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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1178.105407" ID-Pensoft-Pub="1313-2970-1178-165" ID-Pensoft-UUID="6A62359BD3655B84ADDECBD8DD441F3C" ID-ZooBank="49C77DFC4F094786AFCBC83F0B745F0C" ModsDocID="1313-2970-1178-165" checkinTime="1694161514751" checkinUser="pensoft" docAuthor="Cho, Kyuhee, Kim, Jong Guk &amp; Lee, Jimin" docDate="2023" docId="E7D46341203F58EFB13D055A37536DC4" docLanguage="en" docName="ZooKeys 1178: 165-189" docOrigin="ZooKeys 1178" docPubDate="2023-09-07" docSource="http://dx.doi.org/10.3897/zookeys.1178.105407" docTitle="Cerviniella bisegmenta Cho &amp; Kim &amp; Lee 2023, sp. nov." docType="treatment" docUuid="78650D35-782E-409D-9FE6-4FA038DF9680" docUuidSource="ZooBank" docVersion="1" id="6A62359BD3655B84ADDECBD8DD441F3C" lastPageNumber="165" masterDocId="6A62359BD3655B84ADDECBD8DD441F3C" masterDocTitle="Two new species of Cerviniella Smirnov, 1946 (Copepoda, Harpacticoida, Aegisthidae) from the Yellow Sea, Korea" masterLastPageNumber="189" masterPageNumber="165" pageNumber="165" updateTime="1694161514751" updateUser="pensoft">
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<mods:title id="F22FB3F1F65737BEFE64508994A62039">Two new species of Cerviniella Smirnov, 1946 (Copepoda, Harpacticoida, Aegisthidae) from the Yellow Sea, Korea</mods:title>
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<mods:name id="2193C85F5CB21F5BD605266B85B27D6F" type="personal">
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<mods:namePart id="E61FC417950BB749E578826D6B25237D">Cho, Kyuhee</mods:namePart>
<mods:nameIdentifier id="C01AD8EFEFE430DEB0D921503150DAFD" type="ORCID">https://orcid.org/0000-0002-5006-5713</mods:nameIdentifier>
<mods:affiliation id="6D6D863A3EAECAE60192B464CECE8BD6">Ocean Climate Response &amp; Ecosystem Research Department, Korea Institute of Ocean Science &amp; Technology, Busan 49111, Republic of Korea</mods:affiliation>
</mods:name>
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<mods:namePart id="ADA1DC41744383E1EB1E5AF24C91765E">Kim, Jong Guk</mods:namePart>
<mods:nameIdentifier id="9B781084545207C9E5C306D5ACA9AE15" type="ORCID">https://orcid.org/0000-0001-5299-9838</mods:nameIdentifier>
<mods:affiliation id="B512535C36349587B2475CACCEF7C0BB">Division of Zoology, Honam National Institute of Biological Resources, Mokpo 58762, Republic of Korea</mods:affiliation>
</mods:name>
<mods:name id="80AFF7C85145F221B98D0D3B2B7315D0" type="personal">
<mods:role id="7CEB03B6FD9F5D8EEBDBA5A4AC62514A">
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<mods:namePart id="D6A2D11D2FC3D8C3EE76D4B83D3A5F74">Lee, Jimin</mods:namePart>
<mods:nameIdentifier id="9B130582A33C9A4C7D5492809D35F0C9" type="ORCID">https://orcid.org/0000-0001-9004-8275</mods:nameIdentifier>
<mods:affiliation id="B6A56B3A1D6212D079F20811E7CED4E9">Ocean Climate Response &amp; Ecosystem Research Department, Korea Institute of Ocean Science &amp; Technology, Busan 49111, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="CEFC226ECB007E5EEB171AE8AC52E16C" type="email">leejm@kiost.ac</mods:nameIdentifier>
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<mods:typeOfResource id="69EA96FC7C3FA33A5EBAFBC4CBFFFC09">text</mods:typeOfResource>
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<mods:date id="212E2F73FE47F64F8B4D4845895C2E87">2023</mods:date>
<mods:detail id="0C30A7282C9AB803F8BE34888C42C7C5" type="pubDate">
<mods:number id="9CD31DE939132ED7D49A0D50DA081E25">2023-09-07</mods:number>
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<mods:number id="E9ED460AB5B4E4467C3B04EB63EF3D39">1178</mods:number>
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<mods:classification id="DF9438DC3EA2760104F1F654B798E991">journal article</mods:classification>
<mods:identifier id="0BC3A2F4445D2C9D42E8455B730524D0" type="DOI">http://dx.doi.org/10.3897/zookeys.1178.105407</mods:identifier>
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<treatment id="E7D46341203F58EFB13D055A37536DC4" LSID="urn:lsid:zoobank.org:act:78650D35-782E-409D-9FE6-4FA038DF9680" httpUri="http://treatment.plazi.org/id/E7D46341203F58EFB13D055A37536DC4" lastPageNumber="165" pageId="0" pageNumber="165">
<subSubSection id="8C6CE48821A8DB59678937410D4A5609" pageId="0" pageNumber="165" type="nomenclature">
<paragraph id="AA794C054D1629746A19B19290519725" pageId="0" pageNumber="165">
<taxonomicName id="E6AD6454348ACE17D5DCD9D0FE53BCD3" LSID="https://zoobank.org/78650D35-782E-409D-9FE6-4FA038DF9680" authority="Cho &amp; Kim &amp; Lee, 2023" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta" status="sp. nov.">Cerviniella bisegmenta</taxonomicName>
<taxonomicNameLabel id="789F2CD7099E96005928A94C0C7D7865" pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="ED1CDCA82AB6EFD1934AD77FB6BC1146" pageId="0" pageNumber="165" type="description">
<paragraph id="600E682A5B5B252780DC1BA07AF67706" pageId="0" pageNumber="165">
<figureCitation id="D95D9EC1021E2A052F2A2B08957F1B74" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cerviniella bisegmenta sp. nov. Female. Holotype (MABIK CR 00253873) A habitus, dorsal view B habitus, lateral view; Paratype (MABIK CR 00253868) C rostrum. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165">Figs 1</figureCitation>
<figureCitation id="663797562D4E28A234684B5E45DA6CBA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A antennule B antenna C mandible D maxillule. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165">, 2</figureCitation>
<figureCitation id="3D670801B5A5C5ACFD6C34A28DECAF94" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">, 3</figureCitation>
<figureCitation id="22B33A3B89ED4E94165DF983050A8BA3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="62FBDC4A39C7AFDF5BF7CC1D5EBDAF59" pageId="0" pageNumber="165" type="type locality">
<paragraph id="865CA0BAB6B35A8A3EEDCD086CE4DA4C" pageId="0" pageNumber="165">Type locality.</paragraph>
<paragraph id="7BB220BAE98F38AB2DDC13FB6C88B604" pageId="0" pageNumber="165">
The Yellow Sea;
<geoCoordinate id="66F5152E02C1E358348EC039E78F349F" degrees="34" direction="north" minutes="59" orientation="latitude" precision="1" seconds="40.14" value="34.994484">34°59'40.14&quot;N</geoCoordinate>
,
<geoCoordinate id="FEC6D6CB67BFCC899AD48D774EECD188" degrees="125" direction="east" minutes="00" orientation="longitude" precision="1" seconds="2.82" value="125.000786">125°00'2.82&quot;E</geoCoordinate>
; 88 m depth.
</paragraph>
</subSubSection>
<subSubSection id="C697E029A69C557F90276F9AC94C4FF0" pageId="0" pageNumber="165" type="type material">
<paragraph id="57E1E56A8A15D34920B49B9F6B84E870" pageId="0" pageNumber="165">Type material.</paragraph>
<paragraph id="26998736E0B5F4BAB672C7A31FE214B4" pageId="0" pageNumber="165">
<emphasis id="83972D2925EEF4506FAF1A36BA359482" bold="true" pageId="0" pageNumber="165">
<emphasis id="724CA11AE8D2A9AE41FCF76EAE2DBBDD" italics="true" pageId="0" pageNumber="165">Holotype</emphasis>
.
</emphasis>
One ♀ preserved in a vial with 80% ethanol (MABIK CR00253873); collected from the type locality, 20 April 2019.
<emphasis id="70F5B38253B6340D924B72501AFD2F3E" bold="true" pageId="0" pageNumber="165">
<emphasis id="7A03AE3D54939202E065AC7962F91D4F" italics="true" pageId="0" pageNumber="165">Paratypes</emphasis>
.
</emphasis>
One ♀ (MABIK CR00253868) dissected on 11 slides, three ♀♀ (MABIK CR00253869-00253871) each dissected on 10 slides, six ♀♀ (MABIK CR00253874) and eight ♀♀ (MInRB-Hr88-L001) preserved in a vial with 80% ethanol, collection data as in holotype; S.L. Kim leg.
</paragraph>
</subSubSection>
<subSubSection id="505A7052A274808BD705D59BE3FACC26" pageId="0" pageNumber="165" type="description">
<paragraph id="01350D91773495743EF240D574924E50" pageId="0" pageNumber="165">Description.</paragraph>
<paragraph id="47D49508985C2B82E8ED12080162BA27" pageId="0" pageNumber="165">
<emphasis id="284331B534ED61019F6659A405084C2E" bold="true" pageId="0" pageNumber="165">Female</emphasis>
(based on the holotype and paratypes).
<emphasis id="A7E43101314E1B95962611A19C8A737F" bold="true" italics="true" pageId="0" pageNumber="165">Body</emphasis>
length from anterior margin of rostrum to posterior margin of caudal rami (paratype, MABIK CR00253868, in lateral view, telescoping of somites not considered) 797
<normalizedToken id="E787CB56DF7FB649FBCAF964986365E1" originalValue="μm">μm</normalizedToken>
(range: 694-797
<normalizedToken id="0AF50118D140000819BFA957D644E1C9" originalValue="μm">μm</normalizedToken>
,
<emphasis id="241EEF3637430274838543EAC81D7CB2" italics="true" pageId="0" pageNumber="165">n</emphasis>
= 11, holotype: 765
<normalizedToken id="2CC2818AF5BFCCBB19A907B0CBAE1E6A" originalValue="μm">μm</normalizedToken>
).
</paragraph>
<paragraph id="5CAE4E3BBCFB2962E3D4BF44E74BD35A" pageId="0" pageNumber="165">
<emphasis id="AC99C61CD9B698DA3A3D59BFE68143B8" bold="true" italics="true" pageId="0" pageNumber="165">Habitus</emphasis>
(Fig.
<figureCitation id="C534B2D428FF21773C9FFCBFBF7C503F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cerviniella bisegmenta sp. nov. Female. Holotype (MABIK CR 00253873) A habitus, dorsal view B habitus, lateral view; Paratype (MABIK CR 00253868) C rostrum. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165">1A, B</figureCitation>
) subcylindrical, gradually tapering towards posterior caudal rami, with unclear separation between prosome and urosome.
<emphasis id="133A89B4659401E991FB8622A7CDCE8E" bold="true" italics="true" pageId="0" pageNumber="165">Prosome</emphasis>
(Fig.
<figureCitation id="301F0006C8F4CB4B091D558E5207F0BA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cerviniella bisegmenta sp. nov. Female. Holotype (MABIK CR 00253873) A habitus, dorsal view B habitus, lateral view; Paratype (MABIK CR 00253868) C rostrum. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165">1A, B</figureCitation>
) slightly longer than urosome, comprising cephalothorax with completely fused first pedigerous somite, and three free pedigerous somites; posterior margins finely serrated. Cephalothorax bell-shaped, slightly longer than wide, ~ 28% of body length; integument covered with several sensilla, numerous minute pits and striped pattern (discernible under high resolution); rim with anastomosing patterns. P2-bearing somite longer than following two prosomites, with well-developed pleural area. Pleural areas of P3- and P4-bearing somites with pointed posterolateral corners.
</paragraph>
<caption id="45120CDB3457C14C9E4B3067792FA03C" doi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165" start="Figure 1" startId="F1">
<paragraph id="3D8F9D82FB6B0F8061682167EF3BFACC" pageId="0" pageNumber="165">
<emphasis id="7EF69D6074FAFAA9C7FCF9D20BC25F45" bold="true" pageId="0" pageNumber="165">Figure 1.</emphasis>
<taxonomicName id="831E8C128A92698EADA56C4F340CBEB2" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="732D9A1B69CFFD03996BD190E2B9890C" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
sp. nov. Female. Holotype (MABIK CR00253873)
<emphasis id="FE510E534BC53B8DAA1F0EBCC4239AED" bold="true" pageId="0" pageNumber="165">A</emphasis>
habitus, dorsal view
<emphasis id="349A5DD3769A534ACCADF3DEEA22F36E" bold="true" pageId="0" pageNumber="165">B</emphasis>
habitus, lateral view; Paratype (MABIK CR00253868)
<emphasis id="D00EABC980BFE3A3A04381B849CDFF56" bold="true" pageId="0" pageNumber="165">C</emphasis>
rostrum. Scale bars are in
<normalizedToken id="ED6505B2E9C4F65E2BFCF26882400683" originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id="7EEADE952E53F7D41959F48330EA93E8" pageId="0" pageNumber="165">
<emphasis id="43B748F6C011CA760D553B3A82C4F087" bold="true" italics="true" pageId="0" pageNumber="165">Urosome</emphasis>
(Figs
<figureCitation id="450C1B65A80509C6D3912024A15C363A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cerviniella bisegmenta sp. nov. Female. Holotype (MABIK CR 00253873) A habitus, dorsal view B habitus, lateral view; Paratype (MABIK CR 00253868) C rostrum. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165">1A, B</figureCitation>
,
<figureCitation id="2D31F69FCD964C9FDC38D457D1F044CA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3D</figureCitation>
) comprising P5-bearing somite, genital double-somite, and three free abdominal somites; integument of tergites covered with minute spinules and posterior margins finely serrated except for anal somite. Genital somite and third urosomite separated dorsally and laterally, but completely fused ventrally, forming genital double-somite (Figs
<figureCitation id="943C5C83EE294536437958E1CD575B23" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cerviniella bisegmenta sp. nov. Female. Holotype (MABIK CR 00253873) A habitus, dorsal view B habitus, lateral view; Paratype (MABIK CR 00253868) C rostrum. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165">1A, B</figureCitation>
,
<figureCitation id="B5697F95A8D6F88987386C2772A74A7A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3D</figureCitation>
), with one pair of hook-like lateral projections on original genital somite; ventral surface with striations. Genital apertures (Fig.
<figureCitation id="9941047A720BADEB1BC9191AA5056F8C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3D</figureCitation>
) located far anteriorly, closed off by a single plate, on both sides with an outer vestigial seta and an inner bare seta, representing P6 (Fig.
<figureCitation id="819922C386C2EFDF8E67C9D8B5876CCD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4E</figureCitation>
; based on holotype). Copulatory pore located in median depression at level of gonopores. Anal somite (Figs
<figureCitation id="FA8927C4B5687095AE02F264871BF8A4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cerviniella bisegmenta sp. nov. Female. Holotype (MABIK CR 00253873) A habitus, dorsal view B habitus, lateral view; Paratype (MABIK CR 00253868) C rostrum. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165">1A</figureCitation>
,
<figureCitation id="E46086997F10305EF614DDA12FDB6232" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3D</figureCitation>
) as long as two preceding urosomites combined, ~ 2
<normalizedToken id="E4B15C4009201AD5E65600C6D58B7E3E" originalValue="×">x</normalizedToken>
as long as the caudal rami, with one pair of sensilla; semicircular operculum ornamented with minute spinules; anal sinus wide.
</paragraph>
<paragraph id="73F7308FFED0EED5747E7AFD3A367D60" pageId="0" pageNumber="165">
<emphasis id="B80249A61898A17CC8A1A790AFD23D45" bold="true" italics="true" pageId="0" pageNumber="165">Caudal rami</emphasis>
(Figs
<figureCitation id="3C419C76E90420CB835E012263F1460F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cerviniella bisegmenta sp. nov. Female. Holotype (MABIK CR 00253873) A habitus, dorsal view B habitus, lateral view; Paratype (MABIK CR 00253868) C rostrum. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165">1A, B</figureCitation>
,
<figureCitation id="FE4DAA6C7717DD8B9BC2C6BC9AAF038B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3D</figureCitation>
) cylindrical, ~ 1.7
<normalizedToken id="A4D1B697C7A05DAE474EF9E164CCA617" originalValue="×">x</normalizedToken>
as long as wide; surface covered with fine spinules; with one pore on ventro-distal surface; with seven setae: ventro-lateral seta I pinnate, short, inserted in proximal third of ramus, dorso-lateral seta II pinnate, twice as long as seta I; seta III pinnate, arising from outer ventro-distal corner, as long as caudal ramus; terminal setae IV and V well-developed, with internal fracture plane proximally, both setae fused basally, of which seta V longest, ~ 4
<normalizedToken id="1FB107AD0147B81F8CB78D0D9FF5B370" originalValue="×">x</normalizedToken>
as long as seta IV; seta VI shortest, spiniform, located on ventro-posterior margin; tri-articulate seta VII plumose, issuing from dorsal surface subdistally.
</paragraph>
<paragraph id="650E5DBFC371A43DAD7749690CC4E36C" pageId="0" pageNumber="165">
<emphasis id="89C7D960B9469229FEACB6494485430A" bold="true" italics="true" pageId="0" pageNumber="165">Rostrum</emphasis>
(Fig.
<figureCitation id="E2D1EEB1E6901D393164577F674EF3A2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cerviniella bisegmenta sp. nov. Female. Holotype (MABIK CR 00253873) A habitus, dorsal view B habitus, lateral view; Paratype (MABIK CR 00253868) C rostrum. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure1" httpUri="https://binary.pensoft.net/fig/902792" pageId="0" pageNumber="165">1A, C</figureCitation>
) completely fused to cephalothorax, triangular, subdistally with paired sensilla and one ventral tube pore; anterior tip slightly concave.
</paragraph>
<paragraph id="0EFC4E297EA5B7EEF5CE809D722AC2A8" pageId="0" pageNumber="165">
<emphasis id="42F77F5ED10E1AEC5B790F29AFDADA5E" bold="true" italics="true" pageId="0" pageNumber="165">Antennule</emphasis>
(Fig.
<figureCitation id="2216CD6473031C0FFB268B5BE7D93203" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A antennule B antenna C mandible D maxillule. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165">2A</figureCitation>
) robust, short, five-segmented, covered with diminutive dots (or denticles) as shown in Fig.
<figureCitation id="3B7FC8732AF7CC85C256E0A58EAFF7CC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A antennule B antenna C mandible D maxillule. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165">2A</figureCitation>
. First segment with a blunt conical process on outer distal corner. Second segment longest, with a distal peduncle bearing an aesthetasc fused to a seta. Forth segment shortest. Fifth segment distally with a small aesthetasc fused to a pinnate seta. Armature formula: 1 - [1 pinnate], 2 - [12 pinnate + 2 pinnate spine + (1 pinnate + ae)], 3-[3 pinnate], 4-[3 pinnate], 5-[6 pinnate + (1 pinnate + ae)].
</paragraph>
<caption id="DF7AE0C474A6992F4293D7AED1C3EE5F" doi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165" start="Figure 2" startId="F2">
<paragraph id="7BA53F7A46AB0B2F0275B2437951E377" pageId="0" pageNumber="165">
<emphasis id="3C547C8A707C6525CDFC2D03F7C6E3CD" bold="true" pageId="0" pageNumber="165">Figure 2.</emphasis>
<taxonomicName id="9502A3FB40E6E603822F978C841568ED" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="729F4B714200EE3EB204ED2E15EDA559" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
sp. nov. Female. Paratype (MABIK CR00253868)
<emphasis id="B68B0B148296EABEBD8F290C897188C2" bold="true" pageId="0" pageNumber="165">A</emphasis>
antennule
<emphasis id="85AAACDDC96D0D15C3A094054693FFFF" bold="true" pageId="0" pageNumber="165">B</emphasis>
antenna
<emphasis id="7D58ADA9D5D94B60AE0EB0E00BE5E87E" bold="true" pageId="0" pageNumber="165">C</emphasis>
mandible
<emphasis id="CD5FE88779F22D4E28F844CF5DD6ED14" bold="true" pageId="0" pageNumber="165">D</emphasis>
maxillule. Scale bars are in
<normalizedToken id="2A3C16119D9F35CEB35A70BCE085C5AE" originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id="F07925351A382F3B57E586284ECC58AC" pageId="0" pageNumber="165">
<emphasis id="47BA0CE3F978810FFFDD15E8D7C1ED16" bold="true" italics="true" pageId="0" pageNumber="165">Antenna</emphasis>
(Fig.
<figureCitation id="DF2E6469E0CA920EDFFAB4B51679CEC6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A antennule B antenna C mandible D maxillule. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165">2B</figureCitation>
) three-segmented, comprising coxa, allobasis, and one-segmented endopod. Coxa small (not figured). Allobasis elongated, covered with various-sized spinules on surface, and with long spinules along abexopodal margin and a patch of denticles near base of exopod as shown in Fig.
<figureCitation id="D27AA76BCB7A5F9753ABB67592966C32" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A antennule B antenna C mandible D maxillule. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165">2B</figureCitation>
; with one subdistal abexopodal seta. Endopod distinctly shorter than allobasis, with long spinules on inner and outer margins and rows of minute spinules along inner distal margin; lateral armature comprising one plumose and two pinnate elements; distal armature consisting of four serrate spines and three setae, of which outermost spine fused to one seta. Exopod two-segmented, proximal segment longer than distal segment, with two setae; distal segment with two lateral and three apical setae.
</paragraph>
<paragraph id="5BEE57ABC3757AFB1AD8E1E06969B14A" pageId="0" pageNumber="165">
<emphasis id="F7DF90BF95DE10DF084F826AA9AF16D9" bold="true" italics="true" pageId="0" pageNumber="165">Mandible</emphasis>
(Fig.
<figureCitation id="AE5CE5DC916926830DB0DDE2787B2B25" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A antennule B antenna C mandible D maxillule. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165">2C</figureCitation>
). Coxa proximally with rows of spinules; gnathobase well-developed, with several multi-cuspidate teeth and one pinnate seta; outermost tooth largest; with two rows of minute spinules proximally and one row of spinules subdistally. Palp biramous, consisting of basis, three-segmented exopod, and one-segmented endopod; basis with four plumose setae distally, covered with spinules on surface, and with one row of setules on lateral margin. Exopod smaller than endopod; exp-1 and exp-2 each with one plumose lateral seta, and exp-3 with three plumose (one lateral and two apical) setae. Endopod with two lateral and five apical plumose setae, of which two apical setae fused at base (indicated by arrowhead in Fig.
<figureCitation id="E1B7A0BCD9B15EA3E3175135AF7680E4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A antennule B antenna C mandible D maxillule. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165">2C</figureCitation>
).
</paragraph>
<paragraph id="B2B2C57504C32D09E7696DE633AF58B2" pageId="0" pageNumber="165">
<emphasis id="35C8B9A95D73E788DD0585E68760F0D5" bold="true" italics="true" pageId="0" pageNumber="165">Paragnaths</emphasis>
(Fig.
<figureCitation id="C4590B9E320085F5B66D9103B43722DA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3A</figureCitation>
) with well-developed chitinized lobes; distal and lateral margins covered with numerous spinules; posterior face with five strong spinules and one row of tiny spinules.
</paragraph>
<caption id="07A168597D1B9336E7251F711A9B4F44" doi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165" start="Figure 3" startId="F3">
<paragraph id="78FCFAADE66BF2A83E4CFC96B7A67BC0" pageId="0" pageNumber="165">
<emphasis id="75A888BFA178B4BBC480AE69C7EABEB4" bold="true" pageId="0" pageNumber="165">Figure 3.</emphasis>
<taxonomicName id="06A09F59615EC51EA8E0A16FED6F56CD" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="7D02CCE6A66C65630BB7626799B2B95A" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
sp. nov. Female. Paratype (MABIK CR00253868)
<emphasis id="E9E51C4A32DC89476193DBB5C7E7F345" bold="true" pageId="0" pageNumber="165">A</emphasis>
paragnaths, anterior
<emphasis id="28966EB9E0F1DFF499498410227B0121" bold="true" pageId="0" pageNumber="165">B</emphasis>
maxilla
<emphasis id="F278F38C0D777D2A9FC166B11875BED1" bold="true" pageId="0" pageNumber="165">C</emphasis>
maxilliped
<emphasis id="7E8204679156A3D5778E47BDBC89C0F8" bold="true" pageId="0" pageNumber="165">D</emphasis>
urosome, ventral view. Scale bars are in
<normalizedToken id="2A03FAA1B570EF599C62686A45FF74CA" originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id="BA5697B64FBDEC44121DAAAB081D26B3" pageId="0" pageNumber="165">
<emphasis id="D24163AE5B34577C28DA1736B45DFB46" bold="true" italics="true" pageId="0" pageNumber="165">Maxillule</emphasis>
(Fig.
<figureCitation id="32911F33629241E7D0B35947B48BD759" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A antennule B antenna C mandible D maxillule. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure2" httpUri="https://binary.pensoft.net/fig/902793" pageId="0" pageNumber="165">2D</figureCitation>
). Praecoxa ornamented with outer and subdistal spinules; arthrite well-developed, with two juxtaposed anterior setae, ten distal elements, and two pinnate posterior setae; posterior surface with patch of spinules. Coxa ornamented with outer spinules; cylindrical endite with one pinnate and five bare setae. Basis and endopod fused, with spinules on anterior and posterior surface; distal margin with 13 setae. Exopod represented by three pinnate setae.
</paragraph>
<paragraph id="EFDF975AE9AC26A98BA5579E0D2CD477" pageId="0" pageNumber="165">
<emphasis id="E7CDBD0AD3DA1274EE17D02A9851E0C8" bold="true" italics="true" pageId="0" pageNumber="165">Maxilla</emphasis>
(Fig.
<figureCitation id="7B1C94CB12409B4AAB723E13D188E37F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3B</figureCitation>
). Syncoxa (damaged) large, with one row of stout spinules and one row of setules along outer margin, and five groups of spinules on anterior and posterior surfaces; with four endites: proximal praecoxal endite one-segmented, with two pinnate and two bare setae distally; distal praecoxal endite small, incorporated basally into syncoxa, with one pinnate and two bare setae distally; proximal coxal endite large, with one row of spinules subdistally, and one bare and two pinnate setae distally; distal coxal endite cylindrical, with one row of spinules subdistally, and one pinnate spine and two setae (one bare and one pinnate) distally. Allobasis large, with one stout spine and two setae distally; inner part drawn out into a curved strong claw accompanied by one stout pinnate spine and two setae. Endopod small, three-segmented; first segment with two bare setae; second segment with one bare and one geniculate seta; distal segment with one geniculate and three bare setae.
</paragraph>
<paragraph id="F45FF2BC7625F6D06E6D09AFE8C5149E" pageId="0" pageNumber="165">
<emphasis id="3D8792E49C253CA857A87F386FCB22F3" bold="true" italics="true" pageId="0" pageNumber="165">Maxilliped</emphasis>
(Fig.
<figureCitation id="EBD3BA340A6F021C753382677DD0FD73" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3C</figureCitation>
) three-segmented, composed of protopod, and two-segmented endopod. Protopod elongate, with two rows of long outer setules, one row of posterior spinules; with four endites ornamented with inner and anterior spinules: three syncoxal endites represented proximal to distal by one pinnate spine and one pinnate seta, one pinnate spine and two plumose setae, and one pinnate spine and one plumose seta; basal endite represented by one pinnate spine and one plumose seta. Endopod small; first segment with one row of outer spinules and one plumose subdistal seta; second segment with two pinnate distal spines and two pinnate lateral setae, of which proximal one plumose proximally.
</paragraph>
<paragraph id="C522B6D0A527A2850F624BFFEDBF6BF3" pageId="0" pageNumber="165">
<emphasis id="4C3A7CE7CFCEB2950C55C91BA9F94C9E" bold="true" italics="true" pageId="0" pageNumber="165">P1</emphasis>
(Fig.
<figureCitation id="E2EF45959CFC545F2D5E58146F161404" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4A</figureCitation>
). Intercoxal sclerite transversely elongated and narrow, with surface reticulation distally. Praecoxa with spinules on anterior surface. Coxa with various-sized spinules and setules on anterior surface. Basis larger than coxa, with one anterior pore and several rows of anterior spinules; with one long plumose outer and one pinnate inner seta; rami set far away from each other. Exopod one-segmented, with a row of setules on inner margin, and rows of spinules on outer proximal margin and around bases of outer setae; with three inner, two apical, and five outer setae. Endopod one-segmented, with one anterior pore medially, setules along inner and outer margins, and anterior spinules around distal margin; with one inner, one outer, and two apical setae.
</paragraph>
<caption id="1A82C9A88EEEE606D005417532584EE4" doi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165" start="Figure 4" startId="F4">
<paragraph id="B2FA4D002FDEB161DFD253DB7F5F8A80" pageId="0" pageNumber="165">
<emphasis id="189A9E8A15F66321C7054740A77B2F94" bold="true" pageId="0" pageNumber="165">Figure 4.</emphasis>
<taxonomicName id="375C9348445480363A2E47F54DA27312" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="7D0BFE058BFA4D157CD431670B7EEE0E" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
sp. nov. Female. Paratype (MABIK CR00253868)
<emphasis id="7DFEBC5E0F1E0E641265BD1D1BA1F4B2" bold="true" pageId="0" pageNumber="165">A</emphasis>
P1, anterior
<emphasis id="5E7665410E122D6513BB08927E888BB7" bold="true" pageId="0" pageNumber="165">B</emphasis>
P2, anterior
<emphasis id="C0999B2BCC67E210265CBC313B35ACA9" bold="true" pageId="0" pageNumber="165">C</emphasis>
P3, anterior
<emphasis id="C14049E31D56EA9C2F272E54250D3849" bold="true" pageId="0" pageNumber="165">D</emphasis>
P4, anterior
<emphasis id="AEB83D10D62564BCC44FC355499C46A4" bold="true" pageId="0" pageNumber="165">D</emphasis>
' P4, showing abnormal setae on the other pair; Holotype (MABIK CR00253873)
<emphasis id="464D1637DBD05E4A1353F8BC445D1744" bold="true" pageId="0" pageNumber="165">E</emphasis>
P6; Paratype (MABIK CR00253869)
<emphasis id="84FA2F800017208E251E4C4DD8EF1A9A" bold="true" pageId="0" pageNumber="165">F</emphasis>
P2exp
<emphasis id="2C957A8BDFE7B2CB1D467169FA597948" bold="true" pageId="0" pageNumber="165">G</emphasis>
P2enp-2. Scale bars are in
<normalizedToken id="FD99AD999990A38E59696C0EAFB70EB9" originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id="301542F0884342F3F6B46380ED179461" pageId="0" pageNumber="165">
<emphasis id="32D35C38621E5D998D1C4876610397D4" bold="true" italics="true" pageId="0" pageNumber="165">P2</emphasis>
(Fig.
<figureCitation id="E7532C29B9C3DE15596D42E128959612" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4B</figureCitation>
) larger than other thoracic legs. Intercoxal sclerite well-developed, cordiform; anterior surface with conspicuous reticulation, distally with one pair of rows of long setules; distal margin concave. Praecoxa small, with minute spinules along distal margin. Coxa with several rows of various-sized anterior spinules and three rows of long outer spinules. Basis with one anterior pore and numerous rows of various-sized anterior spinules; distal margin with one small acute process between rami; plumose outer seta shorter than those of P1 and P3. Exopod one-segmented, with one anterior pore distally and several rows of setules along inner and outer margins; with five serrate outer spines, increasing in size distally, one serrate apical spines, one plumose apical seta, and four plumose inner setae; outer and distal margins with four acute and three small processes near base of spines. Endopod two-segmented, shorter than exopod; enp-1 with three outer rows of setules and two distal rows of minute spinules, outer margin drawn out into a chitinous process bearing weakly concave tip and inner margin with one plumose seta; enp-2 longer than preceding one, ~ 1.6
<normalizedToken id="2226161C7022B521A016B9AD9BA32186" originalValue="×">x</normalizedToken>
as long as wide, with one outer, two apical, and four inner pinnate setae, outer and distal margins with two acute processes.
</paragraph>
<paragraph id="42C18B0E408BF769C7183355A44F457B" pageId="0" pageNumber="165">
<emphasis id="B7052C289B0C12C2E3413FEB163E9A4E" bold="true" italics="true" pageId="0" pageNumber="165">P3</emphasis>
(Fig.
<figureCitation id="38F4DBE55BAA85DAD7A09E80FCA17111" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4C</figureCitation>
) smaller than P2. Intercoxal sclerite well-developed, subrectangular, with surface reticulation; distal margin concave, with long setules. Praecoxa small, with two distal rows of minute spinules. Coxa with numerous rows of various-sized anterior spinules. Basis with numerous minute anterior spinules and row of inner setules, and one small acute process between rami; anterior pore larger than that of P2; outer seta plumose and ~ 3
<normalizedToken id="00B7001936695700D237E6C93442CC66" originalValue="×">x</normalizedToken>
as long as outer margin of basis. Exopod one-segmented, with one row of outer spinules and several rows of outer and inner setules; outer margin with two blunt and five acute processes; with five serrate outer spines, increasing in size distally, one serrate apical spine and one pinnate apical seta, and three plumose inner setae. Endopod two-segmented, distinctly shorter than exopod; enp-1 with three groups of outer spinules and one plumose inner seta, outer corner drawn out into a chitinous blunt process; enp-2 oval, ~ 2.3
<normalizedToken id="8AD52779B099694EBC2391B343D46B61" originalValue="×">x</normalizedToken>
as long as wide, with one plumose apical seta, outer and inner margins with row of setules.
</paragraph>
<paragraph id="040A118387569A2E7CD8BEC23A2366D7" pageId="0" pageNumber="165">
<emphasis id="730206D3C2D45250CBAF0BA302FC6AE1" bold="true" italics="true" pageId="0" pageNumber="165">P4</emphasis>
(Fig.
<figureCitation id="15032BB529BE822B606302DD500B659E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4D</figureCitation>
) rudimentary, uniramous, comprising outer setophore and exopod, fused basally to fourth pedigerous somite, covered with various-sized spinules; outer setophore cylindrical, longer than exopod, with long plumose seta. Exopod one-segmented, with one apical and one outer setae. Endopod absent.
</paragraph>
<paragraph id="9FBB274848B2341DBEA2E8DF591BE1EB" pageId="0" pageNumber="165">Armature formulae of P1-P4 as follows:</paragraph>
<paragraph id="E72636FFF1890750FACC89B36D9E328C" pageId="0" pageNumber="165">
<table id="A49BB4BFDB25354F3C1EDEAA9B151642" pageId="0" pageNumber="165">
<tr id="76FAD90347CA1F040CF7ADDF8B727D08" pageId="0" pageNumber="165">
<th id="DC24807CD1B9B5AEB33297F27B44FD9C" colspan="1" pageId="0" pageNumber="165" rowspan="1">Leg</th>
<th id="A005C8C5E73564DA4AD0B74D2A31812E" colspan="1" pageId="0" pageNumber="165" rowspan="1">Exopod</th>
<th id="3D8003E7C9CACE61CE6C218B6C5E9A63" colspan="1" pageId="0" pageNumber="165" rowspan="1">Endopod</th>
</tr>
<tr id="BC1F26DA8F4E1EC2019DC60008232496" pageId="0" pageNumber="165">
<td id="E021C039F41ED599BA6C6325D4192B30" colspan="1" pageId="0" pageNumber="165" rowspan="1">P1</td>
<td id="E726B065ABAB64E4A965C7A8C98B9962" colspan="1" pageId="0" pageNumber="165" rowspan="1">325</td>
<td id="B0BE986B4F5E79BEDB331776FFEACCFB" colspan="1" pageId="0" pageNumber="165" rowspan="1">121</td>
</tr>
<tr id="038E8D495EE0B36DD41AA983CCEFFD59" pageId="0" pageNumber="165">
<td id="58CDFD54404387E832AB37383496C5BA" colspan="1" pageId="0" pageNumber="165" rowspan="1">P2</td>
<td id="394ADDDFD1BA17393C8282C66B1E0879" colspan="1" pageId="0" pageNumber="165" rowspan="1">425</td>
<td id="C03D2768E9ADB95B94FF4DB96AB1CEE0" colspan="1" pageId="0" pageNumber="165" rowspan="1">1.[2-4]21</td>
</tr>
<tr id="49925EED157B246EA3337597856BBFAF" pageId="0" pageNumber="165">
<td id="AA7D6B2C02342375D982506C6BD44483" colspan="1" pageId="0" pageNumber="165" rowspan="1">P3</td>
<td id="94723CE43E01F7EC26A8EF7B571A7CAA" colspan="1" pageId="0" pageNumber="165" rowspan="1">325</td>
<td id="EBDDE1766C2CAAED552AC283E585AA26" colspan="1" pageId="0" pageNumber="165" rowspan="1">1.010</td>
</tr>
<tr id="ED376F5F98E1C4AB03633251BF05DBD9" pageId="0" pageNumber="165">
<td id="F443DC73151F1B862BA22D8846D13628" colspan="1" pageId="0" pageNumber="165" rowspan="1">P4</td>
<td id="9AAA35E6267DADFAE9294ED57670618C" colspan="1" pageId="0" pageNumber="165" rowspan="1">011</td>
<td id="AC2CE269E1B1C52143E938027DA254D7" colspan="1" pageId="0" pageNumber="165" rowspan="1">absent</td>
</tr>
</table>
</paragraph>
<paragraph id="20228EFCBA02EFA15251F99AEA13E736" pageId="0" pageNumber="165">
<emphasis id="E10D2743AB07BF2CED91E30FE8BD5423" bold="true" italics="true" pageId="0" pageNumber="165">P5</emphasis>
(Fig.
<figureCitation id="BDFDBDE1929633828383B817137CA8FA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A paragnaths, anterior B maxilla C maxilliped D urosome, ventral view. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure3" httpUri="https://binary.pensoft.net/fig/902794" pageId="0" pageNumber="165">3D</figureCitation>
) bilobate as in P4, fused to somite, covered with minute spinules; outer setophore cylindrical, with one plumose seta. Exopod slightly exceeding setophore, with one plumose outer and one plumose apical seta, the latter ~ 2.5
<normalizedToken id="EE66E54A119515076881ECDD372C0DEA" originalValue="×">x</normalizedToken>
longer than outer seta.
</paragraph>
<paragraph id="4B6F57C016663F8C4F527F21A7DCD288" pageId="0" pageNumber="165">
<emphasis id="0DBBFA2B316951A49966EF6F103F7F46" bold="true" pageId="0" pageNumber="165">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="37E2124C8939BF753F7E5A96B0F1722A" pageId="0" pageNumber="165" type="variability">
<paragraph id="4303BCE267E4CB5A808DD90B86858591" pageId="0" pageNumber="165">Variability.</paragraph>
<paragraph id="E986CB11F6977C8478DC7951E59E6939" pageId="0" pageNumber="165">
The morphological variation in
<taxonomicName id="BD6B4234216AF31E1EE0544C169E24C7" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="8439A244D2E8FBBA31821033C9C62545" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
sp. nov. appears in the armature formula of thoracopods. P2enp-2 presumably has four inner setae in the normal condition, but the P2enp-2 of some specimens (four of 18 specimens) has two or three inner setae.
</paragraph>
</subSubSection>
<subSubSection id="CB9C424415F5162DF84B98370ECE8F2E" pageId="0" pageNumber="165" type="abnormality">
<paragraph id="08CBA2C2C8851CDD52AA8F2CB4BC0658" pageId="0" pageNumber="165">Abnormality.</paragraph>
<paragraph id="BC751CD725550993BD21F7C9BEA5521D" pageId="0" pageNumber="165">
Abnormal exopod and endopod of P2 were observed in one specimen (paratype, MABIK CR00253869) (Fig.
<figureCitation id="0C5E2608DD1A6D2C9AFA4F994F2D0615" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4F, G</figureCitation>
). In comparison with the normal condition of P2 (Fig.
<figureCitation id="0D38FA8D187A4D0E8CE5C6D8D22B98F4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4B</figureCitation>
), the abnormal exopod has a seta in place of the proximal fourth outer spine and a spine in place of the proximal third inner seta (Fig.
<figureCitation id="1F9910F56AFBBEBF53F333203D1943B2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4F</figureCitation>
, indicated by arrowheads), and the enp-2 lacks a small pointed protrusion on either side of the two apical setae (Fig.
<figureCitation id="B81372D49578A52E2162D85A3DB78017" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4G</figureCitation>
, indicated by arrowheads). In addition, paratype (MABIK CR00253868) displayed two setae (the normal condition) on one of the exopods of P4 (Fig.
<figureCitation id="EB240FFF73481C995B78A4A5FAB35F9B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4D</figureCitation>
) and three setae on the other exopod (Fig.
<figureCitation id="CEBF8241E8851C22ECEBA026F95451EC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerviniella bisegmenta sp. nov. Female. Paratype (MABIK CR 00253868) A P 1, anterior B P 2, anterior C P 3, anterior D P 4, anterior D ' P 4, showing abnormal setae on the other pair; Holotype (MABIK CR 00253873) E P 6; Paratype (MABIK CR 00253869) F P 2 exp G P 2 enp- 2. Scale bars are in μm." figureDoi="10.3897/zookeys.1178.105407.figure4" httpUri="https://binary.pensoft.net/fig/902795" pageId="0" pageNumber="165">4D</figureCitation>
').
</paragraph>
</subSubSection>
<subSubSection id="6495DF794B13D682A980BDFD527CD896" pageId="0" pageNumber="165" type="etymology">
<paragraph id="497B28908B586D782FF2EFBBA0831392" pageId="0" pageNumber="165">Etymology.</paragraph>
<paragraph id="D7739BCC54D39DD6F64C5924F6BD08B6" pageId="0" pageNumber="165">
The specific name
<taxonomicName id="7006D0F541D4BD40ABBF10F0AE378773" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="C154B15479F22446DB2CC619C2AE0651" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
is derived from a combination of the Latin prefix
<emphasis id="25AD7B9CB913D9D22B63D2BF18D131C6" italics="true" pageId="0" pageNumber="165">bi</emphasis>
-, meaning 'having two
<normalizedToken id="537A3812C4BBB8AC072F75FFF3D6DF4C" originalValue="parts">parts'</normalizedToken>
and the Latin noun
<emphasis id="1D0CA362BCEF800ACDC11EF3D2AADA26" italics="true" pageId="0" pageNumber="165">segmentum</emphasis>
, meaning
<normalizedToken id="DD0B9D13716482C9180FF2FAFA870EE0" originalValue="cutting">'cutting'</normalizedToken>
or
<normalizedToken id="0582F5AD2FB28419D066A22C3F31FD6A" originalValue="piece">'piece'</normalizedToken>
, and refers to the two-segmented antennary exopod, which is an autapomorphy of this species. It is a noun in the nominative plural.
</paragraph>
</subSubSection>
<subSubSection id="578BE8DFE8C4211B0AA2B72D111B9B4C" pageId="0" pageNumber="165" type="remarks">
<paragraph id="ECF93B9897BBF92B613082B7B6701ED5" pageId="0" pageNumber="165">Remarks.</paragraph>
<paragraph id="4527DCE63A5B0BF772F2B37152D8DAD5" pageId="0" pageNumber="165">
Based on the presence or absence of the P4 endopod,
<bibRefCitation id="4C02AC5CF9A8318EEB35BC40CAD1ABBF" author="Apostolov, A" journalOrPublisher="Libra Scorp, Bourgas" pageId="0" pageNumber="165" refId="B1" refString="Apostolov, A, 2011. BIOICE programme. Les Harpacticoides marins (Crustacea, Copepoda) d'Islande. Libra Scorp, Bourgas" title="BIOICE programme. Les Harpacticoides marins (Crustacea, Copepoda) d'Islande." year="2011">Apostolov (2011)</bibRefCitation>
subdivided the genus
<taxonomicName id="9AB914212799A948835BAA59793384A9" authorityName="Smirnov" authorityYear="1946" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="F076504F2261E0EE0CF266FACE6D4BC3" italics="true" pageId="0" pageNumber="165">Cerviniella</emphasis>
</taxonomicName>
into the
<taxonomicName id="13E1AB5EBF32572CF1672575C91C832D" authorityName="Por" authorityYear="1969" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella brodskayae" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="brodskayae">
<emphasis id="D9D30670FCC1B4B81B5E0EBF76CDBB43" italics="true" pageId="0" pageNumber="165">Cerviniella brodskayae</emphasis>
</taxonomicName>
and
<taxonomicName id="F3720B1D2DD77985348170B90B1607C7" authorityName="Smirnov" authorityYear="1946" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella mirabilipes" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="mirabilipes">
<emphasis id="4C55CF73C764805289121F7AE56FC76C" italics="true" pageId="0" pageNumber="165">Cerviniella mirabilipes</emphasis>
</taxonomicName>
groups.
<taxonomicName id="2EB5A4B158415740B24E793AFE2EBDDD" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="B3A6F7D3490DDBA8CFB10D113ABEB36C" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
sp. nov. lacks the ramus and can be classified within the
<taxonomicName id="1AB0A9E705E1A1138DACB17686E58BCE" authorityName="Smirnov" authorityYear="1946" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella mirabilipes" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="mirabilipes">
<emphasis id="408122E68E72142B5D215F2BA38734AE" italics="true" pageId="0" pageNumber="165">Cerviniella mirabilipes</emphasis>
</taxonomicName>
group, which includes
<taxonomicName id="DE84D2C29AD0CA0D3C791EAA4C31670C" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="0F9C183FAD712C1258C4FAE3D5E3899A" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
,
<taxonomicName id="E5031812B3F8417638D7EA80AC4D0A20" lsidName="C. lagarderei" pageId="0" pageNumber="165" rank="species" species="lagarderei">
<emphasis id="AA0045FFEDF681F88F7938439EC2B955" italics="true" pageId="0" pageNumber="165">C. lagarderei</emphasis>
</taxonomicName>
,
<taxonomicName id="234B1F03204345ADD75EB5C3DE7CD3AD" lsidName="C. langi" pageId="0" pageNumber="165" rank="species" species="langi">
<emphasis id="BA65D426AFE7F185E48F98B0FC74DC86" italics="true" pageId="0" pageNumber="165">C. langi</emphasis>
</taxonomicName>
,
<taxonomicName id="B461D5AFB6BCD010C25C2C11C1D94E6F" lsidName="C. mirabilipes" pageId="0" pageNumber="165" rank="species" species="mirabilipes">
<emphasis id="7787C8E88E0FED0CC7C19E5A4634EF13" italics="true" pageId="0" pageNumber="165">C. mirabilipes</emphasis>
</taxonomicName>
, and
<taxonomicName id="49C8C0B189C119DA9A0A588DD91CE23E" lsidName="C. talpa" pageId="0" pageNumber="165" rank="species" species="talpa">
<emphasis id="4388BDFB3E05102E303344B6E0B4AAE7" italics="true" pageId="0" pageNumber="165">C. talpa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0043A6EBAD18AB7A9C6A99E9FFA3EBB4" pageId="0" pageNumber="165">
The segmentation of the female antennule is useful for differentiating among species of
<taxonomicName id="870FFF3829220CEFD6EC11E0E5E42220" authorityName="Smirnov" authorityYear="1946" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="F12179B5F72B3A0639C93B90F3692E10" italics="true" pageId="0" pageNumber="165">Cerviniella</emphasis>
</taxonomicName>
.
<taxonomicName id="AEBCDF0E15755C32225C215D02178AFE" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="98A8286FDA3AC9129744F93CF4391BB3" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
sp. nov. has an advanced five-segmented antennule, shared in
<taxonomicName id="6D3A7443FDEA107F892239D19B783841" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="3F1A00B5ECE72A4B41F5668E88C22966" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
and
<taxonomicName id="C22573A673CC34748C4DF116AAA8B036" lsidName="C. hitoshii" pageId="0" pageNumber="165" rank="species" species="hitoshii">
<emphasis id="E0C100B80913BE5162FF8AC8E927865C" italics="true" pageId="0" pageNumber="165">C. hitoshii</emphasis>
</taxonomicName>
, but the latter species is in the
<taxonomicName id="078810E26880F4E20C218954CC54A663" authorityName="Por" authorityYear="1969" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella brodskayae" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="brodskayae">
<emphasis id="4DEB1690600E11C57A2ED14C9876AEF3" italics="true" pageId="0" pageNumber="165">Cerviniella brodskayae</emphasis>
</taxonomicName>
group. The other 11
<taxonomicName id="4A886C8EDCB768C2A9287564D8832E75" authorityName="Smirnov" authorityYear="1946" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="9765E8C8FE753764598F2668F370BB16" italics="true" pageId="0" pageNumber="165">Cerviniella</emphasis>
</taxonomicName>
species have a six- or seven-segmented antennule (see discussion below).
</paragraph>
<paragraph id="2E8EC1F5FA3DC2244A98F2BB8AEAE18D" pageId="0" pageNumber="165">
<taxonomicName id="8F03B8C19E97451A6773F48319648C41" authorityName="Cho &amp; Kim &amp; Lee" authorityYear="2023" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella bisegmenta" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="bisegmenta">
<emphasis id="20D182A9D718A9A6C36FD6C9C0117804" italics="true" pageId="0" pageNumber="165">Cerviniella bisegmenta</emphasis>
</taxonomicName>
sp. nov. can be easily distinguished differs in several characters from the same group,
<taxonomicName id="E8260EA164E929E076CD6F02AF142CCD" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="E80491928AE4A5608D4E0FEC4A198B0F" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
. First, the antenna of
<taxonomicName id="E60874F8C73E07D69FFC13F7A068484A" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="815EB3496DCD7D68C7E081908C961D5F" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov. has a two-segmented exopod, compared with the four-segmented antennary exopod of
<taxonomicName id="D84597C4931B1F74D6F437F518BCAD1B" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="4924554B48BC67336C02A029E87190B6" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
. Second, the mandibular endopod of
<taxonomicName id="6595B0C9FE2418C47216F4B552B533D4" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="49DCA51F5871407E0CB60851E7F5BD2C" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov. has two lateral setae, whereas the mandibular endopod of
<taxonomicName id="F86A21243546B447866789D0B0145C3E" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="57A411A80976CDE70E1E833A8F5A37F7" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
has three lateral setae. Third, the basis and endopod of the maxillule of
<taxonomicName id="80883BDDC8DE94D5E8CDFE6C7E789359" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="E090DD76F102F297E1732C1FBA4F232C" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov. have a total of 13 setae, compared with 14 setae in
<taxonomicName id="099DC17675E8A283544783F2E92EA25E" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="F200421C82E660DEC60236BA0684067E" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
. Fourth, the syncoxa of the maxilliped of
<taxonomicName id="A6DF8C83CC24867F352719BA97F6C6BD" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="F2FAA47B047843DA8F26D6B2CB343103" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov. has nine setae and spines, compared with the seven elements of
<taxonomicName id="33A5EF373142BFF6994BF1C57AD0F327" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="93885A1C3C7B6215EF3DC80D9DAE23F3" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
. Fifth, the P1 endopod of
<taxonomicName id="68FE41F7C10FD1F6C55B166842C8B989" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="710F7F7B600A392AC7ABF8A6E2E3866F" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov. has four setae, compared with six setae in
<taxonomicName id="C84D22EC8A9A0A4FEEFED10107BB5E0F" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="EFAA5F5DBC809264FDCAD299EDE8223E" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
. Sixth, the P3 exopod of
<taxonomicName id="DAE4E698AA5F61749C1751AA9B23932A" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="C881EB229E66106454ECA7BA2B8A6990" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov. has three inner setae, compared with two inner setae in
<taxonomicName id="469EB964C175F35F26FF17E907E45E8D" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="CD29C415F55A35BDCD50BE07DACF05C3" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
. Finally, the caudal ramus of
<taxonomicName id="3777F23249B39173E7BE9C62BF11A735" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="5FF955BF34F8649E08207D15D287EF22" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov. is short with a 1.7-fold length relative to width, compared with a 3.5-fold length for
<taxonomicName id="9E2615F446DFED5B8246BB8CF1B9EC63" lsidName="C. danae" pageId="0" pageNumber="165" rank="species" species="danae">
<emphasis id="21668138F2C9576B30663C86D3D18DA8" italics="true" pageId="0" pageNumber="165">C. danae</emphasis>
</taxonomicName>
. The first and seventh aforementioned characteristics provide conclusive evidence for the identification of
<taxonomicName id="C3BA0C36D4D1DCF0D9AA615F97B2A5A6" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="525B597EC1279269E90C337DE9A762C1" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov. In all species of
<taxonomicName id="B23204BA12A53718285CF607A35FCBD7" authorityName="Smirnov" authorityYear="1946" class="Copepoda" family="Aegisthidae" genus="Cerviniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerviniella" order="Harpacticoida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="6B9670BEC72A4E56BFCF090A5569BC0D" italics="true" pageId="0" pageNumber="165">Cerviniella</emphasis>
</taxonomicName>
, except
<taxonomicName id="6A0236AD95945C11A54085A08865EF24" lsidName="C. bisegmenta" pageId="0" pageNumber="165" rank="species" species="bisegmenta">
<emphasis id="11BE41F9008A2C84E6F9B44934D1F4AF" italics="true" pageId="0" pageNumber="165">C. bisegmenta</emphasis>
</taxonomicName>
sp. nov., the antennary exopod is four-segmented, and the length of the caudal rami is&gt; 3-fold to width and longer than the length of the anal somite.
</paragraph>
</subSubSection>
</treatment>
</document>