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<mods:title id="043CF00D02AB643CBDAA8377F78DD430">Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus</mods:title>
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<emphasis id="B942E03AFFB8FFE3969641CFFDCBF92D" bold="true" box="[151,589,1741,1767]" pageId="5" pageNumber="306">
Key to males of the
<taxonomicName id="4C3647ABFFB8FFE3978C41CFFE7EF92D" box="[397,504,1741,1767]" class="Insecta" family="Scirtidae" genus="Nothocyphon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="lindensis">
<emphasis id="B942E03AFFB8FFE3978C41CFFE7EF92D" bold="true" box="[397,504,1741,1767]" italics="true" pageId="5" pageNumber="306">lindensis</emphasis>
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-group
</emphasis>
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</paragraph>
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<keyStep id="30C2248DFFB8FFE396964016FA1DF8A9" pageId="5" pageNumber="306">
<paragraph id="8B893C28FFB8FFE396964016FA1DF88C" blockId="5.[151,1436,1812,2038]" pageId="5" pageNumber="306">
<keyLead id="30C79F1DFFB8FFE396964016FA1DF88C" pageId="5" pageNumber="306">
1 A large hollow cone-like spine between penis and S9 (e.g.,
<figureCitation id="130D20ADFFB8FFE395174016FCE3F8E3" box="[790,869,1812,1833]" captionStart="FIGURES 8 13" captionStartId="8.[151,264,1751,1774]" captionTargetBox="[203,1395,345,1719]" captionTargetId="figure@8.[181,1405,337,1730]" captionTargetPageId="8" captionText="FIGURES 8 13. Nothocyphon lindensis - group, male genitalia. N. lindensis: 8, ventral view of cleared protracted genitalia; 9, the same, dorsal view; 10, oblique ventral view of the erect genitalia, trigonium deflexed. — N. ypsilon: 11, ventral view of tegmen, penis and S 9; 12, penis tip, ventral, enlarged; 13, dorsal view of cleared protracted genitalia; 8, 9, 11, 13 to the same scale, 10 not to scale. Abbreviations: hs, hollow spine; pa, pala; pd, parameroid; pe, paramere; te, tegmen; tr, trigonium." httpUri="https://zenodo.org/record/240980/files/figure.png" pageId="5" pageNumber="306">Figs. 12</figureCitation>
,
<figureCitation id="130D20ADFFB8FFE3956F4016FC0FF8E3" box="[878,905,1812,1833]" captionStart="FIGURES 17 24" captionStartId="12.[151,264,1821,1844]" captionTargetBox="[253,1313,212,1764]" captionTargetId="figure@12.[234,1329,193,1781]" captionTargetPageId="12" captionText="FIGURES 17 24. Nothocyphon lindensis - group, N. taurus (17 20) and N. banksiae (21 24). 17, dorsal view of male terminalia as extracted from abdomen, superimposed components seen by transparency; semi-diagrammatic, setae of S 8 and microtrichia on T 8 not shown; 18, tips of penis and parameres of another specimen, ventral view; 19, T 9 and S 9 superimposed, caudal edge of T 9 and base of S 9 are transparent, the hollow spine lies between them; 20, prehensor; 21, T 8 and the (inverted) S 8; 22, T 9; 23, S 9; 24, dorsal view of penis and superimposed tegmen and parameres. 18 20 and 21 23 to the same scales, respectively. Abbreviations: aT 9, apodeme of T 9; hs, hollow spine; pd, parameroid; pe, paramere; tr, trigonium." httpUri="https://zenodo.org/record/240982/files/figure.png" pageId="5" pageNumber="306">19</figureCitation>
). Yellow or light brown beetles with a brown spot close to the suture near the top of elytra (
<figureCitation id="130D20ADFFB8FFE394334033FD00F88C" box="[562,646,1841,1862]" captionStart="FIGURES 165 173" captionStartId="56.[151,264,1921,1944]" captionTargetBox="[234,1328,219,1863]" captionTargetId="figure@56.[223,1341,193,1875]" captionTargetPageId="56" captionText="FIGURES 165 173. Habitus of Nothocyphon spp.: 165, N. lindensis; 166, N. taurus; 167, N. taeniatus; 168, N. denticulatus; 169, N. donnabuangi; 170, N. imitator; 171, N. frater; 172, N. frater, lateral; 173, N. taurus, lateral. To similar sizes, not to scale." httpUri="https://zenodo.org/record/240999/files/figure.png" pageId="5" pageNumber="306">Fig. 165</figureCitation>
), rarely more extended maculae (
<figureCitation id="130D20ADFFB8FFE395C44033FB9FF88C" box="[965,1049,1841,1862]" captionStart="FIGURES 165 173" captionStartId="56.[151,264,1921,1944]" captionTargetBox="[234,1328,219,1863]" captionTargetId="figure@56.[223,1341,193,1875]" captionTargetPageId="56" captionText="FIGURES 165 173. Habitus of Nothocyphon spp.: 165, N. lindensis; 166, N. taurus; 167, N. taeniatus; 168, N. denticulatus; 169, N. donnabuangi; 170, N. imitator; 171, N. frater; 172, N. frater, lateral; 173, N. taurus, lateral. To similar sizes, not to scale." httpUri="https://zenodo.org/record/240999/files/figure.png" pageId="5" pageNumber="306">Fig. 166</figureCitation>
)............................... 2
</keyLead>
</paragraph>
<paragraph id="8B893C28FFB8FFE39696404CFA1DF8A9" blockId="5.[151,1436,1812,2038]" box="[151,1435,1870,1891]" pageId="5" pageNumber="306">
<keyLead id="30C79F1DFFB8FFE39696404CFA1DF8A9" box="[151,1435,1870,1891]" pageId="5" pageNumber="306">1' Groups of slender spines between penis and S9 (Figs. 26, 29). Brown beetles...................................... 5</keyLead>
</paragraph>
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<paragraph id="8B893C28FFB8FFE39696406EFA1AF84B" blockId="5.[151,1436,1812,2038]" box="[151,1436,1900,1921]" pageId="5" pageNumber="306">
<keyLead id="30C79F1DFFB8FFE39696406EFA1AF84B" box="[151,1436,1900,1921]" pageId="5" pageNumber="306">
2 Parameres enveloping the penis, one or two huge curved spines along their medioventral edge (
<figureCitation id="130D20ADFFB8FFE39272406EFB63F84B" box="[1139,1253,1900,1921]" captionStart="FIGURES 8 13" captionStartId="8.[151,264,1751,1774]" captionTargetBox="[203,1395,345,1719]" captionTargetId="figure@8.[181,1405,337,1730]" captionTargetPageId="8" captionText="FIGURES 8 13. Nothocyphon lindensis - group, male genitalia. N. lindensis: 8, ventral view of cleared protracted genitalia; 9, the same, dorsal view; 10, oblique ventral view of the erect genitalia, trigonium deflexed. — N. ypsilon: 11, ventral view of tegmen, penis and S 9; 12, penis tip, ventral, enlarged; 13, dorsal view of cleared protracted genitalia; 8, 9, 11, 13 to the same scale, 10 not to scale. Abbreviations: hs, hollow spine; pa, pala; pd, parameroid; pe, paramere; te, tegmen; tr, trigonium." httpUri="https://zenodo.org/record/240980/files/figure.png" pageId="5" pageNumber="306">Figs. 10, 11</figureCitation>
)............. 3
</keyLead>
</paragraph>
<paragraph id="8B893C28FFB8FFE39696408BFA1DF854" blockId="5.[151,1436,1812,2038]" box="[151,1435,1929,1950]" pageId="5" pageNumber="306">
<keyLead id="30C79F1DFFB8FFE39696408BFA1DF854" box="[151,1435,1929,1950]" pageId="5" pageNumber="306">2' Parameres slender, lying alongside penis, not surrounding it.................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C2248DFFB8FFE3969640A4FA1AF813" pageId="5" pageNumber="306">
<paragraph id="8B893C28FFB8FFE3969640A4FA1DF871" blockId="5.[151,1436,1812,2038]" box="[151,1435,1957,1979]" pageId="5" pageNumber="306">
<keyLead id="30C79F1DFFB8FFE3969640A4FA1DF871" box="[151,1435,1957,1979]" pageId="5" pageNumber="306">
3 Apex of trigonium entire, with spines in irregular dorsoventral rows (
<figureCitation id="130D20ADFFB8FFE3957840A4FC67F871" box="[889,993,1958,1979]" captionStart="FIGURES 8 13" captionStartId="8.[151,264,1751,1774]" captionTargetBox="[203,1395,345,1719]" captionTargetId="figure@8.[181,1405,337,1730]" captionTargetPageId="8" captionText="FIGURES 8 13. Nothocyphon lindensis - group, male genitalia. N. lindensis: 8, ventral view of cleared protracted genitalia; 9, the same, dorsal view; 10, oblique ventral view of the erect genitalia, trigonium deflexed. — N. ypsilon: 11, ventral view of tegmen, penis and S 9; 12, penis tip, ventral, enlarged; 13, dorsal view of cleared protracted genitalia; 8, 9, 11, 13 to the same scale, 10 not to scale. Abbreviations: hs, hollow spine; pa, pala; pd, parameroid; pe, paramere; te, tegmen; tr, trigonium." httpUri="https://zenodo.org/record/240980/files/figure.png" pageId="5" pageNumber="306">Figs. 810</figureCitation>
)................
<taxonomicName id="4C3647ABFFB8FFE392B140A4FA1DF871" authority="Blackburn" authorityName="Blackburn" box="[1200,1435,1957,1979]" class="Insecta" family="Scirtidae" genus="Nothocyphon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="lindensis">
<emphasis id="B942E03AFFB8FFE392B140A4FAA7F871" box="[1200,1313,1957,1979]" italics="true" pageId="5" pageNumber="306">N. lindensis</emphasis>
(Blackburn)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B893C28FFB8FFE3969640C6FA1AF813" blockId="5.[151,1436,1812,2038]" box="[151,1436,1987,2009]" pageId="5" pageNumber="306">
<keyLead id="30C79F1DFFB8FFE3969640C6FA1AF813" box="[151,1436,1987,2009]" pageId="5" pageNumber="306">
3' Trigonium forked, the hollow spine visible between the short branches (
<figureCitation id="130D20ADFFB8FFE3959340C6FB83F813" box="[914,1029,1988,2009]" captionStart="FIGURES 8 13" captionStartId="8.[151,264,1751,1774]" captionTargetBox="[203,1395,345,1719]" captionTargetId="figure@8.[181,1405,337,1730]" captionTargetPageId="8" captionText="FIGURES 8 13. Nothocyphon lindensis - group, male genitalia. N. lindensis: 8, ventral view of cleared protracted genitalia; 9, the same, dorsal view; 10, oblique ventral view of the erect genitalia, trigonium deflexed. — N. ypsilon: 11, ventral view of tegmen, penis and S 9; 12, penis tip, ventral, enlarged; 13, dorsal view of cleared protracted genitalia; 8, 9, 11, 13 to the same scale, 10 not to scale. Abbreviations: hs, hollow spine; pa, pala; pd, parameroid; pe, paramere; te, tegmen; tr, trigonium." httpUri="https://zenodo.org/record/240980/files/figure.png" pageId="5" pageNumber="306">Figs. 12, 13</figureCitation>
)....................
<taxonomicName id="4C3647ABFFB8FFE392FF40C6FAD9F813" box="[1278,1375,1987,2009]" class="Insecta" family="Scirtidae" genus="Nothocyphon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="ypsilon" status="sp. nov.">
<emphasis id="B942E03AFFB8FFE392FF40C6FAD9F813" box="[1278,1375,1987,2009]" italics="true" pageId="5" pageNumber="306">N. ypsilon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2715D41FFB8FFE3936740C1FA1AF813" box="[1382,1436,1987,2009]" pageId="5" pageNumber="306" rank="species">
<emphasis id="B942E03AFFB8FFE3936740C1FA1AF813" bold="true" box="[1382,1436,1987,2009]" pageId="5" pageNumber="306">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C2248DFFB8FFE0969640E3FA1AFF67" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
<paragraph id="8B893C28FFB8FFE3969640E3FA1AF83C" blockId="5.[151,1436,1812,2038]" box="[151,1436,2016,2038]" pageId="5" pageNumber="306">
<keyLead id="30C79F1DFFB8FFE3969640E3FA1AF83C" box="[151,1436,2016,2038]" pageId="5" pageNumber="306">
4 Trigonium truncate, short, with curved lateral horns (
<figureCitation id="130D20ADFFB8FFE394DC40E3FCA4F83C" box="[733,802,2017,2038]" captionStart="FIGURES 17 24" captionStartId="12.[151,264,1821,1844]" captionTargetBox="[253,1313,212,1764]" captionTargetId="figure@12.[234,1329,193,1781]" captionTargetPageId="12" captionText="FIGURES 17 24. Nothocyphon lindensis - group, N. taurus (17 20) and N. banksiae (21 24). 17, dorsal view of male terminalia as extracted from abdomen, superimposed components seen by transparency; semi-diagrammatic, setae of S 8 and microtrichia on T 8 not shown; 18, tips of penis and parameres of another specimen, ventral view; 19, T 9 and S 9 superimposed, caudal edge of T 9 and base of S 9 are transparent, the hollow spine lies between them; 20, prehensor; 21, T 8 and the (inverted) S 8; 22, T 9; 23, S 9; 24, dorsal view of penis and superimposed tegmen and parameres. 18 20 and 21 23 to the same scales, respectively. Abbreviations: aT 9, apodeme of T 9; hs, hollow spine; pd, parameroid; pe, paramere; tr, trigonium." httpUri="https://zenodo.org/record/240982/files/figure.png" pageId="5" pageNumber="306">Fig. 18</figureCitation>
).......................................
<taxonomicName id="4C3647ABFFB8FFE3930740E3FAD9F83C" box="[1286,1375,2017,2038]" class="Insecta" family="Scirtidae" genus="Nothocyphon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="taurus" status="sp. nov.">
<emphasis id="B942E03AFFB8FFE3930740E3FAD9F83C" box="[1286,1375,2017,2038]" italics="true" pageId="5" pageNumber="306">N. taurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2715D41FFB8FFE3936740E2FA1AF83C" box="[1382,1436,2016,2038]" pageId="5" pageNumber="306" rank="species">
<emphasis id="B942E03AFFB8FFE3936740E2FA1AF83C" bold="true" box="[1382,1436,2016,2038]" pageId="5" pageNumber="306">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B893C28FFBBFFE096964795FA1AFF67" blockId="6.[151,1436,151,290]" box="[151,1436,151,173]" pageId="6" pageNumber="307">
<keyLead id="30C79F1DFFBBFFE096964795FA1AFF67" box="[151,1436,151,173]" pageId="6" pageNumber="307">
4' Trigonium deeply excised, each lobe medially with many spinules (
<figureCitation id="130D20ADFFBBFFE095694795FC28FF66" box="[872,942,151,172]" captionStart="FIGURES 17 24" captionStartId="12.[151,264,1821,1844]" captionTargetBox="[253,1313,212,1764]" captionTargetId="figure@12.[234,1329,193,1781]" captionTargetPageId="12" captionText="FIGURES 17 24. Nothocyphon lindensis - group, N. taurus (17 20) and N. banksiae (21 24). 17, dorsal view of male terminalia as extracted from abdomen, superimposed components seen by transparency; semi-diagrammatic, setae of S 8 and microtrichia on T 8 not shown; 18, tips of penis and parameres of another specimen, ventral view; 19, T 9 and S 9 superimposed, caudal edge of T 9 and base of S 9 are transparent, the hollow spine lies between them; 20, prehensor; 21, T 8 and the (inverted) S 8; 22, T 9; 23, S 9; 24, dorsal view of penis and superimposed tegmen and parameres. 18 20 and 21 23 to the same scales, respectively. Abbreviations: aT 9, apodeme of T 9; hs, hollow spine; pd, parameroid; pe, paramere; tr, trigonium." httpUri="https://zenodo.org/record/240982/files/figure.png" pageId="6" pageNumber="307">Fig. 24</figureCitation>
).........................
<taxonomicName id="4C3647ABFFBBFFE092EF479AFAD9FF67" box="[1262,1375,151,173]" class="Insecta" family="Scirtidae" genus="Nothocyphon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="banksiae" status="sp. nov.">
<emphasis id="B942E03AFFBBFFE092EF479AFAD9FF67" box="[1262,1375,151,173]" italics="true" pageId="6" pageNumber="307">N. banksiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2715D41FFBBFFE093674795FA1AFF67" box="[1382,1436,151,173]" pageId="6" pageNumber="307" rank="species">
<emphasis id="B942E03AFFBBFFE093674795FA1AFF67" bold="true" box="[1382,1436,151,173]" pageId="6" pageNumber="307">n. sp.</emphasis>
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<paragraph id="8B893C28FFBBFFE0969647B7FA1AFF22" blockId="6.[151,1436,151,290]" pageId="6" pageNumber="307">
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5 Notch in trigonium deeper than wide. Each lobe of S9 with a rounded notch. A single triangular patch of spines, parameres with strong teeth (Fig. 26)..............................................................
<taxonomicName id="4C3647ABFFBBFFE092C547D0FAE6FF2D" box="[1220,1376,209,231]" class="Insecta" family="Scirtidae" genus="Nothocyphon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="donnabuangi" status="sp. nov.">
<emphasis id="B942E03AFFBBFFE092C547D0FAE6FF2D" box="[1220,1376,209,231]" italics="true" pageId="6" pageNumber="307">N. donnabuangi</emphasis>
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<emphasis id="B942E03AFFBBFFE0936747D0FA1AFF22" bold="true" box="[1382,1436,210,232]" pageId="6" pageNumber="307">n. sp.</emphasis>
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5' Notch of trigonium shallower, wider than deep. Caudal lobes of S9 not notched. Paired spinule patches, parameres with fine teeth (Fig. 29)...........................................................................
<taxonomicName id="4C3647ABFFBBFFE092F7460FFAD9FEE8" box="[1270,1375,269,290]" class="Insecta" family="Scirtidae" genus="Nothocyphon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="imitator" status="sp. nov.">
<emphasis id="B942E03AFFBBFFE092F7460FFAD9FEE8" box="[1270,1375,269,290]" italics="true" pageId="6" pageNumber="307">N. imitator</emphasis>
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<emphasis id="B942E03AFFBBFFE09367460EFA1AFEE8" bold="true" box="[1382,1436,268,290]" pageId="6" pageNumber="307">n. sp.</emphasis>
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