223 lines
17 KiB
XML
223 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.46.32823" ID-GBIF-Dataset="e546c2a6-4668-4253-b2e2-b462f23dab8c" ID-PMC="PMC6379322" ID-Pensoft-Pub="1314-4049-46-1" ID-PubMed="30787668" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-46-1" ModsDocOrigin="MycoKeys 46" ModsDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" checkinTime="1555333436509" checkinUser="pensoft" docAuthor="Cripps, Cathy L., Eberhardt, Ursula, Schuetz, Nicole, Beker, Henry J., Vera S. Evenson, & Horak, Egon" docDate="2019" docId="636F1235B51D2019AC5B643B052803B5" docLanguage="en" docName="MycoKeys 46: 1-54" docOrigin="MycoKeys 46" docSource="http://dx.doi.org/10.3897/mycokeys.46.32823" docTitle="Hebeloma excedens Sacc., Syll. Fung. 5: 806 1887" docType="treatment" docVersion="4" lastPageNumber="35" masterDocId="AC18AA047D78FFBFD06BFFFDFF948577" masterDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" masterLastPageNumber="54" masterPageNumber="1" pageNumber="32" updateTime="1668136151891" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Hebeloma in the Rocky Mountain Alpine Zone</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cripps, Cathy L.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Eberhardt, Ursula</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schuetz, Nicole</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beker, Henry J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vera S. Evenson,</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Horak, Egon</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>46</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.46.32823</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.46.32823</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-46-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="156202108" LSID="urn:lsid:plazi:treatment:636F1235B51D2019AC5B643B052803B5" httpUri="http://treatment.plazi.org/id/636F1235B51D2019AC5B643B052803B5" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32">
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<subSubSection pageId="31" pageNumber="32" type="nomenclature">
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<paragraph pageId="31" pageNumber="32">
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12.
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<taxonomicName authority="Peck) Sacc., Syll. Fung. 5: 806 (1887" authorityYear="1887" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma excedens" order="Agaricales" pageId="31" pageNumber="32" phylum="Basidiomycota" rank="species" species="excedens">Hebeloma excedens (Peck) Sacc., Syll. Fung. 5: 806 (1887)</taxonomicName>
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Figures 6B, 18, 23 (12)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="32" pageNumber="33" type="etymology">
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<paragraph pageId="32" pageNumber="33">
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<pageBreakToken pageId="32" pageNumber="33" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="32" pageNumber="33">For the pileus cuticle which can exceed the lamellae.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
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<paragraph pageId="32" pageNumber="33">Description.</paragraph>
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<paragraph lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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Cortina present. Pileus 10-25 mm in diameter, shallow convex, campanulate, then almost applanate, slight umbo or not, viscid or greasy, medium cocoa brown to orange caramel in center and pale brown on most of the pileus, with
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<pageBreakToken pageId="33" pageNumber="34" start="start">or</pageBreakToken>
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without white tissue at margin, or with whitish rim; margin originally described as extending beyond the lamellae. Pileus thin-fleshed. Lamellae sinuate, subdecurrent, narrow, becoming broader and eroded, very pale, cream with pinkish buff tint, L = 32-48 plus lamellulae. Stipe 30-50
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<normalizedToken originalValue="×">x</normalizedToken>
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2-4 mm, equal, slightly curved, pale cream, silky, pruinose above ring zone, more dingy brown below but still pale, with a golden brown fibrils in zones, blackening towards base. Context whitish in pileus and stipe apex and yellowish brown in lower stipe down to blackish at base; stipe tough, rubbery. Odor: raphanoid or none. Exsiccate: small, pale buff overall, base of stipe dark in some.
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</paragraph>
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<paragraph pageId="33" pageNumber="34">
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Basidiospores yellow brown, elliptical, a few slightly ovoid, no big apiculus, not guttulate, looks almost smooth to very slightly rough even under high magnification (O1), not or only very slightly dextrinoid (D0,D1), and no perispore loosening (P0), 7-11
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<normalizedToken originalValue="×">x</normalizedToken>
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5-6.5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, on average 9.1
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<normalizedToken originalValue="×">x</normalizedToken>
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5.8
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, Q = 1.55. Basidia 20-30
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<normalizedToken originalValue="×">x</normalizedToken>
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6-9
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, clavate, four-spored mostly. Pleurocystidia absent. Cheilocystidia cylindrical in the upper part and slightly swollen to more swollen at the base, rarely fully cylindrical, 30-60
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long
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<normalizedToken originalValue="×">x</normalizedToken>
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4-7
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<normalizedToken originalValue="µm">µm</normalizedToken>
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at apex, 4-6.5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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in middle, and 6-10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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wide at base, some septate. Epicutis thickness 65-200
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, with some encrusted hyphae.
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</paragraph>
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<caption pageId="33" pageNumber="34">
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<paragraph pageId="33" pageNumber="34">
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Figure 18.
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma excedens" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="excedens">Hebeloma excedens</taxonomicName>
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, CLC1685 and ZT9830.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="rocky mountain ecology">
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<paragraph pageId="33" pageNumber="34">Rocky Mountain ecology.</paragraph>
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<paragraph pageId="33" pageNumber="34">
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In alpine with shrub willow
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix glauca" order="Malpighiales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="glauca">Salix glauca</taxonomicName>
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, Colorado.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="materials_examined">
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<paragraph pageId="33" pageNumber="34">Rocky Mountain specimens examined.</paragraph>
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<paragraph pageId="33" pageNumber="34">
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U.S.A. COLORADO: San Juan County, San Juan Mountains. U.S. Basin, 3658 m, with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix glauca" order="Malpighiales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="glauca">Salix glauca</taxonomicName>
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, 8 Aug 2001, CLC1685 (MONT), C. Cripps. Sawatch Range, Independence Pass, 14 Aug 1999 with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., ZT7475 (ETH), E. Horak; 12 Aug 1999 with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., ZT8136 (ETH), E. Horak; 14 Aug 2001 with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix glauca" order="Malpighiales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="glauca">Salix glauca</taxonomicName>
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and
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<taxonomicName lsidName="S. planifolia" pageId="33" pageNumber="34" rank="species" species="planifolia">S. planifolia</taxonomicName>
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, ZT9830 (ETH), E. Horak; 3760 m, with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix glauca" order="Malpighiales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="glauca">Salix glauca</taxonomicName>
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, 13 Aug 2001, CLC1732 (MONT), C. Cripps. Front Range, Loveland Pass, 7 Aug 1999 with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., ZT8074 (ETH), E. Horak.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="materials_examined">
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<paragraph pageId="33" pageNumber="34">Other specimens examined.</paragraph>
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<paragraph pageId="33" pageNumber="34">
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NEW YORK: Saratoga at approx. 100 m, with
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="genus">Pinus</taxonomicName>
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sp. on sandy soil in woodland, Oct 1870, NYS-F-001123, C.H. Peck (holotype).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="discussion">
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<paragraph pageId="33" pageNumber="34">Discussion.</paragraph>
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<paragraph lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma excedens" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="excedens">Hebeloma excedens</taxonomicName>
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was not treated by
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<bibRefCitation pageId="33" pageNumber="34">Beker et al. (2016)</bibRefCitation>
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. The type of
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<taxonomicName lsidName="H. excedens" pageId="33" pageNumber="34" rank="species" species="excedens">H. excedens</taxonomicName>
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fits in with the majority of the RM
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<taxonomicName lsidName="H. excedens" pageId="33" pageNumber="34" rank="species" species="excedens">H. excedens</taxonomicName>
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collections, but the species cannot be clearly separated from
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<taxonomicName lsidName="H. mesophaeum" pageId="33" pageNumber="34" rank="species" species="mesophaeum">H. mesophaeum</taxonomicName>
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(Fig. 6B). Looking at absolute differ
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<pageBreakToken pageId="34" pageNumber="35" start="start">ences</pageBreakToken>
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, the intraspecific variation of the
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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sample (RM + type = 7 sequences) is 0-8 [0-1] bp, whereas the variation in the sample between
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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and
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<taxonomicName lsidName="H. mesophaeum" pageId="34" pageNumber="35" rank="species" species="mesophaeum">H. mesophaeum</taxonomicName>
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is 2-11 [0-4] bp. In terms of absolute differences, the type of
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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is 5-8 [0-1] bp different from other collections referred to this species, but as Fig. 6B shows it is not strongly differentiated from other members of
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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, if ambiguous positions are treated as missing data as in networks or equated to their constituting bases as in the ML tree. In terms of absolute differences, the type of
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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is 5-11 [0-3] bp away from the
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<taxonomicName lsidName="H. mesophaeum" pageId="34" pageNumber="35" rank="species" species="mesophaeum">H. mesophaeum</taxonomicName>
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sequences of the sample. Thus, within the limited support ITS data can give in this case, we do consider the species identification of the RM
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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collections as molecularly supported. Until the question of the distinctness and delimitation of this species can be clarified, we prefer to treat it as an independent taxon.
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma pubescens" order="Agaricales" pageId="34" pageNumber="35" phylum="Basidiomycota" rank="species" species="pubescens">Hebeloma pubescens</taxonomicName>
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Beker & U. Eberh. is another species from the
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<taxonomicName lsidName="H. mesophaeum" pageId="34" pageNumber="35" rank="species" species="mesophaeum">H. mesophaeum</taxonomicName>
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complex that might occur in the sampled habitats of the Rocky Mountains and is close to
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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in Fig. 6B. Based on a small sample (3 sequences available for
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<taxonomicName lsidName="H. pubescens" pageId="34" pageNumber="35" rank="species" species="pubescens">H. pubescens</taxonomicName>
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; 7 sequences for
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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), the species vary 5-10 [1-3] bp in their ITS region.
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma excedens" order="Agaricales" pageId="34" pageNumber="35" phylum="Basidiomycota" rank="species" species="excedens">Hebeloma excedens</taxonomicName>
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was first described by North American mycologist C.H. Peck; the species, with its lageniform to ventricose cheilocystidia and small elliptical, almost smooth, indextrinoid spores belongs to
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<taxonomicName infraspecific-rank="sect." lsidName="H." pageId="34" pageNumber="35" rank="section" section="Hebeloma">H. sect. Hebeloma</taxonomicName>
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. It is closely allied with
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma mesophaeum" order="Agaricales" pageId="34" pageNumber="35" phylum="Basidiomycota" rank="species" species="mesophaeum">Hebeloma mesophaeum</taxonomicName>
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, with which we believe it has often been confused. Separating these two taxa morphologically is rather difficult, but it does appear that the pileus of
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<taxonomicName lsidName="H. excedens" pageId="34" pageNumber="35" rank="species" species="excedens">H. excedens</taxonomicName>
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may be more evenly colored, less yellow brown, less brown in the center, and it was originally described as having a cuticle that extended beyond the lamellae. The stipe surface appears to have fibrils arranged in zones, in contrast to that of
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<taxonomicName lsidName="H. mesophaeum" pageId="34" pageNumber="35" rank="species" species="mesophaeum">H. mesophaeum</taxonomicName>
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. However, further work is required before we can have confidence that these characters are consistently different.
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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We have examined a number of collections from North America that are morphologically and molecularly consistent with this taxon. Based on these studies it would appear that
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma excedens" order="Agaricales" pageId="34" pageNumber="35" phylum="Basidiomycota" rank="species" species="excedens">Hebeloma excedens</taxonomicName>
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is widespread across North America and occurs in a wide variety of habitats.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |