treatments-xml/data/03/BB/96/03BB9669FFAFFFF7D1E2F954E0974989.xml
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<document id="F8303A63C88A8B356223A63B7F097375" ID-DOI="10.11646/zootaxa.5406.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10618214" ID-ZooBank="F7670D48-6ED9-436D-97A6-0841E03A203E" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707132336199" checkinUser="plazi" docAuthor="Salnitska, Maria, Sheykin, Sergey &amp; Semerikova, Daria" docDate="2024" docId="03BB9669FFAFFFF7D1E2F954E0974989" docLanguage="en" docName="zootaxa.5406.2.4.pdf" docOrigin="Zootaxa 5406 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5406.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euphanias sergeevae Salnitska 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="292" masterDocId="FF82EE11FFADFFF2D175FFBEE2724970" masterDocTitle="Description of a new species Euphanias sergreevae sp. nov. (Coleoptera: Staphylinidae: Oxytelinae) from Western Siberia, Russia" masterLastPageNumber="296" masterPageNumber="288" pageNumber="290" updateTime="1707333198505" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="FCFF790571B09DC4F8B5D85D978B1DB0">Description of a new species Euphanias sergreevae sp. nov. (Coleoptera: Staphylinidae: Oxytelinae) from Western Siberia, Russia</mods:title>
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<mods:namePart id="40DB0FAC0C3D70F92BBE6850B26DC38C">Salnitska, Maria</mods:namePart>
<mods:affiliation id="755EDBAEFE8675C38ACF445A2AA0657E">X-BIO Institute, University of Tyumen, 25 Lenina Str, Tyumen, 625003, Russia</mods:affiliation>
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<mods:namePart id="5F422A628C0BF8940BCB56B32CE795E8">Sheykin, Sergey</mods:namePart>
<mods:affiliation id="6692DCB337C651D4E6C3F399FCC22390">X-BIO Institute, University of Tyumen, 25 Lenina Str, Tyumen, 625003, Russia</mods:affiliation>
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<mods:namePart id="3B1600247E448CBF19BD457308598B8D">Semerikova, Daria</mods:namePart>
<mods:affiliation id="24CEEA26973FFD1F69FAD68F2F79420D">X-BIO Institute, University of Tyumen, 25 Lenina Str, Tyumen, 625003, Russia</mods:affiliation>
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<mods:date id="5AAB13DC6DF8DBFF385CA3B0E01983D0">2024</mods:date>
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<treatment id="03BB9669FFAFFFF7D1E2F954E0974989" ID-DOI="http://doi.org/10.5281/zenodo.10618216" ID-Zenodo-Dep="10618216" LSID="urn:lsid:plazi:treatment:03BB9669FFAFFFF7D1E2F954E0974989" httpUri="http://treatment.plazi.org/id/03BB9669FFAFFFF7D1E2F954E0974989" lastPageId="5" lastPageNumber="292" pageId="2" pageNumber="290">
<subSubSection id="C30874F4FFAFFFF0D1E2F954E0BB4E75" box="[151,713,1770,1797]" pageId="2" pageNumber="290" type="nomenclature">
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<taxonomicName id="4C125CFCFFAFFFF0D1E2F954E02C4E74" authority="Salnitska, 2023" authorityName="Salnitska" authorityYear="2023" box="[151,606,1770,1797]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="290" phylum="Arthropoda" rank="species" species="sergeevae" status="sp. nov.">
<emphasis id="B966FB6DFFAFFFF0D1E2F954E3EA4E75" bold="true" box="[151,408,1770,1797]" italics="true" pageId="2" pageNumber="290">Euphanias sergeevae</emphasis>
Salnitska, 2023
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,
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(
<figureCitation id="13293BFAFFAFFFF0D1EAF8B1E2944E59" box="[159,230,1807,1833]" captionStart="FIGURE 1" captionStartId="1.[151,250,967,991]" captionTargetBox="[151,1436,181,943]" captionTargetId="figure-34@1.[151,1436,181,943]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Euphanias species." figureDoi="http://doi.org/10.5281/zenodo.10618218" httpUri="https://zenodo.org/record/10618218/files/figure.png" pageId="2" pageNumber="290">Figs 1</figureCitation>
,
<figureCitation id="13293BFAFFAFFFF0D181F8B1E36A4E59" box="[244,280,1807,1833]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="2" pageNumber="290">3A</figureCitation>
,
<figureCitation id="13293BFAFFAFFFF0D051F8B1E3464E59" box="[292,308,1807,1833]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="2" pageNumber="290">4</figureCitation>
,
<figureCitation id="13293BFAFFAFFFF0D035F8B0E33C4E59" box="[320,334,1806,1833]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="2" pageNumber="290">5</figureCitation>
)
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<emphasis id="B966FB6DFFAFFFF0D1E2F8E9E33C4E00" bold="true" box="[151,334,1878,1905]" pageId="2" pageNumber="290">Type material.</emphasis>
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: “
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,
<collectingRegion id="49D6E99DFFAFFFF0D31CF8E9E1604E01" box="[617,786,1879,1905]" country="Russia" name="Tyumen'" pageId="2" pageNumber="290">Tyumen Prov.</collectingRegion>
,
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vill.,
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,
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,
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,
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<collectingDate id="EFE8F857FFAFFFF0D1E2F8C5E36B4EE5" box="[151,281,1915,1941]" pageId="2" pageNumber="290" value="2023-06-21">21.VI.2023</collectingDate>
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, leaf litter along flooded shore of lake / mostly with
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<emphasis id="B966FB6DFFAFFFF0D21AF8C5E1964EE5" box="[879,996,1915,1941]" italics="true" pageId="2" pageNumber="290">Salicornia</emphasis>
sp.
</taxonomicName>
, leg.
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. Krivosheeva,
<collectorName id="26E742A9FFAFFFF0D472F8C5E7EB4EE5" box="[1287,1433,1915,1941]" pageId="2" pageNumber="290">M. Salnitska</collectorName>
,
<collectorName id="26E742A9FFAFFFF0D1E2F821E3254EC9" box="[151,343,1951,1977]" pageId="2" pageNumber="290">E. Kargapoltseva</collectorName>
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a/
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_
<collectionCode id="ED03BFBAFFAFFFF0D3D4F821E0A94EC9" box="[673,731,1951,1977]" pageId="2" pageNumber="290">LISP</collectionCode>
_000248” (male,
<collectionCode id="ED03BFBAFFAFFFF0D2E9F821E1864EC9" box="[924,1012,1951,1977]" pageId="2" pageNumber="290">UTMN</collectionCode>
).
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<typeStatus id="54A999DDFFAFFFF0D573F821E6F04EC9" box="[1030,1154,1951,1977]" pageId="2" pageNumber="290" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same data but “
<collectionCode id="ED03BFBAFFAFFFF0D44CF821E7FC4EC9" box="[1337,1422,1951,1977]" pageId="2" pageNumber="290">UTMN</collectionCode>
_
<collectionCode id="ED03BFBAFFAFFFF0D1E2F87DE2A34EAD" box="[151,209,1987,2013]" pageId="2" pageNumber="290">LISP</collectionCode>
_000247” [specimen covered by the thin layer of gold after SEM] (
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,
<collectionCode id="ED03BFBAFFAFFFF0D562F87DE61D4EAD" box="[1047,1135,1987,2013]" pageId="2" pageNumber="290">UTMN</collectionCode>
);
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<collectionCode id="ED03BFBAFFAFFFF0D5FFF87DE6AC4EAD" box="[1162,1246,1987,2013]" pageId="2" pageNumber="290">UTMN</collectionCode>
_
<collectionCode id="ED03BFBAFFAFFFF0D598F87DE7554EAD" box="[1261,1319,1987,2013]" pageId="2" pageNumber="290">LISP</collectionCode>
_ 000249” (
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,
<collectionCode id="ED03BFBAFFAEFFF1D057FF29E30F49C1" box="[290,381,151,177]" pageId="3" pageNumber="291">NHMD</collectionCode>
);
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<materialsCitation id="3B7A2D22FFAEFFF1D0EDFF29E0F14989" collectingDate="2021-08-10" collectingDateMax="2023-06-21" collectingDateMin="2021-08-10" collectionCode="V, UTMN, LISP, NHMD" collectorName="E.V. Sergeeva" country="Russia" latitude="55.34" location="Tavolzhan Lake" longLatPrecision="63" longitude="70.151" municipality="Tavolzhan" pageId="3" pageNumber="291" specimenCode="WS23-08" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Tyumen'" typeStatus="holotype">
<collectingRegion id="49D6E99DFFAEFFF1D0D7FF29E03F49C1" box="[418,589,151,177]" country="Russia" name="Tyumen'" pageId="3" pageNumber="291">Tyumen Prov.</collectingRegion>
, Sladkovskiy Distr., Alexandrovka vill.,
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,
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,
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<collectingDate id="EFE8F857FFAEFFF1D407FF29E35949A5" pageId="3" pageNumber="291" value="2021-08-10" valueMax="2021-08-11" valueMin="2021-08-10">10 11.VIII.2021</collectingDate>
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, E.
<collectionCode id="ED03BFBAFFAEFFF1D03BFF05E31649A5" box="[334,356,187,213]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="291" type="Museum">V</collectionCode>
.
<collectorName id="26E742A9FFAEFFF1D01AFF05E3A749A5" box="[367,469,187,213]" pageId="3" pageNumber="291">Sergeeva</collectorName>
leg. [in Russian] / territory of hunting farm “Tavolzhan”, salt meadows [in Russian]/
<collectionCode id="ED03BFBAFFAEFFF1D1E2FF60E2994989" box="[151,235,222,249]" pageId="3" pageNumber="291">UTMN</collectionCode>
_
<collectionCode id="ED03BFBAFFAEFFF1D18CFF60E3404989" box="[249,306,222,249]" pageId="3" pageNumber="291">LISP</collectionCode>
_000250” (
<specimenCount id="9D14ECF6FFAEFFF1D0C7FF61E0664989" box="[434,532,223,249]" count="1" pageId="3" pageNumber="291" type="female">1 female</specimenCount>
,
<collectionCode id="ED03BFBAFFAEFFF1D36AFF61E0054989" box="[543,631,223,249]" pageId="3" pageNumber="291">UTMN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30874F4FFAEFFF6D1B2FEBCE35A4C09" lastPageId="4" lastPageNumber="292" pageId="3" pageNumber="291" type="description">
<paragraph id="8BAD277FFFAEFFF1D1B2FEBCE29048F9" blockId="3.[151,1437,150,393]" pageId="3" pageNumber="291">
<emphasis id="B966FB6DFFAEFFF1D1B2FEBCE0E2486C" bold="true" box="[199,656,258,285]" pageId="3" pageNumber="291">Description. Measurements and ratios</emphasis>
(n = 4): HL: 0.400.43 (0.41); HW: 0.460.51 (0.47); PL: 0.460.51 (0.48); PW: 0.600.69 (0.63); EL: 0.400.46 (0.44); EW: 0.860.91 (0.89); FB: 1.491.66 (1.55); HL/HW: 0.83 0.94 (0.88); PL/PW: 0.730.81 (0.76); EL/EW: 1.401.57 (1.51); PL/EL: 0.670.81 (0.73); PW/EW: 0.660.73 (0.71).
</paragraph>
<caption id="DF6D77F7FFAEFFF1D1E2FB67E3BF4C64" ID-DOI="http://doi.org/10.5281/zenodo.10618224" ID-Zenodo-Dep="10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="3" pageNumber="291" startId="3.[151,250,1241,1265]" targetBox="[177,1436,417,1217]" targetPageId="3" targetType="figure">
<paragraph id="8BAD277FFFAEFFF1D1E2FB67E3BF4C64" blockId="3.[151,1436,1241,1301]" pageId="3" pageNumber="291">
<emphasis id="B966FB6DFFAEFFF1D1E2FB67E3674D81" bold="true" box="[151,277,1241,1265]" pageId="3" pageNumber="291">FIGURE 3.</emphasis>
Habitus of
<taxonomicName id="4C125CFCFFAEFFF1D0E4FB67E0004D81" box="[401,626,1241,1265]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="291" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B966FB6DFFAEFFF1D0E4FB67E38F4D81" box="[401,509,1241,1265]" italics="true" pageId="3" pageNumber="291">Euphanias</emphasis>
species. A
</taxonomicName>
:
<taxonomicName id="4C125CFCFFAEFFF1D308FB64E08D4D81" authority="Salnitska &amp; Sheykin &amp; Semerikova, 2024" authorityName="Salnitska &amp; Sheykin &amp; Semerikova" authorityYear="2024" box="[637,767,1242,1265]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="291" phylum="Arthropoda" rank="species" species="sergeevae" status="sp. nov.">
<emphasis id="B966FB6DFFAEFFF1D308FB64E08D4D81" box="[637,767,1242,1265]" italics="true" pageId="3" pageNumber="291">E. sergeevae</emphasis>
</taxonomicName>
<emphasis id="B966FB6DFFAEFFF1D272FB67E1284D81" bold="true" box="[775,858,1241,1265]" pageId="3" pageNumber="291">
<taxonomicNameLabel id="A2554616FFAEFFF1D272FB67E1284D81" box="[775,858,1241,1265]" pageId="3" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); B:
<taxonomicName id="4C125CFCFFAEFFF1D28EFB64E6064D81" authorityName="Blinstein" authorityYear="1976" box="[1019,1140,1242,1265]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="291" phylum="Arthropoda" rank="species" species="pusanovi">
<emphasis id="B966FB6DFFAEFFF1D28EFB64E6064D81" box="[1019,1140,1242,1265]" italics="true" pageId="3" pageNumber="291">E. pusanovi</emphasis>
</taxonomicName>
(?holotype); C:
<taxonomicName id="4C125CFCFFAEFFF1D455FB64E7EE4D81" authorityName="Bernhauer" authorityYear="1912" box="[1312,1436,1241,1265]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="291" phylum="Arthropoda" rank="species" species="pliginskii">
<emphasis id="B966FB6DFFAEFFF1D455FB64E7EE4D81" box="[1312,1436,1241,1265]" italics="true" pageId="3" pageNumber="291">E. pliginskii</emphasis>
</taxonomicName>
(holotype). Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption id="DF6D77F7FFAEFFF1D1E2F8ACE6E84E3F" ID-DOI="http://doi.org/10.5281/zenodo.10618226" ID-Zenodo-Dep="10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="3" pageNumber="291" startId="3.[151,250,1810,1834]" targetBox="[170,1420,1335,1780]" targetPageId="3" targetType="figure">
<paragraph id="8BAD277FFFAEFFF1D1E2F8ACE6E84E3F" blockId="3.[151,1436,1810,1871]" pageId="3" pageNumber="291">
<emphasis id="B966FB6DFFAEFFF1D1E2F8ACE3604E5A" bold="true" box="[151,274,1810,1834]" pageId="3" pageNumber="291">FIGURE 4.</emphasis>
Scanning electron micrograph of
<taxonomicName id="4C125CFCFFAEFFF1D31CF8ADE09A4E5A" authorityName="Salnitska &amp; Sheykin &amp; Semerikova" authorityYear="2024" box="[617,744,1811,1834]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="291" phylum="Arthropoda" rank="species" species="sergeevae">
<emphasis id="B966FB6DFFAEFFF1D31CF8ADE09A4E5A" box="[617,744,1811,1834]" italics="true" pageId="3" pageNumber="291">E. sergeevae</emphasis>
</taxonomicName>
s
<emphasis id="B966FB6DFFAEFFF1D38DF8ACE14C4E5A" bold="true" box="[760,830,1810,1834]" pageId="3" pageNumber="291">p. nov.</emphasis>
A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm.
</paragraph>
</caption>
<paragraph id="8BAD277FFFAEFFF1D1B2F82DE08C4E85" blockId="3.[151,1436,1939,2038]" pageId="3" pageNumber="291">
Body black, appendages vaguely paler, legs vaguely brownish, mandibles dark brown with apical portions yellowish, maxillary palps dark brown, labial palps yellowish (
<figureCitation id="13293BFAFFAEFFF1D209F809E1A94EA1" box="[892,987,1975,2001]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="3" pageNumber="291">Fig. 3A</figureCitation>
). Body surfaces deeply punctured and covered with numerous plumose whitish setae (
<figureCitation id="13293BFAFFAEFFF1D3D9F865E0834E85" box="[684,753,2011,2037]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="3" pageNumber="291">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BAD277FFFA9FFF6D1B2FF29E15948A1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
Head transverse (HL/HW: 0.88), widest at eyes, slightly shorter than pronotum; epistomal suture barely visible, disk with strongly developed carina running across midline from strongly pronounced supraantennal prominences to the level of posterior margin of eye, deep invagination present between carina and eye; surface deeply punctured and covered by scattered plumose setae (
<figureCitation id="13293BFAFFA9FFF6D32DFEBDE0C6486D" box="[600,692,259,285]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D3CAFEBDE174486D" box="[703,774,259,285]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4A, D</figureCitation>
). Eyes vaguely convex, temples as long as or slightly longer than eyes (temples/eyes: 1.04), with small setae between facets (
<figureCitation id="13293BFAFFA9FFF6D200FE99E1A34831" box="[885,977,295,321]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">Fig. 4D</figureCitation>
). Antennae moderately long, three times longer than head; segments 18 distinctly longer than wide, 35 gradually increasing in length, 811 as long as or wider than long forming a club; first segment covered by scattered short and flattened whitish setae, 25 covered by rare short hair-like setae, 611 by short and long hair like setae (
<figureCitation id="13293BFAFFA9FFF6D2FCFE2DE19448DD" box="[905,998,403,429]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D287FE2DE64B48DD" box="[1010,1081,403,429]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4AC</figureCitation>
). Labrum vaguely bilobed (
<figureCitation id="13293BFAFFA9FFF6D41AFE2DE2CC48A1" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">Fig. 4D</figureCitation>
). Mouth appendages bearing numerous setae (
<figureCitation id="13293BFAFFA9FFF6D3B1FE09E16D48A1" box="[708,799,439,465]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8BAD277FFFA9FFF6D1B2FE65E0BC4BD9" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
Pronotum transverse, distinctly wider than long (PL/PW: 0.76), widest behind midline at pronounced lateral portions, shorter (PL/EL: 0.73) and narrower (PW/EW: 0.71) than elytra; of hexagonal shape (
<figureCitation id="13293BFAFFA9FFF6D5BBFE41E75A4B69" box="[1230,1320,511,537]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Fig. 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D443FE41E72B4B69" box="[1334,1369,511,537]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4A</figureCitation>
) with smoothly rounded angles; apical margin forming camber line, posterior portion strongly narrowing basad, lateral portions explanate, middle portion with carina of specific shape bearing barely visible middle depression, with deep invaginations between carina and lateral margins extending from apical to posterior margin (
<figureCitation id="13293BFAFFA9FFF6D594FDD5E74C4BF5" box="[1249,1342,619,645]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D43FFDD5E7E24BF5" box="[1354,1424,619,645]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4A, E</figureCitation>
). Punctation and setation same as on head (
<figureCitation id="13293BFAFFA9FFF6D31FFD31E0B34BD9" box="[618,705,655,681]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="8BAD277FFFA9FFF6D1B2FD0DE3604AF1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
Single elytron longer than wide (EL/EW: 1.51), combined elytra longer (PL/EL: 0.73) and wider (PW/EW: 0.71).) than pronotum, slightly expanding posteriad; anterior angles obtuse, posterior angles with deep and round excision; bearing three carina, 1st and 3rd not reaching posterior margin and gradually ending in second half of elytra, 2
<superScript id="7C678A37FFA9FFF6D182FCA0E3754A5C" attach="left" box="[247,263,798,812]" fontSize="6" pageId="4" pageNumber="292">nd</superScript>
extended along entire length and gradually turns into posterior margin (
<figureCitation id="13293BFAFFA9FFF6D523FCA1E6C44A49" box="[1110,1206,799,825]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D5B3FCA1E77F4A49" box="[1222,1293,799,825]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4A, F</figureCitation>
). Surface of deep punctures larger than that on pronotum and elytra, covered with whitish plumose setae (
<figureCitation id="13293BFAFFA9FFF6D5DAFCFDE7774A2D" box="[1199,1285,835,861]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">Fig. 4F</figureCitation>
). Wings fully developed.
</paragraph>
<paragraph id="8BAD277FFFA9FFF6D1B2FC35E6A74A9D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
Abdomen shorter than forebody and slightly wider than elytral width combined, gradually narrowing basad; with numerous sparsely scattered whitish plumose setae, surface between setae shiny and unpunctured, microsculpture absent or vaguely visible (
<figureCitation id="13293BFAFFA9FFF6D0C8FC6DE0684A9D" box="[445,538,979,1005]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D353FC6DE01C4A9D" box="[550,622,979,1005]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4A, G</figureCitation>
). Apical margin of tergite VIII without palisade fringe.
</paragraph>
<paragraph id="8BAD277FFFA9FFF6D1B2FC49E2BA4DD1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
Male. Genital segments: sternite VIII with posterior margin almost straight and bearing minute setae along almost all of its length (
<figureCitation id="13293BFAFFA9FFF6D0D4FBA5E3894D45" box="[417,507,1050,1077]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="4" pageNumber="292">Fig. 5D</figureCitation>
), tergite IX triangular with elongated arm extending basad, bearing minute setae at apex (
<figureCitation id="13293BFAFFA9FFF6D1AEFB81E3414D29" box="[219,307,1086,1113]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="4" pageNumber="292">Fig. 5A</figureCitation>
), sternite IX elongated with square and obtusely pointed apex, posterior margin densely covered with short setae (
<figureCitation id="13293BFAFFA9FFF6D06BFBDDE3044D0D" box="[286,374,1122,1149]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="4" pageNumber="292">Fig. 5C</figureCitation>
), tergite X triangular with sharp apex covered with minute setae and with ventral struts (
<figureCitation id="13293BFAFFA9FFF6D41AFBDDE2C94DD1" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="4" pageNumber="292">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BAD277FFFA9FFF6D1B2FB15E6CD4C41" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
Aedeagus (
<figureCitation id="13293BFAFFA9FFF6D030FB15E3EE4DB5" box="[325,412,1194,1221]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="4" pageNumber="292">Fig. 5C</figureCitation>
) rather short, median lobe (in lateral or dorsal view) wide at base with large basal pressure plate, gradually narrowing apicad to obtusely pointed apex, apical margin covered with numerous microsetae; parameres parallel-sided, slightly but distinctly reaching beyond apex of median lobe (in lateral or dorsal view), sparsely covered with pores rarely bearing microsetae; internal sac with trident shaped sclerite.
</paragraph>
<paragraph id="8BAD277FFFA9FFF6D1B2FA85E35A4C09" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
Female. Sternite VIII triangular and pointed basad with shallow setae at posterior margin (
<figureCitation id="13293BFAFFA9FFF6D5DDFA85E68F4C25" box="[1192,1277,1338,1365]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="4" pageNumber="292">Fig. 5E</figureCitation>
), spermatheca as in
<figureCitation id="13293BFAFFA9FFF6D1A7FAE1E3564C09" box="[210,292,1374,1401]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="4" pageNumber="292">
Fig.
<specimenCount id="9D14ECF6FFA9FFF6D073FAE0E3564C09" box="[262,292,1374,1401]" count="5" pageId="4" pageNumber="292" type="generic">5F</specimenCount>
</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30874F4FFA9FFF6D1B2FA3CE7F94CB1" pageId="4" pageNumber="292" type="distribution">
<paragraph id="8BAD277FFFA9FFF6D1B2FA3CE7F94CB1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
<emphasis id="B966FB6DFFA9FFF6D1B2FA3CE3864CED" bold="true" box="[199,500,1410,1437]" pageId="4" pageNumber="292">Distribution and ecology.</emphasis>
<taxonomicName id="4C125CFCFFA9FFF6D089FA3DE0984CEC" authorityName="Salnitska &amp; Sheykin &amp; Semerikova" authorityYear="2024" box="[508,746,1411,1437]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="sergeevae">
<emphasis id="B966FB6DFFA9FFF6D089FA3DE0984CEC" box="[508,746,1411,1437]" italics="true" pageId="4" pageNumber="292">Euphanias sergeevae</emphasis>
</taxonomicName>
is hitherto known only from the flooded and sandy shores of the Tavolzhan salt lake in
<collectingRegion id="49D6E99DFFA9FFF6D0CEFA19E0F34CB1" box="[443,641,1447,1473]" country="Russia" name="Tyumen'" pageId="4" pageNumber="292">Tyumen Province</collectingRegion>
and the habitat is mostly covered with
<taxonomicName id="4C125CFCFFA9FFF6D54CFA19E6A74CB1" box="[1081,1237,1447,1473]" class="Magnoliopsida" family="Amaranthaceae" genus="Salicornia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="292" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B966FB6DFFA9FFF6D54CFA19E6DC4CB1" box="[1081,1198,1447,1473]" italics="true" pageId="4" pageNumber="292">Salicornia</emphasis>
sp.
</taxonomicName>
(
<figureCitation id="13293BFAFFA9FFF6D596FA19E7584CB1" box="[1251,1322,1447,1473]" captionStart="FIGURE 1" captionStartId="1.[151,250,967,991]" captionTargetBox="[151,1436,181,943]" captionTargetId="figure-34@1.[151,1436,181,943]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Euphanias species." figureDoi="http://doi.org/10.5281/zenodo.10618218" httpUri="https://zenodo.org/record/10618218/files/figure.png" pageId="4" pageNumber="292">Figs 1</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D44DFA19E70F4CB1" box="[1336,1405,1447,1473]" captionStart="FIGURE 2" captionStartId="1.[151,250,1955,1979]" captionTargetBox="[151,1436,1016,1931]" captionTargetId="figure-44@1.[151,1436,1016,1931]" captionTargetPageId="1" captionText="FIGURE 2. Tavolzhan salt lake. A: satellite image, orange dots indicate localities where the type specimens were collected; B and bottom row: lake shore with Salicornia sp." figureDoi="http://doi.org/10.5281/zenodo.10618220" httpUri="https://zenodo.org/record/10618220/files/figure.png" pageId="4" pageNumber="292">2A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30874F4FFA9FFF6D1B2FA75E77E4F5D" pageId="4" pageNumber="292" type="etymology">
<paragraph id="8BAD277FFFA9FFF6D1B2FA75E77E4F5D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="292">
<emphasis id="B966FB6DFFA9FFF6D1B2FA75E3384C95" bold="true" box="[199,330,1483,1509]" pageId="4" pageNumber="292">Etymology.</emphasis>
The species is named after Mrs. Elena Sergeeva (Tobolsk complex scientific station of the Ural Branch of the Russian Academy of Sciences, Tobolsk,
<collectingCountry id="F30567EFFFA9FFF6D3DBFA51E0894F79" box="[686,763,1519,1545]" name="Russia" pageId="4" pageNumber="292">Russia</collectingCountry>
), who collected the first specimen of
<taxonomicName id="4C125CFCFFA9FFF6D5DFFA51E74A4F78" authority="Salnitska &amp; Sheykin &amp; Semerikova, 2024" authorityName="Salnitska &amp; Sheykin &amp; Semerikova" authorityYear="2024" box="[1194,1336,1519,1544]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="sergeevae" status="sp. nov.">
<emphasis id="B966FB6DFFA9FFF6D5DFFA51E74A4F78" box="[1194,1336,1519,1544]" italics="true" pageId="4" pageNumber="292">E. sergeevae</emphasis>
</taxonomicName>
<emphasis id="B966FB6DFFA9FFF6D437FA51E7EE4F79" bold="true" box="[1346,1436,1519,1545]" pageId="4" pageNumber="292">
<taxonomicNameLabel id="A2554616FFA9FFF6D437FA51E7EE4F79" box="[1346,1436,1519,1545]" pageId="4" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<typeStatus id="54A999DDFFA9FFF6D1EAF9ADE3734F5D" box="[159,257,1555,1581]" pageId="4" pageNumber="292" type="paratype">paratype</typeStatus>
, UTMN_LISP_000250) from Tavolzhan lake and drew our attention to this interesting finding.
</paragraph>
</subSubSection>
<subSubSection id="C30874F4FFA9FFF7D1B2F988E0974989" lastPageId="5" lastPageNumber="293" pageId="4" pageNumber="292" type="diagnosis">
<paragraph id="8BAD277FFFA9FFF7D1B2F988E0974989" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,249]" lastPageId="5" lastPageNumber="293" pageId="4" pageNumber="292">
<emphasis id="B966FB6DFFA9FFF6D1B2F988E3BC4F20" bold="true" box="[199,462,1590,1616]" pageId="4" pageNumber="292">Differential diagnosis.</emphasis>
The new species can be distinguished from other members of the genus
<taxonomicName id="4C125CFCFFA9FFF6D474F989E70B4F21" authorityName="Fairmaire &amp; Laboulbene" authorityYear="1856" box="[1281,1401,1591,1617]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B966FB6DFFA9FFF6D474F989E70B4F21" box="[1281,1401,1591,1617]" italics="true" pageId="4" pageNumber="292">Euphanias</emphasis>
</taxonomicName>
by the following combination of characters: piceous, only tarsomeres and mouth appendages slightly lighter, antennae moderately short, distinctly shorter than forebody (
<figureCitation id="13293BFAFFA9FFF6D3ACF9C1E1454FE9" box="[729,823,1663,1689]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D231F9C1E1184FE9" box="[836,874,1663,1689]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4A</figureCitation>
), first segment of antennae more than three times as long as wide, segments 35 gradually increasing in length (
<figureCitation id="13293BFAFFA9FFF6D21AF91DE1A24FCD" box="[879,976,1699,1725]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D2AAF91DE6544FCD" box="[991,1062,1699,1725]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4AC</figureCitation>
), pronotum almost hexagonal in shape with strongly pronounced angles and with weak medial furrow throughout its length, 1
<superScript id="7C678A37FFA9FFF6D5A7F979E6AF4FA5" attach="left" box="[1234,1245,1735,1749]" fontSize="6" pageId="4" pageNumber="292">st</superScript>
and 3
<superScript id="7C678A37FFA9FFF6D456F978E7434FA4" attach="left" box="[1315,1329,1734,1748]" fontSize="6" pageId="4" pageNumber="292">rd</superScript>
ridges of elytra not reaching elytral posterior margin and extending through half of its length (
<figureCitation id="13293BFAFFA9FFF6D529F955E6B34E75" box="[1116,1217,1771,1797]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs. 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D5BBF955E7604E75" box="[1230,1298,1771,1797]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4A, F</figureCitation>
), pronotum, elytra and abdomen covered with brush like plumose setae (
<figureCitation id="13293BFAFFA9FFF6D24EF8B1E1E54E59" box="[827,919,1807,1833]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">Fig. 4H</figureCitation>
). According to the only existing identification key for
<taxonomicName id="4C125CFCFFA9FFF6D19AF88DE3CD4E3D" box="[239,447,1843,1869]" pageId="4" pageNumber="292">
<emphasis id="B966FB6DFFA9FFF6D19AF88DE3154E3D" box="[239,359,1843,1869]" italics="true" pageId="4" pageNumber="292">Euphanias</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="EF835A8EFFA9FFF6D0B8F88DE00A4E3D" author="Blinstein, S. Ya." box="[461,632,1843,1869]" pageId="4" pageNumber="292" pagination="78 - 81" refId="ref4541" refString="Blinstein, S. Ya. (1976) Novyi i maloizvestnye vidy stafilinid (Coleoptera, Staphylinidae) iz severo-zapadnogo Prichernomor'ia i Prisivash'ia. Vestnik Zoologii, 5, 78 - 81. [in Russian]" type="journal article" year="1976">Blinstein 1976</bibRefCitation>
),
<taxonomicName id="4C125CFCFFA9FFF6D3FEF88DE16B4E3C" authority="Salnitska &amp; Sheykin &amp; Semerikova, 2024" authorityName="Salnitska &amp; Sheykin &amp; Semerikova" authorityYear="2024" box="[651,793,1843,1868]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="sergeevae" status="sp. nov.">
<emphasis id="B966FB6DFFA9FFF6D3FEF88DE16B4E3C" box="[651,793,1843,1868]" italics="true" pageId="4" pageNumber="292">E. sergeevae</emphasis>
</taxonomicName>
<emphasis id="B966FB6DFFA9FFF6D254F88DE1084E3D" bold="true" box="[801,890,1843,1869]" pageId="4" pageNumber="292">
<taxonomicNameLabel id="A2554616FFA9FFF6D254F88DE1084E3D" box="[801,890,1843,1869]" pageId="4" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
reaches the couplet comprising
<taxonomicName id="4C125CFCFFA9FFF6D590F88DE71E4E3D" authorityName="Bernhauer" authorityYear="1912" box="[1253,1388,1843,1869]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="pliginskii">
<emphasis id="B966FB6DFFA9FFF6D590F88DE71E4E3D" box="[1253,1388,1843,1869]" italics="true" pageId="4" pageNumber="292">E. pliginskii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C125CFCFFA9FFF6D1E2F8E9E3694E00" authorityName="Blinstein" authorityYear="1976" box="[151,283,1879,1904]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="pusanovi">
<emphasis id="B966FB6DFFA9FFF6D1E2F8E9E3694E00" box="[151,283,1879,1904]" italics="true" pageId="4" pageNumber="292">E. pusanovi</emphasis>
</taxonomicName>
, sharing the character state of the first antennomere, which is three times as long as wide. Our examination of
<taxonomicName id="4C125CFCFFA9FFF6D1C3F8C5E34F4EE5" authorityName="Bernhauer" authorityYear="1912" box="[182,317,1915,1941]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="pliginskii">
<emphasis id="B966FB6DFFA9FFF6D1C3F8C5E34F4EE5" box="[182,317,1915,1941]" italics="true" pageId="4" pageNumber="292">E. pliginskii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C125CFCFFA9FFF6D001F8C5E38A4EE4" authorityName="Blinstein" authorityYear="1976" box="[372,504,1915,1940]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="pusanovi">
<emphasis id="B966FB6DFFA9FFF6D001F8C5E38A4EE4" box="[372,504,1915,1940]" italics="true" pageId="4" pageNumber="292">E. pusanovi</emphasis>
</taxonomicName>
<typeStatus id="54A999DDFFA9FFF6D375F8C5E0494EE5" box="[512,571,1915,1941]" pageId="4" pageNumber="292">types</typeStatus>
and additional materials revealed that
<taxonomicName id="4C125CFCFFA9FFF6D29BF8C5E6004EE4" authorityName="Blinstein" authorityYear="1976" box="[1006,1138,1915,1940]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="pusanovi">
<emphasis id="B966FB6DFFA9FFF6D29BF8C5E6004EE4" box="[1006,1138,1915,1940]" italics="true" pageId="4" pageNumber="292">E. pusanovi</emphasis>
</taxonomicName>
can be distinguished from both species by its darker and uniform coloration, less pronounced eyes, different shape of pronotum with weak medial furrow (
<figureCitation id="13293BFAFFA9FFF6D033F87DE3D14EAD" box="[326,419,1987,2013]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="4" pageNumber="292">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA9FFF6D0DAF87DE0644EAD" box="[431,534,1987,2013]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="4" pageNumber="292">4A, D, E</figureCitation>
); from
<taxonomicName id="4C125CFCFFA9FFF6D31CF87DE09F4EAC" authorityName="Blinstein" authorityYear="1976" box="[617,749,1987,2012]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="pusanovi">
<emphasis id="B966FB6DFFA9FFF6D31CF87DE09F4EAC" box="[617,749,1987,2012]" italics="true" pageId="4" pageNumber="292">E. pusanovi</emphasis>
</taxonomicName>
by the sparser punctuation, brush-like plumose setae, shorter antennae and more transverse segment 6, more sharply tapering apex of abdomen more straight and not curved parameres (
<figureCitation id="13293BFAFFA8FFF7D06CFF05E30449A5" box="[281,374,187,213]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="5" pageNumber="293">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA8FFF7D0F6FF05E05C49A5" box="[387,558,187,213]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="5" pageNumber="293">4A, B, C, G, H</figureCitation>
,
<figureCitation id="13293BFAFFA8FFF7D34CFF04E02D49A5" box="[569,607,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1405,190,1919]" captionTargetId="figure-17@6.[182,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Genital segments of E. sergeevae sp. nov. A: tergite IX; B: tergite X; C: sternite IX; D: male sternite VIII; E: female sternite VIII; F: spermatheca. Aedeagus. G: E. sergeevae sp. nov.; H: E. pliginskii. Scale bars. AF: 0.5 mm; G, H: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10618228" httpUri="https://zenodo.org/record/10618228/files/figure.png" pageId="5" pageNumber="293">5G</figureCitation>
); from
<taxonomicName id="4C125CFCFFA8FFF7D3C7FF05E14B49A5" authorityName="Bernhauer" authorityYear="1912" box="[690,825,187,213]" class="Insecta" family="Staphylinidae" genus="Euphanias" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="293" phylum="Arthropoda" rank="species" species="pliginskii">
<emphasis id="B966FB6DFFA8FFF7D3C7FF05E14B49A5" box="[690,825,187,213]" italics="true" pageId="5" pageNumber="293">E. pliginskii</emphasis>
</taxonomicName>
by the shorter and weaker 1
<superScript id="7C678A37FFA8FFF7D502FF04E6F549B8" attach="left" box="[1143,1159,186,200]" fontSize="6" pageId="5" pageNumber="293">nd</superScript>
and 3
<superScript id="7C678A37FFA8FFF7D5BCFF04E6A549B8" attach="left" box="[1225,1239,186,200]" fontSize="6" pageId="5" pageNumber="293">rd</superScript>
elytral ridges not reaching posterior margin of elytra (
<figureCitation id="13293BFAFFA8FFF7D359FF61E0FB4989" box="[556,649,223,249]" captionStart="FIGURE 3" captionStartId="3.[151,250,1241,1265]" captionTargetBox="[177,1436,417,1217]" captionTargetId="figure-111@3.[151,1436,417,1217]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Euphanias species. A: E. sergeevae sp. nov. (holotype); B: E. pusanovi (?holotype); C: E. pliginskii (holotype). Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10618224" httpUri="https://zenodo.org/record/10618224/files/figure.png" pageId="5" pageNumber="293">Figs 3A</figureCitation>
,
<figureCitation id="13293BFAFFA8FFF7D3E0FF61E0AA4989" box="[661,728,223,249]" captionStart="FIGURE 4" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[170,1420,1335,1780]" captionTargetId="figure-138@3.[151,1436,1325,1789]" captionTargetPageId="3" captionText="FIGURE 4. Scanning electron micrograph of E. sergeevae sp. nov. A: habitus; B, C: antennae; D: head; E: pronotum; F: elytra; G: abdomen; H: plumose brush-like setae. Scale bars. A: 500 µm; B: 50 µm; CG: 100 µm; H: 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10618226" httpUri="https://zenodo.org/record/10618226/files/figure.png" pageId="5" pageNumber="293">4A, F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>