288 lines
36 KiB
XML
288 lines
36 KiB
XML
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<mods:title id="6A97D0544EE19181E66481DECED79D5F">On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<treatment id="03FD87C7EC127A15FF6CFAB8734BFB22" ID-DOI="http://doi.org/10.5281/zenodo.12731012" ID-GBIF-Taxon="235046006" ID-Zenodo-Dep="12731012" LSID="urn:lsid:plazi:treatment:03FD87C7EC127A15FF6CFAB8734BFB22" httpUri="http://treatment.plazi.org/id/03FD87C7EC127A15FF6CFAB8734BFB22" lastPageId="5" lastPageNumber="589" pageId="3" pageNumber="587">
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<subSubSection id="C34E655AEC127A13FF6CFAB87368FA93" box="[151,637,1370,1397]" pageId="3" pageNumber="587" type="nomenclature">
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<paragraph id="8BEB36D1EC127A13FF6CFAB87368FA93" blockId="3.[151,637,1370,1433]" box="[151,637,1370,1397]" pageId="3" pageNumber="587">
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<heading id="D0A381BDEC127A13FF6CFAB87368FA93" bold="true" box="[151,637,1370,1397]" fontSize="11" level="1" pageId="3" pageNumber="587" reason="1">
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<taxonomicName id="4C544D52EC127A13FF6CFAB87368FA93" ID-CoL="736J9" authority="Coiffait, 1977" authorityName="Coiffait" authorityYear="1977" box="[151,637,1370,1397]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="emodensa">
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<emphasis id="B920EAC3EC127A13FF6CFAB87368FA93" bold="true" box="[151,637,1370,1397]" pageId="3" pageNumber="587">
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<emphasis id="B920EAC3EC127A13FF6CFAB870D9FA92" bold="true" box="[151,460,1370,1397]" italics="true" pageId="3" pageNumber="587">Mannerheimia emodensa</emphasis>
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<bibRefCitation id="EFC54B20EC127A13FE28FAB87368FA93" author="Coiffait, H." box="[467,637,1370,1396]" pageId="3" pageNumber="587" pagination="243 - 275" refId="ref4812" refString="Coiffait, H. (1977) Staphylinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112, 243 - 275. [1976]" type="journal article" year="1977">Coiffait, 1977</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E655AEC127A13FF6CFA9D7020FA7E" box="[151,309,1407,1433]" pageId="3" pageNumber="587" type="description">
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<paragraph id="8BEB36D1EC127A13FF6CFA9D7020FA7E" blockId="3.[151,637,1370,1433]" box="[151,309,1407,1433]" pageId="3" pageNumber="587">
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(
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<figureCitation id="136F2A54EC127A13FF64FA9D71F2FA7E" box="[159,231,1407,1433]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="3" pageNumber="587">Figs 3</figureCitation>
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,
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<figureCitation id="136F2A54EC127A13FF0FFA9D7038FA7E" box="[244,301,1407,1433]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="3" pageNumber="587">9–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E655AEC127A13FF6CFA2172FAFA3C" box="[151,1007,1475,1499]" pageId="3" pageNumber="587" type="reference_group">
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<paragraph id="8BEB36D1EC127A13FF6CFA2172FAFA3C" blockId="3.[151,1007,1475,1499]" box="[151,1007,1475,1499]" pageId="3" pageNumber="587">
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<treatmentCitationGroup id="AB4411FFEC127A13FF6CFA2172FAFA3C" box="[151,1007,1475,1499]" pageId="3" pageNumber="587">
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<treatmentCitation id="0AF510C0EC127A13FF6CFA217377FA3C" author="Coiffait, H." box="[151,610,1475,1499]" page="266" pageId="3" pageNumber="587" year="1977">
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<taxonomicName id="4C544D52EC127A13FF6CFA217377FA3C" ID-CoL="736J9" authority="Coiffait, 1977: 266" authorityName="Coiffait" authorityPageNumber="266" authorityYear="1977" box="[151,610,1475,1499]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="emodensa">
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<emphasis id="B920EAC3EC127A13FF6CFA217082FA3C" box="[151,407,1475,1499]" italics="true" pageId="3" pageNumber="587">Mannerheimia emodensa</emphasis>
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<bibRefCitation id="EFC54B20EC127A13FE65FA217377FA3C" author="Coiffait, H." box="[414,610,1475,1499]" pageId="3" pageNumber="587" pagination="243 - 275" refId="ref4812" refString="Coiffait, H. (1977) Staphylinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112, 243 - 275. [1976]" type="journal article" year="1977">Coiffait, 1977: 266</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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;
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<treatmentCitation id="0AF510C0EC127A13FD97FA21723CFA3C" author="Coiffait, H." box="[620,809,1475,1499]" page="150" pageId="3" pageNumber="587" year="1982">
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<bibRefCitation id="EFC54B20EC127A13FD97FA21723CFA3C" author="Coiffait, H." box="[620,809,1475,1499]" pageId="3" pageNumber="587" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait 1982: 150</bibRefCitation>
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</treatmentCitation>
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,
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<treatmentCitation id="0AF510C0EC127A13FCCFFA2172FAFA3C" author="Shavrin, A. V." box="[820,1007,1475,1499]" page="307" pageId="3" pageNumber="587" year="2021">
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||
<bibRefCitation id="EFC54B20EC127A13FCCFFA2172FAFA3C" author="Shavrin, A. V." box="[820,1007,1475,1499]" pageId="3" pageNumber="587" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021: 307</bibRefCitation>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E655AEC127A13FF6CF9EB7215F8A4" pageId="3" pageNumber="587" type="materials_examined">
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<paragraph id="8BEB36D1EC127A13FF6CF9EB75B4F930" blockId="3.[151,1436,1545,2040]" pageId="3" pageNumber="587">
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<emphasis id="B920EAC3EC127A13FF6CF9EB70D1F9C4" bold="true" box="[151,452,1545,1571]" pageId="3" pageNumber="587">Type material examined.</emphasis>
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<typeStatus id="54EF8873EC127A13FE35F9EB7323F9C4" box="[462,566,1545,1571]" pageId="3" pageNumber="587" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D52FD58EC127A13FDC4F9E87341F9C3" box="[575,596,1546,1572]" count="1" pageId="3" pageNumber="587" type="male">♂</specimenCount>
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(
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<figureCitation id="136F2A54EC127A13FD9DF9EB73B8F9C4" box="[614,685,1545,1571]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="3" pageNumber="587">Fig. 3</figureCitation>
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; dissected prior to the present study: the aedeagus and the apical genital segment are glued on the same card with the specimen): ‘Umg.Alm Darghari | b. Maharigaon, 4000m’ <printed>, ‘Gebiet von Jumla | Westnepal,lg.H.Franz’ <printed>, ‘TYPE’ <red, printed>, ‘
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<taxonomicName id="4C544D52EC127A13FB88F9B37409F98C" authorityName="Maklin" authorityYear="1880" box="[1139,1308,1617,1643]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="genus">Mannerheimia</taxonomicName>
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| emodensa | H. COIFFAIT det. 19[printed]76’ <handwritten in black> (
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<collectionCode id="ED45AE14EC127A13FCC5F9977299F968" box="[830,908,1653,1679]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="587">NMW</collectionCode>
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).
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<typeStatus id="54EF8873EC127A13FC64F9947505F977" box="[927,1040,1654,1680]" pageId="3" pageNumber="587" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D52FD58EC127A13FBE7F9977549F977" box="[1052,1116,1653,1680]" count="3" pageId="3" pageNumber="587" type="male">3 ♂♂</specimenCount>
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(
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<specimenCount id="9D52FD58EC127A13FB90F9977412F968" box="[1131,1287,1653,1679]" count="1" pageId="3" pageNumber="587" type="generic">one specimen</specimenCount>
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dissected),
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<specimenCount id="9D52FD58EC127A13FA75F99771D6F953" count="2" pageId="3" pageNumber="587" type="female">2 ♀♀</specimenCount>
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: first two labels as that in the
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<typeStatus id="54EF8873EC127A13FDEAF97B7363F954" box="[529,630,1689,1715]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
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, with additional red printed label: ‘PARATYPE’ (
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<collectionCode id="ED45AE14EC127A13FB64F97B75F8F954" box="[1183,1261,1689,1715]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="587">NMW</collectionCode>
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). All types with additional printed label: ‘
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<taxonomicName id="4C544D52EC127A13FE48F95F7342F930" authorityName="Maklin" authorityYear="1880" box="[435,599,1725,1751]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="genus">
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<emphasis id="B920EAC3EC127A13FE48F95F7342F930" box="[435,599,1725,1751]" italics="true" pageId="3" pageNumber="587">Mannerheimia</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="4C544D52EC127A13FD90F95F726AF930" authority="Coiffait, 1977" authorityName="Coiffait" authorityYear="1977" box="[619,895,1725,1751]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="emodensa">
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<emphasis id="B920EAC3EC127A13FD90F95F73CEF930" box="[619,731,1725,1751]" italics="true" pageId="3" pageNumber="587">emodensa</emphasis>
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||
<bibRefCitation id="EFC54B20EC127A13FD19F95F726AF930" author="Coiffait, H." box="[738,895,1725,1751]" pageId="3" pageNumber="587" pagination="243 - 275" refId="ref4812" refString="Coiffait, H. (1977) Staphylinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112, 243 - 275. [1976]" type="journal article" year="1977">Coiffait, 1977</bibRefCitation>
|
||
</taxonomicName>
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| Shavrin A.
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<collectionCode id="ED45AE14EC127A13FBF3F95C750BF93F" box="[1032,1054,1726,1752]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="587" type="Museum">V</collectionCode>
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. det. 2024’.
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</paragraph>
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<paragraph id="8BEB36D1EC127A13FF3CF9037215F8A4" blockId="3.[151,1436,1545,2040]" pageId="3" pageNumber="587">
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||
<materialsCitation id="3B3C3C8CEC127A13FF3CF90375FAF91C" ID-GBIF-Occurrence="4905213303" box="[199,1263,1761,1788]" collectionCode="NMW" country="Nepal" pageId="3" pageNumber="587" specimenCount="12" specimenCount-female="3" specimenCount-male="9">
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||
<emphasis id="B920EAC3EC127A13FF3CF90373BFF91C" bold="true" box="[199,682,1761,1787]" pageId="3" pageNumber="587">
|
||
<typeStatus id="54EF8873EC127A13FF3CF9037320F91C" box="[199,565,1761,1787]" isOtherTypeStatus="true" pageId="3" pageNumber="587">Additional material examined</typeStatus>
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||
.
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<collectingCountry id="F3437641EC127A13FDBFF90373B0F91C" box="[580,677,1761,1787]" name="Nepal" pageId="3" pageNumber="587">NEPAL</collectingCountry>
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||
:
|
||
</emphasis>
|
||
<specimenCount id="9D52FD58EC127A13FD4FF90373EDF91B" box="[692,760,1761,1788]" count="9" pageId="3" pageNumber="587" type="male">9 ♂♂</specimenCount>
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||
,
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<specimenCount id="9D52FD58EC127A13FCFCF903725EF91B" box="[775,843,1761,1788]" count="3" pageId="3" pageNumber="587" type="female">3 ♀♀</specimenCount>
|
||
: same data as the holotype (
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||
<collectionCode id="ED45AE14EC127A13FB62F90375F2F91C" box="[1177,1255,1761,1787]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="587">NMW</collectionCode>
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||
)
|
||
</materialsCitation>
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||
;
|
||
<materialsCitation id="3B3C3C8CEC127A13FB06F903754AF8F8" ID-GBIF-Occurrence="4905213302" collectionCode="NMW" collectorName="H. Franz" country="Nepal" pageId="3" pageNumber="587" specimenCount="1" specimenCount-male="1">
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||
<specimenCount id="9D52FD58EC127A13FB06F9037438F91B" box="[1277,1325,1761,1788]" count="1" pageId="3" pageNumber="587" type="male">1 ♂</specimenCount>
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||
: ‘
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||
<collectorName id="26A15307EC127A13FABBF903701DF8F8" pageId="3" pageNumber="587">Dzunda Khola-Tal</collectorName>
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b.
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<collectorName id="26A15307EC127A13FED5F8E77060F8F8" box="[302,373,1797,1823]" pageId="3" pageNumber="587">Talphi</collectorName>
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3000-3500m’, ‘
|
||
<collectorName id="26A15307EC127A13FDD2F8E77269F8F8" box="[553,892,1797,1824]" pageId="3" pageNumber="587">Gebiet von Jumla Westnepal</collectorName>
|
||
,lg.
|
||
<collectorName id="26A15307EC127A13FC63F8E772EDF8F8" box="[920,1016,1797,1823]" pageId="3" pageNumber="587">H.Franz’</collectorName>
|
||
(
|
||
<collectionCode id="ED45AE14EC127A13FBF2F8E77542F8F8" box="[1033,1111,1797,1823]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="587">NMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3C3C8CEC127A13FB90F8E773E9F8A4" ID-GBIF-Occurrence="4905213307" collectionCode="NMW" collectorName="H. Franz" country="Nepal" location="Gebiet des Rarasees Westnepal" pageId="3" pageNumber="587" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9D52FD58EC127A13FB90F8E7758CF8C7" box="[1131,1177,1797,1824]" count="1" pageId="3" pageNumber="587" type="female">1 ♀</specimenCount>
|
||
: ‘
|
||
<collectorName id="26A15307EC127A13FB50F8E77436F8F8" box="[1195,1315,1797,1823]" pageId="3" pageNumber="587">Dampelek</collectorName>
|
||
b.
|
||
<collectorName id="26A15307EC127A13FAB2F8E7748DF8F8" box="[1353,1432,1797,1823]" pageId="3" pageNumber="587">Jumla’</collectorName>
|
||
, ‘
|
||
<location id="8E8B600AEC127A13FF60F8CB70EAF8A4" LSID="urn:lsid:plazi:treatment:03FD87C7EC127A15FF6CFAB8734BFB22:8E8B600AEC127A13FF60F8CB70EAF8A4" box="[155,511,1833,1860]" country="Nepal" name="Gebiet des Rarasees Westnepal" pageId="3" pageNumber="587">Gebiet des Rarasees Westnepal</location>
|
||
, lg.
|
||
<collectorName id="26A15307EC127A13FDD5F8C87382F8A4" box="[558,663,1833,1860]" pageId="3" pageNumber="587">H. Franz’</collectorName>
|
||
(
|
||
<collectionCode id="ED45AE14EC127A13FD5DF8CB73E1F8A4" box="[678,756,1833,1859]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="587">NMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E655AEC127A15FF3CF8AF7050FC42" lastPageId="5" lastPageNumber="589" pageId="3" pageNumber="587" type="description">
|
||
<paragraph id="8BEB36D1EC127A13FF3CF8AF75EFF848" blockId="3.[151,1436,1545,2040]" pageId="3" pageNumber="587">
|
||
<emphasis id="B920EAC3EC127A13FF3CF8AF7060F880" bold="true" box="[199,373,1869,1895]" pageId="3" pageNumber="587">Redescription.</emphasis>
|
||
Measurements (n=20): HW: 0.47–0.53; HL: 0.31–0.33; OL: 0.14–0.15; AL (
|
||
<typeStatus id="54EF8873EC127A13FB1DF8AF745AF880" box="[1254,1359,1869,1895]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
|
||
): 0.85; PL: 0.33–0.37; PW: 0.61–0.68; ESL: 0.74–0.78; EW: 0.90–1.05; MTbL (
|
||
<typeStatus id="54EF8873EC127A13FC20F8937551F86C" box="[987,1092,1905,1931]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
|
||
): 0.52; MTrL (
|
||
<typeStatus id="54EF8873EC127A13FB15F8937442F86C" box="[1262,1367,1905,1931]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
|
||
): 0.30 (MTrL 1–4: 0.17; MTrL 5: 0.13); AW: 0.95–1.03; AedL: 0.36–0.40; BL: 1.98–2.65 (
|
||
<typeStatus id="54EF8873EC127A13FBBDF87775BBF848" box="[1094,1198,1941,1967]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
|
||
: 2.37).
|
||
</paragraph>
|
||
<paragraph id="8BEB36D1EC127A14FF3CF85B738DFE4A" blockId="3.[151,1436,1545,2040]" lastBlockId="4.[151,1437,150,429]" lastPageId="4" lastPageNumber="588" pageId="3" pageNumber="587">
|
||
Habitus as in
|
||
<figureCitation id="136F2A54EC127A13FE90F85B70A7F834" box="[363,434,1977,2003]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="3" pageNumber="587">Fig. 3</figureCitation>
|
||
. Body yellow-brown to reddish-brown, with distinctly darker head (abdomen of some specimens slightly darker than forebody; lateral and basal portions of pronotum and apical portion of elytra slightly paler in
|
||
<typeStatus id="54EF8873EC157A14FF0AFF757046FF56" box="[241,339,151,177]" pageId="4" pageNumber="588" type="holotype">holotype</typeStatus>
|
||
and in some
|
||
<typeStatus id="54EF8873EC157A14FE1DFF757342FF56" box="[486,599,151,177]" pageId="4" pageNumber="588" type="paratype">paratypes</typeStatus>
|
||
); antennomeres 5–11 or 6–11 brown; mouthparts, antennomeres 1–4 or 1–5 and paratergites yellow. Forebody without microsculpture; abdomen with fine and dense transverse microreticulation. Head with moderately fine and sparse punctation, distinctly denser in middle; punctation of pronotum denser, larger and deeper than that in middle portion of head, finer and sparser in middle, and sometimes with narrow impunctate portion in middle or in mediobasal third; scutellum with one to four fine punctures in middle or without them; punctation of elytra denser, slightly larger and deeper than that on pronotum, punctures in middle of each elytron forming five to six vague and tangled rows of serial punctures, but some specimens with markedly coarse surface without longitudinal rows between punctures.
|
||
</paragraph>
|
||
<caption id="DF2B6659EC157A14FF6CF8407593F839" ID-DOI="http://doi.org/10.5281/zenodo.12731014" ID-Zenodo-Dep="12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="4" pageNumber="588" startId="4.[151,264,1954,1978]" targetBox="[185,1404,459,1918]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BEB36D1EC157A14FF6CF8407593F839" blockId="4.[151,1437,1954,2015]" pageId="4" pageNumber="588">
|
||
<emphasis id="B920EAC3EC157A14FF6CF840705DF85D" bold="true" box="[151,328,1954,1978]" pageId="4" pageNumber="588">FIGURES 6–10.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="4C544D52EC157A14FE2DF840737EF85D" authorityName="Maklin" authorityYear="1880" box="[470,619,1954,1978]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EAC3EC157A14FE2DF840737EF85D" box="[470,619,1954,1978]" italics="true" pageId="4" pageNumber="588">Mannerheimia</emphasis>
|
||
</taxonomicName>
|
||
(Figs 6–7:
|
||
<taxonomicName id="4C544D52EC157A14FD1AF8417294F85D" authorityName="Coiffait" authorityYear="1982" box="[737,897,1954,1978]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="588" phylum="Arthropoda" rank="species" species="cachemirica">
|
||
<emphasis id="B920EAC3EC157A14FD1AF8417294F85D" box="[737,897,1954,1978]" italics="true" pageId="4" pageNumber="588">M. cachemirica</emphasis>
|
||
</taxonomicName>
|
||
(holotype); Fig. 8:
|
||
<taxonomicName id="4C544D52EC157A14FBB1F84175BEF85D" authorityName="Coiffait" authorityYear="1982" box="[1098,1195,1955,1978]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="588" phylum="Arthropoda" rank="species" species="vicina">
|
||
<emphasis id="B920EAC3EC157A14FBB1F84175BEF85D" box="[1098,1195,1955,1978]" italics="true" pageId="4" pageNumber="588">M. vicina</emphasis>
|
||
</taxonomicName>
|
||
(holotype); Figs. 9–10:
|
||
<taxonomicName id="4C544D52EC157A14FF6CF825700BF839" authorityName="Coiffait" authorityYear="1977" box="[151,286,1990,2014]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="588" phylum="Arthropoda" rank="species" species="emodensa">
|
||
<emphasis id="B920EAC3EC157A14FF6CF825700BF839" box="[151,286,1990,2014]" italics="true" pageId="4" pageNumber="588">M. emodensa</emphasis>
|
||
</taxonomicName>
|
||
(Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFF757312FEA6" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">Head 1.5–1.6 times as broad as long, slightly elevated in middle, with very short and sometimes indistinct anteocellar foveae as long as diameter of ocellus. Antennomere 3 narrower and slightly shorter than 2, 4 distinctly shorter than 3 and slightly longer than broad, 5–6 indistinctly broader than 4, 7–8 slightly longer and distinctly broader than 6, 9–10 slightly longer and broader than 8, apical antennomere about as long as two preceding antennomeres or slightly shorter.</paragraph>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFEA97271FE4A" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">Pronotum 1.8 times as broad as long, 1.2 times as broad as head, widest slightly above middle, evenly narrowed toward obtuse posterior angles, sometimes with indistinctly sinuate laterobasal margins; lateral portions slightly impressed, distinctly broader and deeper in laterobasal portions.</paragraph>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFE55750DFE36" blockId="5.[151,1437,150,1221]" box="[199,1048,439,465]" pageId="5" pageNumber="589">Elytra 1.2–1.3 times broader than long, slightly or strongly widened apicad.</paragraph>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFE397389FE12" blockId="5.[151,1437,150,1221]" box="[199,668,475,501]" pageId="5" pageNumber="589">Metatarsus 1.7 times as long as metatibia.</paragraph>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFE1D708DFDDA" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">Abdomen about as broad as elytra, with two small oval or transverse tomentose wing-folding patches in middle of abdominal tergite V.</paragraph>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFDA5753DFD2A" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">
|
||
Male. Apical margin of abdominal tergite VIII straight or slightly sinuate. Apical margin of abdominal sternite VIII widely sinuate. Aedeagus with wide basal portion, gradually narrowed apically, with relatively narrow median lobe and small rounded apex; parameres with wide apical portions, slightly exceeding apex of median lobe; internal sac narrow and moderately long (
|
||
<figureCitation id="136F2A54EC147A15FDF5FD517347FD2A" box="[526,594,691,717]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="5" pageNumber="589">Fig. 9</figureCitation>
|
||
). Lateral aspect of aedeagus as in
|
||
<figureCitation id="136F2A54EC147A15FC29FD517531FD2A" box="[978,1060,691,717]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="5" pageNumber="589">Fig. 10</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFD35709EFCF2" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">Female. Apical margin of abdominal tergite VIII straight or somewhat rounded. Apical margin of abdominal sternite VIII rounded.</paragraph>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFCFC7050FC42" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">
|
||
<emphasis id="B920EAC3EC147A15FF3CFCFC70A1FCDE" bold="true" box="[199,436,798,825]" pageId="5" pageNumber="589">Comparative notes.</emphasis>
|
||
Based on the general shape of the body and the aedeagus,
|
||
<taxonomicName id="4C544D52EC147A15FB95FCFD7412FCDE" authorityName="Coiffait" authorityYear="1977" box="[1134,1287,799,825]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="species" species="emodensa">
|
||
<emphasis id="B920EAC3EC147A15FB95FCFD7412FCDE" box="[1134,1287,799,825]" italics="true" pageId="5" pageNumber="589">M. emodensa</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C544D52EC147A15FF6CFCA1705CFCBA" authorityName="Coiffait" authorityYear="1982" box="[151,329,835,861]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="species" species="cachemirica">
|
||
<emphasis id="B920EAC3EC147A15FF6CFCA1705CFCBA" box="[151,329,835,861]" italics="true" pageId="5" pageNumber="589">M. cachemirica</emphasis>
|
||
</taxonomicName>
|
||
, from which can be distinguished by the slightly broader elytra, larger and denser punctation on the pronotum and elytra, slightly shorter preapical antennomeres, and longer parameres with significantly broader apical portions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E655AEC147A15FF3CFC4C7012FBF6" pageId="5" pageNumber="589" type="distribution">
|
||
<paragraph id="8BEB36D1EC147A15FF3CFC4C758EFC2E" blockId="5.[151,1437,150,1221]" box="[199,1179,942,969]" pageId="5" pageNumber="589">
|
||
<emphasis id="B920EAC3EC147A15FF3CFC4C7074FC2F" bold="true" box="[199,353,942,968]" pageId="5" pageNumber="589">Distribution.</emphasis>
|
||
<taxonomicName id="4C544D52EC147A15FE9CFC4D7396FC2E" authorityName="Coiffait" authorityYear="1977" box="[359,643,943,969]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="species" species="emodensa">
|
||
<emphasis id="B920EAC3EC147A15FE9CFC4D7396FC2E" box="[359,643,943,969]" italics="true" pageId="5" pageNumber="589">Mannerheimia emodensa</emphasis>
|
||
</taxonomicName>
|
||
is known from four localities in western
|
||
<collectingCountry id="F3437641EC147A15FBAAFC4D758DFC2E" box="[1105,1176,943,969]" name="Nepal" pageId="5" pageNumber="589">Nepal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB36D1EC147A15FF3CFC307012FBF6" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">
|
||
<emphasis id="B920EAC3EC147A15FF3CFC30705EFC0B" bold="true" box="[199,331,978,1004]" pageId="5" pageNumber="589">Bionomics.</emphasis>
|
||
Specimens were collected at altitudes from
|
||
<quantity id="4CAC9B34EC147A15FCB4FC31751CFC0A" box="[847,1033,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="589" unit="m" value="3500.0" valueMax="4000.0" valueMin="3000.0">3000 to 4000 m</quantity>
|
||
a.s.l. Detailed bionomical data are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E655AEC147A15FF3CFBF9734BFB22" pageId="5" pageNumber="589" type="discussion">
|
||
<paragraph id="8BEB36D1EC147A15FF3CFBF9734BFB22" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">
|
||
<emphasis id="B920EAC3EC147A15FF3CFBF9702EFBD2" bold="true" box="[199,315,1051,1077]" pageId="5" pageNumber="589">Remarks.</emphasis>
|
||
<taxonomicName id="4C544D52EC147A15FEB8FBF9734AFBD2" authorityName="Coiffait" authorityYear="1977" box="[323,607,1051,1077]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="species" species="emodensa">
|
||
<emphasis id="B920EAC3EC147A15FEB8FBF9734AFBD2" box="[323,607,1051,1077]" italics="true" pageId="5" pageNumber="589">Mannerheimia emodensa</emphasis>
|
||
</taxonomicName>
|
||
based on several specimens was originally described from four localities in “..région de Jumla”, western
|
||
<collectingCountry id="F3437641EC147A15FE0CFBDD732BFBBE" box="[503,574,1087,1113]" name="Nepal" pageId="5" pageNumber="589">Nepal</collectingCountry>
|
||
. I studied the
|
||
<typeStatus id="54EF8873EC147A15FD23FBDD722FFBBE" box="[728,826,1087,1113]" pageId="5" pageNumber="589" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D52FD58EC147A15FC94FBDD7519FBBE" box="[879,1036,1087,1113]" count="5" pageId="5" pageNumber="589" type="generic" typeStatus="paratype">five paratypes</specimenCount>
|
||
from NMW, and several specimens without
|
||
<typeStatus id="54EF8873EC147A15FF0EFB817040FB9A" box="[245,341,1123,1149]" pageId="5" pageNumber="589" type="paratype">paratype</typeStatus>
|
||
label, some of which are apparently belonging to the type serie (see Additional material section above).
|
||
<bibRefCitation id="EFC54B20EC147A15FF09FB65708FFB46" author="Coiffait, H." box="[242,410,1159,1185]" pageId="5" pageNumber="589" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait (1982)</bibRefCitation>
|
||
included it to the key of
|
||
<taxonomicName id="4C544D52EC147A15FD4DFB65724FFB46" authorityName="Maklin" authorityYear="1880" box="[694,858,1159,1185]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EAC3EC147A15FD4DFB65724FFB46" box="[694,858,1159,1185]" italics="true" pageId="5" pageNumber="589">Mannerheimia</emphasis>
|
||
</taxonomicName>
|
||
of the Himalayan Region.
|
||
<bibRefCitation id="EFC54B20EC147A15FB70FB657421FB46" author="Shavrin, A. V." box="[1163,1332,1159,1185]" pageId="5" pageNumber="589" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin (2021)</bibRefCitation>
|
||
included this species to the key of
|
||
<taxonomicName id="4C544D52EC147A15FE49FB497343FB22" authorityName="Maklin" authorityYear="1880" box="[434,598,1195,1221]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EAC3EC147A15FE49FB497343FB22" box="[434,598,1195,1221]" italics="true" pageId="5" pageNumber="589">Mannerheimia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |