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<document id="0254F1035E32D2240EFABEACF77349FE" ID-DOI="10.11646/zootaxa.5352.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8406992" ID-ZooBank="3F9D6C62-9F76-4663-B9E2-77EAF716C278" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1696427295170" checkinUser="plazi" docAuthor="Donnellan, S. C., Catullo, R. A., Rowley, J. J. L., Doughty, P., Price, L., Hines, H. B. &amp; Richards, S. J." docDate="2023" docId="515EC77E1C1E5A00FF4CFE7FFE3A9653" docLanguage="en" docName="zootaxa.5352.1.3.pdf" docOrigin="Zootaxa 5352 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5352.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Litoria ridibunda Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="103" masterDocId="AD67BF061C045A1EFFDBFFA4FF9F910B" masterDocTitle="Revision of Litoria rothii (Anura: Pelodryadidae) from northern Australia" masterLastPageNumber="108" masterPageNumber="73" pageNumber="99" updateTime="1696427851397" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A039306F4D8108AAFD75BEE2452665D9">Revision of Litoria rothii (Anura: Pelodryadidae) from northern Australia</mods:title>
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<mods:namePart id="326EE2C6D6C1F402FD3E84FEB4827F70">Donnellan, S. C.</mods:namePart>
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<mods:affiliation id="5210BC7BC6EA1ED7F9B8464066319F32">Australian Museum Research Institute, Australian Museum, 1 William St, Sydney, NSW, 2010, Australia Jodi. Rowley @ austmus. gov. au; https: // orcid. org / 0000 - 0002 - 2011 - 9143 &amp; Centre for Ecosystem Science, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, NSW, 2052, Australia</mods:affiliation>
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<mods:affiliation id="0AF202EA79F55EDFC0A8BBBD6A1CE441">Collections &amp; Research, Western Australian Museum, 49 Kew St, Welshpool, WA, 6106 Australia paul. doughty @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 0631 - 0571</mods:affiliation>
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<mods:affiliation id="058E20390C2B0C86E43C4617641B9126">Queensland Parks and Wildlife Service and Partnerships, Department of Environment and Science, PO Box 64, Bellbowrie, Qld, 4070, Australia &amp; Honorary Research Fellow, Biodiversity, Queensland Museum, PO Box 3300, South Brisbane, Qld, 4101, Australia</mods:affiliation>
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<taxonomicName id="1EF70DEB1C1E5A04FF4CFE7FFEF690FE" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[151,361,474,501]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="26" pageNumber="99" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C1E5A04FF4CFE7FFEF690FE" bold="true" box="[151,361,474,501]" italics="true" pageId="26" pageNumber="99">Litoria ridibunda</emphasis>
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<paragraph id="D94876681C1E5A04FF4CFE5BFE469312" blockId="26.[151,968,474,609]" box="[151,473,511,537]" pageId="26" pageNumber="99">Western Laughing Tree Frog</paragraph>
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<paragraph id="D94876681C1E5A04FF4CFD87FE8B9336" blockId="26.[151,968,474,609]" box="[151,276,547,573]" pageId="26" pageNumber="99">
<figureCitation id="41CC6AED1C1E5A04FF4CFD87FF419336" box="[151,222,547,573]" captionStart="FIGURE 4" captionStartId="16.[151,250,1839,1863]" captionTargetBox="[324,1261,181,1814]" captionTargetId="figure-20@16.[324,1262,181,1814]" captionTargetPageId="16" captionText="FIGURE 4. Posterior thighs showing diagnostic patterns. Litoria ridibunda sp. nov.—A) NTM R38969, Groote Eylandt, NT (Jodi Rowley); B) no voucher, Lawn Hill, Qld 18.65°S 138.55°E (Harry Hines); C) WAM R174146, King Edward River, WA (Ryan Ellis); D) AMS R184860, Bradshaw, NT (Jodi Rowley); E) NTM R38968, Groote Eylandt, NT (Jodi Rowley); F) AMS R188538, Groote Eylandt, NT (Jodi Rowley). Litoria rothii—G) QM J84287, 5 km SE Miriam Vale, Qld (Harry Hines); H) QM J86621, 15km ESE Biggenden, Qld (Harry Hines); I) SAMA R63683, 41 km SE Normanton, Qld; J) SAMA R20230, 13 km SW Townsville, Qld; K) SAMA R20231, 13 km SW Townsville, Qld. In life AH, preserved specimen IK." figureDoi="http://doi.org/10.5281/zenodo.8407000" httpUri="https://zenodo.org/record/8407000/files/figure.png" pageId="26" pageNumber="99">Figs 4</figureCitation>
,
<figureCitation id="41CC6AED1C1E5A04FF31FD87FF669336" box="[234,249,547,573]" captionStart="FIGURE 8" captionStartId="26.[151,250,1878,1902]" captionTargetBox="[151,1435,741,1853]" captionTargetId="figure-168@26.[151,1435,741,1853]" captionTargetPageId="26" captionText="FIGURE 8. Holotype of L. ridibunda sp. nov. (NTM R36917) in life: A) latero-dorsal view (Stephen Richards), B) laterodorsal view of head (Stephen Richards), C) posterior of thigh (Stephen Richards), D) groin (Stephen Richards), E) plantar view of foot, F) palmar view of hand." figureDoi="http://doi.org/10.5281/zenodo.8407008" httpUri="https://zenodo.org/record/8407008/files/figure.png" pageId="26" pageNumber="99">8</figureCitation>
,
<figureCitation id="41CC6AED1C1E5A04FEDDFD87FE8B9336" box="[262,276,547,573]" captionStart="FIGURE 9" captionStartId="27.[151,250,1908,1932]" captionTargetBox="[168,1419,181,1884]" captionTargetId="figure-17@27.[168,1419,181,1884]" captionTargetPageId="27" captionText="FIGURE 9. Photos in life of Litoria ridibunda sp. nov. from A) Wyndham, WA (Marion Anstis); B) WAM R167789, Surveyors Pool, Mitchell Plateau, WA (Paul Doughty); C) Red Cone Creek, WA (Paul Doughty); D) Channel Island Road, outer Darwin, NT (Stephen Richards); E) Ringwood, NT (Stephen Richards); F) Fogg Dam, NT (Matt Greenlees); G) Mitchell Plateau, WA (Jen Francis)." figureDoi="http://doi.org/10.5281/zenodo.8407010" httpUri="https://zenodo.org/record/8407010/files/figure.png" pageId="26" pageNumber="99">9</figureCitation>
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<materialsCitation id="699F7C351C1E5A04FF4CFD2BFCB493C6" collectingDate="2013-01-03" collectionCode="NTM" collectorName="Stephen J Richards." country="Australia" county="Darwin" location="Channel Island Road" municipality="Channel Island" pageId="26" pageNumber="99" specimenCode="R36917" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
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.
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<collectionCode id="BFE6EEAD1C1E5A04FECBFD2BFED193A2" box="[272,334,655,681]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="26" pageNumber="99">NTM</collectionCode>
<specimenCode id="8951DE131C1E5A04FE88FD2BFE3193A2" box="[339,430,655,681]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="99">R36917</specimenCode>
, an adult male collected at
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Road
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, outer
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,
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,
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(-12.58; 130.95) on
<date id="AD4950A81C1E5A04FEA3FD17FDBC93C6" box="[376,547,691,717]" pageId="26" pageNumber="99" value="2013-01-03">
<collectingDate id="BD0DA9401C1E5A04FEA3FD17FDBC93C6" box="[376,547,691,717]" pageId="26" pageNumber="99" value="2013-01-03">3 January 2013</collectingDate>
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by
<collectorName id="740213BE1C1E5A04FD96FD17FCB493C6" box="[589,811,691,717]" pageId="26" pageNumber="99">Stephen J Richards.</collectorName>
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<paragraph id="D94876681C1E5A04FF4CF8F2FE4096BD" blockId="26.[151,1436,1878,1974]" pageId="26" pageNumber="99">
<emphasis id="EB83AA7A1C1E5A04FF4CF8F2FE8A9665" bold="true" box="[151,277,1878,1902]" pageId="26" pageNumber="99">FIGURE 8.</emphasis>
Holotype of
<taxonomicName id="1EF70DEB1C1E5A04FE79F8F3FDBC9665" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[418,547,1878,1902]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="26" pageNumber="99" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C1E5A04FE79F8F3FDBC9665" box="[418,547,1878,1902]" italics="true" pageId="26" pageNumber="99">L. ridibunda</emphasis>
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<taxonomicNameLabel id="F0B017011C1E5A04FDF0F8F2FDE19665" box="[555,638,1878,1902]" pageId="26" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
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(NTM R36917) in life:
<emphasis id="EB83AA7A1C1E5A04FCA4F8F2FC069665" bold="true" box="[895,921,1878,1902]" pageId="26" pageNumber="99">A)</emphasis>
latero-dorsal view (Stephen Richards),
<emphasis id="EB83AA7A1C1E5A04FAE3F8F2FACD9665" bold="true" box="[1336,1362,1878,1902]" pageId="26" pageNumber="99">B)</emphasis>
laterodorsal view of head (Stephen Richards),
<emphasis id="EB83AA7A1C1E5A04FDEAF8DEFDD39699" bold="true" box="[561,588,1914,1938]" pageId="26" pageNumber="99">C)</emphasis>
posterior of thigh (Stephen Richards),
<emphasis id="EB83AA7A1C1E5A04FC0EF8DEFC789699" bold="true" box="[981,999,1914,1938]" pageId="26" pageNumber="99">D</emphasis>
) groin (Stephen Richards),
<emphasis id="EB83AA7A1C1E5A04FADBF8DEFA8E9699" bold="true" box="[1280,1297,1914,1938]" pageId="26" pageNumber="99">E</emphasis>
) plantar view of foot,
<emphasis id="EB83AA7A1C1E5A04FF33F83AFF6896BD" bold="true" box="[232,247,1950,1974]" pageId="26" pageNumber="99">F</emphasis>
) palmar view of hand.
</paragraph>
</caption>
<caption id="8D8826E01C1F5A05FF4CF8D0FEBD96F3" ID-DOI="http://doi.org/10.5281/zenodo.8407010" ID-Zenodo-Dep="8407010" httpUri="https://zenodo.org/record/8407010/files/figure.png" pageId="27" pageNumber="100" startId="27.[151,250,1908,1932]" targetBox="[168,1419,181,1884]" targetPageId="27" targetType="figure">
<paragraph id="D94876681C1F5A05FF4CF8D0FEBD96F3" blockId="27.[151,1436,1908,2041]" pageId="27" pageNumber="100">
<emphasis id="EB83AA7A1C1F5A05FF4CF8D0FE8D9687" bold="true" box="[151,274,1908,1932]" pageId="27" pageNumber="100">FIGURE 9.</emphasis>
Photos in life of
<taxonomicName id="1EF70DEB1C1F5A05FE66F8D1FDF29687" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[445,621,1908,1932]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="27" pageNumber="100" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C1F5A05FE66F8D1FDF29687" box="[445,621,1908,1932]" italics="true" pageId="27" pageNumber="100">Litoria ridibunda</emphasis>
</taxonomicName>
<emphasis id="EB83AA7A1C1F5A05FDA8F8D1FD5D9686" bold="true" box="[627,706,1909,1933]" pageId="27" pageNumber="100">
<taxonomicNameLabel id="F0B017011C1F5A05FDA8F8D1FD5D9686" box="[627,706,1909,1933]" pageId="27" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<emphasis id="EB83AA7A1C1F5A05FD24F8D0FC859687" bold="true" box="[767,794,1908,1932]" pageId="27" pageNumber="100">A)</emphasis>
Wyndham, WA (Marion Anstis);
<emphasis id="EB83AA7A1C1F5A05FBB0F8D0FB1A9687" bold="true" box="[1131,1157,1908,1932]" pageId="27" pageNumber="100">B)</emphasis>
WAM R167789, Surveyors Pool, Mitchell Plateau, WA (Paul Doughty);
<emphasis id="EB83AA7A1C1F5A05FD85F83CFDE696BB" bold="true" box="[606,633,1944,1968]" pageId="27" pageNumber="100">C)</emphasis>
Red Cone Creek, WA (Paul Doughty);
<emphasis id="EB83AA7A1C1F5A05FBD7F83CFBB996BB" bold="true" box="[1036,1062,1944,1968]" pageId="27" pageNumber="100">D)</emphasis>
Channel Island Road, outer Darwin, NT (Stephen Richards);
<emphasis id="EB83AA7A1C1F5A05FE4AF818FE3596DF" bold="true" box="[401,426,1980,2004]" pageId="27" pageNumber="100">E)</emphasis>
Ringwood, NT (Stephen Richards);
<emphasis id="EB83AA7A1C1F5A05FCF8F818FCA496DF" bold="true" box="[803,827,1980,2004]" pageId="27" pageNumber="100">F)</emphasis>
Fogg Dam, NT (Matt Greenlees);
<emphasis id="EB83AA7A1C1F5A05FB44F818FB2596DF" bold="true" box="[1183,1210,1980,2004]" pageId="27" pageNumber="100">G)</emphasis>
Mitchell Plateau, WA (Jen Francis).
</paragraph>
</caption>
<paragraph id="D94876681C185A02FF1CFF32FCD491BA" blockId="28.[151,1437,150,1977]" box="[199,843,150,177]" pageId="28" pageNumber="101">
<emphasis id="EB83AA7A1C185A02FF1CFF32FE6391BB" bold="true" box="[199,508,150,176]" pageId="28" pageNumber="101">Other material examined.</emphasis>
See Supplementary
<tableCitation id="947543D31C185A02FD39FF33FCD891BA" box="[738,839,151,177]" captionStart="TABLE 2" captionStartId="13.[151,239,582,608]" captionText="TABLE 2. Apomorphic diagnostic sites in the mitochondrial ND4 alignment of the Litoria peronii Group. Sites are numbered with reference to the Nyctimystes kubori ND4 and flanking tRNAs (GenBank accession JX564879). Bolding indicates apomorphic character states polarised with the remaining members of the Litoria peronii Group as the outgroups." pageId="28" pageNumber="101">Table S2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="91ED25E31C185A02FF1CFF1EFC03921E" pageId="28" pageNumber="101" type="diagnosis">
<paragraph id="D94876681C185A02FF1CFF1EFB36906E" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">
<emphasis id="EB83AA7A1C185A02FF1CFF1EFEDD91DF" bold="true" box="[199,322,186,212]" pageId="28" pageNumber="101">Diagnosis.</emphasis>
Distinguished from other Australian
<taxonomicName id="1EF70DEB1C185A02FD33FF1FFCA991DF" authorityName="Tschudi" authorityYear="1838" box="[744,822,187,212]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="genus">
<emphasis id="EB83AA7A1C185A02FD33FF1FFCA991DF" box="[744,822,187,212]" italics="true" pageId="28" pageNumber="101">Litoria</emphasis>
</taxonomicName>
(except other members of the
<taxonomicName id="1EF70DEB1C185A02FB4AFF1FFA9F91DF" box="[1169,1280,187,212]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="peronii">
<emphasis id="EB83AA7A1C185A02FB4AFF1FFA9F91DF" box="[1169,1280,187,212]" italics="true" pageId="28" pageNumber="101">L. peronii</emphasis>
</taxonomicName>
Group,
<emphasis id="EB83AA7A1C185A02FA85FF1FFA0391DF" box="[1374,1436,187,212]" italics="true" pageId="28" pageNumber="101">sensu</emphasis>
<bibRefCitation id="BD660B991C185A02FF4CFF7BFE0F91F2" author="Tyler, M. J. &amp; Davies, M. M." box="[151,400,223,249]" pageId="28" pageNumber="101" pagination="1 - 47" refId="ref22917" refString="Tyler, M. J. &amp; Davies, M. M. (1978) Species-groups within the Australopapuan hylid frog genus Litoria Tschudi. Australian Journal of Zoology, Supplemental Series 27, 1 - 47. https: // doi. org / 10.1071 / AJZS 063" type="book chapter" year="1978">Tyler &amp; Davies 1978</bibRefCitation>
) by a combination of: vomerine teeth present; distinct tympanum; grey to brown dorsal colouration with dark variegations (
<figureCitation id="41CC6AED1C185A02FDE8FEA7FDE59016" box="[563,634,259,285]" captionStart="FIGURE 9" captionStartId="27.[151,250,1908,1932]" captionTargetBox="[168,1419,181,1884]" captionTargetId="figure-17@27.[168,1419,181,1884]" captionTargetPageId="27" captionText="FIGURE 9. Photos in life of Litoria ridibunda sp. nov. from A) Wyndham, WA (Marion Anstis); B) WAM R167789, Surveyors Pool, Mitchell Plateau, WA (Paul Doughty); C) Red Cone Creek, WA (Paul Doughty); D) Channel Island Road, outer Darwin, NT (Stephen Richards); E) Ringwood, NT (Stephen Richards); F) Fogg Dam, NT (Matt Greenlees); G) Mitchell Plateau, WA (Jen Francis)." figureDoi="http://doi.org/10.5281/zenodo.8407010" httpUri="https://zenodo.org/record/8407010/files/figure.png" pageId="28" pageNumber="101">Fig. 9</figureCitation>
); black shin; posterior thigh colouration comprising variously shaped yellow to orange or reddish spots on a muted dark brown to black background (
<figureCitation id="41CC6AED1C185A02FBE4FE83FB17904A" box="[1087,1160,295,321]" captionStart="FIGURE 4" captionStartId="16.[151,250,1839,1863]" captionTargetBox="[324,1261,181,1814]" captionTargetId="figure-20@16.[324,1262,181,1814]" captionTargetPageId="16" captionText="FIGURE 4. Posterior thighs showing diagnostic patterns. Litoria ridibunda sp. nov.—A) NTM R38969, Groote Eylandt, NT (Jodi Rowley); B) no voucher, Lawn Hill, Qld 18.65°S 138.55°E (Harry Hines); C) WAM R174146, King Edward River, WA (Ryan Ellis); D) AMS R184860, Bradshaw, NT (Jodi Rowley); E) NTM R38968, Groote Eylandt, NT (Jodi Rowley); F) AMS R188538, Groote Eylandt, NT (Jodi Rowley). Litoria rothii—G) QM J84287, 5 km SE Miriam Vale, Qld (Harry Hines); H) QM J86621, 15km ESE Biggenden, Qld (Harry Hines); I) SAMA R63683, 41 km SE Normanton, Qld; J) SAMA R20230, 13 km SW Townsville, Qld; K) SAMA R20231, 13 km SW Townsville, Qld. In life AH, preserved specimen IK." figureDoi="http://doi.org/10.5281/zenodo.8407000" httpUri="https://zenodo.org/record/8407000/files/figure.png" pageId="28" pageNumber="101">Fig. 4</figureCitation>
); fingers with moderate webbing; fully webbed feet; and moderately expanded terminal discs on all digits (
<figureCitation id="41CC6AED1C185A02FBEDFEEFFBE1906E" box="[1078,1150,331,357]" captionStart="FIGURE 8" captionStartId="26.[151,250,1878,1902]" captionTargetBox="[151,1435,741,1853]" captionTargetId="figure-168@26.[151,1435,741,1853]" captionTargetPageId="26" captionText="FIGURE 8. Holotype of L. ridibunda sp. nov. (NTM R36917) in life: A) latero-dorsal view (Stephen Richards), B) laterodorsal view of head (Stephen Richards), C) posterior of thigh (Stephen Richards), D) groin (Stephen Richards), E) plantar view of foot, F) palmar view of hand." figureDoi="http://doi.org/10.5281/zenodo.8407008" httpUri="https://zenodo.org/record/8407008/files/figure.png" pageId="28" pageNumber="101">Figs 8</figureCitation>
,
<figureCitation id="41CC6AED1C185A02FB50FEEFFB04906E" box="[1163,1179,331,357]" captionStart="FIGURE 9" captionStartId="27.[151,250,1908,1932]" captionTargetBox="[168,1419,181,1884]" captionTargetId="figure-17@27.[168,1419,181,1884]" captionTargetPageId="27" captionText="FIGURE 9. Photos in life of Litoria ridibunda sp. nov. from A) Wyndham, WA (Marion Anstis); B) WAM R167789, Surveyors Pool, Mitchell Plateau, WA (Paul Doughty); C) Red Cone Creek, WA (Paul Doughty); D) Channel Island Road, outer Darwin, NT (Stephen Richards); E) Ringwood, NT (Stephen Richards); F) Fogg Dam, NT (Matt Greenlees); G) Mitchell Plateau, WA (Jen Francis)." figureDoi="http://doi.org/10.5281/zenodo.8407010" httpUri="https://zenodo.org/record/8407010/files/figure.png" pageId="28" pageNumber="101">9</figureCitation>
).
</paragraph>
<paragraph id="D94876681C185A02FF1CFECBFA0B938E" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">
Can be distinguished from
<emphasis id="EB83AA7A1C185A02FE2BFECBFB819082" box="[496,1054,367,393]" italics="true" pageId="28" pageNumber="101">
<taxonomicName id="1EF70DEB1C185A02FE2BFECBFD049082" box="[496,667,367,393]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="amboinensis">L. amboinensis</taxonomicName>
,
<taxonomicName id="1EF70DEB1C185A02FD7FFECBFCDE9082" box="[676,833,367,393]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="darlingtoni">L. darlingtoni</taxonomicName>
,
<taxonomicName id="1EF70DEB1C185A02FC91FECBFC239083" baseAuthorityName="Boulenger" baseAuthorityYear="1897" box="[842,956,367,392]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="everetti">L. everetti</taxonomicName>
,
<taxonomicName id="1EF70DEB1C185A02FC1EFECBFB819082" box="[965,1054,367,393]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="rothii">L. rothii</taxonomicName>
</emphasis>
and
<taxonomicName id="1EF70DEB1C185A02FB89FECBFB379082" authorityName="Martin, Watson, Gartside, Littlejohn &amp; Loftus-Hills" authorityYear="1979" box="[1106,1192,367,393]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="tyleri">
<emphasis id="EB83AA7A1C185A02FB89FECBFB379082" box="[1106,1192,367,393]" italics="true" pageId="28" pageNumber="101">L. tyleri</emphasis>
</taxonomicName>
by the colour patterns on the posterior thigh in life.
<taxonomicName id="1EF70DEB1C185A02FE33FE37FD3290A6" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[488,685,403,429]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C185A02FE33FE37FD3290A6" box="[488,685,403,429]" italics="true" pageId="28" pageNumber="101">Litoria ridibunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0B017011C185A02FD6DFE37FC8C90A6" box="[694,787,403,429]" pageId="28" pageNumber="101" rank="species">
<emphasis id="EB83AA7A1C185A02FD6DFE37FC9390A6" bold="true" box="[694,780,403,429]" pageId="28" pageNumber="101">sp. nov</emphasis>
.
</taxonomicNameLabel>
typically has a matt black or dark grey background with yellow, orange or reddish coloured well-spaced patches or linear arrangements of coalesced patches often on the lower margin of the posterior thigh. The matt black or dark grey background provides less contrast than is seen
<taxonomicName id="1EF70DEB1C185A02FA5DFE7FFF4B9312" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="rothii">
<emphasis id="EB83AA7A1C185A02FA5DFE7FFF4B9312" italics="true" pageId="28" pageNumber="101">L. rothii</emphasis>
</taxonomicName>
.
<taxonomicName id="1EF70DEB1C185A02FF39FE5BFE5D9312" box="[226,450,511,537]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="amboinensis">
<emphasis id="EB83AA7A1C185A02FF39FE5BFE5D9312" box="[226,450,511,537]" italics="true" pageId="28" pageNumber="101">Litoria amboinensis</emphasis>
</taxonomicName>
typically has a cream and brown pattern.
<emphasis id="EB83AA7A1C185A02FC4FFE5BFB679312" box="[916,1272,511,537]" italics="true" pageId="28" pageNumber="101">
<taxonomicName id="1EF70DEB1C185A02FC4FFE5BFBF89312" box="[916,1127,511,537]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="darlingtoni">Litoria darlingtoni</taxonomicName>
and
<taxonomicName id="1EF70DEB1C185A02FB45FE5BFB679312" box="[1182,1272,511,537]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="rothii">L. rothii</taxonomicName>
</emphasis>
typically have an immaculate glossy black background with yellow or occasionally white patches with highly discrete margins, providing a strong contrast between the dark background and the coloured patches.
<taxonomicName id="1EF70DEB1C185A02FB94FDE3FB67936B" baseAuthorityName="Boulenger" baseAuthorityYear="1897" box="[1103,1272,583,608]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="everetti">
<emphasis id="EB83AA7A1C185A02FB94FDE3FB67936B" box="[1103,1272,583,608]" italics="true" pageId="28" pageNumber="101">Litoria everetti</emphasis>
</taxonomicName>
has an orange marbled pattern.
<taxonomicName id="1EF70DEB1C185A02FE8DFDCFFE7A938E" authorityName="Martin, Watson, Gartside, Littlejohn &amp; Loftus-Hills" authorityYear="1979" box="[342,485,619,645]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="tyleri">
<emphasis id="EB83AA7A1C185A02FE8DFDCFFE7A938E" box="[342,485,619,645]" italics="true" pageId="28" pageNumber="101">Litoria tyleri</emphasis>
</taxonomicName>
typically has a predominately orange coloration with small dark blotches and spots.
</paragraph>
<paragraph id="D94876681C185A02FF1CFD2BFB4093C6" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">
The calls of
<taxonomicName id="1EF70DEB1C185A02FE8AFD2BFE4293A2" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[337,477,655,681]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C185A02FE8AFD2BFE4293A2" box="[337,477,655,681]" italics="true" pageId="28" pageNumber="101">L. ridibunda</emphasis>
</taxonomicName>
<emphasis id="EB83AA7A1C185A02FE3FFD2BFDA293A2" bold="true" box="[484,573,655,681]" pageId="28" pageNumber="101">
<taxonomicNameLabel id="F0B017011C185A02FE3FFD2BFDA293A2" box="[484,573,655,681]" pageId="28" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are longer and contain more notes per call (average of 0.5
<geoCoordinate id="BCC310AF1C185A02FB0DFD2AFA9693A2" box="[1238,1289,654,681]" degrees="1.2" direction="south" orientation="latitude" pageId="28" pageNumber="101" precision="5555" value="-1.2">1.2 s</geoCoordinate>
call duration and 918 notes per call) than
<taxonomicName id="1EF70DEB1C185A02FE39FD17FDA293C6" box="[482,573,691,717]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="rothii">
<emphasis id="EB83AA7A1C185A02FE39FD17FDA293C6" box="[482,573,691,717]" italics="true" pageId="28" pageNumber="101">L. rothii</emphasis>
</taxonomicName>
(average of 0.3
<geoCoordinate id="BCC310AF1C185A02FD20FD17FCAF93C6" box="[763,816,691,717]" degrees="0.8" direction="south" orientation="latitude" pageId="28" pageNumber="101" precision="5555" value="-0.8">0.8 s</geoCoordinate>
call duration and 510 notes per call).
</paragraph>
<paragraph id="D94876681C185A02FF1CFD73FC03921E" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">
<taxonomicName id="1EF70DEB1C185A02FF1CFD73FE1493FA" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[199,395,727,753]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C185A02FF1CFD73FE1493FA" box="[199,395,727,753]" italics="true" pageId="28" pageNumber="101">Litoria ridibunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0B017011C185A02FE48FD73FE7093FA" box="[403,495,727,753]" pageId="28" pageNumber="101" rank="species">
<emphasis id="EB83AA7A1C185A02FE48FD73FE7793FA" bold="true" box="[403,488,727,753]" pageId="28" pageNumber="101">sp. nov</emphasis>
.
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="1EF70DEB1C185A02FCF8FD73FCE093FA" box="[803,895,727,753]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="rothii">
<emphasis id="EB83AA7A1C185A02FCF8FD73FCE093FA" box="[803,895,727,753]" italics="true" pageId="28" pageNumber="101">L. rothii</emphasis>
</taxonomicName>
and
<taxonomicName id="1EF70DEB1C185A02FC6CFD73FBB793FB" baseAuthorityName="Boulenger" baseAuthorityYear="1897" box="[951,1064,727,752]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="28" pageNumber="101" phylum="Chordata" rank="species" species="everetti">
<emphasis id="EB83AA7A1C185A02FC6CFD73FBB793FB" box="[951,1064,727,752]" italics="true" pageId="28" pageNumber="101">L. everetti</emphasis>
</taxonomicName>
by apomorphic nucleotide states at 31 and 26 sites, respectively, in the
<emphasis id="EB83AA7A1C185A02FD9AFD5FFDE9921E" box="[577,630,763,789]" italics="true" pageId="28" pageNumber="101">ND4</emphasis>
gene alignment (
<tableCitation id="947543D31C185A02FCECFD5FFC10921E" box="[823,911,763,789]" captionStart="TABLE 2" captionStartId="13.[151,239,582,608]" captionText="TABLE 2. Apomorphic diagnostic sites in the mitochondrial ND4 alignment of the Litoria peronii Group. Sites are numbered with reference to the Nyctimystes kubori ND4 and flanking tRNAs (GenBank accession JX564879). Bolding indicates apomorphic character states polarised with the remaining members of the Litoria peronii Group as the outgroups." pageId="28" pageNumber="101">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="91ED25E31C185A00FF1CFCBAFD48971F" lastPageId="30" lastPageNumber="103" pageId="28" pageNumber="101" type="description">
<paragraph id="D94876681C185A02FF1CFCBAFBE19256" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">
<emphasis id="EB83AA7A1C185A02FF1CFCBAFEF09233" bold="true" box="[199,367,798,824]" pageId="28" pageNumber="101">Dimensions of</emphasis>
<materialsCitation id="699F7C351C185A02FEAEFCBBFBE79256" collectionCode="SVL" pageId="28" pageNumber="101" specimenCode="SVL 46.3" specimenCount="1" typeStatus="holotype">
<emphasis id="EB83AA7A1C185A02FEAEFCBBFDB59233" bold="true" box="[373,554,798,825]" pageId="28" pageNumber="101">
<typeStatus id="064CC8CA1C185A02FEAEFCBBFE439232" box="[373,476,799,825]" pageId="28" pageNumber="101">holotype</typeStatus>
(mm).
</emphasis>
<specimenCode id="8951DE131C185A02FDEAFCBBFD009232" box="[561,671,799,825]" collectionCode="SVL" pageId="28" pageNumber="101">SVL 46.3</specimenCode>
; TL 22.7, HL 14.9; HW 13.7; EN 4.1; IND 3.5; ED 4.5; IOD 9.4; TD 3.4; FLL 9.9; Fin3L 12.6; Fin3DW 2.6; PpFin3 1.4; Toe4L 19.9; Toe4DW 2; PpToe4 4 1
</materialsCitation>
.
</paragraph>
<paragraph id="D94876681C185A02FF1CFCC2FBAF95CE" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">
<emphasis id="EB83AA7A1C185A02FF1CFCC2FE7A928A" bold="true" box="[199,485,870,897]" pageId="28" pageNumber="101">
Description of
<typeStatus id="064CC8CA1C185A02FEACFCC3FE7F928A" box="[375,480,871,897]" pageId="28" pageNumber="101">holotype</typeStatus>
.
</emphasis>
Head approximately as long as broad (HL/HW = 1.08), and approximately one third snout to vent length (HL/SVL = 0.32). Snout blunt when viewed from above and in profile. Nostrils more lateral than superior, closer to snout than to eye. Distance between eye and naris approximately equal to internarial span (EN/ IND = 1.17). Canthus rostralis gently curved. Eye relatively large, its diameter equal to eye to naris distance (ED/EN = 1.09). Pupil horizontal when constricted. Tympanum distinct, and oval with long axis vertical. Tympanum length three quarters eye diameter (TD/ED = 0.75). Moderately developed supratympanic fold, glandular in appearance, that partially obscures tympanic region. Vomeropalatines forming two short, prominently raised ridges or, occasionally, round clumps, each 0.51.0 mm wide, located midway between choanae. Each elevation has 26 (most commonly 46) small but distinct teeth. Tongue broadly oval to nearly round with shallow posterior notch. Vocal slits long, lateral, extending from approximately angle of jaws halfway to anterior tip of jaw.
</paragraph>
<paragraph id="D94876681C185A02FF1CFB6BFE6F94CA" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">Fingers long, broad, with moderate webbing between digits 2 and 3 and basal between 3 and 4. Subarticular tubercles prominent. Terminal discs expanded, extending well beyond lateral extremities of penultimate phalanx. Fingers in order of length 3&gt;4&gt;2&gt;1. Hindlimb length moderate (TL/SVL = 0.49). Toes in order of length 4&gt;5=3&gt;2&gt;1. Webbing reaches nearly to disc on toe 4 and to disc on other toes. Subarticular tubercles prominent. Inner metatarsal tubercle absent. Terminal toe discs expanded, extending well beyond lateral extremities of penultimate phalanx. Dorsum lightly shagreened. Abdomen, undersurface of thighs, submandibular area and lateral aspect of body mildly granular. Pectoral fold present.</paragraph>
<paragraph id="D94876681C185A02FF1CFA6EFCF89792" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">
<emphasis id="EB83AA7A1C185A02FF1CFA6EFEA094EF" bold="true" box="[199,319,1482,1508]" pageId="28" pageNumber="101">Variation.</emphasis>
Assessment of morphological variation is based on 85 individuals (
<tableCitation id="947543D31C185A02FB9DFA6FFB0094EE" box="[1094,1183,1483,1509]" captionStart="TABLE 4" captionStartId="21.[151,239,151,175]" captionText="TABLE 4. Measurement summary for adult Litoria everetti, L. ridibunda sp. nov. and L. rothii. Figures are mean ± SD and range." pageId="28" pageNumber="101">Table 4</tableCitation>
, Supplementary
<tableCitation id="947543D31C185A02FA85FA6FFF279702" captionStart="TABLE 2" captionStartId="13.[151,239,582,608]" captionText="TABLE 2. Apomorphic diagnostic sites in the mitochondrial ND4 alignment of the Litoria peronii Group. Sites are numbered with reference to the Nyctimystes kubori ND4 and flanking tRNAs (GenBank accession JX564879). Bolding indicates apomorphic character states polarised with the remaining members of the Litoria peronii Group as the outgroups." pageId="28" pageNumber="101">Table S2</tableCitation>
). Mean SVL: females =
<quantity id="1E0FDB8D1C185A02FE0BFA4BFDAA9702" box="[464,565,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="4.77" pageId="28" pageNumber="101" unit="mm" value="47.7">47.7 mm</quantity>
(n=16), males =
<quantity id="1E0FDB8D1C185A02FD22FA4BFCC19702" box="[761,862,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="4.42" pageId="28" pageNumber="101" unit="mm" value="44.2">44.2 mm</quantity>
(n=69). Head length equal to head width (HW/HL = 0.91.1) and approximately one-third of SVL (HL/SVL = 0.30.4). Distance between eye and naris equal to internarial span (EN/IND = 0.71.7). Eye relatively large but variable in diameter (ED/EN = 0.81.8). Tympanum length variable (TD/ED = 0.61.1). Moderate webbing between fingers 3 and 4. Hindlimb length moderate (TL/ SVL = 0.40.6). Indistinct canthus rostralis in some individuals.
</paragraph>
<paragraph id="D94876681C185A02FF1CF907FDDC97EA" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">Dorsum either smooth, mildly granular or with low and infrequent tubercles. Limbs with low tubercles and mildly granular or sometimes smooth.</paragraph>
<paragraph id="D94876681C185A02FF1CF94EFD3196B2" blockId="28.[151,1437,150,1977]" pageId="28" pageNumber="101">
<emphasis id="EB83AA7A1C185A02FF1CF94EFEF5960F" bold="true" box="[199,362,1770,1796]" pageId="28" pageNumber="101">Colour in life.</emphasis>
Posterior thigh mostly matt black to dark grey, with yellow marking varying from a few blotches to transverse patches with diffuse margins occupying between 2070% of the posterior thigh (
<figureCitation id="41CC6AED1C185A02FB11F8ABFA8E9622" box="[1226,1297,1807,1833]" captionStart="FIGURE 4" captionStartId="16.[151,250,1839,1863]" captionTargetBox="[324,1261,181,1814]" captionTargetId="figure-20@16.[324,1262,181,1814]" captionTargetPageId="16" captionText="FIGURE 4. Posterior thighs showing diagnostic patterns. Litoria ridibunda sp. nov.—A) NTM R38969, Groote Eylandt, NT (Jodi Rowley); B) no voucher, Lawn Hill, Qld 18.65°S 138.55°E (Harry Hines); C) WAM R174146, King Edward River, WA (Ryan Ellis); D) AMS R184860, Bradshaw, NT (Jodi Rowley); E) NTM R38968, Groote Eylandt, NT (Jodi Rowley); F) AMS R188538, Groote Eylandt, NT (Jodi Rowley). Litoria rothii—G) QM J84287, 5 km SE Miriam Vale, Qld (Harry Hines); H) QM J86621, 15km ESE Biggenden, Qld (Harry Hines); I) SAMA R63683, 41 km SE Normanton, Qld; J) SAMA R20230, 13 km SW Townsville, Qld; K) SAMA R20231, 13 km SW Townsville, Qld. In life AH, preserved specimen IK." figureDoi="http://doi.org/10.5281/zenodo.8407000" httpUri="https://zenodo.org/record/8407000/files/figure.png" pageId="28" pageNumber="101">Fig. 4</figureCitation>
). Vent same colour as surrounding dorsal colours. Fore of thighs and groin with same colour and pattern as posterior thigh but separated from posterior thigh pattern by intrusion of dorsal colour along top surface of thigh. Immaculate muted black continues along outer margin of leg and sometimes proximally onto dorsal surface of foot. Axilla with immaculate black or black and yellow blotches.
</paragraph>
<caption id="8D8826E01C195A03FF4CFBC1FE689406" ID-DOI="http://doi.org/10.5281/zenodo.8407012" ID-Zenodo-Dep="8407012" httpUri="https://zenodo.org/record/8407012/files/figure.png" pageId="29" pageNumber="102" startId="29.[151,250,1125,1149]" targetBox="[160,1431,190,1097]" targetPageId="29" targetType="figure">
<paragraph id="D94876681C195A03FF4CFBC1FE689406" blockId="29.[151,1437,1125,1293]" pageId="29" pageNumber="102">
<emphasis id="EB83AA7A1C195A03FF4CFBC1FEBE9576" bold="true" box="[151,289,1125,1149]" pageId="29" pageNumber="102">FIGURE 10.</emphasis>
Call variation in
<taxonomicName id="1EF70DEB1C195A03FE0CFBC2FD169576" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[471,649,1125,1149]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="29" pageNumber="102" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C195A03FE0CFBC2FD169576" box="[471,649,1125,1149]" italics="true" pageId="29" pageNumber="102">Litoria ridibunda</emphasis>
</taxonomicName>
<emphasis id="EB83AA7A1C195A03FD4BFBC1FD7D9576" bold="true" box="[656,738,1125,1149]" pageId="29" pageNumber="102">
<taxonomicNameLabel id="F0B017011C195A03FD4BFBC1FD7D9576" box="[656,738,1125,1149]" pageId="29" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="EB83AA7A1C195A03FCCDFBC2FCEE9576" box="[790,881,1125,1149]" italics="true" pageId="29" pageNumber="102">
<taxonomicName id="1EF70DEB1C195A03FCCDFBC2FCF29576" box="[790,877,1125,1149]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="29" pageNumber="102" phylum="Chordata" rank="species" species="rothii">L. rothii</taxonomicName>
.
</emphasis>
Advertisement calls displayed as a 1.5 s waveform of relative amplitude (Rel. amp.) over time and corresponding spectrogram of frequency (Freq.) over time. The recorder of each call is listed in Table 5.
<emphasis id="EB83AA7A1C195A03FE55FB09FE3F95CE" bold="true" box="[398,416,1197,1221]" pageId="29" pageNumber="102">A</emphasis>
)
<taxonomicName id="1EF70DEB1C195A03FE6BFB0AFDFD95CE" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[432,610,1197,1221]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="29" pageNumber="102" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C195A03FE6BFB0AFDFD95CE" box="[432,610,1197,1221]" italics="true" pageId="29" pageNumber="102">Litoria ridibunda</emphasis>
</taxonomicName>
<emphasis id="EB83AA7A1C195A03FDB2FB09FD4495CE" bold="true" box="[617,731,1197,1221]" pageId="29" pageNumber="102">
<taxonomicNameLabel id="F0B017011C195A03FDB2FB09FD2495CE" box="[617,699,1197,1221]" pageId="29" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
—i
</emphasis>
) MacArthur, NT,
<emphasis id="EB83AA7A1C195A03FC4DFB09FC3B95CE" bold="true" box="[918,932,1197,1221]" pageId="29" pageNumber="102">ii</emphasis>
) Mornington Sanctuary, WA,
<emphasis id="EB83AA7A1C195A03FB00FB09FB6F95CE" bold="true" box="[1243,1264,1197,1221]" pageId="29" pageNumber="102">iii</emphasis>
) FrogID capture 380982, The Gardens, Darwin, NT;
<emphasis id="EB83AA7A1C195A03FDD2FB75FD8595E2" bold="true" box="[521,538,1233,1257]" pageId="29" pageNumber="102">B</emphasis>
)
<taxonomicName id="1EF70DEB1C195A03FDF2FB76FDE395E2" box="[553,636,1233,1257]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="29" pageNumber="102" phylum="Chordata" rank="species" species="rothii">
<emphasis id="EB83AA7A1C195A03FDF2FB76FDE395E2" box="[553,636,1233,1257]" italics="true" pageId="29" pageNumber="102">L. rothii</emphasis>
</taxonomicName>
<emphasis id="EB83AA7A1C195A03FD4DFB75FD0295E2" bold="true" box="[662,669,1233,1257]" pageId="29" pageNumber="102">i</emphasis>
) Georgetown, Qld,
<emphasis id="EB83AA7A1C195A03FCB1FB75FCE795E2" bold="true" box="[874,888,1233,1257]" pageId="29" pageNumber="102">ii</emphasis>
) FrogID capture 84855, Caravonica, Qld,
<emphasis id="EB83AA7A1C195A03FAF6FB75FADD95E2" bold="true" box="[1325,1346,1233,1257]" pageId="29" pageNumber="102">iii</emphasis>
) FrogID capture 225994, Lower Tully, Qld.
</paragraph>
</caption>
<paragraph id="D94876681C195A03FF1CFAFDFBDB9709" blockId="29.[151,1437,1368,2043]" pageId="29" pageNumber="102">
Dorsum and head vary from uniformly bone to light brown with some degree of mottling with light or dark brown spots to large patches (
<figureCitation id="41CC6AED1C195A03FE36FAD8FDA8949D" box="[493,567,1404,1430]" captionStart="FIGURE 8" captionStartId="26.[151,250,1878,1902]" captionTargetBox="[151,1435,741,1853]" captionTargetId="figure-168@26.[151,1435,741,1853]" captionTargetPageId="26" captionText="FIGURE 8. Holotype of L. ridibunda sp. nov. (NTM R36917) in life: A) latero-dorsal view (Stephen Richards), B) laterodorsal view of head (Stephen Richards), C) posterior of thigh (Stephen Richards), D) groin (Stephen Richards), E) plantar view of foot, F) palmar view of hand." figureDoi="http://doi.org/10.5281/zenodo.8407008" httpUri="https://zenodo.org/record/8407008/files/figure.png" pageId="29" pageNumber="102">Figs 8</figureCitation>
,
<figureCitation id="41CC6AED1C195A03FD9EFAD8FDCA949D" box="[581,597,1404,1430]" captionStart="FIGURE 9" captionStartId="27.[151,250,1908,1932]" captionTargetBox="[168,1419,181,1884]" captionTargetId="figure-17@27.[168,1419,181,1884]" captionTargetPageId="27" captionText="FIGURE 9. Photos in life of Litoria ridibunda sp. nov. from A) Wyndham, WA (Marion Anstis); B) WAM R167789, Surveyors Pool, Mitchell Plateau, WA (Paul Doughty); C) Red Cone Creek, WA (Paul Doughty); D) Channel Island Road, outer Darwin, NT (Stephen Richards); E) Ringwood, NT (Stephen Richards); F) Fogg Dam, NT (Matt Greenlees); G) Mitchell Plateau, WA (Jen Francis)." figureDoi="http://doi.org/10.5281/zenodo.8407010" httpUri="https://zenodo.org/record/8407010/files/figure.png" pageId="29" pageNumber="102">9</figureCitation>
). Upper surfaces of limbs with same colour and pattern as dorsum. Wrist sometimes with transverse black mark. Head with thin black stripe, varying in prominence, beginning at posterior margin of eye and continuous across the top of the tympanum to black patch in axilla. Lips usually immaculate. Small black spots sparsely present in some individuals on snout or canthus rostralis.
</paragraph>
<paragraph id="D94876681C195A03FF1CF9A8FE779765" blockId="29.[151,1437,1368,2043]" pageId="29" pageNumber="102">Chin and chest plain white, lower abdomen and ventral surface of limbs sometimes suffused with yellow or orange tones, otherwise plain white. Throat in calling males suffused with black. Upper iris brownish-red to reddish, lower iris grey to light brown.</paragraph>
<paragraph id="D94876681C195A03FF1CF9DCFC499685" blockId="29.[151,1437,1368,2043]" pageId="29" pageNumber="102">
<emphasis id="EB83AA7A1C195A03FF1CF9DCFDFC9799" bold="true" box="[199,611,1656,1682]" pageId="29" pageNumber="102">Habitat and reproductive biology.</emphasis>
Habitat notes from WAM specimens indicate a wide habitat preference within Kimberley populations: water bodies such as swamps with paperbarks and
<taxonomicName id="1EF70DEB1C195A03FBE0F938FB3497BD" box="[1083,1195,1692,1718]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="29" pageNumber="102" phylum="Tracheophyta" rank="genus">
<emphasis id="EB83AA7A1C195A03FBE0F938FB3497BD" box="[1083,1195,1692,1718]" italics="true" pageId="29" pageNumber="102">Pandanus</emphasis>
</taxonomicName>
, as well as vegetated creeklines, rocky creeks and waterfalls in sandstone or basalt, seasonally flooded grasslands and near human habitation such as shower blocks. Males will call from the edges of ponds or from vegetation up to
<quantity id="1E0FDB8D1C195A03FAC7F940FAF997F4" box="[1308,1382,1764,1791]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="29" pageNumber="102" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">12 m</quantity>
(PD pers. obs.). Some individuals observed basking on the ground or in vegetation in full sun during the day (PD pers. obs.). The majority (&gt;70%) of FrogID records of the species as documented by users were from lentic waterbodies including ponds and flooded areas. The species was recorded commonly in suburban backyards and rural areas (&gt;60% of FrogID records of the species) and buildings (
<bibRefCitation id="BD660B991C195A03FCD0F8D0FC569685" author="Tyler, M. J. &amp; Crook, G. A. &amp; Davies, M." box="[779,969,1908,1935]" pageId="29" pageNumber="102" pagination="415 - 440" refId="ref22967" refString="Tyler, M. J., Crook, G. A. &amp; Davies, M. (1983) Reproductive biology of the frogs of the Magela Creek System, Northern Territory. Records of the South Australian Museum, 18, 415 - 440." type="journal article" year="1983">
Tyler
<emphasis id="EB83AA7A1C195A03FC95F8D1FCE09685" box="[846,895,1908,1934]" italics="true" pageId="29" pageNumber="102">et al</emphasis>
. 1983
</bibRefCitation>
).
</paragraph>
<paragraph id="D94876681C195A03FF1CF83CFCD896F1" blockId="29.[151,1437,1368,2043]" pageId="29" pageNumber="102">
The peak calling season for the species is the Australian Monsoon Tropics summer wet season from October to February (&gt;84% of all FrogID records of the species), but
<taxonomicName id="1EF70DEB1C195A03FC9CF819FC4C96DD" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[839,979,1980,2006]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="29" pageNumber="102" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C195A03FC9CF819FC4C96DD" box="[839,979,1980,2006]" italics="true" pageId="29" pageNumber="102">L. ridibunda</emphasis>
</taxonomicName>
<emphasis id="EB83AA7A1C195A03FC00F818FBAA96DD" bold="true" box="[987,1077,1980,2006]" pageId="29" pageNumber="102">
<taxonomicNameLabel id="F0B017011C195A03FC00F818FBAA96DD" box="[987,1077,1980,2006]" pageId="29" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has been recorded calling in all months of the year (&lt;3% of all FrogID records in MayJuly).
</paragraph>
<caption id="8D8826E01C1A5A00FF4CFBCBFBE69413" ID-DOI="http://doi.org/10.5281/zenodo.8407014" ID-Zenodo-Dep="8407014" httpUri="https://zenodo.org/record/8407014/files/figure.png" pageId="30" pageNumber="103" startId="30.[151,250,1135,1159]" targetBox="[151,1424,189,1099]" targetPageId="30" targetType="figure">
<paragraph id="D94876681C1A5A00FF4CFBCBFBE69413" blockId="30.[151,1437,1135,1304]" pageId="30" pageNumber="103">
<emphasis id="EB83AA7A1C1A5A00FF4CFBCBFE80958C" bold="true" box="[151,287,1135,1159]" pageId="30" pageNumber="103">FIGURE 11.</emphasis>
Variation in advertisement calls traits in
<taxonomicName id="1EF70DEB1C1A5A00FD1AFBD4FCEC958C" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[705,883,1135,1159]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="30" pageNumber="103" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C1A5A00FD1AFBD4FCEC958C" box="[705,883,1135,1159]" italics="true" pageId="30" pageNumber="103">Litoria ridibunda</emphasis>
</taxonomicName>
<emphasis id="EB83AA7A1C1A5A00FCA2FBD4FC559583" bold="true" box="[889,970,1136,1160]" pageId="30" pageNumber="103">
<taxonomicNameLabel id="F0B017011C1A5A00FCA2FBD4FC559583" box="[889,970,1136,1160]" pageId="30" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="EB83AA7A1C1A5A00FC26FBD4FBC8958C" box="[1021,1111,1135,1159]" italics="true" pageId="30" pageNumber="103">
<taxonomicName id="1EF70DEB1C1A5A00FC26FBD4FBCC958C" box="[1021,1107,1135,1159]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="30" pageNumber="103" phylum="Chordata" rank="species" species="rothii">L. rothii</taxonomicName>
.
</emphasis>
<emphasis id="EB83AA7A1C1A5A00FB80FBCBFBE9958C" bold="true" box="[1115,1142,1135,1159]" pageId="30" pageNumber="103">A)</emphasis>
Boxplots of
<emphasis id="EB83AA7A1C1A5A00FB20FBCBFA95958C" bold="true" box="[1275,1290,1135,1159]" pageId="30" pageNumber="103">i)</emphasis>
average notes per call,
<emphasis id="EB83AA7A1C1A5A00FF2EFB37FE9495A0" bold="true" box="[245,267,1171,1195]" pageId="30" pageNumber="103">ii)</emphasis>
average call duration (s),
<emphasis id="EB83AA7A1C1A5A00FDF8FB37FDDE95A0" bold="true" box="[547,577,1171,1195]" pageId="30" pageNumber="103">iii)</emphasis>
average note repetition rate (notes/s) and
<emphasis id="EB83AA7A1C1A5A00FBDBFB37FB8395A0" bold="true" box="[1024,1052,1171,1195]" pageId="30" pageNumber="103">iv)</emphasis>
average dominant frequency and
<emphasis id="EB83AA7A1C1A5A00FA58FB37FA0295A0" bold="true" box="[1411,1437,1171,1195]" pageId="30" pageNumber="103">B)</emphasis>
average number of notes per call over the distribution of the species (approximate distribution of
<taxonomicName id="1EF70DEB1C1A5A00FBAFFB1CFB5995C4" box="[1140,1222,1207,1231]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="30" pageNumber="103" phylum="Chordata" rank="species" species="rothii">
<emphasis id="EB83AA7A1C1A5A00FBAFFB1CFB5995C4" box="[1140,1222,1207,1231]" italics="true" pageId="30" pageNumber="103">L. rothii</emphasis>
</taxonomicName>
prior to this revision from
<bibRefCitation id="BD660B991C1A5A00FF0BFB7FFE3A95FF" author="Cutajar, T. P. &amp; Portway, C. &amp; Gillard, G. L. &amp; Rowley, J. J. L." box="[208,421,1243,1268]" pageId="30" pageNumber="103" pagination="1 - 48" refId="ref18994" refString="Cutajar, T. P., Portway, C., Gillard, G. L. &amp; Rowley, J. J. L. (2022) Australian frog atlas: Species' distribution maps informed by the FrogID dataset. Technical Reports of the Australian Museum Online. Sydney, 36, 1 - 48. https: // doi. org / 10.3853 / j. 1835 - 4211.36.2022.1789" type="journal article" year="2022">
Cutajar
<emphasis id="EB83AA7A1C1A5A00FEF8FB78FECE95F8" box="[291,337,1243,1267]" italics="true" pageId="30" pageNumber="103">et al</emphasis>
. [2022]
</bibRefCitation>
), hashed line represents approximate contact for the distributions of the two species. Note that the number of notes per call analysis included an additional 17 individuals (Supplementary Table S3).
</paragraph>
</caption>
<paragraph id="D94876681C1A5A00FF1CFACEFD48971F" blockId="30.[151,1437,1386,1880]" pageId="30" pageNumber="103">
<bibRefCitation id="BD660B991C1A5A00FF1CFACEFEFC948F" author="Anstis, M. A." box="[199,355,1386,1412]" pageId="30" pageNumber="103" refId="ref18086" refString="Anstis, M. A. (2013) Frogs and tadpoles of Australia. New Holland Publishers, Sydney, 829 pp." type="book" year="2013">Anstis (2013)</bibRefCitation>
described the morphology of the larvae of
<taxonomicName id="1EF70DEB1C1A5A00FC85FACEFC72948F" authority="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards, 2023" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[862,1005,1386,1412]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="30" pageNumber="103" phylum="Chordata" rank="species" species="ridibunda" status="sp. nov.">
<emphasis id="EB83AA7A1C1A5A00FC85FACEFC72948F" box="[862,1005,1386,1412]" italics="true" pageId="30" pageNumber="103">L. ridibunda</emphasis>
</taxonomicName>
<emphasis id="EB83AA7A1C1A5A00FC2CFACEFBCB948F" bold="true" box="[1015,1108,1386,1412]" pageId="30" pageNumber="103">
<taxonomicNameLabel id="F0B017011C1A5A00FC2CFACEFBCB948F" box="[1015,1108,1386,1412]" pageId="30" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Mitchell Plateau, WA, Kununurra, WA, and Marrara and Micket Creeks near Darwin, NT in a composite description that also included larvae of
<taxonomicName id="1EF70DEB1C1A5A00FED3FA16FEF994C7" box="[264,358,1458,1484]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="30" pageNumber="103" phylum="Chordata" rank="species" species="rothii">
<emphasis id="EB83AA7A1C1A5A00FED3FA16FEF994C7" box="[264,358,1458,1484]" italics="true" pageId="30" pageNumber="103">L. rothii</emphasis>
</taxonomicName>
from
<collectingRegion id="1B33B88A1C1A5A00FE69FA16FDA994C7" box="[434,566,1458,1484]" country="Australia" name="Queensland" pageId="30" pageNumber="103">Queensland</collectingRegion>
without noting any significant differences among the samples.
<bibRefCitation id="BD660B991C1A5A00FAC0FA16FF7D94FB" author="Tyler, M. J. &amp; Crook, G. A. &amp; Davies, M." pageId="30" pageNumber="103" pagination="415 - 440" refId="ref22967" refString="Tyler, M. J., Crook, G. A. &amp; Davies, M. (1983) Reproductive biology of the frogs of the Magela Creek System, Northern Territory. Records of the South Australian Museum, 18, 415 - 440." type="journal article" year="1983">
Tyler
<emphasis id="EB83AA7A1C1A5A00FABAFA16FA0294C7" box="[1377,1437,1458,1484]" italics="true" pageId="30" pageNumber="103">et al.</emphasis>
(1983)
</bibRefCitation>
described the breeding biology, eggs and larval morphology and development of the species (as
<taxonomicName id="1EF70DEB1C1A5A00FAC7FA72FAE994FB" box="[1308,1398,1494,1520]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="30" pageNumber="103" phylum="Chordata" rank="species" species="rothii">
<emphasis id="EB83AA7A1C1A5A00FAC7FA72FAE994FB" box="[1308,1398,1494,1520]" italics="true" pageId="30" pageNumber="103">L. rothii</emphasis>
</taxonomicName>
) in the Magela Creek system in western Arnhem Land.
</paragraph>
</subSubSection>
<subSubSection id="91ED25E31C1A5A00FF1CF9B9FE81961B" pageId="30" pageNumber="103" type="distribution">
<paragraph id="D94876681C1A5A00FF1CF9B9FE71978B" blockId="30.[151,1437,1386,1880]" pageId="30" pageNumber="103">
<emphasis id="EB83AA7A1C1A5A00FF1CF9B9FEC6973C" bold="true" box="[199,345,1565,1591]" pageId="30" pageNumber="103">Distribution</emphasis>
. From the Kimberley region across northern
<collectingCountry id="A1E036F81C1A5A00FCB3F9BAFC509733" box="[872,975,1566,1592]" name="Australia" pageId="30" pageNumber="103">Australia</collectingCountry>
north of about
<geoCoordinate id="BCC310AF1C1A5A00FB53F9BAFB209733" box="[1160,1215,1566,1592]" degrees="19" direction="south" orientation="latitude" pageId="30" pageNumber="103" precision="55555" value="-19.0">19°S</geoCoordinate>
east to Mt Isa and Normanton. Apparently introduced populations in Port Hedland and Karratha in the Pilbara region of WA (
<bibRefCitation id="BD660B991C1A5A00FAF7F9E6FEA2978B" author="Gaikhorst, G. &amp; Foster, J." pageId="30" pageNumber="103" pagination="197 - 210" refId="ref19681" refString="Gaikhorst, G. &amp; Foster, J. (2022) Green Tree Frogs (Litoria caerulea) on the hop: observations on established populations in the Pilbara. Western Australian Naturalist, 32, 197 - 210." type="journal article" year="2022">Gaikhorst &amp; Foster 2022</bibRefCitation>
; PD pers. obs.).
</paragraph>
<paragraph id="D94876681C1A5A00FF1CF92EFE81961B" blockId="30.[151,1437,1386,1880]" pageId="30" pageNumber="103">
<materialsCitation id="699F7C351C1A5A00FF1CF92EFE85961B" collectionCode="IBRA" county="Dampierland" location="North Kimberley" municipality="Central Kimberley" pageId="30" pageNumber="103" specimenCount="1" stateProvince="Victoria">
It occurs in the following
<collectionCode id="BFE6EEAD1C1A5A00FE26F92EFDA197AF" box="[509,574,1674,1700]" pageId="30" pageNumber="103">IBRA</collectionCode>
regions:
<collectingCounty id="30290EE41C1A5A00FD74F92EFCDB97AF" box="[687,836,1674,1700]" pageId="30" pageNumber="103">Dampierland</collectingCounty>
,
<collectingMunicipality id="392CEC121C1A5A00FC88F92EFBB697AF" box="[851,1065,1674,1700]" pageId="30" pageNumber="103">Central Kimberley</collectingMunicipality>
,
<location id="DC2820B31C1A5A00FBE3F92EFB6297AF" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FBE3F92EFB6297AF" box="[1080,1277,1674,1700]" county="Dampierland" municipality="Central Kimberley" name="North Kimberley" pageId="30" pageNumber="103" stateProvince="Victoria">North Kimberley</location>
,
<location id="DC2820B31C1A5A00FAD7F92EFAA797AF" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FAD7F92EFAA797AF" box="[1292,1336,1674,1700]" county="Dampierland" municipality="Central Kimberley" name="Ord" pageId="30" pageNumber="103" stateProvince="Victoria">Ord</location>
<collectingRegion id="1B33B88A1C1A5A00FA99F92EFA0397AF" box="[1346,1436,1674,1700]" country="Australia" name="Victoria" pageId="30" pageNumber="103">Victoria</collectingRegion>
<location id="DC2820B31C1A5A00FF4CF90AFF4C97C3" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FF4CF90AFF4C97C3" box="[151,211,1710,1736]" county="Dampierland" municipality="Central Kimberley" name="Plain" pageId="30" pageNumber="103" stateProvince="Victoria">Plain</location>
,
<collectingRegion id="1B33B88A1C1A5A00FF06F90AFEA897C3" box="[221,311,1710,1736]" country="Australia" name="Victoria" pageId="30" pageNumber="103">Victoria</collectingRegion>
<location id="DC2820B31C1A5A00FEE6F90AFE2C97C3" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FEE6F90AFE2C97C3" box="[317,435,1710,1736]" county="Dampierland" municipality="Central Kimberley" name="Bonaparte" pageId="30" pageNumber="103" stateProvince="Victoria">Bonaparte</location>
,
<location id="DC2820B31C1A5A00FE66F90AFDA297C3" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FE66F90AFDA297C3" box="[445,573,1710,1736]" county="Dampierland" municipality="Central Kimberley" name="Daly Basin" pageId="30" pageNumber="103" stateProvince="Victoria">Daly Basin</location>
,
<location id="DC2820B31C1A5A00FD9CF90AFD5B97C3" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FD9CF90AFD5B97C3" box="[583,708,1710,1736]" county="Dampierland" municipality="Central Kimberley" name="Pine Creek" pageId="30" pageNumber="103" stateProvince="Victoria">Pine Creek</location>
,
<location id="DC2820B31C1A5A00FD15F90AFCE097C3" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FD15F90AFCE097C3" box="[718,895,1710,1736]" county="Dampierland" municipality="Central Kimberley" name="Darwin Coastal" pageId="30" pageNumber="103" stateProvince="Victoria">Darwin Coastal</location>
,
<location id="DC2820B31C1A5A00FC52F90AFB8697C3" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FC52F90AFB8697C3" box="[905,1049,1710,1736]" county="Dampierland" municipality="Central Kimberley" name="Tiwi Coburg" pageId="30" pageNumber="103" stateProvince="Victoria">Tiwi Coburg</location>
,
<location id="DC2820B31C1A5A00FBFAF90AFB4797C3" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FBFAF90AFB4797C3" box="[1057,1240,1710,1736]" county="Dampierland" municipality="Central Kimberley" name="Arnhem Plateau" pageId="30" pageNumber="103" stateProvince="Victoria">Arnhem Plateau</location>
,
<location id="DC2820B31C1A5A00FB38F90AFA0797C3" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FB38F90AFA0797C3" box="[1251,1432,1710,1736]" county="Dampierland" municipality="Central Kimberley" name="Central Arnhem" pageId="30" pageNumber="103" stateProvince="Victoria">Central Arnhem</location>
,
<location id="DC2820B31C1A5A00FF4CF976FEA097E7" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FF4CF976FEA097E7" box="[151,319,1746,1772]" county="Dampierland" municipality="Central Kimberley" name="Arnhem Coast" pageId="30" pageNumber="103" stateProvince="Victoria">Arnhem Coast</location>
,
<location id="DC2820B31C1A5A00FE90F976FE7F97E7" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FE90F976FE7F97E7" box="[331,480,1746,1772]" county="Dampierland" municipality="Central Kimberley" name="Sturt Plateau" pageId="30" pageNumber="103" stateProvince="Victoria">Sturt Plateau</location>
,
<location id="DC2820B31C1A5A00FE36F976FD7097E7" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FE36F976FD7097E7" box="[493,751,1746,1772]" county="Dampierland" municipality="Central Kimberley" name="Mitchell Grass Downs" pageId="30" pageNumber="103" stateProvince="Victoria">Mitchell Grass Downs</location>
,
<location id="DC2820B31C1A5A00FD27F976FCFA97E7" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FD27F976FCFA97E7" box="[764,869,1746,1772]" county="Dampierland" municipality="Central Kimberley" name="Gulf Fall" pageId="30" pageNumber="103" stateProvince="Victoria">Gulf Fall</location>
and
<location id="DC2820B31C1A5A00FC45F976FC6197E7" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FC45F976FC6197E7" box="[926,1022,1746,1772]" county="Dampierland" municipality="Central Kimberley" name="Uplands" pageId="30" pageNumber="103" stateProvince="Victoria">Uplands</location>
,
<location id="DC2820B31C1A5A00FBD0F976FB0097E7" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FBD0F976FB0097E7" box="[1035,1183,1746,1772]" county="Dampierland" municipality="Central Kimberley" name="Gulf Coastal" pageId="30" pageNumber="103" stateProvince="Victoria">Gulf Coastal</location>
,
<location id="DC2820B31C1A5A00FB70F976FAF997E7" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FB70F976FAF997E7" box="[1195,1382,1746,1772]" county="Dampierland" municipality="Central Kimberley" name="Mount Isa Inlier" pageId="30" pageNumber="103" stateProvince="Victoria">Mount Isa Inlier</location>
, and
<location id="DC2820B31C1A5A00FF4CF952FE85961B" LSID="urn:lsid:plazi:treatment:515EC77E1C1E5A00FF4CFE7FFE3A9653:DC2820B31C1A5A00FF4CF952FE85961B" box="[151,282,1782,1808]" county="Dampierland" municipality="Central Kimberley" name="Gulf Plains" pageId="30" pageNumber="103" stateProvince="Victoria">Gulf Plains</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="91ED25E31C1A5A00FF1CF8BEFE3A9653" pageId="30" pageNumber="103" type="etymology">
<paragraph id="D94876681C1A5A00FF1CF8BEFE3A9653" blockId="30.[151,1437,1386,1880]" pageId="30" pageNumber="103">
<emphasis id="EB83AA7A1C1A5A00FF1CF8BEFED1963F" bold="true" box="[199,334,1818,1844]" pageId="30" pageNumber="103">Etymology.</emphasis>
The specific epithet, a Latin adjective, refers to the nature of the male advertisement call— “laughing” =
<taxonomicName id="1EF70DEB1C1A5A00FEEBF89AFE019653" authorityName="Donnellan &amp; Catullo &amp; Rowley &amp; Doughty &amp; Price &amp; Hines &amp; Richards" authorityYear="2023" box="[304,414,1854,1880]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="30" pageNumber="103" phylum="Chordata" rank="species" species="ridibunda">
<emphasis id="EB83AA7A1C1A5A00FEEBF89AFE019653" box="[304,414,1854,1880]" italics="true" pageId="30" pageNumber="103">ridibunda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>