EU746429</td>
|
||
<td box="[734,847,330,351]" gridcol="4" gridrow="3" pageId="2" pageNumber="84">MF693105</td>
|
||
<td box="[870,1429,330,351]" gridcol="5" gridrow="3" pageId="2" pageNumber="84">Algar de Marradinhas II, Concelho de Alcanena, Portugal</td>
|
||
</tr>
|
||
<tr box="[83,1429,365,387]" gridrow="4" pageId="2" pageNumber="84">
|
||
<th box="[83,355,365,387]" gridcol="0" gridrow="4" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="di Caporiacco" baseAuthorityYear="1934" box="[83,258,367,387]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="menozzii">
|
||
<emphasis box="[83,258,367,387]" italics="true" pageId="2" pageNumber="84">Domitius menozzii</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,365,387]" gridcol="1" gridrow="4" pageId="2" pageNumber="84">D213</td>
|
||
<td box="[462,575,365,387]" gridcol="2" gridrow="4" pageId="2" pageNumber="84">MK860151a</td>
|
||
<td box="[598,711,365,387]" gridcol="3" gridrow="4" pageId="2" pageNumber="84">MK860133a</td>
|
||
<td box="[734,847,365,387]" gridcol="4" gridrow="4" pageId="2" pageNumber="84">MK860142a</td>
|
||
<td box="[870,1429,365,387]" gridcol="5" gridrow="4" pageId="2" pageNumber="84">Tanna da Suja, Prati di Bavari, Liguria, Italy</td>
|
||
</tr>
|
||
<tr box="[83,1429,402,448]" gridrow="5" pageId="2" pageNumber="84">
|
||
<th box="[83,355,402,448]" gridcol="0" gridrow="5" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[83,326,404,424]" pageId="2" pageNumber="84">
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[83,239,404,424]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis bold="true" box="[83,239,404,424]" italics="true" pageId="2" pageNumber="84">Domitius culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[245,326,404,424]" pageId="2" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<td box="[378,439,402,448]" gridcol="1" gridrow="5" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[382,434,404,424]" pageId="2" pageNumber="84">D555</emphasis>
|
||
</td>
|
||
<td box="[462,575,402,448]" gridcol="2" gridrow="5" pageId="2" pageNumber="84">MK860152a</td>
|
||
<td box="[598,711,402,448]" gridcol="3" gridrow="5" pageId="2" pageNumber="84">MK860134 a</td>
|
||
<td box="[734,847,402,448]" gridcol="4" gridrow="5" pageId="2" pageNumber="84">MK860143a</td>
|
||
<td box="[870,1429,402,448]" gridcol="5" gridrow="5" pageId="2" pageNumber="84">Tana delle Fate di Coreglia Antelminelli, Coreglia Antelminelli, Toscana, Italy</td>
|
||
</tr>
|
||
<tr box="[83,1429,465,485]" gridrow="6" pageId="2" pageNumber="84">
|
||
<th box="[83,355,465,485]" gridcol="0" gridrow="6" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Brignoli" baseAuthorityYear="1979" box="[83,263,465,485]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis box="[83,263,465,485]" italics="true" pageId="2" pageNumber="84">Domitius sbordonii</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,465,485]" gridcol="1" gridrow="6" pageId="2" pageNumber="84">Dsbo</td>
|
||
<td box="[462,575,465,485]" gridcol="2" gridrow="6" pageId="2" pageNumber="84">MF693110</td>
|
||
<td box="[598,711,465,485]" gridcol="3" gridrow="6" pageId="2" pageNumber="84">MF693116</td>
|
||
<td box="[734,847,465,485]" gridcol="4" gridrow="6" pageId="2" pageNumber="84">MF693102</td>
|
||
<td box="[870,1429,465,485]" gridcol="5" gridrow="6" pageId="2" pageNumber="84">Tana degli orchetti, Supino, Lazio, Italy</td>
|
||
</tr>
|
||
<tr box="[83,1429,500,522]" gridrow="7" pageId="2" pageNumber="84">
|
||
<th box="[83,355,500,522]" gridcol="0" gridrow="7" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Pavesi" baseAuthorityYear="1873" box="[83,299,502,522]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[83,299,502,522]" italics="true" pageId="2" pageNumber="84">Domitius speluncarum</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,500,522]" gridcol="1" gridrow="7" pageId="2" pageNumber="84">D557</td>
|
||
<td box="[462,575,500,522]" gridcol="2" gridrow="7" pageId="2" pageNumber="84">MK860153a</td>
|
||
<td box="[598,711,500,522]" gridcol="3" gridrow="7" pageId="2" pageNumber="84">MK860135a</td>
|
||
<td box="[734,847,500,522]" gridcol="4" gridrow="7" pageId="2" pageNumber="84">MK860144a</td>
|
||
<td box="[870,1429,500,522]" gridcol="5" gridrow="7" pageId="2" pageNumber="84">Tana di Magnano, Canigiano, Lucca, Toscana, Italy</td>
|
||
</tr>
|
||
<tr box="[83,1429,537,559]" gridrow="8" pageId="2" pageNumber="84">
|
||
<th box="[83,355,537,559]" gridcol="0" gridrow="8" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[83,269,539,559]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="cellulanus">
|
||
<emphasis box="[83,269,539,559]" italics="true" pageId="2" pageNumber="84">Nesticus cellulanus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,537,559]" gridcol="1" gridrow="8" pageId="2" pageNumber="84">N214</td>
|
||
<td box="[462,575,537,559]" gridcol="2" gridrow="8" pageId="2" pageNumber="84">MK860154a</td>
|
||
<td box="[598,711,537,559]" gridcol="3" gridrow="8" pageId="2" pageNumber="84">MK860136a</td>
|
||
<td box="[734,847,537,559]" gridcol="4" gridrow="8" pageId="2" pageNumber="84">MK860145a</td>
|
||
<td box="[870,1429,537,559]" gridcol="5" gridrow="8" pageId="2" pageNumber="84">Cave of Koufovouno, Didimoticho, Thrace, Greece</td>
|
||
</tr>
|
||
<tr box="[83,1429,574,596]" gridrow="9" pageId="2" pageNumber="84">
|
||
<th box="[83,355,574,596]" gridcol="0" gridrow="9" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="López-Pancorbo, Kunt & Ribera" baseAuthorityYear="2013" box="[83,318,576,596]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="dimensis">
|
||
<emphasis box="[83,318,576,596]" italics="true" pageId="2" pageNumber="84">Kryptonesticus dimensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,574,596]" gridcol="1" gridrow="9" pageId="2" pageNumber="84">K566</td>
|
||
<td box="[462,575,574,596]" gridcol="2" gridrow="9" pageId="2" pageNumber="84">MK860155a</td>
|
||
<td box="[598,711,574,596]" gridcol="3" gridrow="9" pageId="2" pageNumber="84">MK860137a</td>
|
||
<td box="[734,847,574,596]" gridcol="4" gridrow="9" pageId="2" pageNumber="84">MK860146a</td>
|
||
<td box="[870,1429,574,596]" gridcol="5" gridrow="9" pageId="2" pageNumber="84">Dim cave, Antalya, Turkey</td>
|
||
</tr>
|
||
<tr box="[83,1429,611,633]" gridrow="10" pageId="2" pageNumber="84">
|
||
<th box="[83,355,611,633]" gridcol="0" gridrow="10" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1880" box="[83,305,613,633]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="eremita">
|
||
<emphasis box="[83,305,613,633]" italics="true" pageId="2" pageNumber="84">Kryptonesticus eremita</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,611,633]" gridcol="1" gridrow="10" pageId="2" pageNumber="84">K211</td>
|
||
<td box="[462,575,611,633]" gridcol="2" gridrow="10" pageId="2" pageNumber="84">MK860156a</td>
|
||
<td box="[598,711,611,633]" gridcol="3" gridrow="10" pageId="2" pageNumber="84">MK860138a</td>
|
||
<td box="[734,847,611,633]" gridcol="4" gridrow="10" pageId="2" pageNumber="84">MK860147a</td>
|
||
<td box="[870,1429,611,633]" gridcol="5" gridrow="10" pageId="2" pageNumber="84">Grotta di Ponte Subiolo, Mori, Veneto, Italy</td>
|
||
</tr>
|
||
<tr box="[83,1429,647,669]" gridrow="11" pageId="2" pageNumber="84">
|
||
<th box="[83,355,647,669]" gridcol="0" gridrow="11" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Kulczyński" baseAuthorityYear="1894" box="[83,355,649,669]" class="Arachnida" family="Nesticidae" genus="Carpathonesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="fodinarum">
|
||
<emphasis box="[83,355,649,669]" italics="true" pageId="2" pageNumber="84">Carpathonesticus fodinarum</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,647,669]" gridcol="1" gridrow="11" pageId="2" pageNumber="84">C162</td>
|
||
<td box="[462,575,647,669]" gridcol="2" gridrow="11" pageId="2" pageNumber="84">MK860157a</td>
|
||
<td box="[598,711,647,669]" gridcol="3" gridrow="11" pageId="2" pageNumber="84">MK860139a</td>
|
||
<td box="[734,847,647,669]" gridcol="4" gridrow="11" pageId="2" pageNumber="84">MK860148a</td>
|
||
<td box="[870,1429,647,669]" gridcol="5" gridrow="11" pageId="2" pageNumber="84">Small cave along the river, Sighistel, Bihor, Romania</td>
|
||
</tr>
|
||
<tr box="[83,1429,684,730]" gridrow="12" pageId="2" pageNumber="84">
|
||
<th box="[83,355,684,730]" gridcol="0" gridrow="12" pageId="2" pageNumber="84">
|
||
<taxonomicName box="[83,345,686,706]" class="Arachnida" family="Nesticidae" genus="Carpathonesitcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="lotriensis">
|
||
<emphasis box="[83,345,686,706]" italics="true" pageId="2" pageNumber="84">Carpathonesitcus lotriensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,684,730]" gridcol="1" gridrow="12" pageId="2" pageNumber="84">C166</td>
|
||
<td box="[462,575,684,730]" gridcol="2" gridrow="12" pageId="2" pageNumber="84">MK860158a</td>
|
||
<td box="[598,711,684,730]" gridcol="3" gridrow="12" pageId="2" pageNumber="84">MK860140a</td>
|
||
<td box="[734,847,684,730]" gridcol="4" gridrow="12" pageId="2" pageNumber="84">MK860149a</td>
|
||
<td box="[870,1429,684,730]" gridcol="5" gridrow="12" pageId="2" pageNumber="84">Humid and shadowed cliff near Lazaret village, Sibiu, Romania</td>
|
||
</tr>
|
||
<tr box="[83,1429,747,767]" gridrow="13" pageId="2" pageNumber="84">
|
||
<th box="[83,355,747,767]" gridcol="0" gridrow="13" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1907" box="[83,343,747,767]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="obcaecatus">
|
||
<emphasis box="[83,343,747,767]" italics="true" pageId="2" pageNumber="84">Typhlonesticus obcaecatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,747,767]" gridcol="1" gridrow="13" pageId="2" pageNumber="84">Tobc</td>
|
||
<td box="[462,575,747,767]" gridcol="2" gridrow="13" pageId="2" pageNumber="84">KF939309</td>
|
||
<td box="[598,711,747,767]" gridcol="3" gridrow="13" pageId="2" pageNumber="84">EU746437</td>
|
||
<td box="[734,847,747,767]" gridcol="4" gridrow="13" pageId="2" pageNumber="84">MF693109</td>
|
||
<td box="[870,1429,747,767]" gridcol="5" gridrow="13" pageId="2" pageNumber="84">Cueva del Molino de Aso, Boltana, Huesca, Spain</td>
|
||
</tr>
|
||
<tr box="[83,1429,784,804]" gridrow="14" pageId="2" pageNumber="84">
|
||
<th box="[83,355,784,804]" gridcol="0" gridrow="14" pageId="2" pageNumber="84">
|
||
<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1931" box="[83,308,784,804]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="idriacus">
|
||
<emphasis box="[83,308,784,804]" italics="true" pageId="2" pageNumber="84">Typhlonesticus idriacus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,784,804]" gridcol="1" gridrow="14" pageId="2" pageNumber="84">T167</td>
|
||
<td box="[462,575,784,804]" gridcol="2" gridrow="14" pageId="2" pageNumber="84">MG201050</td>
|
||
<td box="[598,711,784,804]" gridcol="3" gridrow="14" pageId="2" pageNumber="84">MG200521</td>
|
||
<td box="[734,847,784,804]" gridcol="4" gridrow="14" pageId="2" pageNumber="84">MG201227</td>
|
||
<td box="[870,1429,784,804]" gridcol="5" gridrow="14" pageId="2" pageNumber="84">Grotta Pre Oreak, Nimis, Udine, Friuli Venezia Giulia, Italy</td>
|
||
</tr>
|
||
<tr box="[83,1429,821,841]" gridrow="15" pageId="2" pageNumber="84">
|
||
<th box="[83,355,821,841]" gridcol="0" gridrow="15" pageId="2" pageNumber="84">
|
||
<taxonomicName authorityName="Pantini and Isaia" authorityYear="2019" baseAuthorityName="Brignoli" baseAuthorityYear="1975" box="[83,296,821,841]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="morisii">
|
||
<emphasis box="[83,296,821,841]" italics="true" pageId="2" pageNumber="84">Typhlonesticus morisii</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,821,841]" gridcol="1" gridrow="15" pageId="2" pageNumber="84">Tmor</td>
|
||
<td box="[462,575,821,841]" gridcol="2" gridrow="15" pageId="2" pageNumber="84">KF939311</td>
|
||
<td box="[598,711,821,841]" gridcol="3" gridrow="15" pageId="2" pageNumber="84">KF939308</td>
|
||
<td box="[734,847,821,841]" gridcol="4" gridrow="15" pageId="2" pageNumber="84">∙∙∙</td>
|
||
<td box="[870,1429,821,841]" gridcol="5" gridrow="15" pageId="2" pageNumber="84">Sotterranei del Forte di Vernante, Vernante, Cuneo, Italy</td>
|
||
</tr>
|
||
<tr box="[83,1429,856,878]" gridrow="16" pageId="2" pageNumber="84">
|
||
<th box="[83,355,856,878]" gridcol="0" gridrow="16" pageId="2" pageNumber="84">
|
||
<taxonomicName authorityName="Keyserling" authorityYear="1884" box="[83,303,857,878]" class="Arachnida" family="Nesticidae" genus="Gaucelmus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="augustinus">
|
||
<emphasis box="[83,303,857,878]" italics="true" pageId="2" pageNumber="84">Gaucelmus augustinus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[378,439,856,878]" gridcol="1" gridrow="16" pageId="2" pageNumber="84">G601</td>
|
||
<td box="[462,575,856,878]" gridcol="2" gridrow="16" pageId="2" pageNumber="84">MK860159a</td>
|
||
<td box="[598,711,856,878]" gridcol="3" gridrow="16" pageId="2" pageNumber="84">MK860141a</td>
|
||
<td box="[734,847,856,878]" gridcol="4" gridrow="16" pageId="2" pageNumber="84">MK860150a</td>
|
||
<td box="[870,1429,856,878]" gridcol="5" gridrow="16" pageId="2" pageNumber="84">Climax cave, Bainbridge, Georgia, USA</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="2.[83,219,894,909]" box="[83,219,894,909]" pageId="2" pageNumber="84">
|
||
<tableNote box="[83,219,894,909]" pageId="2" pageNumber="84" targetBox="[83,1429,194,878]" targetPageId="2">
|
||
<superScript attach="left" box="[83,88,894,902]" fontSize="3" pageId="2" pageNumber="84">a</superScript>
|
||
New Sequences.
|
||
</tableNote>
|
||
</paragraph>
|
||
<paragraph blockId="2.[72,1465,959,1243]" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[72,1174,961,986]" pageId="2" pageNumber="84">
|
||
Examined comparative material.
|
||
<taxonomicName authority="(Pavesi, 1873)" baseAuthorityName="Pavesi" baseAuthorityYear="1873" box="[498,980,961,986]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis bold="true" box="[498,790,961,986]" italics="true" pageId="2" pageNumber="84">Domitius speluncarum</emphasis>
|
||
(Pavesi, 1873)
|
||
</taxonomicName>
|
||
: ITALY:
|
||
<emphasis bold="true" box="[1084,1174,961,986]" italics="true" pageId="2" pageNumber="84">Liguria</emphasis>
|
||
</emphasis>
|
||
: 1♂, 1♀, (topotypes), La Spezia Province, Grotta Bocca Lupara cave, 74/Li/SP, 120m a.s.l., 05.III.1969, leg. P.M. Brignoli (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." pageId="2" pageNumber="84" pagination="121 - 229" refId="ref7176" refString="Brignoli, P. M., 1971, Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica, p. 121 - 229." title="Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica" type="book chapter" year="1971">Brignoli, 1971</bibRefCitation>
|
||
);
|
||
<emphasis bold="true" box="[153,260,1025,1050]" italics="true" pageId="2" pageNumber="84">Toscana</emphasis>
|
||
: 1♀, Lucca Province, Garfagnana area, Villa Collemandina Municipality, Canigiano village, Tana di Magnano cave, 162/T/LU, 653m a.s.l., 44.177285°N, 10.38803°E, 03.XI.1967, leg. A. Vigna Taglianti (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." pageId="2" pageNumber="84" pagination="121 - 229" refId="ref7176" refString="Brignoli, P. M., 1971, Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica, p. 121 - 229." title="Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica" type="book chapter" year="1971">Brignoli, 1971</bibRefCitation>
|
||
), 1♀, 01.VIII.1975, leg. P. Magrini (MSNV), 4♀♀, 14.VIII.2015, leg. F. Ballarin and M. Gaiga (MSNV); 1♀, Forno- volasco (not reported in the label but very likely from Grotta del Vento cave), 700m a.s.l., 16.VI.1970, leg. O. Osella (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." box="[186,357,1153,1178]" pageId="2" pageNumber="84" pagination="121 - 229" refId="ref7176" refString="Brignoli, P. M., 1971, Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica, p. 121 - 229." title="Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica" type="book chapter" year="1971">Brignoli, 1971</bibRefCitation>
|
||
); 1♀, Grotta del Buggine cave, 166/T/Lu, 315m a.s.l., 07.X.1967, leg. G. Castellini (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." box="[79,248,1185,1210]" pageId="2" pageNumber="84" pagination="121 - 229" refId="ref7176" refString="Brignoli, P. M., 1971, Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica, p. 121 - 229." title="Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica" type="book chapter" year="1971">Brignoli, 1971</bibRefCitation>
|
||
); 1♀, Massa-Carrara, Buca della Freddana cave, 230/T/Ms, 550m a.s.l., 05.VI.1977, leg. C. Bonzano (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." box="[182,354,1217,1242]" journalOrPublisher="Catalogo dei ragni cavernicoli italiani ": Memorie del Museo Civico di Storia Naturale di Verona" pageId="2" pageNumber="84" pagination="51 - 64" part="4" refId="ref7251" refString="Brignoli, P. M., 1985, Aggiunte e correzioni al " Catalogo dei ragni cavernicoli italiani ": Memorie del Museo Civico di Storia Naturale di Verona, serie 2, v. 4, p. 51 - 64." title="Aggiunte e correzioni al" type="journal article" year="1985">Brignoli, 1985</bibRefCitation>
|
||
); 1♀, Buca del Bacile cave, 226/T/Ms, 10.III.1975, leg. unknown (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." box="[1219,1391,1217,1242]" journalOrPublisher="Catalogo dei ragni cavernicoli italiani ": Memorie del Museo Civico di Storia Naturale di Verona" pageId="2" pageNumber="84" pagination="51 - 64" part="4" refId="ref7251" refString="Brignoli, P. M., 1985, Aggiunte e correzioni al " Catalogo dei ragni cavernicoli italiani ": Memorie del Museo Civico di Storia Naturale di Verona, serie 2, v. 4, p. 51 - 64." title="Aggiunte e correzioni al" type="journal article" year="1985">Brignoli, 1985</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="2.[72,1464,1265,1389]" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[72,794,1267,1293]" pageId="2" pageNumber="84">
|
||
<taxonomicName authority="(di Caporiacco, 1934)" baseAuthorityName="di Caporiacco" baseAuthorityYear="1934" box="[72,601,1267,1293]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="menozzii">
|
||
<emphasis bold="true" box="[72,311,1267,1292]" italics="true" pageId="2" pageNumber="84">Domitius menozzii</emphasis>
|
||
(di Caporiacco, 1934)
|
||
</taxonomicName>
|
||
: ITALY,
|
||
<emphasis bold="true" box="[704,794,1267,1292]" italics="true" pageId="2" pageNumber="84">Liguria</emphasis>
|
||
</emphasis>
|
||
, 1♂, 4♀♀, north-east of Genova town, Tanna da Vulpe cave, 264/Li/GE, 23.XI.1969, leg. A. Vigna Taglianti (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." box="[791,957,1299,1324]" pageId="2" pageNumber="84" pagination="121 - 229" refId="ref7176" refString="Brignoli, P. M., 1971, Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica, p. 121 - 229." title="Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica" type="book chapter" year="1971">Brignoli, 1971</bibRefCitation>
|
||
); 2♂♂, 2♀♀, Creto, Tanna de Fate cave, 17/ Li/GE, 30.X.1971, leg. G. Gardini (MSNV); 2♀♀ (topotypes), Prati di Bavari locality, Tanna da Suja cave, 5/Li/GE, 582m a.s.l., 44.422894°N, 09.035239°E, 30.V.2013, leg. F. Ballarin, A. Trotta, G. Gardini, and S. Zoia.
|
||
</paragraph>
|
||
<paragraph blockId="2.[72,1464,1411,1503]" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[72,695,1413,1438]" pageId="2" pageNumber="84">
|
||
<taxonomicName authority="(Brignoli, 1979)" baseAuthorityName="Brignoli" baseAuthorityYear="1979" box="[72,526,1413,1438]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis bold="true" box="[72,319,1413,1438]" italics="true" pageId="2" pageNumber="84">Domitius sbordonii</emphasis>
|
||
(
|
||
<bibRefCitation author="Brignoli, P. M." box="[335,518,1413,1438]" journalOrPublisher="Quaderni del Museo di Speleologia " V. Rivera" pageId="2" pageNumber="84" pagination="1 - 48" part="10" refId="ref7203" refString="Brignoli, P. M., 1979, Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae):. Quaderni del Museo di Speleologia " V. Rivera ", v. 5, no. 10, p. 1 - 48." title="Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae)" type="journal article" year="1979">Brignoli, 1979</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
: ITALY,
|
||
<emphasis bold="true" box="[627,695,1413,1438]" italics="true" pageId="2" pageNumber="84">Lazio</emphasis>
|
||
</emphasis>
|
||
, 1♂ (holotype), Frosinone Province, Supino, Valle Serena, Grot- ta della Croce cave, 01.IX.1977 leg. V. Sbordoni (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." box="[770,942,1445,1470]" journalOrPublisher="Quaderni del Museo di Speleologia " V. Rivera" pageId="2" pageNumber="84" pagination="1 - 48" part="10" refId="ref7203" refString="Brignoli, P. M., 1979, Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae):. Quaderni del Museo di Speleologia " V. Rivera ", v. 5, no. 10, p. 1 - 48." title="Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae)" type="journal article" year="1979">Brignoli, 1979</bibRefCitation>
|
||
); 1♀ (paratype), 08.II.1976, leg. V. Sbordoni (MSNV) (
|
||
<bibRefCitation author="Brignoli, P. M." box="[182,352,1477,1502]" journalOrPublisher="Quaderni del Museo di Speleologia " V. Rivera" pageId="2" pageNumber="84" pagination="1 - 48" part="10" refId="ref7203" refString="Brignoli, P. M., 1979, Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae):. Quaderni del Museo di Speleologia " V. Rivera ", v. 5, no. 10, p. 1 - 48." title="Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae)" type="journal article" year="1979">Brignoli, 1979</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="2.[72,1465,1525,1887]" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[72,761,1527,1553]" pageId="2" pageNumber="84">
|
||
<taxonomicName authority="(Simon, 1880)" baseAuthorityName="Simon" baseAuthorityYear="1880" box="[72,562,1527,1553]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="eremita">
|
||
<emphasis bold="true" box="[72,374,1527,1552]" italics="true" pageId="2" pageNumber="84">Kryptonesticus eremita</emphasis>
|
||
(Simon, 1880)
|
||
</taxonomicName>
|
||
: ITALY,
|
||
<emphasis bold="true" box="[662,752,1527,1552]" italics="true" pageId="2" pageNumber="84">Liguria</emphasis>
|
||
:
|
||
</emphasis>
|
||
2♂♂ (sub.
|
||
<taxonomicName authorityName="Di Caporiacco" authorityYear="1934" box="[897,1114,1527,1552]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="menozzii">
|
||
<emphasis box="[897,1114,1527,1552]" italics="true" pageId="2" pageNumber="84">Nesticus menozzii</emphasis>
|
||
</taxonomicName>
|
||
), Creto, Tanna de Fate cave, 17/Li/GE, 30.X.1971, leg. G. Gardini (MSNV);
|
||
<emphasis bold="true" box="[617,838,1559,1584]" italics="true" pageId="2" pageNumber="84">Emilia-Romagna:</emphasis>
|
||
2♀♀ Ravenna Province, Riolo Terme, Borgo Rivola, Grotta del Re Tiberio cave, 36/Er/RE. 19.II.1951, Leg. Denis (MSNV) (
|
||
<bibRefCitation author="Zangheri, P." box="[920,1110,1591,1616]" journalOrPublisher="Verona" pageId="2" pageNumber="84" pagination="854" refId="ref8104" refString="Zangheri, P., 1966, Repertorio sistematico e topografico della flora e fauna vivente e fossile della Romagna, Tomo II - Regno Animale. Museo Civico di Storia Naturale di Verona: Memorie Fuori Serie n ° 1, Stamperia Valdonega, Verona, p. 854." title="Repertorio sistematico e topografico della flora e fauna vivente e fossile della Romagna, Tomo II - Regno Animale. Museo Civico di Storia Naturale di Verona: Memorie Fuori Serie n ° 1, Stamperia Valdonega" type="book chapter" year="1966">Zangheri, 1966</bibRefCitation>
|
||
, sub.
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[1183,1448,1591,1616]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[1183,1448,1591,1616]" italics="true" pageId="2" pageNumber="84">Nesticus speluncarum</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<emphasis bold="true" box="[72,188,1623,1648]" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[72,179,1623,1648]" italics="true" pageId="2" pageNumber="84">Toscana</emphasis>
|
||
:
|
||
</emphasis>
|
||
1♀ Garfagnana area, Lucca Province, Coreglia Antelminelli Municipality, Tana delle Fate di Coreglia Antel- minelli cave, 141/T/LU, 260m a.s.l., 44.046336°N, 10.523525°E, 15.VIII.2015, leg. F. Ballarin and M. Gaiga (MSNV); 3♀♀, 24.VI.2017, Leg. F. Ballarin and R. Ballarin (MSNV-MSNVRAr/m 0007); 1♀, 21.VIII.2018, leg. F. Ballarin and D. Avesani (MSNV);
|
||
<emphasis bold="true" box="[283,420,1719,1745]" italics="true" pageId="2" pageNumber="84">Campania:</emphasis>
|
||
1♀, Avellino Province, Bagnoli Irpino, Mt. Piacentini, Grotta Giovannino cave, 16.VI.1956, leg. S.Ruffo (MSNV) (
|
||
<bibRefCitation author="Kritscher, E." box="[330,516,1751,1776]" pageId="2" pageNumber="84" pagination="313 - 320" refId="ref7500" refString="Kritscher, E., 1958, Araneen aus den Picentinischen Bergen: Memorie del Museo Civico di Storia Naturale di Verona, v. 6, p. 313 - 320." title="Araneen aus den Picentinischen Bergen: Memorie del Museo Civico di Storia Naturale di Verona" type="book chapter" year="1958">Kritscher, 1958</bibRefCitation>
|
||
, sub.
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[584,847,1751,1776]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[584,847,1751,1776]" italics="true" pageId="2" pageNumber="84">Nesticus speluncarum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="84" type="etymology">
|
||
<paragraph blockId="2.[72,1465,1525,1887]" box="[72,211,1795,1820]" pageId="2" pageNumber="84">
|
||
<emphasis bold="true" box="[72,211,1795,1820]" pageId="2" pageNumber="84">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="2.[72,1465,1525,1887]" pageId="2" pageNumber="84">The name of the new species is derived from the Etruscan goddess Culsu who, according to the Etruscan mytholo- gy, ruled the cave-like entrance of the underworld. Noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="88" pageId="3" pageNumber="85" type="diagnosis">
|
||
<paragraph blockId="3.[1230,1537,146,1617]" box="[1230,1359,146,171]" pageId="3" pageNumber="85">Diagnosis</paragraph>
|
||
<paragraph blockId="3.[1230,1537,146,1617]" pageId="3" pageNumber="85">Species closely re- lated to D. speluncarum and D. menozzii. Males of D. culsu sp. nov. can be separated from males of all other Italian species of the genus Domitius by the different shape of the apophyses of the para- cymbium (Figs. 1A‒C, 2B,C vs. Fig. 4A‒I). D. culsu sp. nov shows a robust, S-shaped dorsal apophysis 1 in contrast with a large, flat and axe- like Da 1 in D. speluncar- um (Figs. 1A‒C, 2B, C vs. Fig. 4A‒C); a short and stumpy Da 1 in D. me- nozzii (Figs. 1A‒C, 2B, C vs. Fig. 4D‒F) or a long and thread-like Da 1 in D. sbordonii (Figs. 1A‒C, 2B, C vs. Fig. 4G‒I). Addi- tionally, males of D. culsu sp. nov. have a well-de- veloped, triangular medi- an apophysis, absent in males of the other three species (Figs. 1A, 2A vs. Fig. 4A, D, G).</paragraph>
|
||
<paragraph blockId="3.[1230,1537,146,1617]" lastBlockId="4.[72,1465,1120,1694]" lastPageId="4" lastPageNumber="86" pageId="3" pageNumber="85">
|
||
Female D. culsu sp. nov. can be easily distin- guished from female D. speluncarum by the nar- rower, trapezoid-shaped median septum with slant- ing edges, in contrast with the larger, lobate Ms with rounded edges in D. spe- luncarum (Figs. 1E, 2D vs. Fig. 5A). Female D. culsu sp. nov. are separated from female D. morisii by the absence of a bulge on the Ms (clearly visible in
|
||
<emphasis box="[444,558,1624,1649]" italics="true" pageId="3" pageNumber="85">D. morisii</emphasis>
|
||
, Figs. 1E, 2D vs. Fig. 5C). Additionally, they can be distinguished by the different shape of copulatory ducts when the vulva is observed dorsally: with a rather uniform diameter in
|
||
<emphasis box="[1288,1384,1656,1681]" italics="true" pageId="3" pageNumber="85">D. culsu</emphasis>
|
||
sp. nov. and bearing a large, flattened middle trait in
|
||
<emphasis box="[615,727,1688,1713]" italics="true" pageId="3" pageNumber="85">D. morisii</emphasis>
|
||
(Figs. 1G, 2E vs. Fig. 5D). Female
|
||
<emphasis box="[1144,1240,1688,1713]" italics="true" pageId="3" pageNumber="85">D. culsu</emphasis>
|
||
sp. nov. are easily sepa- rated from those of
|
||
<emphasis box="[376,520,1720,1745]" italics="true" pageId="3" pageNumber="85">D. sbordonii</emphasis>
|
||
by the trapezoid-shaped Ms with slanting edges, in contrast with the squared Ms with vertical edges in
|
||
<emphasis box="[349,493,1752,1777]" italics="true" pageId="3" pageNumber="85">D. sbordonii</emphasis>
|
||
(Figs. 1E, 2D vs. Fig. 5E). They can further be distinguished by the different position of spermathecae, located in the lower half of the vulva and below the vulval pockets in
|
||
<emphasis box="[1145,1241,1784,1809]" italics="true" pageId="3" pageNumber="85">D. culsu</emphasis>
|
||
sp. nov., in contrast with S located in the upper half of vulva and above Vp in
|
||
<emphasis box="[761,905,1816,1841]" italics="true" pageId="3" pageNumber="85">D. sbordonii</emphasis>
|
||
(Figs. 1G, 2E vs. Fig. 5F).
|
||
<emphasis bold="true" box="[72,441,1121,1146]" pageId="4" pageNumber="86">Description. Male (holotype).</emphasis>
|
||
Total length 4.19. Carapace: 1.81 long, 1.56 wide.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925080" ID-Zenodo-Dep="3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="3" pageNumber="85" startId="3.[144,209,1504,1528]" targetBox="[144,1198,144,1493]" targetPageId="3">
|
||
<paragraph blockId="3.[144,1199,1504,1579]" box="[144,1199,1504,1528]" pageId="3" pageNumber="85">
|
||
Figure 1.
|
||
<emphasis box="[238,372,1504,1527]" italics="true" pageId="3" pageNumber="85">Nesticus culs</emphasis>
|
||
u sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female,
|
||
</paragraph>
|
||
<paragraph blockId="3.[144,1199,1504,1579]" box="[144,1199,1529,1553]" pageId="3" pageNumber="85">cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view,</paragraph>
|
||
<paragraph blockId="3.[144,1199,1504,1579]" box="[144,1180,1555,1579]" pageId="3" pageNumber="85">the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view.</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925082" ID-Zenodo-Dep="3925082" httpUri="https://zenodo.org/record/3925082/files/figure.png" pageId="4" pageNumber="86" startId="4.[69,134,1045,1069]" targetBox="[80,1447,144,1027]" targetPageId="4">
|
||
<paragraph blockId="4.[69,1464,1045,1095]" pageId="4" pageNumber="86">
|
||
Figure 2. Genitalia of
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[288,436,1046,1069]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="86" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis box="[288,436,1046,1069]" italics="true" pageId="4" pageNumber="86">Nesticus culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[442,522,1045,1069]" pageId="4" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Male palp: A. ventral view; B. dorsal view of the paracymbium; C. retrolateral view; D. female epigyne, ventral view; E vulva, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="4.[72,1465,1120,1694]" pageId="4" pageNumber="86">
|
||
Habitus as in
|
||
<figureCitation box="[273,363,1155,1180]" captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="4" pageNumber="86">Fig. 3A</figureCitation>
|
||
. Carapace uniformly pale yellow with some sparse setae (more reddish while alive, see
|
||
<figureCitation captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="4" pageNumber="86">Fig. 3A</figureCitation>
|
||
). Cephalic region not clearly differentiated from the rest of carapace. Eyes reduced, AM missing, reduced to black maculae. Eye diameters: AM -, AL 0.079, PM 0.080, and PL 0.078. Thoracic grooves and fovea distinct. Mouthparts and sternum uniformly colored as the carapace. Promargin of chelicera with three teeth approximately of the same size, retromargin with several small denticles. Legs uniformly light yellowish. Legs measurements as follows: I 17.40 (4.89, 0.90, 4.78, 4.90, 1.93), II 14.03 (3.92, 0.80, 3.67, 3.76, 1.88), III 10.35 (3.19, 0.69, 2.53, 2.79, 1.15), IV 13.47 (4.34, 0.84, 3.53, 3.49, 1.27). Leg formula: I, II, IV, III. Opisthosoma gray-yellowish colored (lighter than carapace while alive, see
|
||
<figureCitation box="[72,161,1379,1404]" captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="4" pageNumber="86">Fig. 3A</figureCitation>
|
||
), covered with long hairs.
|
||
</paragraph>
|
||
<paragraph blockId="4.[72,1465,1120,1694]" pageId="4" pageNumber="86">
|
||
Palp as in
|
||
<figureCitation box="[232,305,1412,1438]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="4" pageNumber="86">Figs. 1</figureCitation>
|
||
A‒C, 2A-C. Cymbium oval, covered with short, sparse setae, with a tuft of longer hairs in the pro- lateral distal area near the tip. Embolus filamentous, slender in the terminal part. Conductor complex with three distinct processes: Cp 1-3. Cp 1 stocky and roughly triangularly shaped, Cp 2 and Cp 3 located at the distal part of the bulb and diagonally protruding (approx. 2 o’clock seeing the left palp ventrally), their tips curved towards each other (
|
||
<figureCitation box="[1325,1461,1508,1534]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="4" pageNumber="86">Figs. 1A, C</figureCitation>
|
||
, 2A, C). Median apophysis well-developed, shaped as a long, sharp triangle, heading prolaterally (
|
||
<figureCitation box="[1227,1330,1541,1566]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="4" pageNumber="86">Figs. 1A</figureCitation>
|
||
, 2A). Para- cymbium large with well-developed, sclerotized dorsal, distal and ventral processes. Two dorsal apophyses, Da 1-2: Da 1 robust and long, ending sharply, S-shaped when the palp is observed dorsally, Da 2 stocky, triangularly-shaped. Distal apophysis triangularly shaped. Ventral apophysis lobate, dorso-ventrally flattened and heading toward the cym- bium (
|
||
<figureCitation box="[144,219,1668,1694]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="4" pageNumber="86">Figs. 1</figureCitation>
|
||
A‒C, 2A-C).
|
||
</paragraph>
|
||
<paragraph blockId="4.[72,1465,1718,1937]" box="[72,1266,1718,1744]" pageId="4" pageNumber="86">
|
||
<emphasis bold="true" box="[72,465,1719,1744]" pageId="4" pageNumber="86">Female (based on 4 paratypes)</emphasis>
|
||
. Total length 3.65–5.27. Carapace: 1.71–1.98 long, 1.54–1.64 wide.
|
||
</paragraph>
|
||
<paragraph blockId="4.[72,1465,1718,1937]" lastBlockId="6.[1208,1465,146,1709]" lastPageId="6" lastPageNumber="88" pageId="4" pageNumber="86">
|
||
Habitus as in
|
||
<figureCitation box="[261,378,1752,1778]" captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="4" pageNumber="86">Fig. 3B, C</figureCitation>
|
||
. Carapace uniformly yellowish with some sparse setae (often more reddish while alive, see
|
||
<figureCitation captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="4" pageNumber="86">Fig. 3B</figureCitation>
|
||
). Cephalic region not clearly differentiated from the rest of the prosoma.Eyes reduced, AM strongly reduced and bare- ly visible, reduced to small, dark maculae in some specimens. Eye diameters: AM (when present): 0.032, AL: 0.087, PM: 0.078, and PL: 0.77. Thoracic grooves and fovea distinct. Mouthparts and sternum uniformly colored as in the carapace. Teeth of chelicera as in the male. Legs uniformly light yellowish. Leg measurements as follows: I 17.07 (4.38, 1.01, 4.95, 4.90, 1.83), II 13.69 (4.04, 0.92, 3.57, 3.64, 1.52), III 10.46 (3.43, 0.80, 2.50, 2.56, 1.17), IV 13.89 (4.64, 0.92, 3.59, 3.41, 1.33). Leg formula: I, IV, II, III. Opistho- soma yellowish-gray (often lighter colored than carapace while alive, see
|
||
<figureCitation box="[1352,1452,306,331]" captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="6" pageNumber="88">Fig. 3B</figureCitation>
|
||
), covered with long hairs.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925084" ID-Zenodo-Dep="3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="5" pageNumber="87" startId="5.[141,206,1780,1804]" targetBox="[144,1536,144,1766]" targetPageId="5">
|
||
<paragraph blockId="5.[141,1536,1780,1880]" pageId="5" pageNumber="87">
|
||
Figure 3. Habitus and type locality of
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[526,677,1781,1804]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="87" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis box="[526,677,1781,1804]" italics="true" pageId="5" pageNumber="87">Domitius culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[684,764,1780,1804]" pageId="5" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green =
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[882,966,1831,1854]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="87" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis box="[882,966,1831,1854]" italics="true" pageId="5" pageNumber="87">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[974,1057,1831,1855]" pageId="5" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, orange =
|
||
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1880" box="[1165,1402,1831,1855]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="87" phylum="Arthropoda" rank="species" species="eremita">
|
||
<emphasis box="[1165,1402,1831,1855]" italics="true" pageId="5" pageNumber="87">Kryptonesticus eremita</emphasis>
|
||
</taxonomicName>
|
||
, arrow = en- trance of the cave.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[1208,1465,146,1709]" pageId="6" pageNumber="88">
|
||
Epigyne as in
|
||
<figureCitation captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="6" pageNumber="88">Figs. 1E, F</figureCitation>
|
||
, 2D, E. Medi- an septum short, not protruding, shaped as an inverted trape- zoid with a narrower base. Vulval pockets and copulatory ducts externally visible by transparence trough the tegument. Copu- latory openings at the lower, lateral side of median septum. Vul- va as in
|
||
<figureCitation box="[1308,1411,851,877]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="6" pageNumber="88">Figs. 1G</figureCitation>
|
||
, 2E. Spermathecae small and round, located in the lower-half of the vulva, below vulval pockets and being partially covered by them. Vulval pockets wide and rounded, sac-shaped, located above spermathecae. Copulatory ducts with a wider diameter in the ventral trait and narrower in the dor- sal trait, rolling up around the lower part of vulval pockets and reaching spermathe- cae with some turns (
|
||
<figureCitation box="[1215,1313,1492,1517]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="6" pageNumber="88">Figs. 1F</figureCitation>
|
||
, 2E). Insem- ination ducts begin- ning from the lower part of spermathe- cae and following the same course of copu- latory ducts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925086" ID-Zenodo-Dep="3925086" httpUri="https://zenodo.org/record/3925086/files/figure.png" pageId="6" pageNumber="88" startId="6.[72,137,1713,1737]" targetBox="[105,1152,144,1700]" targetPageId="6">
|
||
<paragraph blockId="6.[72,1176,1713,1787]" pageId="6" pageNumber="88">
|
||
Figure 4. Male palps of the Italian
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[433,523,1713,1736]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[433,523,1713,1736]" italics="true" pageId="6" pageNumber="88">Domitius</emphasis>
|
||
</taxonomicName>
|
||
species for comparison.
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[789,952,1713,1736]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[789,952,1713,1736]" italics="true" pageId="6" pageNumber="88">D. speluncarum</emphasis>
|
||
</taxonomicName>
|
||
male palp: A. ventral view; B. dorsal view; C. retrolateral view;
|
||
<taxonomicName authorityName="Di Caporiacco" authorityYear="1934" box="[489,608,1738,1761]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="menozzii">
|
||
<emphasis box="[489,608,1738,1761]" italics="true" pageId="6" pageNumber="88">D. menozzii</emphasis>
|
||
</taxonomicName>
|
||
male palp: D. ventral view; E. dorsal view; F. retrolateral view;
|
||
<taxonomicName baseAuthorityName="Brignoli" baseAuthorityYear="1979" box="[131,254,1764,1787]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis box="[131,254,1764,1787]" italics="true" pageId="6" pageNumber="88">D. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
male palp: G. ventral view; H. dorsal view; I. retrolateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="6" pageNumber="88" type="distribution">
|
||
<paragraph blockId="6.[1208,1464,1731,1833]" box="[1208,1388,1731,1761]" pageId="6" pageNumber="88">
|
||
<heading bold="true" box="[1208,1388,1731,1761]" fontSize="12" level="2" pageId="6" pageNumber="88" reason="0">
|
||
<emphasis bold="true" box="[1208,1388,1731,1761]" pageId="6" pageNumber="88">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[1208,1464,1731,1833]" lastBlockId="7.[144,1537,1589,1938]" lastPageId="7" lastPageNumber="89" pageId="6" pageNumber="88">
|
||
Italy, endemic to the northern Apennines. Known only from the type locality; Tana delle Fate di Coreglia Antelminelli cave (
|
||
<figureCitation box="[1100,1171,1840,1865]" captionStart="Figure 6" captionStartId="8.[72,137,1212,1236]" captionTargetBox="[72,1451,144,1193]" captionTargetId="figure@8.[72,1464,144,1193]" captionTargetPageId="8" captionText="Figure 6. Distribution of the genus Domitius in Italy. Yellow dots D. menozzii, blue dots D. speluncarum; red dot D. culsu sp. nov.; green dot D. sbordonii." figureDoi="http://doi.org/10.5281/zenodo.3925090" httpUri="https://zenodo.org/record/3925090/files/figure.png" pageId="6" pageNumber="88">Fig. 6</figureCitation>
|
||
).
|
||
<subSubSection lastPageId="9" lastPageNumber="91" pageId="7" pageNumber="89" type="biology_ecology">
|
||
<heading bold="true" box="[144,634,1589,1614]" fontSize="10" level="3" pageId="7" pageNumber="89" reason="6">
|
||
<emphasis bold="true" box="[144,634,1589,1614]" pageId="7" pageNumber="89">Biospeleological and ecological notes</emphasis>
|
||
</heading>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925088" ID-Zenodo-Dep="3925088" httpUri="https://zenodo.org/record/3925088/files/figure.png" pageId="7" pageNumber="89" startId="7.[144,209,1513,1537]" targetBox="[144,1536,144,1499]" targetPageId="7">
|
||
<paragraph blockId="7.[144,1537,1513,1562]" pageId="7" pageNumber="89">
|
||
Figure 5. Female epigyne of the Italian
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[543,633,1513,1536]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[543,633,1513,1536]" italics="true" pageId="7" pageNumber="89">Domitius</emphasis>
|
||
</taxonomicName>
|
||
species for comparison.
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[890,1051,1513,1536]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[890,1051,1513,1536]" italics="true" pageId="7" pageNumber="89">D. speluncarum</emphasis>
|
||
</taxonomicName>
|
||
: A. epigyne, ventral view; B. vulva, dorsal view;
|
||
<taxonomicName authorityName="Di Caporiacco" authorityYear="1934" box="[144,263,1539,1562]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="menozzii">
|
||
<emphasis box="[144,263,1539,1562]" italics="true" pageId="7" pageNumber="89">D. menozzii</emphasis>
|
||
</taxonomicName>
|
||
: C. epigyne, ventral view; D. vulva, dorsal view;
|
||
<taxonomicName baseAuthorityName="Brignoli" baseAuthorityYear="1979" box="[755,878,1539,1562]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis box="[755,878,1539,1562]" italics="true" pageId="7" pageNumber="89">D. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
: E. epigyne, ventral view; F. vulva.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[144,1537,1589,1938]" lastBlockId="8.[72,1465,1287,1920]" lastPageId="8" lastPageNumber="90" pageId="7" pageNumber="89">
|
||
The entrance of Tana delle Fate di Coreglia Antelminelli cave (Italian National Caves Registry number: 141/T/LU;
|
||
<figureCitation box="[144,231,1656,1681]" captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="7" pageNumber="89">Fig. 3E</figureCitation>
|
||
) opens in the left bank of the narrow valley of Segone Creek in the Province of Lucca (Toscana region) at an elevation of 260 m a.s.l. The cave occurs in the limestone of the Maiolica formation (lower Tithonian‒lower Aptian, ~150–120 Ma), which is particularly rich in flint nodules. After an initial steep slope (approximately 10 m deep), the cave continues with a long and sub-horizontal spatial development and a general NW‒SE orientation (
|
||
<figureCitation box="[1295,1383,1752,1777]" captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="7" pageNumber="89">Fig. 3F</figureCitation>
|
||
). It branches with several, sub-circular tunnels as a result of ancient groundwater flows. The cave has an estimated total extension of 1100 m, although the deeper segments are still unexplored, as they are either filled with water or ending with sumps. The inner section is generally humid, with mud often covering the bottoms of the tunnels. The cave hosts a rich sub- terranean fauna including some endemic or locally protected species, e.g. the carnivorous land snail
|
||
<taxonomicName box="[1375,1491,1880,1906]" class="Mollusca" family="Mollusca" genus="Oxychilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mollusca" pageId="7" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[1375,1491,1880,1906]" italics="true" pageId="7" pageNumber="89">Oxychilus</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName box="[150,297,1912,1938]" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" pageId="7" pageNumber="89" phylum="Mollusca" rank="class">Gastropoda</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="P.Hesse" authorityYear="1927" box="[309,453,1912,1938]" class="Gastropoda" family="Oxychilidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="89" phylum="Mollusca" rank="family">Oxychilidae</taxonomicName>
|
||
), the cave cricket
|
||
<taxonomicName authority="Minozzi, 1920" authorityName="Minozzi" authorityYear="1920" box="[668,1094,1912,1937]" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="89" phylum="Arthropoda" rank="species" species="laetitiae">
|
||
<emphasis box="[668,919,1912,1937]" italics="true" pageId="7" pageNumber="89">Dolichopoda laetitiae</emphasis>
|
||
Minozzi, 1920
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[1109,1244,1912,1938]" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="89" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
|
||
,
|
||
<taxonomicName box="[1256,1481,1912,1937]" class="Insecta" family="Rhaphidophoridae" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="89" phylum="Arthropoda" rank="family">Rhaphidophoridae</taxonomicName>
|
||
), the blind subterranean beetle
|
||
<taxonomicName authority="Straneo, 1943" authorityName="Straneo" authorityYear="1943" box="[379,840,1287,1313]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="subSpecies" species="apuanus" subSpecies="lanzai">
|
||
<emphasis box="[379,666,1287,1312]" italics="true" pageId="8" pageNumber="90">Duvalius apuanus lanzai</emphasis>
|
||
Straneo, 1943
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[854,991,1287,1313]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
, Trechinae), the Italian cave salamander
|
||
<taxonomicName authority="(Dunn, 1923)" baseAuthorityName="Dunn" baseAuthorityYear="1923" box="[72,495,1319,1344]" class="Amphibia" family="Plethodontidae" genus="Speleomantes" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="italicus">
|
||
<emphasis box="[72,333,1319,1344]" italics="true" pageId="8" pageNumber="90">Speleomantes italicus</emphasis>
|
||
(Dunn, 1923)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[509,628,1319,1344]" class="Amphibia" kingdom="Animalia" pageId="8" pageNumber="90" phylum="Chordata" rank="class">Amphibia</taxonomicName>
|
||
,
|
||
<taxonomicName box="[639,823,1319,1344]" class="Amphibia" family="Plethodontidae" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="90" phylum="Chordata" rank="family">Plethodontidae</taxonomicName>
|
||
), and three species of bats: the greater horseshoe bat
|
||
<taxonomicName authority="(Schreber, 1774)" baseAuthorityName="Schreber" baseAuthorityYear="1774" box="[72,617,1351,1377]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="ferrumequinum">
|
||
<emphasis box="[72,410,1351,1376]" italics="true" pageId="8" pageNumber="90">Rhinolophus ferrumequinum</emphasis>
|
||
(Schreber, 1774)
|
||
</taxonomicName>
|
||
, the lesser horseshoe bat
|
||
<taxonomicName authority="(Bechstein, 1800)" baseAuthorityName="Bechstein" baseAuthorityYear="1800" box="[936,1464,1351,1376]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="hipposideros">
|
||
<emphasis box="[936,1245,1351,1376]" italics="true" pageId="8" pageNumber="90">Rhinolophus hipposideros</emphasis>
|
||
(Bechstein, 1800)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Blumenbach" authorityYear="1779" box="[78,209,1383,1409]" class="Mammalia" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="90" phylum="Chordata" rank="order">Chiroptera</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Gray" authorityYear="1825" box="[220,397,1383,1408]" class="Mammalia" family="Rhinolophidae" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="90" phylum="Chordata" rank="family">Rhinolophidae</taxonomicName>
|
||
), and the common bent-wing bat
|
||
<taxonomicName authority="(Kuhl, 1817)" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[788,1220,1383,1408]" class="Mammalia" family="Miniopteridae" genus="Miniopterus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis box="[788,1072,1383,1408]" italics="true" pageId="8" pageNumber="90">Miniopterus schreibersii</emphasis>
|
||
(Kuhl, 1817)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Blumenbach" authorityYear="1779" box="[1234,1365,1383,1409]" class="Mammalia" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="90" phylum="Chordata" rank="order">Chiroptera</taxonomicName>
|
||
, Miniop- teridae). Other animals known from the cave from the literature (
|
||
<bibRefCitation author="Lanza, B" box="[815,963,1415,1440]" pageId="8" pageNumber="90" pagination="23 - 51" refId="ref7533" refString="Lanza, B, 1961, La fauna cavernicola della Toscana: Rassegna Speleologica Italiana, v. 13, no. 2, p. 23 - 51." title="La fauna cavernicola della Toscana: Rassegna Speleologica Italiana" type="book chapter" year="1961">Lanza, 1961</bibRefCitation>
|
||
) or directly observed by the author include:
|
||
<taxonomicName authority="(Duges, 1828)" baseAuthorityName="Duges" baseAuthorityYear="1828" box="[72,533,1447,1473]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="90" phylum="Annelida" rank="species" species="complanatus">
|
||
<emphasis box="[72,352,1447,1472]" italics="true" pageId="8" pageNumber="90">Octodrilus complanatus</emphasis>
|
||
(Dugès, 1828)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Cognetti, 1901)" baseAuthorityName="Cognetti" baseAuthorityYear="1901" box="[544,918,1447,1473]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="90" phylum="Annelida" rank="species" species="hemiandrus">
|
||
<emphasis box="[544,737,1447,1472]" italics="true" pageId="8" pageNumber="90">O. hemiandrus (</emphasis>
|
||
Cognetti, 1901)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Rosa, 1884)" baseAuthorityName="Rosa" baseAuthorityYear="1884" box="[933,1297,1447,1472]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="90" phylum="Annelida" rank="species" species="transpadanus">
|
||
<emphasis box="[933,1131,1447,1472]" italics="true" pageId="8" pageNumber="90">O. transpadanus</emphasis>
|
||
(Rosa, 1884)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="(Savigny, 1826)" baseAuthorityName="Savigny" baseAuthorityYear="1826" class="Clitellata" family="Lumbricidae" genus="Aporrectodea" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="90" phylum="Annelida" rank="species" species="rosea">
|
||
<emphasis italics="true" pageId="8" pageNumber="90">Aporrectodea rosea</emphasis>
|
||
(Savigny, 1826)
|
||
</taxonomicName>
|
||
(Anellida,
|
||
<taxonomicName box="[538,689,1479,1504]" class="Clitellata" family="Lumbricidae" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="90" phylum="Annelida" rank="family">Lumbricidae</taxonomicName>
|
||
);
|
||
<taxonomicName authority="(Dollfus, 1884)" baseAuthorityName="Dollfus" baseAuthorityYear="1884" box="[708,1176,1479,1504]" class="Malacostraca" family="Philosciidae" genus="Chaetophiloscia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="cellaria">
|
||
<emphasis box="[708,993,1479,1504]" italics="true" pageId="8" pageNumber="90">Chaetophiloscia cellaria</emphasis>
|
||
(Dollfus, 1884)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[1192,1294,1479,1504]" class="Malacostraca" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Isopoda</taxonomicName>
|
||
,
|
||
<taxonomicName box="[1306,1454,1479,1504]" class="Malacostraca" family="Philosciidae" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Philosciidae</taxonomicName>
|
||
),
|
||
<taxonomicName authority="Verhoeff, 1908" authorityName="Verhoeff" authorityYear="1908" box="[72,509,1511,1536]" class="Malacostraca" family="Trichoniscidae" genus="Androniscus" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="dentiger">
|
||
<emphasis box="[72,324,1511,1536]" italics="true" pageId="8" pageNumber="90">Androniscus dentiger</emphasis>
|
||
Verhoeff, 1908
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="(Brian, 1913)" baseAuthorityName="Brian" baseAuthorityYear="1913" box="[571,1011,1511,1536]" class="Malacostraca" family="Trichoniscidae" genus="Spelaeonethes" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="mancinii">
|
||
<emphasis box="[571,855,1511,1536]" italics="true" pageId="8" pageNumber="90">Spelaeonethes mancinii</emphasis>
|
||
(Brian, 1913)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[1026,1128,1511,1536]" class="Malacostraca" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Isopoda</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="G.O.Sars" authorityYear="1899" box="[1138,1315,1511,1536]" class="Malacostraca" family="Trichoniscidae" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Trichoniscidae</taxonomicName>
|
||
);
|
||
<taxonomicName authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="carpathicus">
|
||
<emphasis italics="true" pageId="8" pageNumber="90">Euscorpius carpathicus</emphasis>
|
||
(Linnaeus, 1767)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[438,579,1543,1569]" class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Scorpiones</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Laurie" authorityYear="1896" box="[593,753,1543,1568]" class="Arachnida" family="Euscorpiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Euscorpiidae</taxonomicName>
|
||
);
|
||
<taxonomicName box="[773,874,1543,1568]" class="Arachnida" family="Trogulidae" genus="Trogulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[773,874,1543,1568]" italics="true" pageId="8" pageNumber="90">Trogulus</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName authorityName="Sundevall" authorityYear="1833" box="[938,1056,1543,1569]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Opiliones</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Sundevall" authorityYear="1833" box="[1070,1200,1543,1568]" class="Arachnida" family="Trogulidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Trogulidae</taxonomicName>
|
||
),
|
||
<taxonomicName authorityName="C.L.Koch" authorityYear="1839" box="[1220,1375,1543,1568]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1220,1375,1543,1568]" italics="true" pageId="8" pageNumber="90">Ischyropsalis</emphasis>
|
||
</taxonomicName>
|
||
a
|
||
<emphasis box="[1400,1464,1543,1568]" italics="true" pageId="8" pageNumber="90">damii</emphasis>
|
||
Canestrini, 1873 (
|
||
<taxonomicName authorityName="Sundevall" authorityYear="1833" box="[281,399,1575,1601]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Opiliones</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Simon" authorityYear="1879" box="[411,623,1575,1600]" class="Arachnida" family="Ischyropsalididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Ischyropsalididae</taxonomicName>
|
||
);
|
||
<taxonomicName authority="Latzel, 1882" authorityName="Latzel" authorityYear="1882" box="[640,996,1575,1600]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="tylopus">
|
||
<emphasis box="[640,841,1575,1600]" italics="true" pageId="8" pageNumber="90">Lithobius tylopus</emphasis>
|
||
Latzel, 1882
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[1011,1138,1575,1601]" class="Chilopoda" kingdom="Animalia" pageId="8" pageNumber="90" phylum="Arthropoda" rank="class">Chilopoda</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Newport & May" authorityYear="1844" box="[1149,1284,1575,1600]" class="Chilopoda" family="Lithobiidae" kingdom="Animalia" order="Lithobiomorpha" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Lithobiidae</taxonomicName>
|
||
),
|
||
<taxonomicName authority="(Ocskay, 1832)" baseAuthorityName="Ocskay" baseAuthorityYear="1832" class="Insecta" family="Haglotettigoniidae" genus="Gryllomorpha" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="dalmatina">
|
||
<emphasis italics="true" pageId="8" pageNumber="90">Gryllomorpha dalmatina</emphasis>
|
||
(Ocskay, 1832)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[391,526,1607,1633]" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Laicharting" authorityYear="1781" box="[538,650,1607,1633]" class="Insecta" family="Gryllidae" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Gryllidae</taxonomicName>
|
||
),
|
||
<taxonomicName authorityName="Lucas" authorityYear="1895" box="[668,775,1607,1632]" class="Insecta" family="Erebidae" genus="Hypaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[668,775,1607,1632]" italics="true" pageId="8" pageNumber="90">Hypaena</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[834,982,1607,1632]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
|
||
,
|
||
<taxonomicName box="[994,1120,1607,1632]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Noctuidae</taxonomicName>
|
||
),
|
||
<taxonomicName authorityName="McLachlan" authorityYear="1895" box="[1138,1410,1607,1633]" class="Insecta" family="Limnephilidae" genus="Stenophylax" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="permistus">
|
||
<emphasis box="[1138,1410,1607,1633]" italics="true" pageId="8" pageNumber="90">Stenophylax permistus</emphasis>
|
||
</taxonomicName>
|
||
Mc- Lachlan, 1895 (Tricoptera,
|
||
<taxonomicName box="[389,559,1639,1664]" class="Insecta" family="Limnephilidae" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Limnephilidae</taxonomicName>
|
||
), and a large population of limoniid crane flies (
|
||
<taxonomicName box="[1108,1201,1639,1664]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
|
||
,
|
||
<taxonomicName box="[1211,1346,1639,1664]" class="Insecta" family="Limoniidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Limoniidae</taxonomicName>
|
||
). Near the entrance and in the early section of the cave, numerous spiders were also observed:
|
||
<taxonomicName authority="(Walckenaer, 1830)" baseAuthorityName="Walckenaer" baseAuthorityYear="1830" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="ferox">
|
||
<emphasis box="[1090,1300,1671,1696]" italics="true" pageId="8" pageNumber="90">Amaurobius ferox</emphasis>
|
||
(Walckenaer, 1830)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Ballarin and Pantini, 2017" authorityName="Ballarin and Pantini" authorityYear="2017" box="[153,594,1703,1728]" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="pesarinii">
|
||
<emphasis box="[153,286,1703,1728]" italics="true" pageId="8" pageNumber="90">A. pesarinii</emphasis>
|
||
Ballarin and Pantini, 2017
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[607,780,1703,1728]" class="Arachnida" family="Amaurobiidae" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Amaurobiidae</taxonomicName>
|
||
),
|
||
<taxonomicName authority="(Simon, 1880)" baseAuthorityName="Simon" baseAuthorityYear="1880" box="[796,1241,1703,1729]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="eremita">
|
||
<emphasis box="[796,1069,1703,1728]" italics="true" pageId="8" pageNumber="90">Kryptonesticus eremita</emphasis>
|
||
(Simon, 1880)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Simon" authorityYear="1894" box="[1255,1388,1703,1728]" class="Arachnida" family="Nesticidae" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Nesticidae</taxonomicName>
|
||
),
|
||
<taxonomicName authority="(Latreille, 1804)" baseAuthorityName="Latreille" baseAuthorityYear="1804" class="Arachnida" family="Tetragnathidae" genus="Meta" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="menardi">
|
||
<emphasis italics="true" pageId="8" pageNumber="90">Meta menardi</emphasis>
|
||
(Latreille, 1804)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Scopoli, 1763)" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[375,785,1735,1761]" class="Arachnida" family="Tetragnathidae" genus="Metellina" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="merianae">
|
||
<emphasis box="[375,601,1735,1760]" italics="true" pageId="8" pageNumber="90">Metellina merianae</emphasis>
|
||
(Scopoli, 1763)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Menge" authorityYear="1866" box="[798,984,1735,1760]" class="Arachnida" family="Tetragnathidae" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Tetragnathidae</taxonomicName>
|
||
),
|
||
<taxonomicName authority="(Fuesslin, 1775)" baseAuthorityName="Fuesslin" baseAuthorityYear="1775" box="[1001,1464,1735,1760]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="phalangioides">
|
||
<emphasis box="[1001,1268,1735,1760]" italics="true" pageId="8" pageNumber="90">Pholcus phalangioides</emphasis>
|
||
(Fuesslin, 1775)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[78,203,1767,1792]" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
|
||
), and
|
||
<taxonomicName box="[277,396,1767,1792]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[277,396,1767,1792]" italics="true" pageId="8" pageNumber="90">Tegenaria</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName box="[457,598,1767,1792]" class="Arachnida" family="Agelenidae" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="family">Agelenidae</taxonomicName>
|
||
). The new species was found in the initial segments of the cave, but at some distance from the entrance (
|
||
<figureCitation box="[474,561,1799,1824]" captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="8" pageNumber="90">Fig. 3F</figureCitation>
|
||
). During summer, when the cave was visited, adults, subadults, and juveniles of
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[104,203,1831,1856]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis box="[104,203,1831,1856]" italics="true" pageId="8" pageNumber="90">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[214,309,1831,1856]" pageId="8" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were observed together, with a substantially higher number of adults and subadults during the month of August. Most of the juveniles collected in the cave and bred in captivity became adults after 2‒3 months of captivity, while it took approximately one year for the youngest specimens to reach sexual maturity.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925090" ID-Zenodo-Dep="3925090" httpUri="https://zenodo.org/record/3925090/files/figure.png" pageId="8" pageNumber="90" startId="8.[72,137,1212,1236]" targetBox="[72,1451,144,1193]" targetPageId="8">
|
||
<paragraph blockId="8.[72,1465,1212,1261]" pageId="8" pageNumber="90">
|
||
Figure 6. Distribution of the genus
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[427,517,1212,1235]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[427,517,1212,1235]" italics="true" pageId="8" pageNumber="90">Domitius</emphasis>
|
||
</taxonomicName>
|
||
in Italy. Yellow dots
|
||
<taxonomicName authorityName="Di Caporiacco" authorityYear="1934" box="[751,871,1212,1235]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="menozzii">
|
||
<emphasis box="[751,871,1212,1235]" italics="true" pageId="8" pageNumber="90">D. menozzii</emphasis>
|
||
</taxonomicName>
|
||
, blue dots
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[1010,1171,1212,1235]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[1010,1171,1212,1235]" italics="true" pageId="8" pageNumber="90">D. speluncarum</emphasis>
|
||
</taxonomicName>
|
||
; red dot
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[1289,1373,1212,1235]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis box="[1289,1373,1212,1235]" italics="true" pageId="8" pageNumber="90">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1379,1462,1212,1236]" pageId="8" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; green dot
|
||
<taxonomicName baseAuthorityName="Brignoli" baseAuthorityYear="1979" box="[200,323,1237,1260]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis box="[200,323,1237,1260]" italics="true" pageId="8" pageNumber="90">D. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925092" ID-Zenodo-Dep="3925092" httpUri="https://zenodo.org/record/3925092/files/figure.png" pageId="9" pageNumber="91" startId="9.[144,209,1102,1126]" targetBox="[153,1512,146,1081]" targetPageId="9">
|
||
<paragraph blockId="9.[144,1536,1102,1227]" pageId="9" pageNumber="91">
|
||
Figure 7. Phylogenetic tree of the main European nesticid genera inferred using ML in RAxML. Different colored branches reflect different genera: violet =
|
||
<taxonomicName authorityName="Kulczynski" authorityYear="1914" box="[306,458,1127,1151]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[306,458,1127,1151]" italics="true" pageId="9" pageNumber="91">Typhlonesticus</emphasis>
|
||
</taxonomicName>
|
||
, green =
|
||
<taxonomicName box="[553,732,1127,1150]" class="Arachnida" family="Nesticidae" genus="Carpathonesticus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[553,732,1127,1150]" italics="true" pageId="9" pageNumber="91">Carpathonesticus</emphasis>
|
||
</taxonomicName>
|
||
, yellow =
|
||
<taxonomicName authorityName="Thorell" authorityYear="1869" box="[833,921,1128,1151]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[833,921,1128,1151]" italics="true" pageId="9" pageNumber="91">Nesticus</emphasis>
|
||
</taxonomicName>
|
||
, blue =
|
||
<taxonomicName authorityName="Pavlek and Ribera" authorityYear="2017" box="[1001,1152,1127,1151]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1001,1152,1127,1151]" italics="true" pageId="9" pageNumber="91">Kryptonesticus</emphasis>
|
||
</taxonomicName>
|
||
, red =
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[1222,1311,1128,1151]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1222,1311,1128,1151]" italics="true" pageId="9" pageNumber="91">Domitius</emphasis>
|
||
</taxonomicName>
|
||
. The newly-described species is highlighted in bold. Species distributed in Italy are underlined. Code before each species refers to the origin of the data, see Table 1. Branch lengths are scaled in relation to the number of substitutions per site; numbers at nodes denote bootstrap support according to ML.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="9.[144,1536,1253,1662]" pageId="9" pageNumber="91">
|
||
Two different species of nesticid spiders,
|
||
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1880" box="[689,814,1253,1278]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="eremita">
|
||
<emphasis box="[689,814,1253,1278]" italics="true" pageId="9" pageNumber="91">K. eremita</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[881,980,1253,1278]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis box="[881,980,1253,1278]" italics="true" pageId="9" pageNumber="91">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[991,1090,1253,1278]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, were collected together in the Tana delle Fate di Coreglia Antelminelli cave. These species cover a different spatial distribution within the cave (
|
||
<figureCitation box="[1434,1524,1285,1310]" captionStart="Figure 3" captionStartId="5.[141,206,1780,1804]" captionTargetBox="[144,1536,144,1766]" captionTargetId="figure@5.[144,1536,144,1766]" captionTargetPageId="5" captionText="Figure 3. Habitus and type locality of Domitius culsu sp. nov. A. Habitus of male; B. habitus of female; C. female with prey; D. juvenile in captivity; E. entrance of Tana delle Fate di Coreglia Antelminelli cave; F. map of the cave and detail of the entrance, showing the spatial distribution of the two co-existing nesticid species living inside: green = D. culsu sp. nov., orange = Kryptonesticus eremita, arrow = en- trance of the cave." figureDoi="http://doi.org/10.5281/zenodo.3925084" httpUri="https://zenodo.org/record/3925084/files/figure.png" pageId="9" pageNumber="91">Fig. 3F</figureCitation>
|
||
), coexisting without overlapping despite occupying approximately the same ecological niche. Cohabitant nesticids, in particular involving
|
||
<taxonomicName authorityName="Di Caporiacco" authorityYear="1934" box="[374,513,1349,1374]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="menozzii">
|
||
<emphasis box="[374,513,1349,1374]" italics="true" pageId="9" pageNumber="91">D. menozzii</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[551,739,1349,1374]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[551,739,1349,1374]" italics="true" pageId="9" pageNumber="91">D. speluncarum</emphasis>
|
||
</taxonomicName>
|
||
together with
|
||
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1880" box="[907,1030,1349,1374]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="eremita">
|
||
<emphasis box="[907,1030,1349,1374]" italics="true" pageId="9" pageNumber="91">K. eremita</emphasis>
|
||
</taxonomicName>
|
||
, have been previously observed in several occasions in Italian caves, and sometimes collected at short distances from each other (
|
||
<bibRefCitation author="Brignoli, P. M." box="[1200,1369,1381,1406]" pageId="9" pageNumber="91" pagination="121 - 229" refId="ref7176" refString="Brignoli, P. M., 1971, Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica, p. 121 - 229." title="Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica" type="book chapter" year="1971">Brignoli, 1971</bibRefCitation>
|
||
). However, no clear species overlap are reported within the same cave. Such distinct spatial partition can be explained by the different grade of adaptation to the hypogean environment showed by these arachnids. In fact,
|
||
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1880" box="[1164,1286,1445,1470]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="eremita">
|
||
<emphasis box="[1164,1286,1445,1470]" italics="true" pageId="9" pageNumber="91">K. eremita</emphasis>
|
||
</taxonomicName>
|
||
appears to be a less specialized cave-dweller, lacking extreme morphological adaptations to subterranean life. Therefore, it mostly occurs near the entrance of caves or inside artificial tunnels, including, occasionally, shadowed epigean habitats with constant temperature and high relative humidity (
|
||
<bibRefCitation author="Brignoli, P. M." box="[611,775,1541,1566]" pageId="9" pageNumber="91" pagination="121 - 229" refId="ref7176" refString="Brignoli, P. M., 1971, Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica, p. 121 - 229." title="Note su ragni cavernicoli italiani (Araneae): Fragmenta Entomologica" type="book chapter" year="1971">Brignoli, 1971</bibRefCitation>
|
||
and personal observations by the author). On the other hand, all
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[144,248,1573,1598]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[144,248,1573,1598]" italics="true" pageId="9" pageNumber="91">Domitius</emphasis>
|
||
</taxonomicName>
|
||
species show a greater degree of adaptation to the subterranean habitat, as suggested by reduction of the eyes and body depigmentation. Such strong adaptation allows
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[883,987,1605,1630]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[883,987,1605,1630]" italics="true" pageId="9" pageNumber="91">Domitius</emphasis>
|
||
</taxonomicName>
|
||
to occupy deeper segments of the caves, thus avoiding direct competition with
|
||
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1880" box="[524,646,1637,1662]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="eremita">
|
||
<emphasis box="[524,646,1637,1662]" italics="true" pageId="9" pageNumber="91">K. eremita</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="91" type="conservation">
|
||
<paragraph blockId="9.[144,1536,1681,1934]" box="[144,400,1681,1707]" pageId="9" pageNumber="91">
|
||
<heading bold="true" box="[144,400,1681,1707]" fontSize="10" level="3" pageId="9" pageNumber="91" reason="6">
|
||
<emphasis bold="true" box="[144,400,1681,1707]" pageId="9" pageNumber="91">Conservation Notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="9.[144,1536,1681,1934]" pageId="9" pageNumber="91">
|
||
Since caves are a unique and delicate ecosystem, they are highly susceptible to external disturbance (
|
||
<bibRefCitation author="Culver D. C. & Pipan T." journalOrPublisher="UK: Oxford University Press" pageId="9" pageNumber="91" pagination="256" refId="ref7295" refString="Culver D. C., and Pipan T., 2009, The Biology of Caves and Other Subterranean Habitats: Oxford, UK: Oxford University Press, p. 256." title="The Biology of Caves and Other Subterranean Habitats: Oxford" type="book chapter" year="2009">Culver and Pipan, 2009</bibRefCitation>
|
||
). Its visible entrance and sub-horizontal extension makes Tana delle Fate di Coreglia Antelminelli cave easily accessible to visitors even with limited experience in speleology, and the cave is often used for training purpos- es by local speleological clubs. Although not threatened,
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[830,927,1813,1838]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis box="[830,927,1813,1838]" italics="true" pageId="9" pageNumber="91">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[937,1030,1813,1838]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
should be considered potentially at risk in case of frequent and long-lasting human disturbance due to its strict habitat requirements, its reduced population, and its extremely limited distribution, which appears to be confined to a single cave. Therefore, the new species is a good candidate for species conservation, deserving a place in the list of locally protected species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="11" lastPageNumber="93" pageId="10" pageNumber="92" type="discussion">
|
||
<paragraph blockId="10.[72,1465,146,977]" box="[72,358,146,171]" pageId="10" pageNumber="92">
|
||
<heading bold="true" box="[72,358,146,171]" fontSize="10" level="3" pageId="10" pageNumber="92" reason="6">
|
||
<emphasis bold="true" box="[72,358,146,171]" pageId="10" pageNumber="92">Phylogenetic Analysis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="10.[72,1465,146,977]" pageId="10" pageNumber="92">
|
||
A total of 16 nesticid species were used in this study, including representatives of the main nesticid genera present in Europe and all the species distributed in the Italian peninsula. Taxon sampling comprised the wide majority of
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[1359,1463,214,239]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1359,1463,214,239]" italics="true" pageId="10" pageNumber="92">Domitius</emphasis>
|
||
</taxonomicName>
|
||
species. Only
|
||
<taxonomicName authority="(Ribera and De Mas, 2003)" baseAuthorityName="Ribera and De Mas" baseAuthorityYear="2003" box="[244,697,246,271]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="murgis">
|
||
<emphasis box="[244,359,246,271]" italics="true" pageId="10" pageNumber="92">D. murgis</emphasis>
|
||
(Ribera and De Mas, 2003)
|
||
</taxonomicName>
|
||
from Spain was excluded from the analysis due to the absence of available sequences and fresh samples. The final dataset was formed by 1975 pair bases (bp) distributed as; COI = 1197 bp, 16S = 469 bp, and H3 = 309 bp. The resulting phylogenetic tree is illustrated in
|
||
<figureCitation box="[1115,1214,310,335]" captionStart="Figure 7" captionStartId="9.[144,209,1102,1126]" captionTargetBox="[153,1512,146,1081]" captionTargetId="figure@9.[144,1549,144,1085]" captionTargetPageId="9" captionText="Figure 7. Phylogenetic tree of the main European nesticid genera inferred using ML in RAxML. Different colored branches reflect different genera: violet = Typhlonesticus, green = Carpathonesticus, yellow = Nesticus, blue = Kryptonesticus, red = Domitius. The newly-described species is highlighted in bold. Species distributed in Italy are underlined. Code before each species refers to the origin of the data, see Table 1. Branch lengths are scaled in relation to the number of substitutions per site; numbers at nodes denote bootstrap support according to ML." figureDoi="http://doi.org/10.5281/zenodo.3925092" httpUri="https://zenodo.org/record/3925092/files/figure.png" pageId="10" pageNumber="92">Figure 7</figureCitation>
|
||
, and the uncorrected pairwise distance between the species is reported in
|
||
<tableCitation box="[704,794,342,367]" captionStart="Table 2" captionStartId="10.[77,136,1023,1046]" captionTargetPageId="10" captionText="Table 2. Uncorrected genetic p-distance of the COI partial sequence of the nesticid species discussed in the text. The newly described species is in bold." httpUri="http://table.plazi.org/id/DF6A418C1E68FFC7FFF067D2C6D3FBF6" pageId="10" pageNumber="92" tableUuid="DF6A418C1E68FFC7FFF067D2C6D3FBF6">Table 2</tableCitation>
|
||
. The European nesticids cluster into five different clades corresponding to the main genera
|
||
<taxonomicName box="[481,689,373,398]" class="Arachnida" family="Nesticidae" genus="Carpathonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[481,689,373,398]" italics="true" pageId="10" pageNumber="92">Carpathonesticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[705,809,374,399]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[705,809,374,399]" italics="true" pageId="10" pageNumber="92">Domitius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Pavlek and Ribera" authorityYear="2017" box="[824,1001,374,399]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[824,1001,374,399]" italics="true" pageId="10" pageNumber="92">Kryptonesticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Thorell" authorityYear="1869" box="[1016,1119,374,399]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1016,1119,374,399]" italics="true" pageId="10" pageNumber="92">Nesticus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authorityName="Kulczynski" authorityYear="1914" box="[1185,1361,374,399]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1185,1361,374,399]" italics="true" pageId="10" pageNumber="92">Typhlonesticus</emphasis>
|
||
</taxonomicName>
|
||
, each of them highly supported (bootstrap support value = 100%). Each lineage represents a different and well-defined evolu- tionary line. These results concur with the outcomes of recent morphological and phylogenetic studies on the family
|
||
<taxonomicName authorityName="Simon" authorityYear="1894" box="[72,198,470,495]" class="Arachnida" family="Nesticidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="family">Nesticidae</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Pavlek, M. & Ribera, C." box="[213,513,470,495]" journalOrPublisher="et sp. nov. (Araneae, Nesticidae), with notes on the Mediterranean cave spe- cies: European Journal of Taxonomy" pageId="10" pageNumber="92" pagination="1 - 27" part="262" refId="ref7835" refString="Pavlek, M., and Ribera, C. 2017, Kryptonesticus deelemanae gen. et sp. nov. (Araneae, Nesticidae), with notes on the Mediterranean cave spe- cies: European Journal of Taxonomy, 262, p. 1 - 27, https: // doi. org / 10.5852 / ejt. 2017.262." title="Kryptonesticus deelemanae gen" type="journal article" year="2017">Pavlek and Ribera, 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ribera, C." box="[526,683,470,495]" pageId="10" pageNumber="92" pagination="229 - 240" refId="ref7897" refString="Ribera, C., 2018, A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa, v. 4407, p. 229 - 240, https: // doi. org / 10.11646 / zootaxa. 4407.2.4." title="A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa" type="book chapter" year="2018">Ribera, 2018</bibRefCitation>
|
||
; Ballarin and Li, in prep.), supporting the validity of the newly-es- tablished genera. According to these results,
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[611,715,502,527]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[611,715,502,527]" italics="true" pageId="10" pageNumber="92">Domitius</emphasis>
|
||
</taxonomicName>
|
||
represents the sister lineage of the monophyletic clade formed by the genera
|
||
<taxonomicName box="[242,450,533,558]" class="Arachnida" family="Nesticidae" genus="Carpathonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[242,450,533,558]" italics="true" pageId="10" pageNumber="92">Carpathonesticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Pavlek and Ribera" authorityYear="2017" box="[466,643,534,559]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[466,643,534,559]" italics="true" pageId="10" pageNumber="92">Kryptonesticus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authorityName="Thorell" authorityYear="1869" box="[709,812,534,559]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[709,812,534,559]" italics="true" pageId="10" pageNumber="92">Nesticus</emphasis>
|
||
</taxonomicName>
|
||
, with which it shares a common ancestor. The analysis supports
|
||
<taxonomicName authorityName="Kulczynski" authorityYear="1914" box="[183,359,566,591]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[183,359,566,591]" italics="true" pageId="10" pageNumber="92">Typhlonesticus</emphasis>
|
||
</taxonomicName>
|
||
as a basal clade within the European
|
||
<taxonomicName authorityName="Simon" authorityYear="1894" box="[805,935,566,591]" class="Arachnida" family="Nesticidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="family">Nesticidae</taxonomicName>
|
||
, as also suggested by recent molecular stud- ies (
|
||
<bibRefCitation author="Ballarin, F. & Li, S." box="[119,366,598,623]" pageId="10" pageNumber="92" pagination="577 - 591" refId="ref7121" refString="Ballarin, F., and Li, S., 2018, Diversification in tropics and subtropics following the mid-Miocene climate change: A case study of the spider genus Nesticella: Global Change Biology, v. 24, p. 577 - 591, doi: 10.1111 / gcb. 13958." title="Diversification in tropics and subtropics following the mid-Miocene climate change: A case study of the spider genus Nesticella: Global Change Biology" type="book chapter" year="2018">Ballarin and Li, 2018</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ribera, C." box="[375,531,598,623]" pageId="10" pageNumber="92" pagination="229 - 240" refId="ref7897" refString="Ribera, C., 2018, A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa, v. 4407, p. 229 - 240, https: // doi. org / 10.11646 / zootaxa. 4407.2.4." title="A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa" type="book chapter" year="2018">Ribera, 2018</bibRefCitation>
|
||
). Within
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[629,733,598,623]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[629,733,598,623]" italics="true" pageId="10" pageNumber="92">Domitius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[747,842,598,623]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov.">
|
||
<emphasis box="[747,842,598,623]" italics="true" pageId="10" pageNumber="92">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[850,941,598,623]" pageId="10" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||
shows a closer affinity with the species from the same geographic area; particularly
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[536,723,630,655]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[536,723,630,655]" italics="true" pageId="10" pageNumber="92">D. speluncarum</emphasis>
|
||
</taxonomicName>
|
||
, but also
|
||
<taxonomicName box="[839,971,630,655]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="menozzi">
|
||
<emphasis box="[839,971,630,655]" italics="true" pageId="10" pageNumber="92">D. menozzi</emphasis>
|
||
</taxonomicName>
|
||
. Its position at the far end of the phyloge- netic tree of the genus suggests a more recent origin in comparison with the other congeneric species.
|
||
</paragraph>
|
||
<paragraph blockId="10.[72,1465,146,977]" lastBlockId="11.[144,1536,146,203]" lastPageId="11" lastPageNumber="93" pageId="10" pageNumber="92">
|
||
All the
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[192,296,696,721]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[192,296,696,721]" italics="true" pageId="10" pageNumber="92">Domitius</emphasis>
|
||
</taxonomicName>
|
||
species distributed in the Northern Apennines share a close affinity with species from the Iberian Peninsula. Such close relations also reflected in genital morphology. For instance, all these species share a similar position of spermathecae, located in the lower-half of the vulva, and below the vulval pockets (see
|
||
<figureCitation box="[1231,1333,759,785]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="10" pageNumber="92">Figs. 1G</figureCitation>
|
||
, 5B, D and
|
||
<figureCitation box="[72,150,792,817]" captionStart="Figure 4" captionStartId="6.[72,137,1713,1737]" captionTargetBox="[105,1152,144,1700]" captionTargetId="figure@6.[72,1176,144,1706]" captionTargetPageId="6" captionText="Figure 4. Male palps of the Italian Domitius species for comparison. D. speluncarum male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. menozzii male palp: D. ventral view; E. dorsal view; F. retrolateral view; D. sbordonii male palp: G. ventral view; H. dorsal view; I. retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.3925086" httpUri="https://zenodo.org/record/3925086/files/figure.png" pageId="10" pageNumber="92">Figs. 4</figureCitation>
|
||
A‒E in
|
||
<bibRefCitation author="Ribera, C." box="[236,391,792,817]" pageId="10" pageNumber="92" pagination="229 - 240" refId="ref7897" refString="Ribera, C., 2018, A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa, v. 4407, p. 229 - 240, https: // doi. org / 10.11646 / zootaxa. 4407.2.4." title="A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa" type="book chapter" year="2018">Ribera, 2018</bibRefCitation>
|
||
). On the other hand,
|
||
<taxonomicName baseAuthorityName="Brignoli" baseAuthorityYear="1979" box="[633,775,792,817]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis box="[633,775,792,817]" italics="true" pageId="10" pageNumber="92">D. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
from the Central Apennines appears to be morphologically and genetically separated from all the other species of the genus, including those from Northern Apennines. The differ- ence is highlighted in the peculiar shape of the vulva, being the only
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[891,995,856,881]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[891,995,856,881]" italics="true" pageId="10" pageNumber="92">Domitius</emphasis>
|
||
</taxonomicName>
|
||
species showing simple-coiled internal ducts and spermathecae located in the upper-half of the vulva, over the vulval pockets (see
|
||
<figureCitation box="[1152,1235,888,913]" captionStart="Figure 5" captionStartId="7.[144,209,1513,1537]" captionTargetBox="[144,1536,144,1499]" captionTargetId="figure@7.[144,1536,144,1508]" captionTargetPageId="7" captionText="Figure 5. Female epigyne of the Italian Domitius species for comparison. D. speluncarum: A. epigyne, ventral view; B. vulva, dorsal view; D. menozzii: C. epigyne, ventral view; D. vulva, dorsal view; D. sbordonii: E. epigyne, ventral view; F. vulva." figureDoi="http://doi.org/10.5281/zenodo.3925088" httpUri="https://zenodo.org/record/3925088/files/figure.png" pageId="10" pageNumber="92">Fig. 5F</figureCitation>
|
||
vs.
|
||
<figureCitation box="[1283,1385,887,913]" captionStart="Figure 1" captionStartId="3.[144,209,1504,1528]" captionTargetBox="[144,1198,144,1493]" captionTargetId="figure@3.[144,1198,144,1497]" captionText="Figure 1. Nesticus culsu sp. nov. Male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. female, cephalic region showing the eye pattern; E. epigyne, ventral view; F. epigyne after clearing, ventral view, the schematic course of internal ducts is outlined with a white line; G. vulva after clearing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3925080" httpUri="https://zenodo.org/record/3925080/files/figure.png" pageId="10" pageNumber="92">Figs. 1G</figureCitation>
|
||
, 5B, D and
|
||
<figureCitation box="[123,202,920,945]" captionStart="Figure 4" captionStartId="6.[72,137,1713,1737]" captionTargetBox="[105,1152,144,1700]" captionTargetId="figure@6.[72,1176,144,1706]" captionTargetPageId="6" captionText="Figure 4. Male palps of the Italian Domitius species for comparison. D. speluncarum male palp: A. ventral view; B. dorsal view; C. retrolateral view; D. menozzii male palp: D. ventral view; E. dorsal view; F. retrolateral view; D. sbordonii male palp: G. ventral view; H. dorsal view; I. retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.3925086" httpUri="https://zenodo.org/record/3925086/files/figure.png" pageId="10" pageNumber="92">Figs. 4</figureCitation>
|
||
A‒E in
|
||
<bibRefCitation author="Ribera, C." box="[290,446,920,945]" pageId="10" pageNumber="92" pagination="229 - 240" refId="ref7897" refString="Ribera, C., 2018, A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa, v. 4407, p. 229 - 240, https: // doi. org / 10.11646 / zootaxa. 4407.2.4." title="A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa" type="book chapter" year="2018">Ribera, 2018</bibRefCitation>
|
||
). Upper-positioned spermathechae are also present in several other European nesticid genera such as
|
||
<taxonomicName box="[259,467,951,976]" class="Arachnida" family="Nesticidae" genus="Carpathonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[259,467,951,976]" italics="true" pageId="10" pageNumber="92">Carpathonesticus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[481,630,952,977]" italics="true" pageId="10" pageNumber="92">sensu stricto</emphasis>
|
||
),
|
||
<emphasis box="[652,844,952,977]" italics="true" pageId="10" pageNumber="92">Krypthonesticus</emphasis>
|
||
, and
|
||
<taxonomicName authorityName="Thorell" authorityYear="1869" box="[908,1011,952,977]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[908,1011,952,977]" italics="true" pageId="10" pageNumber="92">Nesticus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[1025,1173,952,977]" italics="true" pageId="10" pageNumber="92">sensu stricto</emphasis>
|
||
). Based on these results it is possible to speculate that
|
||
<taxonomicName baseAuthorityName="Brignoli" baseAuthorityYear="1979" box="[496,640,146,171]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis box="[496,640,146,171]" italics="true" pageId="11" pageNumber="93">D. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
represents a basal element within the genus
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[1171,1275,146,171]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1171,1275,146,171]" italics="true" pageId="11" pageNumber="93">Domitius</emphasis>
|
||
</taxonomicName>
|
||
, possibly still carrying the ancestral characters of the older forebear of the European nesticids.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="DF6A418C1E68FFC7FFF067D2C6D3FBF6" httpUri="http://table.plazi.org/id/DF6A418C1E68FFC7FFF067D2C6D3FBF6" pageId="10" pageNumber="92" targetBox="[78,1451,1087,1939]" targetIsTable="true" targetPageId="10">
|
||
<paragraph blockId="10.[77,1440,1023,1071]" pageId="10" pageNumber="92">Table 2. Uncorrected genetic p-distance of the COI partial sequence of the nesticid species discussed in the text. The newly described species is in bold.</paragraph>
|
||
</caption>
|
||
<paragraph pageId="10" pageNumber="92">
|
||
<table box="[78,1451,1087,1939]" gridcols="17" gridrows="17" pageId="10" pageNumber="92">
|
||
<tr box="[78,1451,1087,1110]" gridrow="0" pageId="10" pageNumber="92">
|
||
<th box="[78,117,1087,1110]" gridcol="0" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[78,117,1087,1110]" pageId="10" pageNumber="92">No.</emphasis>
|
||
</th>
|
||
<th box="[134,392,1087,1110]" gridcol="1" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[220,313,1087,1110]" pageId="10" pageNumber="92">Species</emphasis>
|
||
</th>
|
||
<th box="[418,473,1087,1110]" gridcol="2" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[439,452,1087,1110]" pageId="10" pageNumber="92">1</emphasis>
|
||
</th>
|
||
<th box="[488,543,1087,1110]" gridcol="3" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[509,522,1087,1110]" pageId="10" pageNumber="92">2</emphasis>
|
||
</th>
|
||
<th box="[558,613,1087,1110]" gridcol="4" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[579,592,1087,1110]" pageId="10" pageNumber="92">3</emphasis>
|
||
</th>
|
||
<th box="[628,684,1087,1110]" gridcol="5" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[649,662,1087,1110]" pageId="10" pageNumber="92">4</emphasis>
|
||
</th>
|
||
<th box="[698,752,1087,1110]" gridcol="6" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[719,732,1087,1110]" pageId="10" pageNumber="92">5</emphasis>
|
||
</th>
|
||
<th box="[768,823,1087,1110]" gridcol="7" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[789,802,1087,1110]" pageId="10" pageNumber="92">6</emphasis>
|
||
</th>
|
||
<th box="[838,893,1087,1110]" gridcol="8" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[859,872,1087,1110]" pageId="10" pageNumber="92">7</emphasis>
|
||
</th>
|
||
<th box="[908,963,1087,1110]" gridcol="9" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[929,942,1087,1110]" pageId="10" pageNumber="92">8</emphasis>
|
||
</th>
|
||
<th box="[978,1033,1087,1110]" gridcol="10" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[999,1012,1087,1110]" pageId="10" pageNumber="92">9</emphasis>
|
||
</th>
|
||
<th box="[1048,1103,1087,1110]" gridcol="11" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[1062,1088,1087,1110]" pageId="10" pageNumber="92">10</emphasis>
|
||
</th>
|
||
<th box="[1118,1173,1087,1110]" gridcol="12" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[1133,1157,1087,1110]" pageId="10" pageNumber="92">11</emphasis>
|
||
</th>
|
||
<th box="[1188,1243,1087,1110]" gridcol="13" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[1202,1227,1087,1110]" pageId="10" pageNumber="92">12</emphasis>
|
||
</th>
|
||
<th box="[1258,1313,1087,1110]" gridcol="14" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[1272,1297,1087,1110]" pageId="10" pageNumber="92">13</emphasis>
|
||
</th>
|
||
<th box="[1327,1382,1087,1110]" gridcol="15" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[1342,1367,1087,1110]" pageId="10" pageNumber="92">14</emphasis>
|
||
</th>
|
||
<th box="[1398,1451,1087,1110]" gridcol="16" gridrow="0" pageId="10" pageNumber="92">
|
||
<emphasis bold="true" box="[1412,1437,1087,1110]" pageId="10" pageNumber="92">15</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[78,1451,1126,1146]" gridrow="1" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[78,117,1126,1146]" gridcol="0" gridrow="1" pageId="10" pageNumber="92">1</th>
|
||
<td box="[134,392,1126,1146]" gridcol="1" gridrow="1" pageId="10" pageNumber="92">
|
||
Dbae_
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[199,283,1126,1146]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Domitius</taxonomicName>
|
||
_baeticus
|
||
</td>
|
||
</tr>
|
||
<tr box="[78,1451,1162,1182]" gridrow="2" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[78,117,1162,1182]" gridcol="0" gridrow="2" pageId="10" pageNumber="92">2</th>
|
||
<td box="[134,392,1162,1182]" gridcol="1" gridrow="2" pageId="10" pageNumber="92">
|
||
Dluq_
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[191,277,1162,1182]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Domitius</taxonomicName>
|
||
_luquei
|
||
</td>
|
||
<td box="[418,473,1162,1182]" gridcol="2" gridrow="2" pageId="10" pageNumber="92">0.142</td>
|
||
</tr>
|
||
<tr box="[78,1451,1199,1219]" gridrow="3" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[78,117,1199,1219]" gridcol="0" gridrow="3" pageId="10" pageNumber="92">3</th>
|
||
<td box="[134,392,1199,1219]" gridcol="1" gridrow="3" pageId="10" pageNumber="92">
|
||
Dlus_
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[190,275,1199,1219]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Domitius</taxonomicName>
|
||
_lusitanicus
|
||
</td>
|
||
<td box="[418,473,1199,1219]" gridcol="2" gridrow="3" pageId="10" pageNumber="92">0.135</td>
|
||
<td box="[488,543,1199,1219]" gridcol="3" gridrow="3" pageId="10" pageNumber="92">0.135</td>
|
||
</tr>
|
||
<tr box="[78,1451,1236,1256]" gridrow="4" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[78,117,1236,1256]" gridcol="0" gridrow="4" pageId="10" pageNumber="92">4</th>
|
||
<td box="[134,392,1236,1256]" gridcol="1" gridrow="4" pageId="10" pageNumber="92">
|
||
D213_
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[198,283,1236,1256]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Domitius</taxonomicName>
|
||
_menozzii
|
||
</td>
|
||
<td box="[418,473,1236,1256]" gridcol="2" gridrow="4" pageId="10" pageNumber="92">0.140</td>
|
||
<td box="[488,543,1236,1256]" gridcol="3" gridrow="4" pageId="10" pageNumber="92">0.146</td>
|
||
<td box="[558,613,1236,1256]" gridcol="4" gridrow="4" pageId="10" pageNumber="92">0.119</td>
|
||
</tr>
|
||
<tr box="[78,1451,1273,1317]" gridrow="5" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[78,117,1273,1317]" gridcol="0" gridrow="5" pageId="10" pageNumber="92">5</th>
|
||
<td box="[134,392,1273,1317]" gridcol="1" gridrow="5" pageId="10" pageNumber="92">
|
||
D555_
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[198,283,1273,1293]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Domitius</taxonomicName>
|
||
_culsu
|
||
<taxonomicNameLabel pageId="10" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</td>
|
||
<td box="[418,473,1273,1317]" gridcol="2" gridrow="5" pageId="10" pageNumber="92">0.167</td>
|
||
<td box="[488,543,1273,1317]" gridcol="3" gridrow="5" pageId="10" pageNumber="92">0.144</td>
|
||
<td box="[558,613,1273,1317]" gridcol="4" gridrow="5" pageId="10" pageNumber="92">0.133</td>
|
||
<td box="[628,684,1273,1317]" gridcol="5" gridrow="5" pageId="10" pageNumber="92">0.121</td>
|
||
</tr>
|
||
<tr box="[78,1451,1334,1354]" gridrow="6" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[78,117,1334,1354]" gridcol="0" gridrow="6" pageId="10" pageNumber="92">6</th>
|
||
<td box="[134,392,1334,1354]" gridcol="1" gridrow="6" pageId="10" pageNumber="92">
|
||
Dsbo_
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[197,282,1334,1354]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Domitius</taxonomicName>
|
||
_sbordonii
|
||
</td>
|
||
<td box="[418,473,1334,1354]" gridcol="2" gridrow="6" pageId="10" pageNumber="92">0.181</td>
|
||
<td box="[488,543,1334,1354]" gridcol="3" gridrow="6" pageId="10" pageNumber="92">0.176</td>
|
||
<td box="[558,613,1334,1354]" gridcol="4" gridrow="6" pageId="10" pageNumber="92">0.167</td>
|
||
<td box="[628,684,1334,1354]" gridcol="5" gridrow="6" pageId="10" pageNumber="92">0.181</td>
|
||
<td box="[698,752,1334,1354]" gridcol="6" gridrow="6" pageId="10" pageNumber="92">0.185</td>
|
||
</tr>
|
||
<tr box="[78,1451,1371,1415]" gridrow="7" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[78,117,1371,1415]" gridcol="0" gridrow="7" pageId="10" pageNumber="92">7</th>
|
||
<td box="[134,392,1371,1415]" gridcol="1" gridrow="7" pageId="10" pageNumber="92">
|
||
D557_
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[197,281,1371,1391]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Domitius</taxonomicName>
|
||
_ speluncarum
|
||
</td>
|
||
<td box="[418,473,1371,1415]" gridcol="2" gridrow="7" pageId="10" pageNumber="92">0.162</td>
|
||
<td box="[488,543,1371,1415]" gridcol="3" gridrow="7" pageId="10" pageNumber="92">0.144</td>
|
||
<td box="[558,613,1371,1415]" gridcol="4" gridrow="7" pageId="10" pageNumber="92">0.121</td>
|
||
<td box="[628,684,1371,1415]" gridcol="5" gridrow="7" pageId="10" pageNumber="92">0.121</td>
|
||
<td box="[698,752,1371,1415]" gridcol="6" gridrow="7" pageId="10" pageNumber="92">0.071</td>
|
||
<td box="[768,823,1371,1415]" gridcol="7" gridrow="7" pageId="10" pageNumber="92">0.190</td>
|
||
</tr>
|
||
<tr box="[78,1451,1432,1452]" gridrow="8" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-9="1">
|
||
<th box="[78,117,1432,1452]" gridcol="0" gridrow="8" pageId="10" pageNumber="92">8</th>
|
||
<td box="[134,392,1432,1452]" gridcol="1" gridrow="8" pageId="10" pageNumber="92">
|
||
N214_
|
||
<taxonomicName authorityName="Thorell" authorityYear="1869" box="[197,281,1432,1452]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Nesticus</taxonomicName>
|
||
_cellulanus
|
||
</td>
|
||
<td box="[418,473,1432,1452]" gridcol="2" gridrow="8" pageId="10" pageNumber="92">0.172</td>
|
||
<td box="[488,543,1432,1452]" gridcol="3" gridrow="8" pageId="10" pageNumber="92">0.174</td>
|
||
<td box="[558,613,1432,1452]" gridcol="4" gridrow="8" pageId="10" pageNumber="92">0.151</td>
|
||
<td box="[628,684,1432,1452]" gridcol="5" gridrow="8" pageId="10" pageNumber="92">0.190</td>
|
||
<td box="[698,752,1432,1452]" gridcol="6" gridrow="8" pageId="10" pageNumber="92">0.181</td>
|
||
<td box="[768,823,1432,1452]" gridcol="7" gridrow="8" pageId="10" pageNumber="92">0.176</td>
|
||
<td box="[838,893,1432,1452]" gridcol="8" gridrow="8" pageId="10" pageNumber="92">0.172</td>
|
||
</tr>
|
||
<tr box="[78,1451,1468,1512]" gridrow="9" pageId="10" pageNumber="92" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1">
|
||
<th box="[78,117,1468,1512]" gridcol="0" gridrow="9" pageId="10" pageNumber="92">9</th>
|
||
<td box="[134,392,1468,1512]" gridcol="1" gridrow="9" pageId="10" pageNumber="92">
|
||
K566_
|
||
<taxonomicName authorityName="Pavlek and Ribera" authorityYear="2017" box="[197,342,1468,1488]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Kryptonesticus</taxonomicName>
|
||
_ dimensis
|
||
</td>
|
||
<td box="[418,473,1468,1512]" gridcol="2" gridrow="9" pageId="10" pageNumber="92">0.156</td>
|
||
<td box="[488,543,1468,1512]" gridcol="3" gridrow="9" pageId="10" pageNumber="92">0.151</td>
|
||
<td box="[558,613,1468,1512]" gridcol="4" gridrow="9" pageId="10" pageNumber="92">0.135</td>
|
||
<td box="[628,684,1468,1512]" gridcol="5" gridrow="9" pageId="10" pageNumber="92">0.144</td>
|
||
<td box="[698,752,1468,1512]" gridcol="6" gridrow="9" pageId="10" pageNumber="92">0.165</td>
|
||
<td box="[768,823,1468,1512]" gridcol="7" gridrow="9" pageId="10" pageNumber="92">0.172</td>
|
||
<td box="[838,893,1468,1512]" gridcol="8" gridrow="9" pageId="10" pageNumber="92">0.146</td>
|
||
<td box="[908,963,1468,1512]" gridcol="9" gridrow="9" pageId="10" pageNumber="92">0.121</td>
|
||
</tr>
|
||
<tr box="[78,1451,1529,1573]" gridrow="10" pageId="10" pageNumber="92" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1">
|
||
<th box="[78,117,1529,1573]" gridcol="0" gridrow="10" pageId="10" pageNumber="92">10</th>
|
||
<td box="[134,392,1529,1573]" gridcol="1" gridrow="10" pageId="10" pageNumber="92">
|
||
K211_
|
||
<taxonomicName authorityName="Pavlek and Ribera" authorityYear="2017" box="[194,339,1529,1549]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Kryptonesticus</taxonomicName>
|
||
_ eremita
|
||
</td>
|
||
<td box="[418,473,1529,1573]" gridcol="2" gridrow="10" pageId="10" pageNumber="92">0.149</td>
|
||
<td box="[488,543,1529,1573]" gridcol="3" gridrow="10" pageId="10" pageNumber="92">0.156</td>
|
||
<td box="[558,613,1529,1573]" gridcol="4" gridrow="10" pageId="10" pageNumber="92">0.142</td>
|
||
<td box="[628,684,1529,1573]" gridcol="5" gridrow="10" pageId="10" pageNumber="92">0.156</td>
|
||
<td box="[698,752,1529,1573]" gridcol="6" gridrow="10" pageId="10" pageNumber="92">0.169</td>
|
||
<td box="[768,823,1529,1573]" gridcol="7" gridrow="10" pageId="10" pageNumber="92">0.169</td>
|
||
<td box="[838,893,1529,1573]" gridcol="8" gridrow="10" pageId="10" pageNumber="92">0.162</td>
|
||
<td box="[908,963,1529,1573]" gridcol="9" gridrow="10" pageId="10" pageNumber="92">0.117</td>
|
||
<td box="[978,1033,1529,1573]" gridcol="10" gridrow="10" pageId="10" pageNumber="92">0.078</td>
|
||
</tr>
|
||
<tr box="[78,1451,1590,1634]" gridrow="11" pageId="10" pageNumber="92" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1">
|
||
<th box="[78,117,1590,1634]" gridcol="0" gridrow="11" pageId="10" pageNumber="92">11</th>
|
||
<td box="[134,392,1590,1634]" gridcol="1" gridrow="11" pageId="10" pageNumber="92">
|
||
C162_
|
||
<taxonomicName box="[196,367,1590,1610]" class="Arachnida" family="Nesticidae" genus="Carpathonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Carpathonesticus</taxonomicName>
|
||
_ fodinarum
|
||
</td>
|
||
<td box="[418,473,1590,1634]" gridcol="2" gridrow="11" pageId="10" pageNumber="92">0.169</td>
|
||
<td box="[488,543,1590,1634]" gridcol="3" gridrow="11" pageId="10" pageNumber="92">0.153</td>
|
||
<td box="[558,613,1590,1634]" gridcol="4" gridrow="11" pageId="10" pageNumber="92">0.153</td>
|
||
<td box="[628,684,1590,1634]" gridcol="5" gridrow="11" pageId="10" pageNumber="92">0.176</td>
|
||
<td box="[698,752,1590,1634]" gridcol="6" gridrow="11" pageId="10" pageNumber="92">0.178</td>
|
||
<td box="[768,823,1590,1634]" gridcol="7" gridrow="11" pageId="10" pageNumber="92">0.172</td>
|
||
<td box="[838,893,1590,1634]" gridcol="8" gridrow="11" pageId="10" pageNumber="92">0.167</td>
|
||
<td box="[908,963,1590,1634]" gridcol="9" gridrow="11" pageId="10" pageNumber="92">0.085</td>
|
||
<td box="[978,1033,1590,1634]" gridcol="10" gridrow="11" pageId="10" pageNumber="92">0.089</td>
|
||
<td box="[1048,1103,1590,1634]" gridcol="11" gridrow="11" pageId="10" pageNumber="92">0.108</td>
|
||
</tr>
|
||
<tr box="[78,1451,1651,1695]" gridrow="12" pageId="10" pageNumber="92" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1">
|
||
<th box="[78,117,1651,1695]" gridcol="0" gridrow="12" pageId="10" pageNumber="92">12</th>
|
||
<td box="[134,392,1651,1695]" gridcol="1" gridrow="12" pageId="10" pageNumber="92">C166_Carpathonesitcus_ lotriensis</td>
|
||
<td box="[418,473,1651,1695]" gridcol="2" gridrow="12" pageId="10" pageNumber="92">0.144</td>
|
||
<td box="[488,543,1651,1695]" gridcol="3" gridrow="12" pageId="10" pageNumber="92">0.160</td>
|
||
<td box="[558,613,1651,1695]" gridcol="4" gridrow="12" pageId="10" pageNumber="92">0.146</td>
|
||
<td box="[628,684,1651,1695]" gridcol="5" gridrow="12" pageId="10" pageNumber="92">0.162</td>
|
||
<td box="[698,752,1651,1695]" gridcol="6" gridrow="12" pageId="10" pageNumber="92">0.190</td>
|
||
<td box="[768,823,1651,1695]" gridcol="7" gridrow="12" pageId="10" pageNumber="92">0.176</td>
|
||
<td box="[838,893,1651,1695]" gridcol="8" gridrow="12" pageId="10" pageNumber="92">0.190</td>
|
||
<td box="[908,963,1651,1695]" gridcol="9" gridrow="12" pageId="10" pageNumber="92">0.112</td>
|
||
<td box="[978,1033,1651,1695]" gridcol="10" gridrow="12" pageId="10" pageNumber="92">0.094</td>
|
||
<td box="[1048,1103,1651,1695]" gridcol="11" gridrow="12" pageId="10" pageNumber="92">0.101</td>
|
||
<td box="[1118,1173,1651,1695]" gridcol="12" gridrow="12" pageId="10" pageNumber="92">0.069</td>
|
||
</tr>
|
||
<tr box="[78,1451,1712,1756]" gridrow="13" pageId="10" pageNumber="92" rowspan-14="1" rowspan-15="1" rowspan-16="1">
|
||
<th box="[78,117,1712,1756]" gridcol="0" gridrow="13" pageId="10" pageNumber="92">13</th>
|
||
<td box="[134,392,1712,1756]" gridcol="1" gridrow="13" pageId="10" pageNumber="92">
|
||
Tobc_
|
||
<taxonomicName authorityName="Kulczynski" authorityYear="1914" box="[194,336,1712,1732]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Typhlonesticus</taxonomicName>
|
||
_ obcaecatus
|
||
</td>
|
||
<td box="[418,473,1712,1756]" gridcol="2" gridrow="13" pageId="10" pageNumber="92">0.197</td>
|
||
<td box="[488,543,1712,1756]" gridcol="3" gridrow="13" pageId="10" pageNumber="92">0.178</td>
|
||
<td box="[558,613,1712,1756]" gridcol="4" gridrow="13" pageId="10" pageNumber="92">0.167</td>
|
||
<td box="[628,684,1712,1756]" gridcol="5" gridrow="13" pageId="10" pageNumber="92">0.165</td>
|
||
<td box="[698,752,1712,1756]" gridcol="6" gridrow="13" pageId="10" pageNumber="92">0.178</td>
|
||
<td box="[768,823,1712,1756]" gridcol="7" gridrow="13" pageId="10" pageNumber="92">0.176</td>
|
||
<td box="[838,893,1712,1756]" gridcol="8" gridrow="13" pageId="10" pageNumber="92">0.181</td>
|
||
<td box="[908,963,1712,1756]" gridcol="9" gridrow="13" pageId="10" pageNumber="92">0.149</td>
|
||
<td box="[978,1033,1712,1756]" gridcol="10" gridrow="13" pageId="10" pageNumber="92">0.156</td>
|
||
<td box="[1048,1103,1712,1756]" gridcol="11" gridrow="13" pageId="10" pageNumber="92">0.158</td>
|
||
<td box="[1118,1173,1712,1756]" gridcol="12" gridrow="13" pageId="10" pageNumber="92">0.140</td>
|
||
<td box="[1188,1243,1712,1756]" gridcol="13" gridrow="13" pageId="10" pageNumber="92">0.142</td>
|
||
</tr>
|
||
<tr box="[78,1451,1773,1817]" gridrow="14" pageId="10" pageNumber="92" rowspan-15="1" rowspan-16="1">
|
||
<th box="[78,117,1773,1817]" gridcol="0" gridrow="14" pageId="10" pageNumber="92">14</th>
|
||
<td box="[134,392,1773,1817]" gridcol="1" gridrow="14" pageId="10" pageNumber="92">
|
||
T167_
|
||
<taxonomicName authorityName="Kulczynski" authorityYear="1914" box="[193,335,1773,1793]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Typhlonesticus</taxonomicName>
|
||
_ idriacus
|
||
</td>
|
||
<td box="[418,473,1773,1817]" gridcol="2" gridrow="14" pageId="10" pageNumber="92">0.181</td>
|
||
<td box="[488,543,1773,1817]" gridcol="3" gridrow="14" pageId="10" pageNumber="92">0.174</td>
|
||
<td box="[558,613,1773,1817]" gridcol="4" gridrow="14" pageId="10" pageNumber="92">0.167</td>
|
||
<td box="[628,684,1773,1817]" gridcol="5" gridrow="14" pageId="10" pageNumber="92">0.183</td>
|
||
<td box="[698,752,1773,1817]" gridcol="6" gridrow="14" pageId="10" pageNumber="92">0.192</td>
|
||
<td box="[768,823,1773,1817]" gridcol="7" gridrow="14" pageId="10" pageNumber="92">0.197</td>
|
||
<td box="[838,893,1773,1817]" gridcol="8" gridrow="14" pageId="10" pageNumber="92">0.185</td>
|
||
<td box="[908,963,1773,1817]" gridcol="9" gridrow="14" pageId="10" pageNumber="92">0.142</td>
|
||
<td box="[978,1033,1773,1817]" gridcol="10" gridrow="14" pageId="10" pageNumber="92">0.156</td>
|
||
<td box="[1048,1103,1773,1817]" gridcol="11" gridrow="14" pageId="10" pageNumber="92">0.165</td>
|
||
<td box="[1118,1173,1773,1817]" gridcol="12" gridrow="14" pageId="10" pageNumber="92">0.149</td>
|
||
<td box="[1188,1243,1773,1817]" gridcol="13" gridrow="14" pageId="10" pageNumber="92">0.156</td>
|
||
<td box="[1258,1313,1773,1817]" gridcol="14" gridrow="14" pageId="10" pageNumber="92">0.096</td>
|
||
</tr>
|
||
<tr box="[78,1451,1834,1878]" gridrow="15" pageId="10" pageNumber="92" rowspan-16="1">
|
||
<th box="[78,117,1834,1878]" gridcol="0" gridrow="15" pageId="10" pageNumber="92">15</th>
|
||
<td box="[134,392,1834,1878]" gridcol="1" gridrow="15" pageId="10" pageNumber="92">
|
||
Tmor_
|
||
<taxonomicName authorityName="Kulczynski" authorityYear="1914" box="[196,337,1834,1854]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Typhlonesticus</taxonomicName>
|
||
_ morisii
|
||
</td>
|
||
<td box="[418,473,1834,1878]" gridcol="2" gridrow="15" pageId="10" pageNumber="92">0.174</td>
|
||
<td box="[488,543,1834,1878]" gridcol="3" gridrow="15" pageId="10" pageNumber="92">0.174</td>
|
||
<td box="[558,613,1834,1878]" gridcol="4" gridrow="15" pageId="10" pageNumber="92">0.174</td>
|
||
<td box="[628,684,1834,1878]" gridcol="5" gridrow="15" pageId="10" pageNumber="92">0.181</td>
|
||
<td box="[698,752,1834,1878]" gridcol="6" gridrow="15" pageId="10" pageNumber="92">0.199</td>
|
||
<td box="[768,823,1834,1878]" gridcol="7" gridrow="15" pageId="10" pageNumber="92">0.208</td>
|
||
<td box="[838,893,1834,1878]" gridcol="8" gridrow="15" pageId="10" pageNumber="92">0.181</td>
|
||
<td box="[908,963,1834,1878]" gridcol="9" gridrow="15" pageId="10" pageNumber="92">0.183</td>
|
||
<td box="[978,1033,1834,1878]" gridcol="10" gridrow="15" pageId="10" pageNumber="92">0.160</td>
|
||
<td box="[1048,1103,1834,1878]" gridcol="11" gridrow="15" pageId="10" pageNumber="92">0.183</td>
|
||
<td box="[1118,1173,1834,1878]" gridcol="12" gridrow="15" pageId="10" pageNumber="92">0.176</td>
|
||
<td box="[1188,1243,1834,1878]" gridcol="13" gridrow="15" pageId="10" pageNumber="92">0.174</td>
|
||
<td box="[1258,1313,1834,1878]" gridcol="14" gridrow="15" pageId="10" pageNumber="92">0.117</td>
|
||
<td box="[1327,1382,1834,1878]" gridcol="15" gridrow="15" pageId="10" pageNumber="92">0.124</td>
|
||
</tr>
|
||
<tr box="[78,1451,1894,1939]" gridrow="16" pageId="10" pageNumber="92">
|
||
<th box="[78,117,1894,1939]" gridcol="0" gridrow="16" pageId="10" pageNumber="92">16</th>
|
||
<td box="[134,392,1894,1939]" gridcol="1" gridrow="16" pageId="10" pageNumber="92">
|
||
G601_
|
||
<taxonomicName authorityName="Keyserling" authorityYear="1884" box="[196,308,1894,1914]" class="Arachnida" family="Nesticidae" genus="Gaucelmus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="92" phylum="Arthropoda" rank="genus">Gaucelmus</taxonomicName>
|
||
_ augustinus
|
||
</td>
|
||
<td box="[418,473,1894,1939]" gridcol="2" gridrow="16" pageId="10" pageNumber="92">0.229</td>
|
||
<td box="[488,543,1894,1939]" gridcol="3" gridrow="16" pageId="10" pageNumber="92">0.229</td>
|
||
<td box="[558,613,1894,1939]" gridcol="4" gridrow="16" pageId="10" pageNumber="92">0.227</td>
|
||
<td box="[628,684,1894,1939]" gridcol="5" gridrow="16" pageId="10" pageNumber="92">0.222</td>
|
||
<td box="[698,752,1894,1939]" gridcol="6" gridrow="16" pageId="10" pageNumber="92">0.245</td>
|
||
<td box="[768,823,1894,1939]" gridcol="7" gridrow="16" pageId="10" pageNumber="92">0.243</td>
|
||
<td box="[838,893,1894,1939]" gridcol="8" gridrow="16" pageId="10" pageNumber="92">0.236</td>
|
||
<td box="[908,963,1894,1939]" gridcol="9" gridrow="16" pageId="10" pageNumber="92">0.252</td>
|
||
<td box="[978,1033,1894,1939]" gridcol="10" gridrow="16" pageId="10" pageNumber="92">0.211</td>
|
||
<td box="[1048,1103,1894,1939]" gridcol="11" gridrow="16" pageId="10" pageNumber="92">0.238</td>
|
||
<td box="[1118,1173,1894,1939]" gridcol="12" gridrow="16" pageId="10" pageNumber="92">0.240</td>
|
||
<td box="[1188,1243,1894,1939]" gridcol="13" gridrow="16" pageId="10" pageNumber="92">0.233</td>
|
||
<td box="[1258,1313,1894,1939]" gridcol="14" gridrow="16" pageId="10" pageNumber="92">0.249</td>
|
||
<td box="[1327,1382,1894,1939]" gridcol="15" gridrow="16" pageId="10" pageNumber="92">0.254</td>
|
||
<td box="[1398,1451,1894,1939]" gridcol="16" gridrow="16" pageId="10" pageNumber="92">0.247</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="11.[144,1537,224,746]" box="[144,335,224,255]" pageId="11" pageNumber="93">
|
||
<heading bold="true" box="[144,335,224,255]" fontSize="12" level="2" pageId="11" pageNumber="93" reason="0">
|
||
<emphasis bold="true" box="[144,335,224,255]" pageId="11" pageNumber="93">Conclusions</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="11.[144,1537,224,746]" pageId="11" pageNumber="93">
|
||
All Italian species of the genus
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[541,645,270,295]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[541,645,270,295]" italics="true" pageId="11" pageNumber="93">Domitius</emphasis>
|
||
</taxonomicName>
|
||
appear to be highly adapted to a permanent life in the subterranean environ- ment, showing eye reduction and lack of body pigmentation. They further present a localized distribution, with distinct genetic and morphological differences between the species living in the Northern and Central areas of the Apennines. At the same time a close affinity with the species distributed in the Iberian Peninsula is observed. Such features, to- gether with a high genetic p-distance among the species (
|
||
<tableCitation box="[824,917,398,423]" captionStart="Table 2" captionStartId="10.[77,136,1023,1046]" captionTargetPageId="10" captionText="Table 2. Uncorrected genetic p-distance of the COI partial sequence of the nesticid species discussed in the text. The newly described species is in bold." httpUri="http://table.plazi.org/id/DF6A418C1E68FFC7FFF067D2C6D3FBF6" pageId="11" pageNumber="93" tableUuid="DF6A418C1E68FFC7FFF067D2C6D3FBF6">Table 2</tableCitation>
|
||
), suggests a potentially complex evolutionary history of the genus
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[299,403,430,455]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[299,403,430,455]" italics="true" pageId="11" pageNumber="93">Domitius</emphasis>
|
||
</taxonomicName>
|
||
that still needs to be properly explored (see also
|
||
<bibRefCitation author="Ribera, C." box="[985,1142,430,455]" pageId="11" pageNumber="93" pagination="229 - 240" refId="ref7897" refString="Ribera, C., 2018, A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa, v. 4407, p. 229 - 240, https: // doi. org / 10.11646 / zootaxa. 4407.2.4." title="A new genus of nesticid spiders from western European Peninsulas (Araneae, Nesticidae): Zootaxa" type="book chapter" year="2018">Ribera, 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="11.[144,1537,224,746]" pageId="11" pageNumber="93">
|
||
Because of their apparent similarities in habitus and female genitalia,
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[989,1084,464,489]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov">
|
||
<emphasis box="[989,1084,464,489]" italics="true" pageId="11" pageNumber="93">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1090,1182,464,489]" pageId="11" pageNumber="93">
|
||
<taxonomicNameLabel box="[1090,1182,464,489]" pageId="11" pageNumber="93" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
has previously been mistaken for
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1979" box="[184,374,496,521]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="species" species="speluncarum">
|
||
<emphasis box="[184,374,496,521]" italics="true" pageId="11" pageNumber="93">D. speluncarum</emphasis>
|
||
</taxonomicName>
|
||
and ignored as a distinct species by previous arachnologists. A detailed molecular and morpho- logical analysis of both sexes of
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[529,626,528,553]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov">
|
||
<emphasis box="[529,626,528,553]" italics="true" pageId="11" pageNumber="93">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[634,727,528,553]" pageId="11" pageNumber="93">
|
||
<taxonomicNameLabel box="[634,727,528,553]" pageId="11" pageNumber="93" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
carried out in this work supports the validity of the new species and its close relationship with the other
|
||
<taxonomicName authorityName="Ribera" authorityYear="2018" box="[572,676,560,585]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[572,676,560,585]" italics="true" pageId="11" pageNumber="93">Domitius</emphasis>
|
||
</taxonomicName>
|
||
species from the same geographical area. Its potential susceptibility to external disturbance, and extremely limited distribution, makes
|
||
<taxonomicName authority="Ballarin, 2020" authorityName="Ballarin" authorityYear="2020" box="[891,987,592,617]" class="Arachnida" family="Nesticidae" genus="Domitius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="species" species="culsu" status="sp. nov">
|
||
<emphasis box="[891,987,592,617]" italics="true" pageId="11" pageNumber="93">D. culsu</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[995,1088,592,617]" pageId="11" pageNumber="93">
|
||
<taxonomicNameLabel box="[995,1088,592,617]" pageId="11" pageNumber="93" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
of interest for conservation.
|
||
</paragraph>
|
||
<paragraph blockId="11.[144,1537,224,746]" pageId="11" pageNumber="93">
|
||
Finding a new nesticid species in Italy further suggests that our knowledge on the diversity of the family
|
||
<taxonomicName authorityName="Simon" authorityYear="1894" box="[1410,1536,625,650]" class="Arachnida" family="Nesticidae" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="93" phylum="Arthropoda" rank="family">Nesticidae</taxonomicName>
|
||
in Southern Europe is still far from complete. Further collections along the Italian peninsula will probably lead to the discovery of other highly-specialized nesticid species allowing a deeper and more precise understanding of the spider cave fauna in Italy and in the Mediterranean area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |