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<document id="21E3FFC35685F9557C14F64E77DAE73C" ID-CLB-Dataset="299719" ID-DOI="10.11646/zootaxa.5474.5.8" ID-GBIF-Dataset="09a20124-10af-4376-a2f8-cedb504e1b52" ID-ISSN="1175-5326" ID-Zenodo-Dep="12731004" ID-ZooBank="170DE3E1-360B-4B15-A29B-C1813F04E7CC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720374090561" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="03FD87C7EC147A17FF6CFAF47481FD2A" docLanguage="en" docName="zootaxa.5474.5.8.pdf" docOrigin="Zootaxa 5474 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.5.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mannerheimia obscura Coiffait 1982" docType="treatment" docVersion="5" lastPageNumber="591" masterDocId="FFC4FFBFEC117A10FFFBFFE27115FFE7" masterDocTitle="On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="592" masterPageNumber="584" pageNumber="589" updateTime="1720775253469" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DCE913D947C5C882D014330BB39F9DCC">On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<treatment id="03FD87C7EC147A17FF6CFAF47481FD2A" ID-DOI="http://doi.org/10.5281/zenodo.12731018" ID-GBIF-Taxon="235046005" ID-Zenodo-Dep="12731018" LSID="urn:lsid:plazi:treatment:03FD87C7EC147A17FF6CFAF47481FD2A" httpUri="http://treatment.plazi.org/id/03FD87C7EC147A17FF6CFAF47481FD2A" lastPageId="7" lastPageNumber="591" pageId="5" pageNumber="589">
<subSubSection id="C34E655AEC147A15FF6CFAF47370FAD7" box="[151,613,1302,1329]" pageId="5" pageNumber="589" type="nomenclature">
<paragraph id="8BEB36D1EC147A15FF6CFAF47370FAD7" blockId="5.[151,613,1302,1365]" box="[151,613,1302,1329]" pageId="5" pageNumber="589">
<heading id="D0A381BDEC147A15FF6CFAF47370FAD7" bold="true" box="[151,613,1302,1329]" fontSize="11" level="1" pageId="5" pageNumber="589" reason="1">
<taxonomicName id="4C544D52EC147A15FF6CFAF47370FAD7" ID-CoL="3XVMP" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[151,613,1302,1329]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC147A15FF6CFAF47370FAD7" bold="true" box="[151,613,1302,1329]" pageId="5" pageNumber="589">
<emphasis id="B920EAC3EC147A15FF6CFAF470A6FAD6" bold="true" box="[151,435,1302,1329]" italics="true" pageId="5" pageNumber="589">Mannerheimia obscura</emphasis>
<bibRefCitation id="EFC54B20EC147A15FE40FAF47370FAD7" author="Coiffait, H." box="[443,613,1302,1328]" pageId="5" pageNumber="589" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC147A15FF6CFAD9700FFAB2" box="[151,282,1339,1365]" pageId="5" pageNumber="589" type="description">
<paragraph id="8BEB36D1EC147A15FF6CFAD9700FFAB2" blockId="5.[151,613,1302,1365]" box="[151,282,1339,1365]" pageId="5" pageNumber="589">
(
<figureCitation id="136F2A54EC147A15FF64FAD971FDFAB2" box="[159,232,1339,1365]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="5" pageNumber="589">Figs 4</figureCitation>
, 12)
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC147A15FF6CFA9D757DFA70" box="[151,1128,1407,1431]" pageId="5" pageNumber="589" type="reference_group">
<paragraph id="8BEB36D1EC147A15FF6CFA9D757DFA70" blockId="5.[151,1128,1407,1431]" box="[151,1128,1407,1431]" pageId="5" pageNumber="589">
<treatmentCitationGroup id="AB4411FFEC147A15FF6CFA9D757DFA70" box="[151,1128,1407,1431]" pageId="5" pageNumber="589">
<treatmentCitation id="0AF510C0EC147A15FF6CFA9D735BFA70" author="Coiffait, H." box="[151,590,1407,1431]" page="148" pageId="5" pageNumber="589" year="1982">
<taxonomicName id="4C544D52EC147A15FF6CFA9D735BFA70" ID-CoL="3XVMP" authority="Coiffait, 1982: 148" authorityName="Coiffait" authorityPageNumber="148" authorityYear="1982" box="[151,590,1407,1431]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC147A15FF6CFA9D7091FA70" box="[151,388,1407,1431]" italics="true" pageId="5" pageNumber="589">Mannerheimia obscura</emphasis>
<bibRefCitation id="EFC54B20EC147A15FE71FA9D735BFA70" author="Coiffait, H." box="[394,590,1407,1431]" pageId="5" pageNumber="589" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982: 148</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AF510C0EC147A15FDA2FA9D7200FA70" author="Coiffait, H." box="[601,789,1407,1431]" page="274" pageId="5" pageNumber="589" year="1983">
<bibRefCitation id="EFC54B20EC147A15FDA2FA9D7200FA70" author="Coiffait, H." box="[601,789,1407,1431]" pageId="5" pageNumber="589" pagination="273 - 289" refId="ref4890" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe Entomologique de France, New Series, 19, 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait 1983: 274</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AF510C0EC147A15FCDBFA9D72C8FA70" author="Shavrin, A. V." box="[800,989,1407,1431]" httpUri="http://treatment.plazi.org/id/03A187C108652456FF06FF70FDFA1276" page="307" pageId="5" pageNumber="589" year="2021">
<bibRefCitation id="EFC54B20EC147A15FCDBFA9D72C8FA70" author="Shavrin, A. V." box="[800,989,1407,1431]" pageId="5" pageNumber="589" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021: 307</bibRefCitation>
</treatmentCitation>
, 324, 2022: 4
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC147A15FF6CFA27720CF88C" pageId="5" pageNumber="589" type="materials_examined">
<paragraph id="8BEB36D1EC147A15FF6CFA277500F93C" blockId="5.[151,1437,1477,2008]" pageId="5" pageNumber="589">
<emphasis id="B920EAC3EC147A15FF6CFA2770ABFA38" bold="true" box="[151,446,1477,1503]" pageId="5" pageNumber="589">Type material examined.</emphasis>
<typeStatus id="54EF8873EC147A15FE3FFA277339FA38" box="[452,556,1477,1503]" pageId="5" pageNumber="589" type="holotype">Holotype</typeStatus>
(
<figureCitation id="136F2A54EC147A15FDC0FA27736AFA38" box="[571,639,1477,1503]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="5" pageNumber="589">Fig. 4</figureCitation>
; dissected prior to the present study: the aedeagus and the apical genital segment are glued on the same card with the specimen): ‘Kashmir,Chandanwari | b. Pahalgam,
<quantity id="4CAC9B34EC147A15FB07FA0B749EF9E4" box="[1276,1419,1513,1539]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="5" pageNumber="589" unit="m" value="2950.0" valueMax="3100.0" valueMin="2800.0">2800-3100m</quantity>
| lg.H.Franz,
<date id="FFEA1011EC147A15FEE1F9EF7093F9C0" box="[282,390,1549,1575]" pageId="5" pageNumber="589" value="1977-10">Okt.1977</date>
’ &lt;printed&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘
<taxonomicName id="4C544D52EC147A15FCE9F9EF72AEF9C0" authorityName="Maklin" authorityYear="1880" box="[786,955,1549,1575]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="genus">Mannerheimia</taxonomicName>
| obscura | H. COIFFAIT det. 19[printed]79’ &lt;handwritten in black&gt; (
<collectionCode id="ED45AE14EC147A15FE54F9D370E8F9AC" box="[431,509,1585,1611]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
).
<typeStatus id="54EF8873EC147A15FDEBF9D07394F9AB" box="[528,641,1586,1612]" pageId="5" pageNumber="589" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D52FD58EC147A15FD76F9D373DAF9AB" box="[653,719,1585,1612]" count="2" pageId="5" pageNumber="589" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D52FD58EC147A15FD27F9D3721CF9AB" box="[732,777,1585,1612]" count="1" pageId="5" pageNumber="589" type="female">1 ♀</specimenCount>
: the first label as that in the
<typeStatus id="54EF8873EC147A15FBB5F9D375A5F9AC" box="[1102,1200,1585,1611]" pageId="5" pageNumber="589" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED45AE14EC147A15FB3BF9D3741BF9AC" box="[1216,1294,1585,1611]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
);
<specimenCount id="9D52FD58EC147A15FAD9F9D37477F9AB" box="[1314,1378,1585,1612]" count="4" pageId="5" pageNumber="589" type="male">4 ♂♂</specimenCount>
(
<specimenCount id="9D52FD58EC147A15FA89F9D37014F988" count="1" pageId="5" pageNumber="589" type="generic">one specimen</specimenCount>
dissected),
<specimenCount id="9D52FD58EC147A15FE78F9B770BBF997" box="[387,430,1621,1648]" count="1" pageId="5" pageNumber="589" type="female">1 ♀</specimenCount>
: ‘Kashmir,Zagipal b. | Pahalgam,
<quantity id="4CAC9B34EC147A15FCDEF9B772A1F997" box="[805,948,1621,1648]" metricMagnitude="3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="5" pageNumber="589" unit="m" value="3550.0" valueMax="3600.0" valueMin="3500.0">3500-3600m</quantity>
| lg. H.Franz,
<date id="FFEA1011EC147A15FBABF9B775A9F988" box="[1104,1212,1621,1647]" pageId="5" pageNumber="589" value="1977-10">Okt.1977</date>
’ &lt;printed&gt; (
<collectionCode id="ED45AE14EC147A15FAB9F9B77485F988" box="[1346,1424,1621,1647]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
);
<specimenCount id="9D52FD58EC147A15FF6CF99B71D0F973" box="[151,197,1657,1684]" count="1" pageId="5" pageNumber="589" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D52FD58EC147A15FF29F99B7015F973" box="[210,256,1657,1684]" count="1" pageId="5" pageNumber="589" type="female">1 ♀</specimenCount>
: ‘Kashmir, Pahalgam | lg.H.Franz,
<date id="FFEA1011EC147A15FD6FF99B7215F974" box="[660,768,1657,1683]" pageId="5" pageNumber="589" value="1977-10">Okt.1977</date>
’ &lt;printed&gt; (
<collectionCode id="ED45AE14EC147A15FC77F99B72CFF974" box="[908,986,1657,1683]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
);
<specimenCount id="9D52FD58EC147A15FC14F99B7509F973" box="[1007,1052,1657,1684]" count="1" pageId="5" pageNumber="589" type="female">1 ♀</specimenCount>
: ‘Kashmir, Aru | lg.H.Franz,
<date id="FFEA1011EC147A15FA90F99B71C7F950" pageId="5" pageNumber="589" value="1977-10">Okt. 1977</date>
’ &lt;printed&gt; (
<collectionCode id="ED45AE14EC147A15FEA6F97F70BEF950" box="[349,427,1693,1719]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
). All
<typeStatus id="54EF8873EC147A15FE12F97C7341F95F" box="[489,596,1694,1720]" pageId="5" pageNumber="589" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘PARATYPE’. All types with additional printed label: ‘
<taxonomicName id="4C544D52EC147A15FEC7F92370F5F93C" authorityName="Maklin" authorityYear="1880" box="[316,480,1729,1755]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC3EC147A15FEC7F92370F5F93C" box="[316,480,1729,1755]" italics="true" pageId="5" pageNumber="589">Mannerheimia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C544D52EC147A15FE0FF92373E7F93C" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[500,754,1729,1755]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC147A15FE0FF923735BF93C" box="[500,590,1729,1755]" italics="true" pageId="5" pageNumber="589">obscura</emphasis>
<bibRefCitation id="EFC54B20EC147A15FDAEF92373E7F93C" author="Coiffait, H." box="[597,754,1729,1755]" pageId="5" pageNumber="589" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</taxonomicName>
| Shavrin A.
<collectionCode id="ED45AE14EC147A15FC80F9207284F93B" box="[891,913,1730,1756]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="589" type="Museum">V</collectionCode>
. det. 2024’.
</paragraph>
<paragraph id="8BEB36D1EC147A15FF3CF907720CF88C" blockId="5.[151,1437,1477,2008]" pageId="5" pageNumber="589">
<emphasis id="B920EAC3EC147A15FF3CF9077320F918" bold="true" box="[199,565,1765,1791]" pageId="5" pageNumber="589">Additional material examined.</emphasis>
<materialsCitation id="3B3C3C8CEC147A15FDC7F90771E0F8C4" ID-GBIF-Occurrence="4905213301" collectingDate="1977-10" collectionCode="NMW" collectorName="H. Franz" country="India" location="Pahalgam" municipality="Kashmir" pageId="5" pageNumber="589" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="KASHMIR">
<emphasis id="B920EAC3EC147A15FDC7F9077200F918" bold="true" box="[572,789,1765,1791]" pageId="5" pageNumber="589">
<collectingCountry id="F3437641EC147A15FDC7F9077386F918" box="[572,659,1765,1791]" name="India" pageId="5" pageNumber="589">INDIA</collectingCountry>
:
<collectingRegion id="4990F833EC147A15FD64F9077219F918" box="[671,780,1765,1791]" country="India" name="KASHMIR" pageId="5" pageNumber="589">KASHMIR</collectingRegion>
:
</emphasis>
<specimenCount id="9D52FD58EC147A15FCE6F907725CF8E7" box="[797,841,1765,1792]" count="1" pageId="5" pageNumber="589" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D52FD58EC147A15FCAEF9077282F8E7" box="[853,919,1765,1792]" count="2" pageId="5" pageNumber="589" type="female">2 ♀♀</specimenCount>
: ‘
<collectingMunicipality id="6B8FACABEC147A15FC53F907751AF918" box="[936,1039,1765,1791]" pageId="5" pageNumber="589">Kashmir</collectingMunicipality>
,
<location id="8E8B600AEC147A15FBE1F907759DF918" LSID="urn:lsid:plazi:treatment:03FD87C7EC147A17FF6CFAF47481FD2A:8E8B600AEC147A15FBE1F907759DF918" box="[1050,1160,1765,1791]" country="India" municipality="Kashmir" name="Pahalgam" pageId="5" pageNumber="589" stateProvince="KASHMIR">Pahalgam</location>
lg.
<collectorName id="26A15307EC147A15FB4FF904740FF8E7" box="[1204,1306,1766,1792]" pageId="5" pageNumber="589">H. Franz</collectorName>
,
<date id="FFEA1011EC147A15FADEF907748DF918" box="[1317,1432,1765,1791]" pageId="5" pageNumber="589" value="1977-10">
<collectingDate id="EFAEE9F9EC147A15FADEF907748DF918" box="[1317,1432,1765,1791]" pageId="5" pageNumber="589" value="1977-10">Okt. 1977</collectingDate>
</date>
’ (
<collectionCode id="ED45AE14EC147A15FF64F8EB71F8F8C4" box="[159,237,1801,1827]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3C8CEC147A15FEFAF8EB7080F8A0" ID-GBIF-Occurrence="4905213304" collectionCode="NMW" collectorName="H. Franz'" country="India" elevation="3000" location="Himachal-Pradesh Indien" municipality="Rohtangpass" pageId="5" pageNumber="589" specimenCount="32" specimenCount-female="16" specimenCount-male="16" stateProvince="Himachal Pradesh">
<emphasis id="B920EAC3EC147A15FEFAF8EB70ECF8C4" bold="true" box="[257,505,1801,1827]" pageId="5" pageNumber="589">
<collectingRegion id="4990F833EC147A15FEFAF8EB70E0F8C4" box="[257,501,1801,1827]" country="India" name="Himachal Pradesh" pageId="5" pageNumber="589">HIMACHAL PRADESH</collectingRegion>
:
</emphasis>
<specimenCount id="9D52FD58EC147A15FDFAF8EB7344F8C3" box="[513,593,1801,1828]" count="16" pageId="5" pageNumber="589" type="male">16 ♂♂</specimenCount>
,
<specimenCount id="9D52FD58EC147A15FDA4F8EB73BAF8C3" box="[607,687,1801,1828]" count="16" pageId="5" pageNumber="589" type="female">16 ♀♀</specimenCount>
: ‘
<collectingMunicipality id="6B8FACABEC147A15FD3BF8EB7258F8C4" box="[704,845,1801,1827]" pageId="5" pageNumber="589">Rohtangpass</collectingMunicipality>
S-Hang
<quantity id="4CAC9B34EC147A15FC48F8EB755FF8C3" box="[947,1098,1801,1828]" metricMagnitude="3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="5" pageNumber="589" unit="m" value="3000.0" valueMax="3500.0" valueMin="2500.0">
<elevation id="0079D1E2EC147A15FC48F8EB755FF8C3" box="[947,1098,1801,1828]" metricMagnitude="3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="5" pageNumber="589" unit="m" value="3000.0" valueMax="3500.0" valueMin="2500.0">2500-3500 m</elevation>
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<location id="8E8B600AEC147A15FBA9F8EB7466F8C4" LSID="urn:lsid:plazi:treatment:03FD87C7EC147A17FF6CFAF47481FD2A:8E8B600AEC147A15FBA9F8EB7466F8C4" box="[1106,1395,1801,1827]" country="India" municipality="Rohtangpass" name="Himachal-Pradesh Indien" pageId="5" pageNumber="589" stateProvince="Himachal Pradesh">Himachal-Pradesh Indien</location>
, lg.
<collectorName id="26A15307EC147A15FF6CF8CF71EFF8A0" box="[151,250,1837,1863]" pageId="5" pageNumber="589">H.Franz’</collectorName>
(cSh,
<collectionCode id="ED45AE14EC147A15FEBAF8CF709BF8A0" box="[321,398,1837,1863]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
)
</materialsCitation>
;
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<specimenCount id="9D52FD58EC147A15FE5BF8CF70DEF8AF" box="[416,459,1837,1864]" count="1" pageId="5" pageNumber="589" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D52FD58EC147A15FE2CF8CF7317F8AF" box="[471,514,1837,1864]" count="1" pageId="5" pageNumber="589" type="female">1 ♀</specimenCount>
: ‘
<collectingMunicipality id="6B8FACABEC147A15FDE8F8CF73A4F8A0" box="[531,689,1837,1863]" pageId="5" pageNumber="589">Umg. Katrain</collectingMunicipality>
b. Kulu
<location id="8E8B600AEC147A15FCEAF8CF7525F8A0" LSID="urn:lsid:plazi:treatment:03FD87C7EC147A17FF6CFAF47481FD2A:8E8B600AEC147A15FCEAF8CF7525F8A0" box="[785,1072,1837,1863]" country="India" municipality="Umg. Katrain" name="Himachal-Pradesh Indien" pageId="5" pageNumber="589" stateProvince="HIMACHAL PRADESH">Himachal-Pradesh Indien</location>
, lg.
<collectorName id="26A15307EC147A15FBA4F8CC75D2F8A0" box="[1119,1223,1837,1864]" pageId="5" pageNumber="589">H. Franz’</collectorName>
(
<collectionCode id="ED45AE14EC147A15FB2DF8CF7431F8A0" box="[1238,1316,1837,1863]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3C8CEC147A15FACDF8CF7200F88C" ID-GBIF-Occurrence="4905213305" collectionCode="NMW" collectorName="Franz'" country="Nepal" county="Godawari" location="Nepal" municipality="Kathmandu" pageId="5" pageNumber="589" specimenCount="1" specimenCount-male="1" stateProvince="HIMACHAL PRADESH">
<emphasis id="B920EAC3EC147A15FACDF8CF7489F8A0" bold="true" box="[1334,1436,1837,1863]" pageId="5" pageNumber="589">
<collectingCountry id="F3437641EC147A15FACDF8CF7482F8A0" box="[1334,1431,1837,1863]" name="Nepal" pageId="5" pageNumber="589">NEPAL</collectingCountry>
:
</emphasis>
<specimenCount id="9D52FD58EC147A15FF6CF8B371D1F88B" box="[151,196,1873,1900]" count="1" pageId="5" pageNumber="589" type="male">1 ♂</specimenCount>
: ‘
<collectingCounty id="628A4E5DEC147A15FF2FF8B3705DF88C" box="[212,328,1873,1899]" pageId="5" pageNumber="589">Godawari</collectingCounty>
b.
<collectingMunicipality id="6B8FACABEC147A15FE90F8B370E4F88C" box="[363,497,1873,1899]" pageId="5" pageNumber="589">Kathmandu</collectingMunicipality>
,
<location id="8E8B600AEC147A15FE00F8B37354F88C" LSID="urn:lsid:plazi:treatment:03FD87C7EC147A17FF6CFAF47481FD2A:8E8B600AEC147A15FE00F8B37354F88C" box="[507,577,1873,1899]" country="Nepal" county="Godawari" municipality="Kathmandu" name="Nepal" pageId="5" pageNumber="589" stateProvince="HIMACHAL PRADESH">Nepal</location>
, lg.
<collectorName id="26A15307EC147A15FD9EF8B373BAF88C" box="[613,687,1873,1899]" pageId="5" pageNumber="589">Franz’</collectorName>
(
<collectionCode id="ED45AE14EC147A15FD44F8B37218F88C" box="[703,781,1873,1899]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="589">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC147A17FF3CF89771E3FE82" lastPageId="7" lastPageNumber="591" pageId="5" pageNumber="589" type="description">
<paragraph id="8BEB36D1EC147A15FF3CF89775EFF830" blockId="5.[151,1437,1477,2008]" pageId="5" pageNumber="589">
<emphasis id="B920EAC3EC147A15FF3CF8977060F868" bold="true" box="[199,373,1909,1935]" pageId="5" pageNumber="589">Redescription.</emphasis>
Measurements (n=90): HW: 0.53–0.60; HL: 0.30–0.39; OL: 0.15–0.17; AL (
<typeStatus id="54EF8873EC147A15FB1DF897745AF868" box="[1254,1359,1909,1935]" pageId="5" pageNumber="589" type="holotype">holotype</typeStatus>
): 1.07; PL: 0.48–0.54; PW: 0.83–0.90; ESL: 0.95–0.97; EW: 0.98–1.17; MTbL (
<typeStatus id="54EF8873EC147A15FC20F87B7551F854" box="[987,1092,1945,1971]" pageId="5" pageNumber="589" type="holotype">holotype</typeStatus>
): 0.57; MTrL (
<typeStatus id="54EF8873EC147A15FB15F87B7442F854" box="[1262,1367,1945,1971]" pageId="5" pageNumber="589" type="holotype">holotype</typeStatus>
): 0.31 (MTrL 1–4: 0.19; MTrL 5: 0.12); AW: 1.05–1.25; AedL: 0.66–0.72; BL: 2.55–3.40 (
<typeStatus id="54EF8873EC147A15FBBDF85F75BBF830" box="[1094,1198,1981,2007]" pageId="5" pageNumber="589" type="holotype">holotype</typeStatus>
: 3.37).
</paragraph>
<caption id="DF2B6659EC177A16FF6CFB7172ACFB4C" ID-DOI="http://doi.org/10.5281/zenodo.12731016" ID-Zenodo-Dep="12731016" box="[151,953,1171,1195]" httpUri="https://zenodo.org/record/12731016/files/figure.png" pageId="6" pageNumber="590" startId="6.[151,250,1171,1195]" targetBox="[151,1435,181,1147]" targetPageId="6" targetType="figure">
<paragraph id="8BEB36D1EC177A16FF6CFB7172ACFB4C" blockId="6.[151,953,1171,1195]" box="[151,953,1171,1195]" pageId="6" pageNumber="590">
<emphasis id="B920EAC3EC177A16FF6CFB71700BFB4C" bold="true" box="[151,286,1171,1195]" pageId="6" pageNumber="590">FIGURE 11.</emphasis>
Distribution of
<taxonomicName id="4C544D52EC177A16FE3AFB7173BBFB4C" authorityName="Coiffait" authorityYear="1982" box="[449,686,1171,1195]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="590" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC177A16FE3AFB7173BBFB4C" box="[449,686,1171,1195]" italics="true" pageId="6" pageNumber="590">Mannerheimia obscura</emphasis>
</taxonomicName>
in the Himalayan Region.
</paragraph>
</caption>
<paragraph id="8BEB36D1EC177A16FF3CFB1C7443F9DF" blockId="6.[151,1437,1277,2024]" pageId="6" pageNumber="590">
Body convex, elongate (
<figureCitation id="136F2A54EC177A16FE25FB1C7337FAFF" box="[478,546,1278,1304]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="6" pageNumber="590">Fig. 4</figureCitation>
and
<figureCitation id="136F2A54EC177A16FDA6FB1C73B5FAF0" box="[605,672,1277,1304]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="6" pageNumber="590">
Fig.
<quantity id="4CAC9B34EC177A16FD69FB1F73B5FAF0" box="[658,672,1277,1303]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="590" unit="in" value="6.0">6</quantity>
</figureCitation>
in
<bibRefCitation id="EFC54B20EC177A16FD33FB1C726AFAFF" author="Shavrin, A. V." box="[712,895,1278,1304]" pageId="6" pageNumber="590" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin (2021))</bibRefCitation>
. Body yellowish-brown to reddish-brown, with distinctly darker head (lateral and basal portions of pronotum, laterobasal and apical portions of elytra, and paratergites usually paler); antennomeres 4–11 or 5–11 brown; mouthparts, antennomeres 1–3 or 1–4 and legs yellow. Body without microsculpture except for abdomen with very fine transverse or isodiametric dense microreticulation. Head with irregular and sparse, moderately fine punctation; punctation of pronotum realtively dense, slightly larger and deeper than that in middle portion of head, sparser and finer in middle, sometimes with impunctate portions in middle or in mediobasal third; scutellum with several very fine punctures in middle or without them; punctation of elytra about as that on pronotum, but slightly larger and deeper, sometimes finer along suture, punctures in middle of each elytron forming five to six vague and tangled rows of serial punctures, indistinct in some specimens.
</paragraph>
<paragraph id="8BEB36D1EC177A16FF3CF9A0724AF92F" blockId="6.[151,1437,1277,2024]" pageId="6" pageNumber="590">Head 1.7 times as broad as long, with very short and indistinct anteocellar foveae about as long as diameter of ocellus. Antennomere 3 distinctly longer and narrower than 2, 4 about twice shorter and slightly broader than 3, 5–6 slightly longer and distinctly broader than 4, 7–9 slightly longer and broader than 6, 10 slightly shorter than 9, apical antennomere slightly shorter than two preceding antennomeres.</paragraph>
<paragraph id="8BEB36D1EC177A16FF3CF93070DAF8F7" blockId="6.[151,1437,1277,2024]" pageId="6" pageNumber="590">Pronotum 1.6–1.7 times as broad as long, 1.5 times as broad as head, widest in middle, sometimes with slightly sinuate laterobasal margins.</paragraph>
<paragraph id="8BEB36D1EC177A16FF3CF8F873F4F8BF" blockId="6.[151,1437,1277,2024]" pageId="6" pageNumber="590">Elytra slightly or strongly widened apicad, 1.7–1.9 times as long as pronotum. Surface of elytra in middle with irregular longitudinal elevations in some specimens.</paragraph>
<paragraph id="8BEB36D1EC177A16FF3CF8807499F80F" blockId="6.[151,1437,1277,2024]" pageId="6" pageNumber="590">
Male. Apical margins of abdominal tergite VIII and sternite VIII widely concave. Aedeagus with wide basal portion, gradually narrowed apically toward widely rounded apex; parameres wide, significantly exceeding apex of median lobe, widened apically, with row of small distinct thorns along inner margins of apical lobes; internal sac moderately wide and long (Fig.
<quantity id="4CAC9B34EC177A16FE05F82F7322F80F" box="[510,567,1997,2024]" metricMagnitude="0" metricUnit="m" metricValue="1.5493999999999999" pageId="6" pageNumber="590" unit="in" value="61.0">61 in</quantity>
<bibRefCitation id="EFC54B20EC177A16FDC5F82C73E1F80F" author="Shavrin, A. V." box="[574,756,1998,2024]" pageId="6" pageNumber="590" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin (2021))</bibRefCitation>
. Lateral aspect of aedeagus as in Fig.
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<bibRefCitation id="EFC54B20EC177A16FB27F82C749DF80F" author="Shavrin, A. V." box="[1244,1416,1998,2024]" pageId="6" pageNumber="590" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin (2021)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEB36D1EC167A17FF3CFF757490FF56" blockId="7.[151,1437,151,717]" box="[199,1413,151,177]" pageId="7" pageNumber="591">Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</paragraph>
<paragraph id="8BEB36D1EC167A17FF3CFF5871E3FE82" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="591">
<emphasis id="B920EAC3EC167A17FF3CFF5870A1FF32" bold="true" box="[199,436,186,213]" pageId="7" pageNumber="591">Comparative notes.</emphasis>
<taxonomicName id="4C544D52EC167A17FE45FF5973D2FF32" authorityName="Coiffait" authorityYear="1982" box="[446,711,187,213]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="591" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC167A17FE45FF5973D2FF32" box="[446,711,187,213]" italics="true" pageId="7" pageNumber="591">Mannerheimia obscura</emphasis>
</taxonomicName>
can be easily distinguished from the remaining species of the genus by the completely different external morphology of the aedeagus, with widely rounded apex of the median lobe (somewhat resembles some specimens of widespread
<taxonomicName id="4C544D52EC167A17FCB9FEE175FBFEFA" authority="(Motschulsky, 1860))" authorityName="" baseAuthorityName="Motschulsky" baseAuthorityYear="1860" box="[834,1262,259,285]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="591" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B920EAC3EC167A17FCB9FEE172F8FEFA" box="[834,1005,259,285]" italics="true" pageId="7" pageNumber="591">M. brevipennis</emphasis>
(Motschulsky, 1860))
</taxonomicName>
, and the shape of the apical portion of the parameres with the presence of small thorns along each latero-apical margin (unique feature).
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC167A17FF3CFE8C729DFE12" pageId="7" pageNumber="591" type="distribution">
<paragraph id="8BEB36D1EC167A17FF3CFE8C706DFE4A" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="591">
<emphasis id="B920EAC3EC167A17FF3CFE8C7074FE6F" bold="true" box="[199,353,366,392]" pageId="7" pageNumber="591">Distribution.</emphasis>
<taxonomicName id="4C544D52EC167A17FE91FE8D7364FE6E" authorityName="Coiffait" authorityYear="1982" box="[362,625,367,393]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="591" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC167A17FE91FE8D7364FE6E" box="[362,625,367,393]" italics="true" pageId="7" pageNumber="591">Mannerheimia obscura</emphasis>
</taxonomicName>
is distributed from
<collectingCountry id="F3437641EC167A17FCADFE8D72A0FE6E" box="[854,949,367,393]" name="Pakistan" pageId="7" pageNumber="591">Pakistan</collectingCountry>
and Kashmir to
<collectingRegion id="4990F833EC167A17FB83FE8D7452FE6E" box="[1144,1351,367,393]" country="India" name="Himachal Pradesh" pageId="7" pageNumber="591">Himachal Pradesh</collectingRegion>
(
<collectingCountry id="F3437641EC167A17FAA3FE8D7480FE6E" box="[1368,1429,367,393]" name="India" pageId="7" pageNumber="591">India</collectingCountry>
) and
<collectingCountry id="F3437641EC167A17FF3CFE71701FFE4A" box="[199,266,403,429]" name="Nepal" pageId="7" pageNumber="591">Nepal</collectingCountry>
(
<figureCitation id="136F2A54EC167A17FEE2FE71707FFE4A" box="[281,362,403,429]" captionStart="FIGURE 11" captionStartId="6.[151,250,1171,1195]" captionTargetBox="[151,1435,181,1147]" captionTargetId="figure-16@6.[151,1435,181,1147]" captionTargetPageId="6" captionText="FIGURE 11. Distribution of Mannerheimia obscura in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.12731016" httpUri="https://zenodo.org/record/12731016/files/figure.png" pageId="7" pageNumber="591">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BEB36D1EC167A17FF3CFE54729DFE12" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="591">
<emphasis id="B920EAC3EC167A17FF3CFE54705EFE37" bold="true" box="[199,331,438,464]" pageId="7" pageNumber="591">Bionomics.</emphasis>
Specimens were collected at altitudes from
<quantity id="4CAC9B34EC167A17FCCEFE5572F6FE36" box="[821,995,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.85" metricValueMax="3.6" metricValueMin="2.1" pageId="7" pageNumber="591" unit="m" value="2850.0" valueMax="3600.0" valueMin="2100.0">2100 to 3600 m</quantity>
a.s.l. (
<bibRefCitation id="EFC54B20EC167A17FBD0FE5575D0FE36" author="Shavrin, A. V." box="[1067,1221,439,465]" pageId="7" pageNumber="591" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021</bibRefCitation>
, the present study).
<specimenCount id="9D52FD58EC167A17FF6CFE397022FE12" box="[151,311,475,501]" count="1" pageId="7" pageNumber="591" type="generic">One specimen</specimenCount>
from Kashmir was sifted from moss (
<bibRefCitation id="EFC54B20EC167A17FD1BFE397269FE12" author="Shavrin, A. V." box="[736,892,475,501]" pageId="7" pageNumber="591" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC167A17FF3CFE1D7481FD2A" pageId="7" pageNumber="591" type="discussion">
<paragraph id="8BEB36D1EC167A17FF3CFE1D7481FD2A" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="591">
<emphasis id="B920EAC3EC167A17FF3CFE1D702EFDFE" bold="true" box="[199,315,511,537]" pageId="7" pageNumber="591">Remarks.</emphasis>
<taxonomicName id="4C544D52EC167A17FEB8FE1D735FFDFE" authorityName="Coiffait" authorityYear="1982" box="[323,586,511,537]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="591" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC167A17FEB8FE1D735FFDFE" box="[323,586,511,537]" italics="true" pageId="7" pageNumber="591">Mannerheimia obscura</emphasis>
</taxonomicName>
based on several specimens was originally described from Kashmir,
<collectingCountry id="F3437641EC167A17FAA7FE1D748CFDFE" box="[1372,1433,511,537]" name="India" pageId="7" pageNumber="591">India</collectingCountry>
.
<bibRefCitation id="EFC54B20EC167A17FF6CFDC17028FDDA" author="Coiffait, H." box="[151,317,547,573]" pageId="7" pageNumber="591" pagination="273 - 289" refId="ref4890" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe Entomologique de France, New Series, 19, 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait (1983)</bibRefCitation>
recorded it from “Pahalgam, Shashnag,
<quantity id="4CAC9B34EC167A17FD0CFDC17259FDDA" box="[759,844,547,573]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="591" unit="m" value="3000.0">3000 m</quantity>
” (Kashmir).
<bibRefCitation id="EFC54B20EC167A17FC20FDC17594FDDA" author="Shavrin, A. V." box="[987,1153,547,573]" pageId="7" pageNumber="591" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin (2021)</bibRefCitation>
recorded it from
<collectingCountry id="F3437641EC167A17FAC6FDC17489FDDA" box="[1341,1436,547,573]" name="Pakistan" pageId="7" pageNumber="591">Pakistan</collectingCountry>
and Kashmir, provided the photograph of the habitus (
<bibRefCitation id="EFC54B20EC167A17FD04FDA57289FD86" author="Shavrin, A. V." box="[767,924,583,609]" pageId="7" pageNumber="591" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021</bibRefCitation>
:
<figureCitation id="136F2A54EC167A17FC53FDA572F9FD86" box="[936,1004,583,609]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="7" pageNumber="591">Fig. 6</figureCitation>
) and figures of the aedeagus (
<bibRefCitation id="EFC54B20EC167A17FAB8FDA571C6FD62" author="Shavrin, A. V." pageId="7" pageNumber="591" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021</bibRefCitation>
: Figs. 61–62), and included this species to the key of the genus. Later, the same author recorded additional material from
<collectingCountry id="F3437641EC167A17FEC3FD6D7082FD4E" box="[312,407,655,681]" name="Pakistan" pageId="7" pageNumber="591">Pakistan</collectingCountry>
(
<bibRefCitation id="EFC54B20EC167A17FE5DFD6D7351FD4E" author="Shavrin, A. V." box="[422,580,655,681]" pageId="7" pageNumber="591" pagination="1 - 4" refId="ref4990" refString="Shavrin, A. V. (2022) A new species and new records of the genus Mannerheimia Maklin, 1880 from Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Journal of Insect Biodiversity, 32 (1), 1 - 4. https: // doi. org / 10.12976 / jib / 2022.32.1.1" type="journal article" year="2022">Shavrin 2022</bibRefCitation>
).
<taxonomicName id="4C544D52EC167A17FDACFD6D7248FD4E" authorityName="Coiffait" authorityYear="1982" box="[599,861,655,681]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="591" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC167A17FDACFD6D7248FD4E" box="[599,861,655,681]" italics="true" pageId="7" pageNumber="591">Mannerheimia obscura</emphasis>
</taxonomicName>
is the most widely distributed species of the genus in the Himalayan Region (
<figureCitation id="136F2A54EC167A17FE44FD517304FD2A" box="[447,529,691,717]" captionStart="FIGURE 11" captionStartId="6.[151,250,1171,1195]" captionTargetBox="[151,1435,181,1147]" captionTargetId="figure-16@6.[151,1435,181,1147]" captionTargetPageId="6" captionText="FIGURE 11. Distribution of Mannerheimia obscura in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.12731016" httpUri="https://zenodo.org/record/12731016/files/figure.png" pageId="7" pageNumber="591">Fig. 11</figureCitation>
). It is here recorded from
<collectingRegion id="4990F833EC167A17FCCEFD517517FD2A" box="[821,1026,691,717]" country="India" name="Himachal Pradesh" pageId="7" pageNumber="591">Himachal Pradesh</collectingRegion>
(
<collectingCountry id="F3437641EC167A17FBEBFD517558FD2A" box="[1040,1101,691,717]" name="India" pageId="7" pageNumber="591">India</collectingCountry>
) and
<collectingCountry id="F3437641EC167A17FB70FD5175DBFD2A" box="[1163,1230,691,717]" name="Nepal" pageId="7" pageNumber="591">Nepal</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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