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<document id="369B26086715762D44CC0BE78CADA898" ID-DOI="10.5281/zenodo.10396362" ID-ISSN="1942-1354" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707168658757" checkinUser="felipe" docAuthor="Zhang, Jing, Cong, Qian &amp; Grishin, Nick V." docDate="2023" docId="03810139FFE0BB10C0CAF9E1E125B629" docLanguage="en" docName="InsectaMundi.2023.1026.1-115.pdf" docOrigin="Insecta Mundi 2023 (26)" docSource="http://dx.doi.org/10.5281/zenodo.10396362" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Cobalopsis adictys Zhang &amp; Cong &amp; Grishin 2023, new species" docType="treatment" docVersion="1" lastPageNumber="63" masterDocId="FFB87941FFDFBB50C00AFF56E355B262" masterDocTitle="Supplementary Materials and Appendix" masterLastPageNumber="115" masterPageNumber="1" pageNumber="62" updateTime="1707169288913" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="96E9E3E797B73E9735514DBA7206E1EC">Supplementary Materials and Appendix</mods:title>
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<mods:name id="62DCE49346FCF4E87122295D36EFD8F4" type="personal">
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<mods:namePart id="4246A8DF0911484D791B6DFE18CBCEC0">Zhang, Jing</mods:namePart>
<mods:affiliation id="AC0C0521A22525DF3F9435F6FC655611">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:namePart id="5592DAD6919A199046E67B6C5F6819DC">Cong, Qian</mods:namePart>
<mods:affiliation id="CFE4AE914A596C67FC309AF6F19BCE24">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:namePart id="981F5F6D807669C57E0BD988DBBC6FFE">Grishin, Nick V.</mods:namePart>
<mods:affiliation id="09FAB50E9F5ECA53F86C54A9F5B69694">Departments of Biophysics and Biochemistry University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 9050 USA</mods:affiliation>
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<mods:title id="BD9CB6F30ACAD9E61CD9253DCC8421E5">Insecta Mundi</mods:title>
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<mods:date id="8F907B02D3D4EA3D4974D7A472E16B67">2023</mods:date>
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<heading id="D0DF0743FFE0BB6FC0CAF9E1E1E0B4B6" box="[192,693,1719,1748]" fontSize="12" level="2" pageId="63" pageNumber="62" reason="2">
<emphasis id="B95C6C3DFFE0BB6FC0CAF9E1E1E0B4B6" bold="true" box="[192,693,1719,1748]" pageId="63" pageNumber="62">
<taxonomicName id="4C28CBACFFE0BB6FC0CAF9E1E141B4B6" authority="Grishin" authorityName="Zhang &amp; Cong &amp; Grishin" authorityYear="2023" box="[192,532,1719,1748]" class="Insecta" family="Hesperiidae" genus="Cobalopsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="adictys" status="sp. nov.">
<emphasis id="B95C6C3DFFE0BB6FC0CAF9E1E2FDB4B6" bold="true" box="[192,424,1719,1748]" italics="true" pageId="63" pageNumber="62">Cobalopsis adictys</emphasis>
Grishin
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,
<taxonomicNameLabel id="A26FD146FFE0BB6FC22AF9E1E1E0B4B6" box="[544,693,1719,1748]" pageId="63" pageNumber="62" rank="species">new species</taxonomicNameLabel>
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https://zoobank.org/
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<paragraph id="8B97B02FFFE0BB6FC0CAF9ABE14BB57A" blockId="63.[192,934,1719,1816]" box="[192,542,1789,1816]" pageId="63" pageNumber="62">
(
<figureCitation id="1313ACAAFFE0BB6FC0C2F9ABE252B575" box="[200,263,1789,1815]" captionStart="Figure 6" captionStartId="60.[192,261,1378,1402]" captionTargetBox="[193,1439,259,1336]" captionTargetId="figure-250@60.[192,1440,256,1336]" captionTargetPageId="60" captionText="Figure 6. Phylogenetic trees of Moncina (part 2) inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome. See Fig. 1 legend for other notations." pageId="63" pageNumber="62">Fig. 6</figureCitation>
part, 153154, 380381)
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FFE0BB10C0CAF871E7D8B0CD" lastPageId="64" lastPageNumber="63" pageId="63" pageNumber="62" type="diagnosis">
<paragraph id="8B97B02FFFE0BB10C0CAF871E7B5B3AC" blockId="63.[192,1442,1831,1927]" lastBlockId="64.[192,1442,230,462]" lastPageId="64" lastPageNumber="63" pageId="63" pageNumber="62">
<emphasis id="B95C6C3DFFE0BB6FC0CAF871E2BEB523" bold="true" box="[192,491,1831,1858]" pageId="63" pageNumber="62">Definition and diagnosis.</emphasis>
Phylogenetic trees reveal that a number of specimens identified as
<taxonomicName id="4C28CBACFFE0BB6FC4ECF87EE22EB506" authority="(Godman, 1900)" baseAuthorityName="Godman" baseAuthorityYear="1900" class="Insecta" family="Hesperiidae" genus="Cobalopsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="dictys">
<emphasis id="B95C6C3DFFE0BB6FC4ECF87EE6F7B520" box="[1254,1442,1832,1858]" italics="true" pageId="63" pageNumber="62">Cobalopsis dictys</emphasis>
(Godman, 1900)
</taxonomicName>
(
<typeStatus id="54930E8DFFE0BB6FC183F81CE2EEB506" box="[393,443,1866,1892]" pageId="63" pageNumber="62">type</typeStatus>
locality in
<collectingCountry id="F33FF0BFFFE0BB6FC233F81CE1DAB506" box="[569,655,1866,1892]" name="Mexico" pageId="63" pageNumber="62">Mexico</collectingCountry>
:
<collectingRegion id="49EC7ECDFFE0BB6FC293F81CE055B506" box="[665,768,1866,1892]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="63" pageNumber="62">Veracruz</collectingRegion>
,
<collectingCountry id="F33FF0BFFFE0BB6FC306F81CE0DDB506" box="[780,904,1866,1892]" name="Guatemala" pageId="63" pageNumber="62">Guatemala</collectingCountry>
,
<collectingCountry id="F33FF0BFFFE0BB6FC398F81CE75BB506" box="[914,1038,1866,1892]" name="Costa Rica" pageId="63" pageNumber="62">Costa Rica</collectingCountry>
,
<collectingCountry id="F33FF0BFFFE0BB6FC412F81CE721B506" box="[1048,1140,1866,1892]" name="Panama" pageId="63" pageNumber="62">Panama</collectingCountry>
, illustrated specimen from
<collectingCountry id="F33FF0BFFFE0BB6FC0CBF83AE268B5E4" box="[193,317,1900,1926]" name="Guatemala" pageId="63" pageNumber="62">Guatemala</collectingCountry>
) show prominent genetic differentiation from it (
<figureCitation id="1313ACAAFFE0BB6FC360F83AE0F8B5E4" box="[874,941,1900,1926]" captionStart="Figure 6" captionStartId="60.[192,261,1378,1402]" captionTargetBox="[193,1439,259,1336]" captionTargetId="figure-250@60.[192,1440,256,1336]" captionTargetPageId="60" captionText="Figure 6. Phylogenetic trees of Moncina (part 2) inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome. See Fig. 1 legend for other notations." pageId="63" pageNumber="62">Fig. 6</figureCitation>
): e.g., their COI barcodes differ by 3.3% (22 bp), and therefore represent a new species. This new species keys to “
<taxonomicName id="4C28CBACFF9FBB10C3C1FFB1E70CB362" authorityName="Godman" authorityYear="1900" box="[971,1113,230,256]" class="Insecta" family="Hesperiidae" genus="Papias" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="63" phylum="Arthropoda" rank="species" species="dictys">
<emphasis id="B95C6C3DFF9FBB10C3C1FFB1E70CB362" box="[971,1113,230,256]" italics="true" pageId="64" pageNumber="63">Papias dictys</emphasis>
</taxonomicName>
” (J.36.6) in
<bibRefCitation id="EFB9CDDEFF9FBB10C4E8FFB0E621B363" author="Evans WH" box="[1250,1396,230,257]" pageId="64" pageNumber="63" pagination="54 - 88" refId="ref77169" refString="Evans WH. 1955. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History); London. v + 499 p., pl. 54 - 88." type="book chapter" year="1955">Evans (1955)</bibRefCitation>
but differs from it by comparatively darker ventral forewing and hindwing area by inner margin, vestigial or absent forewing apical spot (
<figureCitation id="1313ACAAFF9FBB10C1BFFE7CE11EB327" box="[437,587,298,325]" captionStart="Figures 149172" captionStartId="102.[192,271,1810,1834]" captionTargetBox="[198,1437,224,1779]" captionTargetId="figure-87@102.[192,1440,221,1781]" captionTargetPageId="102" captionText="Figures 149172. Holotypes, data in text. 149150) Amblyscirtes (Flor) meridus sp. n. 151152) Rectava chiriquensis sp. n. 153154) Cobalopsis adictys sp. n. 155156) Cymaenes melaporphyrus sp. n. 157158) Lerema (Morys) ecuadorica sp. n. 159160) Saturnus obscurior sp. n. 161162) Cantha zoirodicta sp. n. 163164) Cantha meiodicta sp. n. 165166) Phlebodes duplex sp. n. 167168) Lychnuchus (Enosis) valle sp. n. 169170) Eutychide ochoides sp. n. 171172) Dion bora sp. n." pageId="64" pageNumber="63">Fig. 153154</figureCitation>
), smaller tooth on ventral margin of harpe, distal margin of harpe stronger projecting dorsad, and more angled connection between tegumen and uncus in lateral view (
<figureCitation id="1313ACAAFF9FBB10C4EAFE1AE66AB305" box="[1248,1343,332,359]" captionStart="Figures 361390" captionStartId="113.[192,270,1710,1734]" captionTargetBox="[196,1434,221,1688]" captionTargetId="figure-177@113.[192,1440,216,1689]" captionTargetPageId="113" captionText="Figures 361390. Genitalia of holotypes (unless indicated), data in text. 361363) Niconiades victoria sp. n. paratype ♂ NVG-20057B04. 364365) Lancephallus purpurus sp.n.366367) Mnasicles (Remella) ecua sp. n.368369) Amblyscirtes (Amblyscirtes) aeratus sp. n. 370371) Amblyscirtes (Mastor) chrysoplea sp. n. 372374) Amblyscirtes (Mastor) chrysomisa sp. n. 375376) Amblyscirtes (Flor) meridus sp. n. 377379) Rectava chiriquensis sp. n. 380381) Cobalopsis adictys sp. n. 382383) Cymaenes melaporphyrus sp. n. 384385) Lerema (Morys) ecuadorica sp. n. 386387) Saturnus obscurior sp. n. 388390) Cantha zoirodicta sp. n. Views:dorsal (361, 364, 366, 368, 370, 375, 377, 380, 383, 384, 386, 388), left lateral (362 left valva removed, 365, 367, 369, 371, 376, 379, 381, 383, 385, 390), right lateral of left valva (363, 378, 389) and genital capsule, a flipped image" pageId="64" pageNumber="63">Fig. 381</figureCitation>
). Due to the cryptic nature of this species, most reliable identification is achieved by DNA and a combination of the following base pairs is diagnostic in the nuclear genome: aly
<date id="FF9696EFFF9FBB10C354FEC7E095B3CE" box="[862,960,401,428]" pageId="64" pageNumber="63" value="1905-05-02">1905.5.2</date>
:A148G, aly260.3.2:A28T, aly260.3.2:T51C, aly451.31.3:C84T, aly318.41.3:G78A, and COI barcode: A100T, T112C, C343T, T445C, T536C.
</paragraph>
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<emphasis id="B95C6C3DFF9FBB10C0CAFE88E11BB39A" bold="true" box="[192,590,477,504]" pageId="64" pageNumber="63">
Barcode sequence of the
<typeStatus id="54930E8DFF9FBB10C1EAFE88E11CB39A" box="[480,585,478,504]" pageId="64" pageNumber="63" type="holotype">holotype</typeStatus>
.
</emphasis>
Sample NVG-19018H05, GenBank OR837691, 658 base pairs: AACTTTATACTTCATCTTTGGAATTTGAGCCGGAATATTAGGAACTTCTTTAAGATTATTAATTCGAACAGAATTAGGTAATCCTGGATCTTTAATT GGTGATGATCAAATCTATAATACTATTGTAACAGCTCATGCTTTTATTATAATTTTTTTTATAGTTATACCTATTATAATTGGAGGATTTGGAAATT GATTAGTACCCCTAATACTTGGTGCTCCTGATATAGCTTTCCCACGAATAAATAATATAAGTTTTTGAATACTACCACCTTCCTTAATATTACTAAT TTCAAGAAGAATTGTAGAAAATGGGGCAGGTACTGGTTGAACTGTATACCCTCCTCTTTCTTCTAATATTGCACATCAAGGAGCATCTGTTGATTTA GCAATTTTTTCTCTTCATTTAGCTGGTATTTCTTCTATTCTAGGAGCTATTAATTTCATTACTACAATTATTAATATACGAATTAAAAATTTATCTT TTGATCAAATACCTTTATTTGTCTGATCAGTTGGAATTACAGCTTTACTACTACTATTATCTTTACCTGTACTAGCAGGAGCTATTACTATACTTCT TACTGATCGAAATTTAAATACTTCTTTTTTTGATCCTGCAGGAGGAGGAGATCCTATTTTATACCAACATTTATTT
</paragraph>
</subSubSection>
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<paragraph id="8B97B02FFF9FBB10C0CAFD94E634B1C8" blockId="64.[191,1441,706,938]" pageId="64" pageNumber="63">
<emphasis id="B95C6C3DFF9FBB10C0CAFD94E230B0BE" bold="true" box="[192,357,706,732]" pageId="64" pageNumber="63">Type material.</emphasis>
<materialsCitation id="3B40BA72FF9FBB10C160FD94E2D3B104" collectingDate="1981-09-12" collectionCode="DC, USNM, V" collectorName="G. B. Small &amp; Nick &amp; Grishin &amp; QR Code" country="Panama" latitude="8.516666" location="Smithsonian Institution" longLatPrecision="1301" longitude="-81.083336" municipality="National Museum of Natural History" pageId="64" pageNumber="63" specimenCount="2" specimenCount-male="2" stateProvince="Veraguas" typeStatus="holotype">
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<typeStatus id="54930E8DFF9FBB10C160FD94E282B0BE" box="[362,471,706,732]" pageId="64" pageNumber="63" type="holotype">Holotype</typeStatus>
:
</emphasis>
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deposited in the
<collectingMunicipality id="6BF32A55FF9FBB10C2A5FD94E714B0BE" box="[687,1089,706,733]" pageId="64" pageNumber="63">National Museum of Natural History</collectingMunicipality>
,
<location id="8EF7E6F4FF9FBB10C440FD94E605B0BE" LSID="urn:lsid:plazi:treatment:03810139FFE0BB10C0CAF9E1E125B629:8EF7E6F4FF9FBB10C440FD94E605B0BE" box="[1098,1360,706,732]" country="Panama" latitude="8.516666" longLatPrecision="1301" longitude="-81.083336" municipality="National Museum of Natural History" name="Smithsonian Institution" pageId="64" pageNumber="63" stateProvince="Veraguas">Smithsonian Institution</location>
, Washington,
<collectionCode id="ED3928EAFF9FBB10C11DFDB2E215B09C" box="[279,320,740,766]" pageId="64" pageNumber="63">DC</collectionCode>
, USA (
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), illustrated in
<figureCitation id="1313ACAAFF9FBB10C299FDB2E070B09D" box="[659,805,740,767]" captionStart="Figures 149172" captionStartId="102.[192,271,1810,1834]" captionTargetBox="[198,1437,224,1779]" captionTargetId="figure-87@102.[192,1440,221,1781]" captionTargetPageId="102" captionText="Figures 149172. Holotypes, data in text. 149150) Amblyscirtes (Flor) meridus sp. n. 151152) Rectava chiriquensis sp. n. 153154) Cobalopsis adictys sp. n. 155156) Cymaenes melaporphyrus sp. n. 157158) Lerema (Morys) ecuadorica sp. n. 159160) Saturnus obscurior sp. n. 161162) Cantha zoirodicta sp. n. 163164) Cantha meiodicta sp. n. 165166) Phlebodes duplex sp. n. 167168) Lychnuchus (Enosis) valle sp. n. 169170) Eutychide ochoides sp. n. 171172) Dion bora sp. n." pageId="64" pageNumber="63">Fig. 153154</figureCitation>
, bears the following four rectangular labels, three white: [
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:
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| Sante Fe |
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|
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</date>
| leg.
<collectorName id="26DDD5F9FF9FBB10C3E3FC50E702B142" box="[1001,1111,774,800]" pageId="64" pageNumber="63">G.B.Small</collectorName>
], [DNA sample ID: | NVG-19018H05 | c/o
<collectorName id="26DDD5F9FF9FBB10C160FC7FE2CBB121" box="[362,414,809,835]" pageId="64" pageNumber="63">Nick</collectorName>
<collectionCode id="ED3928EAFF9FBB10C1A9FC7FE2E3B121" box="[419,438,809,835]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="64" pageNumber="63" type="Museum">V</collectionCode>
.
<collectorName id="26DDD5F9FF9FBB10C1B4FC7FE147B121" box="[446,530,809,835]" pageId="64" pageNumber="63">Grishin</collectorName>
], [USNMENT | {
<collectorName id="26DDD5F9FF9FBB10C2DFFC7FE06AB121" box="[725,831,809,835]" pageId="64" pageNumber="63">QR Code</collectorName>
} | 01532536], and one red [
<typeStatus id="54930E8DFF9FBB10C467FC7FE7A9B121" box="[1133,1276,809,835]" pageId="64" pageNumber="63" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D2E7BA6FF9FBB10C50BFC7CE643B124" box="[1281,1302,810,838]" count="1" pageId="64" pageNumber="63" type="male"></specimenCount>
|
<taxonomicName id="4C28CBACFF9FBB10C52DFC7FE6F5B121" authorityName="Godman" authorityYear="1900" box="[1319,1440,809,835]" class="Insecta" family="Hesperiidae" genus="Cobalopsis" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="63" phylum="Arthropoda" rank="genus">Cobalopsis</taxonomicName>
|
<taxonomicName id="4C28CBACFF9FBB10C0C7FC1DE221B107" authority="Grishin" authorityName="Zhang &amp; Cong &amp; Grishin" authorityYear="2023" box="[205,372,843,869]" class="Insecta" family="Hesperiidae" genus="Cobalopsis" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="63" phylum="Arthropoda" rank="species" species="adictys">adictys Grishin</taxonomicName>
]
</materialsCitation>
.
<materialsCitation id="3B40BA72FF9FBB10C198FC1DE3A2B1C8" collectingDate="2011-11-17" collectionCode="USNM, GPS" collectorName="de Conservacion" country="Costa Rica" elevation="710" latitude="10.9845" location="Area de Conservacion" longLatPrecision="7" longitude="-85.4248" pageId="64" pageNumber="63" specimenCount="2" specimenCount-male="2" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="B95C6C3DFF9FBB10C198FC1DE15CB107" bold="true" box="[402,521,843,869]" pageId="64" pageNumber="63">
<typeStatus id="54930E8DFF9FBB10C198FC1DE151B107" box="[402,516,843,869]" pageId="64" pageNumber="63" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D2E7BA6FF9FBB10C205FC1DE113B10A" box="[527,582,843,872]" count="2" pageId="64" pageNumber="63" type="male">2♂♂</specimenCount>
from
<collectingCountry id="F33FF0BFFF9FBB10C280FC1DE056B107" box="[650,771,843,869]" name="Costa Rica" pageId="64" pageNumber="63">Costa Rica</collectingCountry>
:
<location id="8EF7E6F4FF9FBB10C307FC1DE754B107" LSID="urn:lsid:plazi:treatment:03810139FFE0BB10C0CAF9E1E125B629:8EF7E6F4FF9FBB10C307FC1DE754B107" box="[781,1025,843,869]" country="Costa Rica" latitude="10.9845" longLatPrecision="7" longitude="-85.4248" name="Area de Conservacion" pageId="64" pageNumber="63" stateProvince="Guanacaste">
Area
<collectorName id="26DDD5F9FF9FBB10C342FC1DE754B107" box="[840,1025,843,869]" pageId="64" pageNumber="63">de Conservación</collectorName>
</location>
<collectingRegion id="49EC7ECDFF9FBB10C402FC1DE7DDB107" box="[1032,1160,843,869]" country="Costa Rica" name="Guanacaste" pageId="64" pageNumber="63">Guanacaste</collectingRegion>
,
<collectingRegion id="49EC7ECDFF9FBB10C498FC1DE604B104" box="[1170,1361,843,870]" country="Costa Rica" name="Guanacaste" pageId="64" pageNumber="63">Guanacaste Prov.</collectingRegion>
, Sector Pitilla [
<collectionCode id="ED3928EAFF9FBB10C106FC3BE236B1E5" box="[268,355,877,903]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="64" pageNumber="63" type="Museum">USNM</collectionCode>
]: NVG-18064B08, 11-SRNP-33187 Sendero Nacho,
<quantity id="4CD01DCAFF9FBB10C39DFC3BE088B1EA" box="[919,989,877,904]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="64" pageNumber="63" unit="m" value="710.0">
<elevation id="0005571CFF9FBB10C39DFC3BE088B1EA" box="[919,989,877,904]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="64" pageNumber="63" unit="m" value="710.0">710 m</elevation>
</quantity>
,
<collectionCode id="ED3928EAFF9FBB10C3EDFC3BE742B1E5" box="[999,1047,877,903]" pageId="64" pageNumber="63">GPS</collectionCode>
<geoCoordinate id="EE1CD6E8FF9FBB10C417FC3BE721B1EA" box="[1053,1140,877,904]" degrees="10.9845" direction="north" orientation="latitude" pageId="64" pageNumber="63" precision="5" value="10.9845">10.9845</geoCoordinate>
,
<geoCoordinate id="EE1CD6E8FF9FBB10C477FC3BE7B0B1EA" box="[1149,1253,877,904]" degrees="85.4248" direction="west" orientation="longitude" pageId="64" pageNumber="63" precision="5" value="-85.4248">85.4248</geoCoordinate>
, eclosed
<date id="FF9696EFFF9FBB10C54EFC3BE3A2B1C8" pageId="64" pageNumber="63" value="2011-11-17">
<collectingDate id="EFD26F07FF9FBB10C54EFC3BE3A2B1C8" pageId="64" pageNumber="63" value="2011-11-17">17-Nov- 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B40BA72FF9FBB10C10BFCD9E609B1C8" box="[257,1372,911,938]" collectingDate="2011-03-03" collectionCode="GPS" country="Costa Rica" elevation="680" latitude="10.9888" longLatPrecision="7" longitude="-85.4234" pageId="64" pageNumber="63" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
NVG-18064C02, 11-SRNP-30141 Sendero Laguna
<quantity id="4CD01DCAFF9FBB10C327FCD9E027B1C8" box="[813,882,911,938]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="64" pageNumber="63" unit="m" value="680.0">
<elevation id="0005571CFF9FBB10C327FCD9E027B1C8" box="[813,882,911,938]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="64" pageNumber="63" unit="m" value="680.0">680 m</elevation>
</quantity>
<collectionCode id="ED3928EAFF9FBB10C37DFCC6E0F2B1C8" box="[887,935,912,938]" pageId="64" pageNumber="63">GPS</collectionCode>
<geoCoordinate id="EE1CD6E8FF9FBB10C3A4FCC6E750B1C8" box="[942,1029,912,938]" degrees="10.9888" direction="north" orientation="latitude" pageId="64" pageNumber="63" precision="5" value="10.9888">10.9888</geoCoordinate>
,
<geoCoordinate id="EE1CD6E8FF9FBB10C405FCD9E723B1C8" box="[1039,1142,911,938]" degrees="85.4234" direction="west" orientation="longitude" pageId="64" pageNumber="63" precision="5" value="-85.4234">85.4234</geoCoordinate>
, eclosed
<date id="FF9696EFFF9FBB10C4DDFCC6E609B1C8" box="[1239,1372,912,938]" pageId="64" pageNumber="63" value="2011-03-03">
<collectingDate id="EFD26F07FF9FBB10C4DDFCC6E609B1C8" box="[1239,1372,912,938]" pageId="64" pageNumber="63" value="2011-03-03">3-Mar-2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B97B02FFF9FBB10C0CAFCECE75FB1B6" blockId="64.[192,1034,953,980]" box="[192,1034,953,980]" pageId="64" pageNumber="63">
<emphasis id="B95C6C3DFF9FBB10C0CAFCECE20EB1B1" bold="true" box="[192,347,953,980]" pageId="64" pageNumber="63">
<typeStatus id="54930E8DFF9FBB10C0CAFCECE3ACB1B6" box="[192,249,954,980]" pageId="64" pageNumber="63">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33FF0BFFF9FBB10C16BFCECE2EBB1B6" box="[353,446,954,980]" name="Panama" pageId="64" pageNumber="63">Panama</collectingCountry>
:
<collectingRegion id="49EC7ECDFF9FBB10C1C2FCECE1CCB1B6" box="[456,665,954,980]" country="Panama" name="Veraguas" pageId="64" pageNumber="63">Veraguas Province</collectingRegion>
,
<taxonomicName id="4C28CBACFF9FBB10C2A9FCECE06AB1B6" authority="Fe, GPS" authorityName="Fe, GPS" box="[675,831,954,980]" class="Insecta" family="Hesperiidae" genus="Nisoniades" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="63" phylum="Arthropoda" rank="species" species="santa">Santa Fe, GPS</taxonomicName>
<geoCoordinate id="EE1CD6E8FF9FBB10C34FFCEFE0C4B1B6" box="[837,913,953,980]" degrees="8.5167" direction="north" orientation="latitude" pageId="64" pageNumber="63" precision="5" value="8.5167">8.5167</geoCoordinate>
,
<geoCoordinate id="EE1CD6E8FF9FBB10C391FCECE753B1B6" box="[923,1030,954,980]" degrees="81.0833" direction="west" orientation="longitude" pageId="64" pageNumber="63" precision="5" value="-81.0833">81.0833</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FF9FBB10C0CAFCB2E7DFB642" pageId="64" pageNumber="63" type="etymology">
<paragraph id="8B97B02FFF9FBB10C0CAFCB2E7DFB642" blockId="64.[192,1440,996,1057]" pageId="64" pageNumber="63">
<emphasis id="B95C6C3DFF9FBB10C0CAFCB2E211B19C" bold="true" box="[192,324,996,1022]" pageId="64" pageNumber="63">Etymology.</emphasis>
The name is derived by combining
<taxonomicName id="4C28CBACFF9FBB10C2C4FCB2E058B19C" authorityName="Godman" authorityYear="1900" box="[718,781,996,1022]" class="Insecta" family="Hesperiidae" genus="Papias" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="63" phylum="Arthropoda" rank="species" species="dictys">
<emphasis id="B95C6C3DFF9FBB10C2C4FCB2E058B19C" box="[718,781,996,1022]" italics="true" pageId="64" pageNumber="63">dictys</emphasis>
</taxonomicName>
with the negating prefix “a”. As a mnemonic, a longer name indicates it is a more southern species than
<taxonomicName id="4C28CBACFF9FBB10C2AEFB51E057B642" authorityName="Godman" authorityYear="1900" box="[676,770,1030,1057]" class="Insecta" family="Hesperiidae" genus="Papias" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="63" phylum="Arthropoda" rank="species" species="dictys">
<emphasis id="B95C6C3DFF9FBB10C2AEFB51E057B642" box="[676,770,1030,1057]" italics="true" pageId="64" pageNumber="63">C. dictys</emphasis>
</taxonomicName>
. The name is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FF9FBB10C0CAFB66E125B629" box="[192,624,1072,1099]" pageId="64" pageNumber="63" type="distribution">
<paragraph id="8B97B02FFF9FBB10C0CAFB66E125B629" blockId="64.[192,624,1072,1099]" box="[192,624,1072,1099]" pageId="64" pageNumber="63">
<emphasis id="B95C6C3DFF9FBB10C0CAFB66E20EB628" bold="true" box="[192,347,1072,1098]" pageId="64" pageNumber="63">Distribution.</emphasis>
<collectingCountry id="F33FF0BFFF9FBB10C16BFB67E28DB628" box="[353,472,1072,1099]" name="Costa Rica" pageId="64" pageNumber="63">Costa Rica</collectingCountry>
and
<collectingCountry id="F33FF0BFFF9FBB10C205FB67E139B629" box="[527,620,1073,1099]" name="Panama" pageId="64" pageNumber="63">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>