170 lines
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170 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-PMC="PMC3689139" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-PubMed="23794925" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="AC758877AFC09AE85D5B21BF766CACD8" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Lonchodraco machaerorhynchus Rodrigues & Kellner, 2013, comb. n." docType="treatment" docVersion="6" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="19" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="18" updateTime="1668155938761" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044965" LSID="urn:lsid:plazi:treatment:AC758877AFC09AE85D5B21BF766CACD8" httpUri="http://treatment.plazi.org/id/AC758877AFC09AE85D5B21BF766CACD8" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<subSubSection pageId="17" pageNumber="18" type="nomenclature">
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<paragraph pageId="17" pageNumber="18">
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<taxonomicName LSID="AC758877-AFC0-9AE8-5D5B-21BF766CACD8" authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Lonchodraco machaerorhynchus" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="machaerorhynchus" status="comb. n.">Lonchodraco machaerorhynchus</taxonomicName>
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<taxonomicNameLabel pageId="17" pageNumber="18">comb. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchodraco machaerorhynchus comb. n. Holotype CAMSM B 54855 (Albian, Cambridge Greensand), fragment of the mandibular symphysis A right lateral view B respective line drawing C dorsal view D respective line drawing E ventral view F respective line drawing. Abbreviations: d - dentary, dcr - dentary crest, sul - sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22073" pageId="17" pageNumber="18">Fig. 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="reference_group">
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<paragraph pageId="17" pageNumber="18">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Ornithocheiridae" genus="Ptenodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="machaerorhynchus" status="comb. n.">
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<normalizedToken originalValue="“Ptenodactylus”">"Ptenodactylus"</normalizedToken>
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machaerorhynchus
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</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B77" refString="Seeley, HG, 1869. Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." title="Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." year="1869">Seeley 1869</bibRefCitation>
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: p. xvi [disclaimed]
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</paragraph>
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<paragraph pageId="17" pageNumber="18">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="machaerorhynvhus" status="comb. n.">Ornithocheirus machaerorhynvhus</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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: p. 113, pl. XII, fig. 1-2
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</paragraph>
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<paragraph pageId="17" pageNumber="18">
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<taxonomicName authority="" authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="subSpecies" species="machaeorhynchus" status="comb. n." subSpecies="sic">Lonchodectes machaeorhynchus [sic]</taxonomicName>
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(Seeley):
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 535
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</paragraph>
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<paragraph pageId="17" pageNumber="18">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="machaerorhynchus" status="comb. n.">Lonchodectes machaerorhynchus</taxonomicName>
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(Seeley):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: p. 195, fig. 12D-E, table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="holotype">
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<paragraph pageId="17" pageNumber="18">Holotype:</paragraph>
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<paragraph pageId="17" pageNumber="18">
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CAMSM B54855, partial mandibular symphysis (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchodraco machaerorhynchus comb. n. Holotype CAMSM B 54855 (Albian, Cambridge Greensand), fragment of the mandibular symphysis A right lateral view B respective line drawing C dorsal view D respective line drawing E ventral view F respective line drawing. Abbreviations: d - dentary, dcr - dentary crest, sul - sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22073" pageId="17" pageNumber="18">Fig. 5A-F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="type locality">
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<paragraph pageId="17" pageNumber="18">Type locality.</paragraph>
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<paragraph pageId="17" pageNumber="18">Cambridge, Cambridgeshire, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="type horizon">
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<paragraph pageId="17" pageNumber="18">Type horizon.</paragraph>
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<paragraph pageId="17" pageNumber="18">Cambridge Greensand (Cenomanian; fossils Albian in age).</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="diagnosis">
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<paragraph pageId="17" pageNumber="18">Diagnosis.</paragraph>
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<paragraph pageId="17" pageNumber="18">Lonchodraconid pterosaur with the following combination of characters that distinguishes it from other members of the clade (autapomorphies are marked with an asterisk): deep dentary crest*; ventral margin of the mandible posterior to the dentary crest ascending in lateral view*; ventral depression located posteriorly to the dentary crest*; wide mandibular groove*; approximately 4.5 alveoli per 3 cm of jaw margin.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
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<paragraph pageId="17" pageNumber="18">Description.</paragraph>
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<paragraph lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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CAMSM B54855 consists of a fragment of the posterior portion of the mandibular symphysis.
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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described the mandible as narrow, with parallel alveolar margins, deep mandibular sulcus, spacing between alveoli equal to the size of their diameters, and with the lateral margins forming a sharp keel, which
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
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interpreted as a dentary crest.
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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mentioned the presence of a suture
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<pageBreakToken pageId="18" pageNumber="19" start="start">with</pageBreakToken>
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the angular bone, which would almost reach the tip of the mandible, as a unique character for this species in comparison to the pterosaurs from Germany. We here interpret this structure as the posterior margin of a dentary crest rather than a suture.
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</paragraph>
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<paragraph pageId="18" pageNumber="19">
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In addition to a deep mandibular groove, CAMSM B54855 shares with
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="giganteus">
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<emphasis italics="true" pageId="18" pageNumber="19">Lonchodraco giganteus</emphasis>
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</taxonomicName>
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small and well-spaced alveoli, without significant size variation. However, it differs from this species in having straight alveolar margins in dorsal view (
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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), whereas the margins diverge in
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="giganteus">
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<emphasis italics="true" pageId="18" pageNumber="19">Lonchodraco giganteus</emphasis>
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</taxonomicName>
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. The mandibular crest of
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="machaerorhynchus">
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<emphasis italics="true" pageId="18" pageNumber="19">Lonchodraco machaerorhynchus</emphasis>
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</taxonomicName>
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is deep and its apex coincident with its terminus, with a ventral depression posterior to it. In
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="giganteus">
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<emphasis italics="true" pageId="18" pageNumber="19">Lonchodraco giganteus</emphasis>
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</taxonomicName>
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, the mandibular crest is short and blade-like, being more restricted to the length of the symphysis; the presence of a depression cannot be determined.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/22073" pageId="18" pageNumber="19" start="Figure 5" startId="F5">
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<paragraph pageId="18" pageNumber="19">
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<emphasis bold="true" pageId="18" pageNumber="19">Figure 5.</emphasis>
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="machaerorhynchus">
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<emphasis italics="true" pageId="18" pageNumber="19">Lonchodraco machaerorhynchus</emphasis>
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</taxonomicName>
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comb. n. Holotype CAMSM B54855 (Albian, Cambridge Greensand), fragment of the mandibular symphysis
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<emphasis bold="true" pageId="18" pageNumber="19">A</emphasis>
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right lateral view
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<emphasis bold="true" pageId="18" pageNumber="19">B</emphasis>
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respective line drawing
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<emphasis bold="true" pageId="18" pageNumber="19">C</emphasis>
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dorsal view
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<emphasis bold="true" pageId="18" pageNumber="19">D</emphasis>
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respective line drawing
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<emphasis bold="true" pageId="18" pageNumber="19">E</emphasis>
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ventral view
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<emphasis bold="true" pageId="18" pageNumber="19">F</emphasis>
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respective line drawing. Abbreviations:
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<emphasis bold="true" pageId="18" pageNumber="19">d</emphasis>
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- dentary,
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<emphasis bold="true" pageId="18" pageNumber="19">dcr</emphasis>
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- dentary crest,
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<emphasis bold="true" pageId="18" pageNumber="19">sul</emphasis>
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- sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |