treatments-xml/data/03/8C/87/038C87863F6EFFD1FF3667DE4DB1625B.xml
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<document id="CD809BB7985A87B9EDD4F1C239282233" ID-CLB-Dataset="45708" ID-DOI="10.5281/zenodo.279573" ID-GBIF-Dataset="7359830e-3896-4dc8-91bc-1c15c4ee2993" ID-ISSN="1175-5326" ID-Zenodo-Dep="279573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460423643735" checkinUser="plazi" docAuthor="Holleman, Wouter &amp; Bogorodsky, Sergey V." docDate="2012" docId="038C87863F6EFFD1FF3667DE4DB1625B" docLanguage="en" docName="zt03152p060.pdf" docOrigin="Zootaxa 3152" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tripterygiidae" docType="key" docVersion="8" lastPageNumber="39" masterDocId="FFB5FFFE3F6CFFD2FFA16231482A6756" masterDocTitle="A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980" masterLastPageNumber="60" masterPageNumber="36" pageNumber="38" updateTime="1698330352715" updateUser="ExternalLinkService">
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<mods:title id="81123ADACA135153D95BD87CBBD4F8BD">A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980</mods:title>
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<treatment id="038C87863F6EFFD1FF3667DE4DB1625B" ID-DOI="http://doi.org/10.5281/zenodo.5626510" ID-GBIF-Taxon="119427992" ID-Zenodo-Dep="5626510" LSID="urn:lsid:plazi:treatment:038C87863F6EFFD1FF3667DE4DB1625B" httpUri="http://treatment.plazi.org/id/038C87863F6EFFD1FF3667DE4DB1625B" lastPageId="3" lastPageNumber="39" pageId="2" pageNumber="38">
<subSubSection id="C33F651B3F6EFFD0FF3667DE4B06615F" box="[151,812,1519,1545]" pageId="2" pageNumber="38" type="nomenclature">
<paragraph id="8B9A36903F6EFFD0FF3667DE4B06615F" blockId="2.[151,812,1519,1545]" box="[151,812,1519,1545]" pageId="2" pageNumber="38">
<heading id="D0D281FC3F6EFFD0FF3667DE4B06615F" bold="true" box="[151,812,1519,1545]" fontSize="11" level="1" pageId="2" pageNumber="38" reason="1">
<emphasis id="B951EA823F6EFFD0FF3667DE4B06615F" bold="true" box="[151,812,1519,1545]" pageId="2" pageNumber="38">
Key to the males of Red Sea species of
<taxonomicName id="4C254D133F6EFFD0FDD967DE4B06615F" ID-CoL="HH8" ID-ENA="57771" box="[632,812,1519,1545]" class="Actinopterygii" family="Tripterygiidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="38" phylum="Chordata" rank="family">Tripterygiidae</taxonomicName>
</emphasis>
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<subSubSection id="C33F651B3F6EFFD1FF3664044DB1625B" lastPageId="3" lastPageNumber="39" pageId="2" pageNumber="38" type="key">
<keyStep id="30D12E353F6EFFD0FF3664044DB66131" pageId="2" pageNumber="38">
<paragraph id="8B9A36903F6EFFD0FF3664044DB6611C" blockId="2.[151,1436,1589,2020]" box="[151,1436,1589,1610]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF3664044DB6611C" box="[151,1436,1589,1610]" pageId="2" pageNumber="38">1a. First dorsal fin with three spines; no scales on head...........................................................2</keyLead>
</paragraph>
<paragraph id="8B9A36903F6EFFD0FF3664634DB66131" blockId="2.[151,1436,1589,2020]" box="[151,1436,1618,1639]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF3664634DB66131" box="[151,1436,1618,1639]" pageId="2" pageNumber="38">
1b. First dorsal fin with four spines; head with scales...........................................
<taxonomicName id="4C254D133F6EFFD0FB6B64634DB66131" ID-CoL="47QBD" box="[1226,1436,1618,1639]" class="Actinopterygii" family="Tripterygiidae" genus="Norfolkia" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="38" phylum="Chordata" rank="species" species="brachylepis">
<emphasis id="B951EA823F6EFFD0FB6B64634DB66131" box="[1226,1436,1618,1639]" italics="true" pageId="2" pageNumber="38">Norfolkia brachylepis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D12E353F6EFFD0FF36645E4DB661F4" pageId="2" pageNumber="38">
<paragraph id="8B9A36903F6EFFD0FF36645E4DB661D2" blockId="2.[151,1436,1589,2020]" box="[151,1436,1647,1668]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF36645E4DB661D2" box="[151,1436,1647,1668]" pageId="2" pageNumber="38">2a. LL divided into anterior section of pored scales and posterior section of notched scales.............................. 4</keyLead>
</paragraph>
<paragraph id="8B9A36903F6EFFD0FF3664BC4DB661F4" blockId="2.[151,1436,1589,2020]" box="[151,1436,1677,1698]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF3664BC4DB661F4" box="[151,1436,1677,1698]" pageId="2" pageNumber="38">2b. LL only of pored scales, ending below second or third dorsal fin................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D12E353F6EFFD0FF36649B4DB160DB" pageId="2" pageNumber="38">
<paragraph id="8B9A36903F6EFFD0FF36649B4DB66041" blockId="2.[151,1436,1589,2020]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF36649B4DB66041" pageId="2" pageNumber="38">
3a. Males red in life, head dark with broad, pale blue mark above lip and on lower preopercle; females translucent greenish; body with indistinct brownish-red bars and scattered blotches; membrane of first dorsal fin of both sexes cream with reddish mark- ings and many micro-melanophores; dentary pores 3 + 12 + 3; underwater males with indistinct brownish-red bars and scat- tered blotches........................................................................
<taxonomicName id="4C254D133F6EFFD0FB6365334DB66041" ID-CoL="3K482" box="[1218,1436,1794,1815]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="38" phylum="Chordata" rank="species" species="steinitzi">
<emphasis id="B951EA823F6EFFD0FB6365334DB66041" box="[1218,1436,1794,1815]" italics="true" pageId="2" pageNumber="38">Helcogramma steinitzi</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8B9A36903F6EFFD0FF36652E4DB160DB" blockId="2.[151,1436,1589,2020]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF36652E4DB160DB" pageId="2" pageNumber="38">
3b. Males dark grey to dark red in life; lower half of head with many melanophores and bright blue line from corner of mouth, angling posteroventrally across cheek onto preopercle; both sexes with distinct, irregular bright red bars; first dorsal-fin mem- brane translucent with some iridescence; dentary pores 4 + 1 + 4; underwater with distinct irregular, bright red bars in both sexes, first dorsal-fin membrane translucent with some iridescence.........................
<taxonomicName id="4C254D133F6EFFD0FB3565494DB160DB" ID-CoL="3K47Q" box="[1172,1435,1912,1933]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="38" phylum="Chordata" rank="species" species="obtusirostris">
<emphasis id="B951EA823F6EFFD0FB3565494DB160DB" box="[1172,1435,1912,1933]" italics="true" pageId="2" pageNumber="38">Helcogramma obtusirostris</emphasis>
</taxonomicName>
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</keyStep>
<keyStep id="30D12E353F6EFFD0FF3665A44DB66091" pageId="2" pageNumber="38">
<paragraph id="8B9A36903F6EFFD0FF3665A44DB660FC" blockId="2.[151,1436,1589,2020]" box="[151,1436,1941,1962]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF3665A44DB660FC" box="[151,1436,1941,1962]" pageId="2" pageNumber="38">4a. Body and anal fin with bars.............................................................................. 5</keyLead>
</paragraph>
<paragraph id="8B9A36903F6EFFD0FF3665834DB66091" blockId="2.[151,1436,1589,2020]" box="[151,1436,1970,1991]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF3665834DB66091" box="[151,1436,1970,1991]" pageId="2" pageNumber="38">4b. Body without bars; anal fin with narrow bars or without bars................................................... 6</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B9A36903F6EFFD0FF3665FE4DB660B2" blockId="2.[151,1436,1589,2020]" box="[151,1436,1999,2020]" pageId="2" pageNumber="38">
<keyLead id="30D495A53F6EFFD0FF3665FE4DB660B2" box="[151,1436,1999,2020]" pageId="2" pageNumber="38">
<keyStep id="30D12E353F6EFFD0FF3665FE4DB660B2" box="[151,1436,1999,2020]" pageId="2" pageNumber="38">5a. Body with four dark bars usually divided below LL and continuing onto anal fin, and two paler bars below pectoral fin; nape</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9A36903F6FFFD1FF4062A64DB66674" blockId="3.[151,1436,151,554]" pageId="3" pageNumber="39">
and belly scaled; yellow and brown in life................................................
<taxonomicName id="4C254D133F6FFFD1FB1662A94DB667FB" ID-CoL="39VYY" box="[1207,1436,152,173]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="clarkae">
<emphasis id="B951EA823F6FFFD1FB1662A94DB667FB" box="[1207,1436,152,173]" italics="true" pageId="3" pageNumber="39">Enneapterygius clarkae</emphasis>
</taxonomicName>
5b. Body with dark bars, darkest across peduncle, with black spots above and often below; anal fin with five oblique black bars, prominent subcutaneous black spots at base of anal fin, and black pre-anal spot; belly without scales; pinkish in life...........................................................................................
<taxonomicName id="4C254D133F6FFFD1FB6762C14DB16653" ID-CoL="39VZ3" box="[1222,1435,240,261]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="destai">
<emphasis id="B951EA823F6FFFD1FB6762C14DB16653" box="[1222,1435,240,261]" italics="true" pageId="3" pageNumber="39">Enneapterygius destai</emphasis>
</taxonomicName>
<keyLead id="30D495A53F6FFFD1FF36633C4DB66674" box="[151,1436,269,290]" pageId="3" pageNumber="39">
<keyStep id="30D12E353F6FFFD1FF36633C4DB66674" box="[151,1436,269,290]" pageId="3" pageNumber="39">6a. Anal fin with narrow bars............................................................................... 7</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9A36903F6FFFD1FF36631B4DB66669" blockId="3.[151,1436,151,554]" box="[151,1436,298,319]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF36631B4DB66669" box="[151,1436,298,319]" pageId="3" pageNumber="39">6b. Anal fin without bars................................................................................... 9</keyLead>
</paragraph>
<keyStep id="30D12E353F6FFFD1FF3663764DB166C1" pageId="3" pageNumber="39">
<paragraph id="8B9A36903F6FFFD1FF3663764DB6660A" blockId="3.[151,1436,151,554]" box="[151,1436,327,348]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF3663764DB6660A" box="[151,1436,327,348]" pageId="3" pageNumber="39">7a. First dorsal fin of males higher than second................................................................. 8</keyLead>
</paragraph>
<paragraph id="8B9A36903F6FFFD1FF3663544DB166C1" blockId="3.[151,1436,151,554]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF3663544DB166C1" pageId="3" pageNumber="39">
7b. First dorsal fin subequal to second; head of males dark olive, body olive green with white blotches, anal fin barred black and white.......................................................................
<taxonomicName id="4C254D133F6FFFD1FBD563B34DB166C1" ID-CoL="39W3K" box="[1140,1435,386,407]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="ventermaculus">
<emphasis id="B951EA823F6FFFD1FBD563B34DB166C1" box="[1140,1435,386,407]" italics="true" pageId="3" pageNumber="39">Enneapterygius ventermaculus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D12E353F6FFFD1FF3663AE4DB6657C" pageId="3" pageNumber="39">
<paragraph id="8B9A36903F6FFFD1FF3663AE4DB66684" blockId="3.[151,1436,151,554]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF3663AE4DB66684" pageId="3" pageNumber="39">
8a. Third dorsal fin with 89 rays; supraoccipital sensory canal C-shaped (A); all pectoral-fin rays simple; in life base of ventral fins and fins white, head and body speckled with orange, first dorsal fin with crimson...........
<taxonomicName id="4C254D133F6FFFD1FB04638C4DB66684" ID-CoL="39VYL" box="[1189,1436,445,466]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="altipinnis">
<emphasis id="B951EA823F6FFFD1FB04638C4DB66684" box="[1189,1436,445,466]" italics="true" pageId="3" pageNumber="39">Enneapterygius altipinnis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9A36903F6FFFD1FF3663EB4DB6657C" blockId="3.[151,1436,151,554]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF3663EB4DB6657C" pageId="3" pageNumber="39">
8b. Third dorsal fin with 1011 rays; supraoccipital sensory canal U-shaped (B); base of ventral fins and proximal part of rays black in males; some pectoral-fin rays divided; in life face red, first dorsal fin yellow and sometimes very tall...............................................................................................
<taxonomicName id="4C254D133F6FFFD1FB1060244DB6657C" ID-CoL="6FBJQ" box="[1201,1436,533,554]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="pusillus">
<emphasis id="B951EA823F6FFFD1FB1060244DB6657C" box="[1201,1436,533,554]" italics="true" pageId="3" pageNumber="39">Enneapterygius pusillus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D12E353F6FFFD1FF3661DF4DB16377" pageId="3" pageNumber="39">
<paragraph id="8B9A36903F6FFFD1FF3661DF4DB16355" blockId="3.[151,1436,1006,1294]" box="[151,1435,1006,1027]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF3661DF4DB16355" box="[151,1435,1006,1027]" pageId="3" pageNumber="39">9a. Nape scaled, anal fin with fewer than 20 rays............................................................... 10</keyLead>
</paragraph>
<paragraph id="8B9A36903F6FFFD1FF36663D4DB16377" blockId="3.[151,1436,1006,1294]" box="[151,1435,1036,1057]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF36663D4DB16377" box="[151,1435,1036,1057]" pageId="3" pageNumber="39">
9b. Nape naked, anal fin with 2122 rays...................................................
<taxonomicName id="4C254D133F6FFFD1FB13663D4DB16377" ID-CoL="39W2K" box="[1202,1435,1036,1057]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B951EA823F6FFFD1FB13663D4DB16377" box="[1202,1435,1036,1057]" italics="true" pageId="3" pageNumber="39">Enneapterygius pallidus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D12E353F6FFFD1FF3666184DB6632F" pageId="3" pageNumber="39">
<paragraph id="8B9A36903F6FFFD1FF3666184DB16368" blockId="3.[151,1436,1006,1294]" box="[151,1435,1065,1086]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF3666184DB16368" box="[151,1435,1065,1086]" pageId="3" pageNumber="39">10a. Dentary pores 2 + 2 + 2................................................................................ 11</keyLead>
</paragraph>
<paragraph id="8B9A36903F6FFFD1FF3666774DB6632F" blockId="3.[151,1436,1006,1294]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF3666774DB6632F" pageId="3" pageNumber="39">
10b. Dentary pores 3 + 1 + 3, males bright yellow in life, head with many melanophores to solid black, females with body scales edged in orange-brown, forming narrow irregular bars; orange bar below eye......................
<taxonomicName id="4C254D133F6FFFD1FB6E66554DB6632F" ID-CoL="39VYK" box="[1231,1436,1124,1145]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="abeli">
<emphasis id="B951EA823F6FFFD1FB6E66554DB6632F" box="[1231,1436,1124,1145]" italics="true" pageId="3" pageNumber="39">Enneapterygius abeli</emphasis>
</taxonomicName>
</keyLead>
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</keyStep>
<keyStep id="30D12E353F6FFFD1FF3666B04DB1625B" pageId="3" pageNumber="39">
<paragraph id="8B9A36903F6FFFD1FF3666B04DB163E5" blockId="3.[151,1436,1006,1294]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF3666B04DB163E5" pageId="3" pageNumber="39">
11a. Small black spot on upper caudal peduncle; no patch of melanophores on dorsum; anterior LL scales 1012; males translucent and red in life, first dorsal fin white to cream, females translucent, scales edged in pink...........
<taxonomicName id="4C254D133F6FFFD1FB0466AF4DB163E5" ID-CoL="39W2F" box="[1189,1435,1182,1203]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B951EA823F6FFFD1FB0466AF4DB163E5" box="[1189,1435,1182,1203]" italics="true" pageId="3" pageNumber="39">Enneapterygius obscurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9A36903F6FFFD1FF36668F4DB1625B" blockId="3.[151,1436,1006,1294]" pageId="3" pageNumber="39">
<keyLead id="30D495A53F6FFFD1FF36668F4DB1625B" pageId="3" pageNumber="39">
11b. No black spot on upper caudal peduncle; small oval patch of melanophores on dorsum between 3rd and 4th dorsal-fin spines; anterior LL scales 810; crimson in life, first dorsal fin crimson anteriorly with gold spots, and white posteriorly, males with rectangular blotch of melanophores on midside, below second dorsal fin.................
<taxonomicName id="4C254D133F6FFFD1FBCE66C84D636258" ID-CoL="39W2X" box="[1135,1353,1273,1294]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="39" phylum="Chordata" rank="species" species="qirmiz" status="sp. nov.">
<emphasis id="B951EA823F6FFFD1FBCE66C84D636258" box="[1135,1353,1273,1294]" italics="true" pageId="3" pageNumber="39">Enneapterygius qirmiz</emphasis>
</taxonomicName>
<emphasis id="B951EA823F6FFFD1FAEE66C94DB1625B" bold="true" box="[1359,1435,1272,1293]" pageId="3" pageNumber="39">
<taxonomicNameLabel id="A26257F93F6FFFD1FAEE66C94DB1625B" box="[1359,1435,1272,1293]" pageId="3" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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