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<document id="9F9D8723B8CE4E470501278E2A44693F" ID-CLB-Dataset="27726" ID-DOI="10.11646/zootaxa.4539.1.1" ID-GBIF-Dataset="e7c7a589-9f74-4546-9a7b-c3192b9659e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615911" ID-ZooBank="2C72727B-79C5-407F-BD92-B12F98196800" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553789058718" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2018" docId="193787A0FFD6FFABF4CBFBD54558CD2D" docLanguage="en" docName="zootaxa.4539.1.1.pdf" docOrigin="Zootaxa 4539 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lydiaster johannae Koehler 1909" docType="treatment" docVersion="9" lastPageNumber="53" masterDocId="E50EFFD8FFE2FF9FF45CFFC04464CA17" masterDocTitle="New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean" masterLastPageNumber="116" masterPageNumber="1" pageNumber="53" updateTime="1698715174966" updateUser="ExternalLinkService">
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<mods:title id="7BB4383C53EE6B802C90F0373E23B6E4">New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean</mods:title>
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<treatment id="193787A0FFD6FFABF4CBFBD54558CD2D" ID-DOI="http://doi.org/10.5281/zenodo.5990777" ID-GBIF-Taxon="154521950" ID-Zenodo-Dep="5990777" LSID="urn:lsid:plazi:treatment:193787A0FFD6FFABF4CBFBD54558CD2D" httpUri="http://treatment.plazi.org/id/193787A0FFD6FFABF4CBFBD54558CD2D" lastPageNumber="53" pageId="52" pageNumber="53">
<subSubSection id="D984653DFFD6FFABF4CBFBD5464ACE38" box="[151,558,1045,1071]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="912136B6FFD6FFABF4CBFBD5464ACE38" blockId="52.[151,558,1045,1104]" box="[151,558,1045,1071]" pageId="52" pageNumber="53">
<heading id="CA6981DAFFD6FFABF4CBFBD5464ACE38" bold="true" box="[151,558,1045,1071]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
<taxonomicName id="569E4D35FFD6FFABF4CBFBD5464ACE38" ID-CoL="6R3F9" authority="Koehler 1909" authorityName="Koehler" authorityYear="1909" box="[151,558,1045,1071]" class="Asteroidea" family="Goniasteridae" genus="Lydiaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="52" pageNumber="53" phylum="Echinodermata" rank="species" species="johannae">
<emphasis id="A3EAEAA4FFD6FFABF4CBFBD5464ACE38" bold="true" box="[151,558,1045,1071]" pageId="52" pageNumber="53">
<emphasis id="A3EAEAA4FFD6FFABF4CBFBD5451BCE38" bold="true" box="[151,383,1045,1071]" italics="true" pageId="52" pageNumber="53">Lydiaster johannae</emphasis>
<bibRefCitation id="F50F4B47FFD6FFABF5DBFBD5464ACE38" author="Koehler, R." box="[391,558,1045,1071]" pageId="52" pageNumber="53" pagination="115 - 131" refId="ref71489" refString="Koehler, R. (1909) An account of the deep-sea Asteroidea collected by the Royal Indian Marine Survey Ship Investigator. Echinoderma of the Indian Museum, Part 5, 115 - 131. https: // doi. org / 10.5962 / bhl. title. 11614" type="book chapter" year="1909">Koehler 1909</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="D984653DFFD6FFABF4CBFBF747AACEA3" pageId="52" pageNumber="53" type="description">
<paragraph id="912136B6FFD6FFABF4CBFBF74559CE47" blockId="52.[151,558,1045,1104]" box="[151,317,1079,1104]" pageId="52" pageNumber="53">
<figureCitation id="09A52A33FFD6FFABF4CBFBF74567CE47" box="[151,259,1079,1104]" captionStart="FIGURE 20" captionStartId="54.[151,250,1908,1930]" captionTargetBox="[151,1436,228,1861]" captionTargetId="figure@54.[151,1436,228,1861]" captionTargetPageId="54" captionText="FIGURE 20. Lydiaster johannae Specimen IE-2013-12128. R=12.5, r=4.6 cm. A. Abactinal surface. Scale bar=3.0 cm. B. Closeup of abactinal plates showing disk-arm transition. Scale bar=0.5 cm. C. Lateral view showing detail on marginal plate surface. Scale bar=0.5. D. Actinal plate surface. Scale bar=3.0 cm. E. Furrow spine, pedicellariae and other adambulacral plate accessories. Scale bar=0.5 cm." httpUri="https://zenodo.org/record/2615966/files/figure.png" pageId="52" pageNumber="53">Figure 20</figureCitation>
AE
</paragraph>
<paragraph id="912136B6FFD6FFABF4CBFB404044CE81" blockId="52.[151,1056,1152,1204]" box="[151,1056,1152,1174]" pageId="52" pageNumber="53">
<bibRefCitation id="F50F4B47FFD6FFABF4CBFB404541CE81" author="Koehler, R." box="[151,293,1152,1174]" pageId="52" pageNumber="53" pagination="115 - 131" refId="ref71489" refString="Koehler, R. (1909) An account of the deep-sea Asteroidea collected by the Royal Indian Marine Survey Ship Investigator. Echinoderma of the Indian Museum, Part 5, 115 - 131. https: // doi. org / 10.5962 / bhl. title. 11614" type="book chapter" year="1909">Koehler 1909</bibRefCitation>
: 91, pls III, fig. 9; VIII, figs, 1,2; VIII, fig.1;
<bibRefCitation id="F50F4B47FFD6FFABF6B2FB40473CCE81" author="Mah, C. L." box="[750,856,1152,1174]" pageId="52" pageNumber="53" pagination="917 - 954" refId="ref72068" refString="Mah, C. L. (2006) Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema, 28 (4), 917 - 954." type="journal article" year="2006">Mah 2006</bibRefCitation>
: 924. (as
<taxonomicName id="569E4D35FFD6FFABF7E5FB40407CCE81" authorityName="Koehler" authorityYear="1909" box="[953,1048,1152,1174]" class="Asteroidea" family="Goniasteridae" genus="Lydiaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="52" pageNumber="53" phylum="Echinodermata" rank="genus">
<emphasis id="A3EAEAA4FFD6FFABF7E5FB40407CCE81" box="[953,1048,1152,1174]" italics="true" pageId="52" pageNumber="53">Lydiaster</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="912136B6FFD6FFABF4CBFB5E47AACEA3" blockId="52.[151,1056,1152,1204]" box="[151,974,1182,1204]" pageId="52" pageNumber="53">
<bibRefCitation id="F50F4B47FFD6FFABF4CBFB5E4555CEA3" author="Halpern, J. A." box="[151,305,1182,1204]" pageId="52" pageNumber="53" pagination="193 - 286" refId="ref70647" refString="Halpern, J. A. (1970) Goniasteridae (Echinodermata: Asteroidea) of the Straits of Florida. Bulletin of Marine Science, 20 (1), 193 - 286." type="journal article" year="1970">Halpern 1970a</bibRefCitation>
: 265; 1970b: 173;
<bibRefCitation id="F50F4B47FFD6FFABF5B1FB5F46FACEA3" author="Clark, A. M." box="[493,670,1182,1204]" pageId="52" pageNumber="53" pagination="187 - 366" refId="ref69077" refString="Clark, A. M. (1993) An index of names of recent Asteroidea-Part 2: Valvatida. E chinoderm Studies, 4, 187 - 366." type="journal article" year="1993">A.M. Clark 1993</bibRefCitation>
: 251 (as
<taxonomicName id="569E4D35FFD6FFABF6A4FB5F47A2CEA3" box="[760,966,1182,1204]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="52" pageNumber="53" phylum="Echinodermata" rank="species" species="johannae">
<emphasis id="A3EAEAA4FFD6FFABF6A4FB5F47A2CEA3" box="[760,966,1182,1204]" italics="true" pageId="52" pageNumber="53">Circeaster johannae</emphasis>
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="D984653DFFD6FFABF4CBFB2140EDCC0D" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="912136B6FFD6FFABF4CBFB2140EDCC0D" blockId="52.[151,1437,1249,1850]" pageId="52" pageNumber="53">
<emphasis id="A3EAEAA4FFD6FFABF4CBFB214575CEED" bold="true" box="[151,273,1249,1274]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
Body stellate (R/r&gt; 2.7) (
<figureCitation id="09A52A33FFD6FFABF664FB2146A7CEED" box="[568,707,1249,1274]" captionStart="FIGURE 20" captionStartId="54.[151,250,1908,1930]" captionTargetBox="[151,1436,228,1861]" captionTargetId="figure@54.[151,1436,228,1861]" captionTargetPageId="54" captionText="FIGURE 20. Lydiaster johannae Specimen IE-2013-12128. R=12.5, r=4.6 cm. A. Abactinal surface. Scale bar=3.0 cm. B. Closeup of abactinal plates showing disk-arm transition. Scale bar=0.5 cm. C. Lateral view showing detail on marginal plate surface. Scale bar=0.5. D. Actinal plate surface. Scale bar=3.0 cm. E. Furrow spine, pedicellariae and other adambulacral plate accessories. Scale bar=0.5 cm." httpUri="https://zenodo.org/record/2615966/files/figure.png" pageId="52" pageNumber="53">Fig. 20A, D</figureCitation>
). Abactinal arm plates similar in size to those on disk (
<figureCitation id="09A52A33FFD6FFABF175FB2141F4CEED" box="[1321,1424,1249,1274]" captionStart="FIGURE 20" captionStartId="54.[151,250,1908,1930]" captionTargetBox="[151,1436,228,1861]" captionTargetId="figure@54.[151,1436,228,1861]" captionTargetPageId="54" captionText="FIGURE 20. Lydiaster johannae Specimen IE-2013-12128. R=12.5, r=4.6 cm. A. Abactinal surface. Scale bar=3.0 cm. B. Closeup of abactinal plates showing disk-arm transition. Scale bar=0.5 cm. C. Lateral view showing detail on marginal plate surface. Scale bar=0.5. D. Actinal plate surface. Scale bar=3.0 cm. E. Furrow spine, pedicellariae and other adambulacral plate accessories. Scale bar=0.5 cm." httpUri="https://zenodo.org/record/2615966/files/figure.png" pageId="52" pageNumber="53">Fig. 20B</figureCitation>
). Granules two to five, and paddle-like pedicellariae with sharp teeth present on disk and arm plates. Abactinal, marginal, actinal surface/peripheral granules, coarse, angular. Marginal plate surface covered with granular spinelets (
<figureCitation id="09A52A33FFD6FFABF555FA8D4515CF71" box="[265,369,1357,1382]" captionStart="FIGURE 20" captionStartId="54.[151,250,1908,1930]" captionTargetBox="[151,1436,228,1861]" captionTargetId="figure@54.[151,1436,228,1861]" captionTargetPageId="54" captionText="FIGURE 20. Lydiaster johannae Specimen IE-2013-12128. R=12.5, r=4.6 cm. A. Abactinal surface. Scale bar=3.0 cm. B. Closeup of abactinal plates showing disk-arm transition. Scale bar=0.5 cm. C. Lateral view showing detail on marginal plate surface. Scale bar=0.5. D. Actinal plate surface. Scale bar=3.0 cm. E. Furrow spine, pedicellariae and other adambulacral plate accessories. Scale bar=0.5 cm." httpUri="https://zenodo.org/record/2615966/files/figure.png" pageId="52" pageNumber="53">Fig. 20C</figureCitation>
). Spinelets more abundant on inferomarginals than on superomarginals. Arms broad, elongate. Actinal plates with one or two of any of the following: enlarged tubercles, paddle-like pedicellariae with sharp teeth, spines with roughened tips (
<figureCitation id="09A52A33FFD6FFABF640FA5546E7CFB9" box="[540,643,1429,1454]" captionStart="FIGURE 20" captionStartId="54.[151,250,1908,1930]" captionTargetBox="[151,1436,228,1861]" captionTargetId="figure@54.[151,1436,228,1861]" captionTargetPageId="54" captionText="FIGURE 20. Lydiaster johannae Specimen IE-2013-12128. R=12.5, r=4.6 cm. A. Abactinal surface. Scale bar=3.0 cm. B. Closeup of abactinal plates showing disk-arm transition. Scale bar=0.5 cm. C. Lateral view showing detail on marginal plate surface. Scale bar=0.5. D. Actinal plate surface. Scale bar=3.0 cm. E. Furrow spine, pedicellariae and other adambulacral plate accessories. Scale bar=0.5 cm." httpUri="https://zenodo.org/record/2615966/files/figure.png" pageId="52" pageNumber="53">Fig. 20E</figureCitation>
). Actinal granules angular, coarse with roughened tips. Furrow spines six to seven, angular in cross-section (
<figureCitation id="09A52A33FFD6FFABF614FA7946CBCFC5" box="[584,687,1465,1490]" captionStart="FIGURE 20" captionStartId="54.[151,250,1908,1930]" captionTargetBox="[151,1436,228,1861]" captionTargetId="figure@54.[151,1436,228,1861]" captionTargetPageId="54" captionText="FIGURE 20. Lydiaster johannae Specimen IE-2013-12128. R=12.5, r=4.6 cm. A. Abactinal surface. Scale bar=3.0 cm. B. Closeup of abactinal plates showing disk-arm transition. Scale bar=0.5 cm. C. Lateral view showing detail on marginal plate surface. Scale bar=0.5. D. Actinal plate surface. Scale bar=3.0 cm. E. Furrow spine, pedicellariae and other adambulacral plate accessories. Scale bar=0.5 cm." httpUri="https://zenodo.org/record/2615966/files/figure.png" pageId="52" pageNumber="53">Fig. 20E</figureCitation>
). Enlarged subambulacral spine laterally adjacent to adambulacral paddle-shaped, pedicellariae with sharp teeth. Enlarged subambulacral granules two to five, polygonal in crosssection present abradial to pedicellariae, adjacent to actinal plate region (from
<bibRefCitation id="F50F4B47FFD6FFABF05AF9C14019CC0D" author="Mah, C. L." box="[1030,1149,1537,1562]" pageId="52" pageNumber="53" pagination="917 - 954" refId="ref72068" refString="Mah, C. L. (2006) Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema, 28 (4), 917 - 954." type="journal article" year="2006">Mah 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D984653DFFD6FFABF49BF9E547D7CCDA" pageId="52" pageNumber="53" type="discussion">
<paragraph id="912136B6FFD6FFABF49BF9E54758CCBD" blockId="52.[151,1437,1249,1850]" pageId="52" pageNumber="53">
<emphasis id="A3EAEAA4FFD6FFABF49BF9E5452CCC29" bold="true" box="[199,328,1573,1598]" pageId="52" pageNumber="53">Comments</emphasis>
.The specimen record herein represents a third occurrence of this species since, the first recorded from
<collectingCountry id="E9897626FFD6FFABF488F989453FCC75" box="[212,347,1609,1634]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
(
<bibRefCitation id="F50F4B47FFD6FFABF536F989458DCC75" author="Mah, C. L." box="[362,489,1609,1634]" pageId="52" pageNumber="53" pagination="917 - 954" refId="ref72068" refString="Mah, C. L. (2006) Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema, 28 (4), 917 - 954." type="journal article" year="2006">Mah. 2006</bibRefCitation>
). A misprint in
<bibRefCitation id="F50F4B47FFD6FFABF6CBF989477FCC75" author="Mah, C. L." box="[663,795,1609,1634]" pageId="52" pageNumber="53" pagination="917 - 954" refId="ref72068" refString="Mah, C. L. (2006) Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema, 28 (4), 917 - 954." type="journal article" year="2006">Mah (2006)</bibRefCitation>
gives incorrect coordinates for the
<taxonomicName id="569E4D35FFD6FFABF0FAF98A414FCC75" box="[1190,1323,1609,1634]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="52" pageNumber="53" phylum="Echinodermata" rank="species" species="johannae">
<emphasis id="A3EAEAA4FFD6FFABF0FAF98A414FCC75" box="[1190,1323,1609,1634]" italics="true" pageId="52" pageNumber="53">L. johannae</emphasis>
</taxonomicName>
specimen (EcAs 11724) specimen listed; correct coordinates are
<geoCoordinate id="F4AA5071FFD6FFABF6A0F9AD470ECC91" box="[764,874,1645,1670]" direction="south" orientation="latitude" pageId="52" pageNumber="53" precision="92" value="-22.261667">22°15.7S</geoCoordinate>
<geoCoordinate id="F4AA5071FFD6FFABF72DF9AD47B2CC91" box="[881,982,1645,1670]" direction="east" orientation="longitude" pageId="52" pageNumber="53" precision="92" value="43.025">43°1.5E</geoCoordinate>
, indicating that the specimen herein was collected not far from EcAs 11724 and from a similar depth.
</paragraph>
<paragraph id="912136B6FFD6FFABF49BF97547D7CCDA" blockId="52.[151,1437,1249,1850]" box="[199,947,1717,1742]" pageId="52" pageNumber="53">
<emphasis id="A3EAEAA4FFD6FFABF49BF975453DCCD9" bold="true" box="[199,345,1717,1742]" pageId="52" pageNumber="53">Occurrence.</emphasis>
Northern Indian Ocean and
<collectingCountry id="E9897626FFD6FFABF6C4F9754745CCD9" box="[664,801,1717,1742]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
,
<quantity id="56669B53FFD6FFABF771F97647CBCCDA" box="[813,943,1718,1742]" metricMagnitude="2" metricUnit="m" metricValue="7.55" metricValueMax="8.1" metricValueMin="7.0" pageId="52" pageNumber="53" unit="m" value="755.0" valueMax="810.0" valueMin="700.0">
700
<specimenCount id="8798FD3FFFD6FFABF739F97647CBCCDA" box="[869,943,1718,1742]" pageId="52" pageNumber="53" type="generic">810 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="D984653DFFD6FFABF49BF9194558CD2D" pageId="52" pageNumber="53" type="description">
<paragraph id="912136B6FFD6FFABF49BF9194558CD2D" blockId="52.[151,1437,1249,1850]" pageId="52" pageNumber="53">
<emphasis id="A3EAEAA4FFD6FFABF49BF91945C4CCE5" bold="true" box="[199,416,1753,1778]" pageId="52" pageNumber="53">Specimen Record.</emphasis>
<materialsCitation id="21F63CEBFFD6FFABF5F4F91A455CCD2D" ID-GBIF-Occurrence="2121242939" collectionCode="IE-" collectorName="Coll. R. von Cosel &amp; V Mascarene Islands &amp; Expedition" country="Mozambique" elevation="700" latitude="-22.3" location="Channel" longLatPrecision="1260" longitude="43.016666" municipality="Mascarene Islands" pageId="52" pageNumber="53" specimenCode="IE-2013-12128" specimenCount="700">
<specimenCode id="C1389ECDFFD6FFABF5F4F91A4633CCE5" box="[424,599,1754,1778]" collectionCode="IE-" pageId="52" pageNumber="53">IE-2013-12128</specimenCode>
.
<collectingCountry id="E9897626FFD6FFABF63EF9194692CCE5" box="[610,758,1753,1778]" name="Mozambique" pageId="52" pageNumber="53">Mozambique</collectingCountry>
<location id="9441606DFFD6FFABF6A2F919473ACCE5" LSID="urn:lsid:plazi:treatment:193787A0FFD6FFABF4CBFBD54558CD2D:9441606DFFD6FFABF6A2F919473ACCE5" box="[766,862,1753,1778]" country="Mozambique" latitude="-22.3" longLatPrecision="1260" longitude="43.016666" municipality="Mascarene Islands" name="Channel" pageId="52" pageNumber="53">Channel</location>
, west coast of
<collectingCountry id="E9897626FFD6FFABF05AF91940EBCCE5" box="[1030,1167,1753,1778]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
,
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,
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,
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<elevation id="1AB3D185FFD6FFABF12EF91A44D4CD02" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="52" pageNumber="53" unit="m" value="700.0">
<quantity id="56669B53FFD6FFABF12EF91A44D4CD02" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="52" pageNumber="53" unit="m" value="700.0">700 m</quantity>
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.
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<collectorName id="3C6B5360FFD6FFABF4E0F93D45EACD01" box="[188,398,1789,1814]" pageId="52" pageNumber="53">Coll. R. von Cosel</collectorName>
aboard R/
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V
<emphasis id="A3EAEAA4FFD6FFABF67DF93E4691CD01" box="[545,757,1789,1814]" italics="true" pageId="52" pageNumber="53">
<collectingMunicipality id="7145ACCCFFD6FFABF67DF93E4691CD01" box="[545,757,1789,1814]" pageId="52" pageNumber="53">Mascarene Islands</collectingMunicipality>
</emphasis>
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III, CREVETTIERE 1986
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, CH 105. 1 wet spec, R=12.5, r=4.6
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>