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<document id="527D815BEDF1525567D6EB245E8DAA67" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F10EFFFB75CDFAFF511B7A50" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras STEPHENS 1833" docType="treatment" docVersion="5" lastPageNumber="220" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="215" updateTime="1699471357828" updateUser="ExternalLinkService">
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<mods:title id="0CBCF8A11231AE288F8FD98B4CF3B048">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<subSubSection id="C34E653CF10EFFFE75CDFAFF570A7A4D" pageId="2" pageNumber="215" type="nomenclature">
<paragraph id="8BEB36B7F10EFFFE75CDFAFF570A7A4D" blockId="2.[189,1436,1327,1569]" pageId="2" pageNumber="215">
3.2 Key to the species of the subgenus
<taxonomicName id="4C544D34F10EFFFE77EBFAFF577A7A77" ID-CoL="8VS6V" authorityName="STEPHENS" authorityYear="1833" box="[718,782,1327,1359]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAA5F10EFFFE77EBFAFF577A7A77" box="[718,782,1327,1359]" italics="true" pageId="2" pageNumber="215">Zyras</emphasis>
</taxonomicName>
of the Oriental Region (sensu
<bibRefCitation id="EFC54B46F10EFFFE71A2FAFF55237A4D" author="SCHULKE, M. &amp; SMETANA, A." pageId="2" pageNumber="215" refId="ref27584" refString="SCHULKE, M. &amp; SMETANA, A. 2015: Staphylinidae: 304 - 1134. - In: LOBL, I. &amp; LOBL, D. (eds): Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea - Staphylinoidea., Revised and updated Edition - Brill, Leiden - ISBN 9789004289925." type="journal volume" year="2015">SCHÜLKE &amp; SMETANA 2015</bibRefCitation>
), exclusive of
<collectingCountry id="F3437627F10EFFFE7724FA8556377A4D" box="[513,579,1365,1397]" name="India" pageId="2" pageNumber="215">India</collectingCountry>
,
<collectingCountry id="F3437627F10EFFFE776BFA8556B17A4D" box="[590,709,1365,1397]" name="Sri Lanka" pageId="2" pageNumber="215">Sri Lanka</collectingCountry>
, and
<collectingCountry id="F3437627F10EFFFE7622FA85570A7A4D" box="[775,894,1365,1397]" name="Myanmar" pageId="2" pageNumber="215">Myanmar</collectingCountry>
</paragraph>
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<subSubSection id="C34E653CF10EFFFE7598FA5A56ED7AF9" pageId="2" pageNumber="215" type="key">
<paragraph id="8BEB36B7F10EFFFE7598FA5A56ED7AF9" blockId="2.[189,1436,1327,1569]" pageId="2" pageNumber="215">
In all, 45 species are currently known from the southeastern Oriental regions (comprising
<collectingCountry id="F3437627F10EFFFE71A8FA59509B7A99" box="[1165,1263,1417,1441]" name="Thailand" pageId="2" pageNumber="215">Thailand</collectingCountry>
,
<collectingCountry id="F3437627F10EFFFE71DEFA5A515B7A99" box="[1275,1327,1418,1441]" name="Laos" pageId="2" pageNumber="215">Laos</collectingCountry>
,
<collectingCountry id="F3437627F10EFFFE701FFA5951EC7A99" box="[1338,1432,1417,1441]" name="Vietnam" pageId="2" pageNumber="215">Vietnam</collectingCountry>
, Peninsular
<collectingCountry id="F3437627F10EFFFE7410FA7955E17AF9" box="[309,405,1449,1473]" name="Malaysia" pageId="2" pageNumber="215">Malaysia</collectingCountry>
, and the Sunda Islands).
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF10EFFFE7598FA1955A47919" pageId="2" pageNumber="215" type="discussion">
<paragraph id="8BEB36B7F10EFFFE7598FA1955A47919" blockId="2.[189,1436,1327,1569]" pageId="2" pageNumber="215">
A positive identification of females without reference material may not always be possible. The same applies to aberrant (e.g., regarding coloration) or nanistic specimens. Two variable species,
<taxonomicName id="4C544D34F10EFFFE76D1FA3A50137939" authorityName="CAMERON" authorityYear="1939" box="[1012,1127,1513,1537]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="gratellus">
<emphasis id="B920EAA5F10EFFFE76D1FA3A50137939" box="[1012,1127,1513,1537]" italics="true" pageId="2" pageNumber="215">Z. gratellus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D34F10EFFFE71BBFA3A514B7939" authorityName="Cameron" authorityYear="1939" box="[1182,1343,1514,1537]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B920EAA5F10EFFFE71BBFA3A514B7939" box="[1182,1343,1514,1537]" italics="true" pageId="2" pageNumber="215">Z. punctipennis</emphasis>
</taxonomicName>
, key out in more than one couplet.
</paragraph>
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<subSubSection id="C34E653CF10EFFFB75E0F992511B7A50" lastPageId="7" lastPageNumber="220" pageId="2" pageNumber="215" type="description">
<key id="FFC56186F10EFFFB75E0F992511B7A50" lastPageId="7" lastPageNumber="220" pageId="2" pageNumber="215">
<keyStep id="30A02E12F10EFFFE75E0F99251E17991" pageId="2" pageNumber="215">
<paragraph id="8BEB36B7F10EFFFE75E0F99251E17962" blockId="2.[197,1429,1602,1705]" box="[197,1429,1602,1626]" pageId="2" pageNumber="215">
<keyLead id="30A59582F10EFFFE75E0F99251E17962" box="[197,1429,1602,1626]" pageId="2" pageNumber="215">
1. Punctation of pronotum
<emphasis id="B920EAA5F10EFFFE74D1F99256687962" box="[500,540,1602,1626]" italics="true" pageId="2" pageNumber="215">and</emphasis>
elytra conspicuously coarse and dense (e.g.,
<figureCitation id="136F2A32F10EFFFE76D0F99250347962" box="[1013,1088,1602,1626]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="2" pageNumber="215">Fig. 30</figureCitation>
). ................................................. 2
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10EFFFE75E0F9A151E17991" blockId="2.[197,1429,1602,1705]" pageId="2" pageNumber="215">
<keyLead id="30A59582F10EFFFE75E0F9A151E17991" pageId="2" pageNumber="215">
– Punctation of pronotum
<emphasis id="B920EAA5F10EFFFE74D0F9A1566979B1" box="[501,541,1649,1673]" italics="true" pageId="2" pageNumber="215">and</emphasis>
elytra not conspicuously coarse and dense (though sometimes very fine and dense, or only the elytra with coarse and dense punctation). ................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10EFFFE75E0F91051E078FE" pageId="2" pageNumber="215">
<paragraph id="8BEB36B7F10EFFFE75E0F91051E17800" blockId="2.[196,1429,1728,1991]" pageId="2" pageNumber="215">
<keyLead id="30A59582F10EFFFE75E0F91051E17800" pageId="2" pageNumber="215">
2. Punctation of pronotum and elytra strongly and conspicuously coarsely granulose (
<figureCitation id="136F2A32F10EFFFE7162F91050E779E0" box="[1095,1171,1728,1752]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="2" pageNumber="215">Fig. 30</figureCitation>
). Body smaller and more slender, of predominantly brown coloration. Antennae reddish (
<figureCitation id="136F2A32F10EFFFE76A3F93057B179C0" box="[902,965,1760,1784]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="2" pageNumber="215">Fig. 7</figureCitation>
). Pronotum nearly as long as broad, strongly convex in cross-section, and with sinuate lateral margins in dorsal view (
<figureCitation id="136F2A32F10EFFFE76D6F8D050347820" box="[1011,1088,1792,1816]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="2" pageNumber="215">Fig. 30</figureCitation>
). Median lobe of aedeagus as in
<figureCitation id="136F2A32F10EFFFE75CFF8F0552B7800" box="[234,351,1824,1848]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="2" pageNumber="215">Figs 66–67</figureCitation>
. Borneo. .......................................................................................................................................
<taxonomicName id="4C544D34F10EFFFE7024F8F051E17800" authorityName="Cameron" authorityYear="1930" box="[1281,1429,1824,1848]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="granulipennis">
<emphasis id="B920EAA5F10EFFFE7024F8F051E17800" bold="true" box="[1281,1429,1824,1848]" italics="true" pageId="2" pageNumber="215">granulipennis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10EFFFE75E1F88051E078FE" blockId="2.[196,1429,1728,1991]" pageId="2" pageNumber="215">
<keyLead id="30A59582F10EFFFE75E1F88051E078FE" pageId="2" pageNumber="215">
– Punctation of pronotum and elytra not distinctly granulose. Body larger and more robust, of predominantly black coloration. Antennae bicoloured, dark-brown to blackish-brown with the apical 1–2 antennomeres yellow. Pronotum distinctly transverse, moderately convex in cross-section; lateral margins not sinuate in dorsal view. Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10EFFFE74F2F87F561F78FF" author="ASSING, V." box="[471,619,1967,1991]" pageId="2" pageNumber="215" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 266–267, 271–272). Borneo, Peninsular
<collectingCountry id="F3437627F10EFFFE7149F87F50B978FF" box="[1132,1229,1967,1991]" name="Malaysia" pageId="2" pageNumber="215">Malaysia</collectingCountry>
. ............
<taxonomicName id="4C544D34F10EFFFE700CF87F51E078FE" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[1321,1428,1967,1990]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B920EAA5F10EFFFE700CF87F51E078FE" bold="true" box="[1321,1428,1967,1990]" italics="true" pageId="2" pageNumber="215">montanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BEB36B7F10EFFFE75E1F80E51E1772E" blockId="2.[196,1429,2014,2070]" pageId="2" pageNumber="215">
<keyLead id="30A59582F10EFFFE75E1F80E51E1772E" pageId="2" pageNumber="215">
3. Forebody with conspicuously dense and fine punctation. Small species; length of forebody
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. Antennae slender, preapical antennomeres not transverse. ............................................................................................................ 4
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BAFF29511B7E08" blockId="3.[159,1391,248,304]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFF29511B7E08" pageId="3" pageNumber="216">– Punctation of forebody not conspicuously dense and fine. Mostly larger species. Preapical antennomeres mostly transverse. ............................................................................................................................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyLead id="30A59582F10FFFFF75BAFE98511B7DA6" pageId="3" pageNumber="216">
<keyStep id="30A02E12F10FFFFF75BAFE98511A7D36" pageId="3" pageNumber="216">
<paragraph id="8BEB36B7F10FFFFF75BAFE98511B7EA7" blockId="3.[159,1391,327,526]" pageId="3" pageNumber="216">
4. Punctation of forebody finer and denser (
<bibRefCitation id="EFC54B46F10FFFFF775EFE9857717E58" author="ASSING, V." box="[635,773,328,352]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 75). Pronotum dark-brown, of similar coloration as head (
<bibRefCitation id="EFC54B46F10FFFFF7422FEB855E67EB8" author="ASSING, V." box="[263,402,360,384]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 75). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10FFFFF7671FEB8579C7EB8" author="ASSING, V." box="[852,1000,360,384]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 239–241).
<collectingRegion id="4990F855F10FFFFF718AFEB751797E47" box="[1199,1293,359,383]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="216">Sulawesi</collectingRegion>
. ............. .............................................................................................................................................................................
<taxonomicName id="4C544D34F10FFFFF71D5FE57511B7EA7" authorityName="ASSING" authorityYear="2017" box="[1264,1391,391,415]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="densissimus">
<emphasis id="B920EAA5F10FFFFF71D5FE57511B7EA7" bold="true" box="[1264,1391,391,415]" italics="true" pageId="3" pageNumber="216">densissimus</emphasis>
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</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BAFE67511A7D36" blockId="3.[159,1391,327,526]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFE67511A7D36" pageId="3" pageNumber="216">
– Punctation of forebody less fine and less dense (
<figureCitation id="136F2A32F10FFFFF77E2FE6757607EF7" box="[711,788,439,463]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 23</figureCitation>
). Pronotum pale-reddish, distinctly contrasting with the dark-brown to blackish head. Median lobe of aedeagus as in
<figureCitation id="136F2A32F10FFFFF7663FE0757C87ED6" box="[838,956,470,495]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="3" pageNumber="216">Figs 59–60</figureCitation>
. Widespread in the Oriental region. ....... ......................................................................................................................................................................................
<taxonomicName id="4C544D34F10FFFFF703AFE26511A7D36" authorityName="Cameron" authorityYear="1943" box="[1311,1390,502,526]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B920EAA5F10FFFFF703AFE26511A7D36" bold="true" box="[1311,1390,502,526]" italics="true" pageId="3" pageNumber="216">bryanti</emphasis>
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<keyStep id="30A02E12F10FFFFF75BAFDF6511B7DA6" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFDF6511B7DA6" pageId="3" pageNumber="216">
5. Antennae short (&lt;
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long) and clavate; antennomeres IV–X transverse, X at least approximately twice as broad as long. Body rather small (length of forebody
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) and of predominantly reddish coloration, with the head reddish to reddish-brown and the pronotum pale reddish. Forebody moderately sparsely or very sparsely punctate. ............................................................................................................................................................................... 6
</keyLead>
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</paragraph>
</keyLead>
<paragraph id="8BEB36B7F10FFFFF75BAFD65511B7DD4" blockId="3.[159,1391,550,749]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFD65511B7DD4" pageId="3" pageNumber="216">– Character combination different. Antennae usually longer and more slender, with antennomere IV weakly transverse at most. Species of similarly small size have at least the head darker. ................................................................ 7</keyLead>
</paragraph>
<keyStep id="30A02E12F10FFFFF75BAFCD4511B7CF3" pageId="3" pageNumber="216">
<paragraph id="8BEB36B7F10FFFFF75BAFCD4511B7C64" blockId="3.[159,1391,772,971]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFCD4511B7C64" pageId="3" pageNumber="216">
6. Antennomere IV disc-shaped, approximately twice as broad as long (
<figureCitation id="136F2A32F10FFFFF76BBFCD4579D7C24" box="[926,1001,772,796]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 17</figureCitation>
). Elytra with very sparse punctation; interstices on average more than twice as broad as diameter of punctures (
<figureCitation id="136F2A32F10FFFFF76F1FCF450557C04" box="[980,1057,804,828]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 22</figureCitation>
). Pronotum small in relation to head, approximately 1.2 times as broad as head (
<figureCitation id="136F2A32F10FFFFF77E7FC94577A7C64" box="[706,782,836,860]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 22</figureCitation>
). Male sexual characters unknown. Borneo. ......
<taxonomicName id="4C544D34F10FFFFF7029FC94511B7C64" authorityName="Pace" authorityYear="2008" box="[1292,1391,836,860]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="pallipyga">
<emphasis id="B920EAA5F10FFFFF7029FC94511B7C64" bold="true" box="[1292,1391,836,860]" italics="true" pageId="3" pageNumber="216">pallipyga</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BAFCA4511B7CF3" blockId="3.[159,1391,772,971]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFCA4511B7CF3" pageId="3" pageNumber="216">
– Antennomere IV noticeably transverse, but not disc-shaped (
<figureCitation id="136F2A32F10FFFFF7676FCA357D47CB3" box="[851,928,883,907]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 16</figureCitation>
). Elytra with moderately dense punctation; interstices on average twice as broad as diameter of punctures at most (
<figureCitation id="136F2A32F10FFFFF7681FC43579B7C93" box="[932,1007,915,939]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 20</figureCitation>
). Pronotum large in relation to head,&gt; 1.35 times as broad as head (
<figureCitation id="136F2A32F10FFFFF772DFC6356217CF3" box="[520,597,947,971]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 20</figureCitation>
). Median lobe of aedeagus as in
<figureCitation id="136F2A32F10FFFFF768EFC6350557CF3" box="[939,1057,947,971]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="3" pageNumber="216">Figs 53–54</figureCitation>
.
<collectingRegion id="4990F855F10FFFFF710FFC64502C7CF3" box="[1066,1112,948,971]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="216">Java</collectingRegion>
. ......................
<taxonomicName id="4C544D34F10FFFFF71D3FC63511B7CF3" authorityName="Cameron" authorityYear="1939" box="[1270,1391,947,971]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B920EAA5F10FFFFF71D3FC63511B7CF3" bold="true" box="[1270,1391,947,971]" italics="true" pageId="3" pageNumber="216">elegantulus</emphasis>
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<keyStep id="30A02E12F10FFFFF75BAFC33511B7B71" pageId="3" pageNumber="216">
<paragraph id="8BEB36B7F10FFFFF75BAFC33511B7B22" blockId="3.[159,1391,994,1097]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFC33511B7B22" pageId="3" pageNumber="216">7. Pronotum reddish or pale-reddish, strongly constrasting with the blackish head and elytra. Abdomen distinctly bicoloured, with segments III–V or III–VI reddish and the posterior segments at least partly blackish. .............. 8</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BAFBE2511B7B71" blockId="3.[159,1391,994,1097]" box="[159,1391,1073,1097]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFBE2511B7B71" box="[159,1391,1073,1097]" pageId="3" pageNumber="216">– Coloration of forebody different. .................................................................................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyLead id="30A59582F10FFFFF75BAFBB1511B7A70" pageId="3" pageNumber="216">
<keyStep id="30A02E12F10FFFFF75BAFBB1511B7A70" pageId="3" pageNumber="216">
<paragraph id="8BEB36B7F10FFFFF75BAFBB1511A7BE0" blockId="3.[159,1391,1120,1352]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFBB1511A7BE0" pageId="3" pageNumber="216">
8. Antennomere XI dark-brown to black (
<bibRefCitation id="EFC54B46F10FFFFF7745FBB1569E7B41" author="ASSING, V." box="[608,746,1121,1145]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 20). Legs of uniformly yellowish coloration. Pronotum strongly transverse, 1.15–1.18 times as broad as long. Elytra with coarser and sparser punctation (
<bibRefCitation id="EFC54B46F10FFFFF71FAFB5151187BA1" author="ASSING, V." box="[1247,1388,1153,1177]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 58). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10FFFFF77A7FB7157FD7B80" author="ASSING, V." box="[642,905,1184,1209]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017: 166–167)</bibRefCitation>
. Widespread (
<bibRefCitation id="EFC54B46F10FFFFF7106FB7150C47B81" author="ASSING, V." box="[1059,1200,1185,1209]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: map 8). .............. .............................................................................................................................................................................
<taxonomicName id="4C544D34F10FFFFF702DFB10511A7BE0" baseAuthorityName="CAMERON" baseAuthorityYear="1925" box="[1288,1390,1216,1240]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B920EAA5F10FFFFF702DFB10511A7BE0" bold="true" box="[1288,1390,1216,1240]" italics="true" pageId="3" pageNumber="216">alternans</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BAFB20511B7A70" blockId="3.[159,1391,1120,1352]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BAFB20511B7A70" pageId="3" pageNumber="216">– Apical 1–3 antennomeres yellow, much paler than antennomeres IV–VII. Legs with at least the apices of the mesoand metafemora slightly to distinctly infuscate. Elytra with denser and finer punctation. Median lobe of aedeagus of different shape. ................................................................................................................................................................ 9</keyLead>
</paragraph>
</keyStep>
</keyLead>
<keyStep id="30A02E12F10FFFFF75BBFA8F511A791E" pageId="3" pageNumber="216">
<paragraph id="8BEB36B7F10FFFFF75BBFA8F511B7A8E" blockId="3.[158,1391,1375,1574]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BBFA8F511B7A8E" pageId="3" pageNumber="216">
9. Profemora and the apical halves of the meso- and metafemora blackish. Antennomere X blackish. Pronotum dark-reddish. Elytra with distinct punctation. Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10FFFFF713FFAAF50D17AAF" author="PACE, R." box="[1050,1189,1407,1431]" pageId="3" pageNumber="216" pagination="427 - 468" refId="ref26635" refString="PACE, R. 1986 a: Aleocharinae della Thailandia e della Birmania riportate da G. de Rougemont (Coleoptera, Staphylinidae). - Bollettino del Museo Civico di Storia Naturale di Verona 11 [1984]: 427 - 468." type="journal article" year="1986">PACE (1986a</bibRefCitation>
: figures 120–121).
<collectingCountry id="F3437627F10FFFFF75E1FA4F55537A8F" box="[196,295,1439,1463]" name="Thailand" pageId="3" pageNumber="216">Thailand</collectingCountry>
. ................................................................................................................................................................
<taxonomicName id="4C544D34F10FFFFF702CFA4E511B7A8E" authorityName="PACE" authorityYear="1986" box="[1289,1391,1438,1462]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="thaiorum">
<emphasis id="B920EAA5F10FFFFF702CFA4E511B7A8E" bold="true" box="[1289,1391,1438,1462]" italics="true" pageId="3" pageNumber="216">thaiorum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BBFA1E511A791E" blockId="3.[158,1391,1375,1574]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BBFA1E511A791E" pageId="3" pageNumber="216">
– Apices of femora only slightly and narrowly infuscate; profemora often uniformly yellowish. Antennomere X and often also antennomere IX yellowish. Pronotum pale-reddish. Elytra with very fine punctation. Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10FFFFF7447F9DE5676791E" author="ASSING, V." box="[354,514,1550,1574]" pageId="3" pageNumber="216" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 274–281). Widespread (
<bibRefCitation id="EFC54B46F10FFFFF767CF9DE5797791E" author="ASSING, V." box="[857,995,1550,1574]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: map 8). .................................
<taxonomicName id="4C544D34F10FFFFF7030F9DE511A791E" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[1301,1390,1550,1574]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B920EAA5F10FFFFF7030F9DE511A791E" bold="true" box="[1301,1390,1550,1574]" italics="true" pageId="3" pageNumber="216">geminus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10FFFFF75BBF9ED511B79BC" pageId="3" pageNumber="216">
<paragraph id="8BEB36B7F10FFFFF75BBF9ED511B796D" blockId="3.[158,1391,1597,1668]" box="[158,1391,1597,1621]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BBF9ED511B796D" box="[158,1391,1597,1621]" pageId="3" pageNumber="216">10. Antennomere XI pale-yellow to dark-yellow. ................................................................................................................ 11</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BBF9BD511B79BC" blockId="3.[158,1391,1597,1668]" box="[158,1391,1644,1668]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BBF9BD511B79BC" box="[158,1391,1644,1668]" pageId="3" pageNumber="216">– Antennomere XI reddish to black. (In cases of doubt regarding this character follow both alternatives.). ......... 26</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10FFFFF75BBF94B511B783A" pageId="3" pageNumber="216">
<paragraph id="8BEB36B7F10FFFFF75BBF94B511B79EB" blockId="3.[158,1391,1691,1794]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BBF94B511B79EB" pageId="3" pageNumber="216">
11. Small (length of forebody approximately
<quantity id="4CAC9B52F10FFFFF775BF94B56A4798B" box="[638,720,1691,1715]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="216" unit="mm" value="2.4">2.4 mm</quantity>
) and micropterous species with short elytra and reduced hind wings; elytra 0.65–0.75 times as long as pronotum. ..................................................................................................... 12
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BBF93B511B783A" blockId="3.[158,1391,1691,1794]" box="[158,1391,1770,1794]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BBF93B511B783A" box="[158,1391,1770,1794]" pageId="3" pageNumber="216">– Macropterous species with longer elytra and long hind wings. Body usually larger. .............................................. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10FFFFF75BBF8CA511B7718" pageId="3" pageNumber="216">
<paragraph id="8BEB36B7F10FFFFF75BBF8CA511A78A9" blockId="3.[158,1391,1817,2081]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BBF8CA511A78A9" pageId="3" pageNumber="216">
12. Antennae uniformly dark-yellowish (
<figureCitation id="136F2A32F10FFFFF7772F8C956E2780A" box="[599,662,1817,1842]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 4</figureCitation>
). Pronotum and elytra with coarse and dense punctation (
<figureCitation id="136F2A32F10FFFFF7033F8C95117780A" box="[1302,1379,1817,1842]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 21</figureCitation>
). Antennomere XI shorter than the combined length of antennomeres IX and X; antennomere X weakly transverse (
<figureCitation id="136F2A32F10FFFFF75E9F8895578784A" box="[204,268,1881,1906]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="3" pageNumber="216">Fig. 4</figureCitation>
). Anterior abdominal tergites dark-brown (
<figureCitation id="136F2A32F10FFFFF77CFF889574D784A" box="[746,825,1881,1906]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="3" pageNumber="216">Fig. 44</figureCitation>
). Mesofemora uniformly yellowish. Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10FFFFF7447F8AA55AA78AA" author="PACE, R." box="[354,478,1914,1938]" pageId="3" pageNumber="216" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE (2008</bibRefCitation>
: figures 98–99). Borneo. ................................................................................
<taxonomicName id="4C544D34F10FFFFF71FEF8A9511A78A9" authorityName="Pace" authorityYear="2008" box="[1243,1390,1913,1937]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B920EAA5F10FFFFF71FEF8A9511A78A9" bold="true" box="[1243,1390,1913,1937]" italics="true" pageId="3" pageNumber="216">kinabaluensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10FFFFF75BBF879511B7718" blockId="3.[158,1391,1817,2081]" pageId="3" pageNumber="216">
<keyLead id="30A59582F10FFFFF75BBF879511B7718" pageId="3" pageNumber="216">
– Antennomeres IV–X dark-brown (
<bibRefCitation id="EFC54B46F10FFFFF7714F87956CE78F9" author="ASSING, V." box="[561,698,1961,1985]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 34). Pronotum and elytra with sparser and finer punctation (
<bibRefCitation id="EFC54B46F10FFFFF75E8F819552378D9" author="ASSING, V." box="[205,343,1993,2017]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 70). Antennomere XI approximately as long as the combined length of antennomeres IX and X; antennomere X distinctly transverse (
<bibRefCitation id="EFC54B46F10FFFFF7742F839569B7739" author="ASSING, V." box="[615,751,2025,2049]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 34). Meso- and metafemora apically infuscate. Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10FFFFF7494F7D956317719" author="ASSING, V." box="[433,581,2057,2081]" pageId="3" pageNumber="216" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 212–213).
<collectingCountry id="F3437627F10FFFFF7628F7D957047719" box="[781,880,2057,2081]" name="Thailand" pageId="3" pageNumber="216">Thailand</collectingCountry>
. ............................................................... ambulans
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E0FF2951E17EB8" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E0FF2951E17E09" blockId="4.[197,1429,248,384]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E0FF2951E17E09" pageId="4" pageNumber="217">13. Legs bicoloured, yellowish with at least the metafemora distinctly infuscate apically. Pronotum brown to black. .............................................................................................................................................................................................. 14</keyLead>
</paragraph>
<paragraph id="8BEB36B7F108FFF875E0FE9851E17EB8" blockId="4.[197,1429,248,384]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E0FE9851E17EB8" pageId="4" pageNumber="217">– Legs uniformly yellowish, apices of meso- and metafemora in one variable large species with reddish pronotum sometimes indistinctly darker. ........................................................................................................................................ 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E1FE4751E07D26" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E1FE4751E17EF6" blockId="4.[196,1429,407,542]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FE4751E17EF6" pageId="4" pageNumber="217">14. Meso- and metafemora apically narrowly infuscate. Elytra with very dense punctation. Antennae not particularly long and slender; antennomere IV indistinctly oblong at most. ................................................................................ 15</keyLead>
</paragraph>
<paragraph id="8BEB36B7F108FFF875E1FE3651E07D26" blockId="4.[196,1429,407,542]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FE3651E07D26" pageId="4" pageNumber="217">
– Meso- and metafemora apically extensively infuscate. Elytra with moderately dense punctation. Antennae long and slender; antennomere IV at least approximately twice as long as broad (species of the
<taxonomicName id="4C544D34F108FFF87142FDD750B57D25" authorityName="CAMERON" authorityYear="1944" box="[1127,1217,518,542]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B920EAA5F108FFF87142FDD750B57D25" box="[1127,1217,518,542]" italics="true" pageId="4" pageNumber="217">Z. nitens</emphasis>
</taxonomicName>
group). .............. 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E1FDE551E07C24" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E1FDE551E17DB4" blockId="4.[196,1429,565,796]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FDE551E17DB4" pageId="4" pageNumber="217">
15. Body black. Antennomeres IX–XI pale-yellow. Elytra with distinctly finer punctation (
<bibRefCitation id="EFC54B46F108FFF871B5FDE551687D75" author="ASSING, V." box="[1168,1308,565,589]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 77). Anterior impressions of tergites III and IV with very fine and scattered non-setiferous punctation (
<bibRefCitation id="EFC54B46F108FFF87023FD8551E57D55" author="ASSING, V." box="[1286,1425,597,621]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 115). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F108FFF8778AFDA557377DB5" author="ASSING, V." box="[687,835,629,653]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 250–251).
<collectingRegion id="4990F855F108FFF8712EFDA5501D7DB5" box="[1035,1129,629,653]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="217">Sulawesi</collectingRegion>
. ..........................
<taxonomicName id="4C544D34F108FFF87039FDA451E17DB4" authorityName="ASSING" authorityYear="2017" box="[1308,1429,628,652]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="densihirtus">
<emphasis id="B920EAA5F108FFF87039FDA451E17DB4" bold="true" box="[1308,1429,628,652]" italics="true" pageId="4" pageNumber="217">densihirtus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F108FFF875E1FD7551E07C24" blockId="4.[196,1429,565,796]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FD7551E07C24" pageId="4" pageNumber="217">
– Head and pronotum brown (
<figureCitation id="136F2A32F108FFF87730FD74562B7D84" box="[533,607,676,700]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="4" pageNumber="217">Fig. 19</figureCitation>
); elytra bicoloured, reddish with the postero-lateral portions infuscate
<figureCitation id="136F2A32F108FFF8701BFD7451FC7D84" box="[1342,1416,676,700]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="4" pageNumber="217">Fig. 19</figureCitation>
); abdomen partly reddish to reddish-brown (
<figureCitation id="136F2A32F108FFF87791FD1457757DE4" box="[692,769,708,732]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="4" pageNumber="217">Fig. 34</figureCitation>
). Elytra with coarse punctation (
<figureCitation id="136F2A32F108FFF87172FD1450D77DE4" box="[1111,1187,708,732]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="4" pageNumber="217">Fig. 19</figureCitation>
). Anterior impressions of tergites III and IV with coarse and dense non-setiferous punctation (
<figureCitation id="136F2A32F108FFF876F3FD3450567DC4" box="[982,1058,740,764]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="4" pageNumber="217">Fig. 34</figureCitation>
). Male sexual characters unknown.
<collectingRegion id="4990F855F108FFF875CFFCD5556C7C24" box="[234,280,773,796]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="217">Java</collectingRegion>
. ..................................................................................................................................................................
<taxonomicName id="4C544D34F108FFF8702FFCD451E07C24" authorityName="Cameron" authorityYear="1939" box="[1290,1428,772,796]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B920EAA5F108FFF8702FFCD451E07C24" bold="true" box="[1290,1428,772,796]" italics="true" pageId="4" pageNumber="217">punctipennis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E1FCE351E17C82" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E1FCE351E07C53" blockId="4.[196,1430,819,954]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FCE351E07C53" pageId="4" pageNumber="217">
16. Pronotum more strongly transverse, approximately 1.2 times as broad as long (
<figureCitation id="136F2A32F108FFF87103FCE350067C73" box="[1062,1138,819,843]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="4" pageNumber="217">Fig. 31</figureCitation>
). Pronotum and elytra with sparser and finer punctation (
<figureCitation id="136F2A32F108FFF87703FC8356067C53" box="[550,626,851,875]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="4" pageNumber="217">Fig. 31</figureCitation>
). Median lobe of aedeagus as in
<figureCitation id="136F2A32F108FFF876ECFC83504B7C53" box="[969,1087,851,875]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="4" pageNumber="217">Figs 81–82</figureCitation>
.
<collectingCountry id="F3437627F108FFF8716DFC8450097C53" box="[1096,1149,852,875]" name="Laos" pageId="4" pageNumber="217">Laos</collectingCountry>
. .....................
<taxonomicName id="4C544D34F108FFF87036FC8351E07C53" authorityName="Assing" authorityYear="2017" box="[1299,1428,851,875]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="rutrilobatus">
<emphasis id="B920EAA5F108FFF87036FC8351E07C53" bold="true" box="[1299,1428,851,875]" italics="true" pageId="4" pageNumber="217">rutrilobatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F108FFF875E1FC5351E17C82" blockId="4.[196,1430,819,954]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FC5351E17C82" pageId="4" pageNumber="217">– Pronotum more slender, approximately 1.1 times as long. Pronotum and elytra with denser and coarser punctation. .............................................................................................................................................................................................. 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E1FC0151E17B41" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E1FC0151E17B11" blockId="4.[196,1429,977,1145]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FC0151E17B11" pageId="4" pageNumber="217">
17. Species of slightly smaller body size and with weakly infuscate apical halves of the meso- and metafemora. Male sternite VIII apically acute and somewhat bifid (
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: figure 221). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F108FFF87063FC21555E7B11" author="ASSING, V." pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 216–217). Peninsular
<collectingCountry id="F3437627F108FFF8774CFBC156BE7B11" box="[617,714,1041,1065]" name="Malaysia" pageId="4" pageNumber="217">Malaysia</collectingCountry>
. ........................................................................................................
<taxonomicName id="4C544D34F108FFF87071FBC151E17B11" authorityName="CAMERON" authorityYear="1944" box="[1364,1429,1041,1065]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B920EAA5F108FFF87071FBC151E17B11" bold="true" box="[1364,1429,1041,1065]" italics="true" pageId="4" pageNumber="217">nitens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F108FFF875E1FB9151E17B41" blockId="4.[196,1429,977,1145]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FB9151E17B41" pageId="4" pageNumber="217">
– Slightly larger species with the apical halves of the meso- and metafemora blackish. Male sternite VIII and median lobe of aedeagus of different shapes. (Male sexual characters of
<taxonomicName id="4C544D34F108FFF876AFFBB1578A7B40" baseAuthorityName="PACE" baseAuthorityYear="1986" box="[906,1022,1120,1144]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="B920EAA5F108FFF876AFFBB1578A7B40" box="[906,1022,1120,1144]" italics="true" pageId="4" pageNumber="217">Z. brignolii</emphasis>
</taxonomicName>
are unknown.). ................................ 18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E1FB4051E07A0E" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E1FB4051E07BD0" blockId="4.[196,1429,1168,1335]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FB4051E07BD0" pageId="4" pageNumber="217">
18. Antennae more slender; antennomeres IX–X distinctly oblong. Antennomere XI pale-yellowish. Male sternite VIII obtusely pointed posteriorly. Ventral process of aedeagus (
<bibRefCitation id="EFC54B46F108FFF876A0FB6057837BF0" author="PACE, R." box="[901,1015,1200,1224]" pageId="4" pageNumber="217" pagination="33 - 78" refId="ref27087" refString="PACE, R. 2003: Aleocharinae della Penisola di Malacca (Malaysia) (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 20 (1): 33 - 78." type="journal article" year="2003">PACE 2003</bibRefCitation>
: figures 87–88) conspicuously broad in ventral view. Peninsular
<collectingCountry id="F3437627F108FFF874CBFB00563A7BD0" box="[494,590,1232,1256]" name="Malaysia" pageId="4" pageNumber="217">Malaysia</collectingCountry>
; Borneo. ....................................................................................................
<taxonomicName id="4C544D34F108FFF8703AFB0051E07BD0" authorityName="Cameron" authorityYear="1943" box="[1311,1428,1232,1256]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="B920EAA5F108FFF8703AFB0051E07BD0" bold="true" box="[1311,1428,1232,1256]" italics="true" pageId="4" pageNumber="217">nigerrimus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F108FFF875E1FB2F51E07A0E" blockId="4.[196,1429,1168,1335]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FB2F51E07A0E" pageId="4" pageNumber="217">
– Antennae less slender; antennomeres IX–X weakly transverse (
<bibRefCitation id="EFC54B46F108FFF876B5FB2F50687A2F" author="ASSING, V." box="[912,1052,1279,1303]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 15). Antennomere XI darkyellowish. Male sexual characters unknown.
<collectingCountry id="F3437627F108FFF877E6FACF57517A0F" box="[707,805,1311,1335]" name="Thailand" pageId="4" pageNumber="217">Thailand</collectingCountry>
;
<collectingCountry id="F3437627F108FFF87615FACF57007A0F" box="[816,884,1311,1335]" name="China" pageId="4" pageNumber="217">China</collectingCountry>
:
<collectingRegion id="4990F855F108FFF87658FACF57A07A0E" box="[893,980,1311,1334]" country="China" name="Yunnan" pageId="4" pageNumber="217">Yunnan</collectingRegion>
. .......................................................
<taxonomicName id="4C544D34F108FFF8701DFACE51E07A0E" baseAuthorityName="PACE" baseAuthorityYear="1986" box="[1336,1428,1310,1334]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="B920EAA5F108FFF8701DFACE51E07A0E" bold="true" box="[1336,1428,1310,1334]" italics="true" pageId="4" pageNumber="217">brignolii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E1FA9E51E17AAD" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E1FA9E51E17A5E" blockId="4.[196,1429,1358,1429]" box="[196,1429,1358,1382]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FA9E51E17A5E" box="[196,1429,1358,1382]" pageId="4" pageNumber="217">
19. Pronotum reddish. Large to very large species (species of the
<taxonomicName id="4C544D34F108FFF87656FA9F57B97A5E" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[883,973,1358,1382]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B920EAA5F108FFF87656FA9F57B97A5E" box="[883,973,1358,1382]" italics="true" pageId="4" pageNumber="217">Z. hirtus</emphasis>
</taxonomicName>
group). ...................................................... 20
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F108FFF875E1FAAD51E17AAD" blockId="4.[196,1429,1358,1429]" box="[196,1429,1405,1429]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FAAD51E17AAD" box="[196,1429,1405,1429]" pageId="4" pageNumber="217">– Pronotum dark-brown to blackish. Species of relatively small to large size. ............................................................. 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E1FA7C51E1794B" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E1FA7C51E0791C" blockId="4.[196,1429,1452,1652]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1FA7C51E0791C" pageId="4" pageNumber="217">
20. Head pale-reddish, of similar coloration as pronotum; elytra dark-brown to blackish with the anterior margin reddish (
<bibRefCitation id="EFC54B46F108FFF8746CFA1C55A07ADC" author="ASSING, V." box="[329,468,1484,1508]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 61); abdomen distinctly bicoloured, with tergites III–V reddish and VI–VIII extensively blackish (
<bibRefCitation id="EFC54B46F108FFF874AAFA3C566C793C" author="ASSING, V." box="[399,536,1516,1540]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 114). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F108FFF876FDFA3C501F793C" author="ASSING, V." box="[984,1131,1516,1540]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 187–188).
<collectingCountry id="F3437627F108FFF8700BFA3C51E4793C" box="[1326,1424,1516,1540]" name="Thailand" pageId="4" pageNumber="217">Thailand</collectingCountry>
; Peninsular
<collectingCountry id="F3437627F108FFF87447F9DC55B7791C" box="[354,451,1548,1572]" name="Malaysia" pageId="4" pageNumber="217">Malaysia</collectingCountry>
. ..............................................................................................................................................
<taxonomicName id="4C544D34F108FFF87010F9DC51E0791C" authorityName="ASSING" authorityYear="2017" box="[1333,1428,1548,1572]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="russiceps">
<emphasis id="B920EAA5F108FFF87010F9DC51E0791C" bold="true" box="[1333,1428,1548,1572]" italics="true" pageId="4" pageNumber="217">russiceps</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F108FFF875E1F9EC51E1794B" blockId="4.[196,1429,1452,1652]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1F9EC51E1794B" pageId="4" pageNumber="217">– Head reddish-brown to black, darker than the pronotum; elytra uniformly reddish, or reddish with the posterolateral portions more or less distinctly infuscate; abdomen not distinctly bicoloured. Larger species. ................ 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F108FFF875E1F95A51E078A9" pageId="4" pageNumber="217">
<paragraph id="8BEB36B7F108FFF875E1F95A51E1783A" blockId="4.[196,1429,1674,1938]" pageId="4" pageNumber="217">
<keyLead id="30A59582F108FFF875E1F95A51E1783A" pageId="4" pageNumber="217">
21. Largest species of the subgenus in the Oriental region; length of forebody 4.0–
<quantity id="4CAC9B52F108FFF87165F95A50E7799A" box="[1088,1171,1674,1698]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="217" unit="mm" value="4.3">4.3 mm</quantity>
. Pronotum slender, less than 1.1 times as broad as long (
<bibRefCitation id="EFC54B46F108FFF87765F97A56BF79FA" author="ASSING, V." box="[576,715,1706,1730]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 76). Elytra with coarse punctation (
<bibRefCitation id="EFC54B46F108FFF871B7F97A516979FA" author="ASSING, V." box="[1170,1309,1706,1730]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 76). Anterior impressions of tergites III–V with coarse and dense non-setiferous punctation (
<bibRefCitation id="EFC54B46F108FFF871AFF91A516779DA" author="ASSING, V." box="[1162,1299,1738,1762]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 113). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F108FFF8770BF93A56B7783A" author="ASSING, V." box="[558,707,1770,1794]" pageId="4" pageNumber="217" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 245–246).
<collectingRegion id="4990F855F108FFF876AFF93A579C783A" box="[906,1000,1770,1794]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="217">Sulawesi</collectingRegion>
. ...........................................................
<taxonomicName id="4C544D34F108FFF87044F93A51E1783A" authorityName="ASSING" authorityYear="2017" box="[1377,1429,1770,1794]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="titan">
<emphasis id="B920EAA5F108FFF87044F93A51E1783A" bold="true" box="[1377,1429,1770,1794]" italics="true" pageId="4" pageNumber="217">titan</emphasis>
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</paragraph>
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– Large species, but less so than
<taxonomicName id="4C544D34F108FFF87709F8CA560E7809" authorityName="ASSING" authorityYear="2017" box="[556,634,1818,1841]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="titan">
<emphasis id="B920EAA5F108FFF87709F8CA560E7809" box="[556,634,1818,1841]" italics="true" pageId="4" pageNumber="217">Z. titan</emphasis>
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; length of forebody less than 4.0 mm. Pronotum more transverse, approximately 1.15 times as broad as long. Elytra with fine and dense punctation. Anterior impressions of tergites III–V with fine and sparse non-setiferous punctation. Median lobe of aedeagus as in
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. Widespread (
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: map 10). ................................................................................................................................
<taxonomicName id="4C544D34F108FFF87026F8AA51E078A9" authorityName="Cameron" authorityYear="1939" box="[1283,1428,1914,1937]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="preangeranus">
<emphasis id="B920EAA5F108FFF87026F8AA51E078A9" bold="true" box="[1283,1428,1914,1937]" italics="true" pageId="4" pageNumber="217">preangeranus</emphasis>
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</paragraph>
</keyStep>
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<paragraph id="8BEB36B7F108FFF875E1F87951E07718" blockId="4.[196,1428,1961,2081]" pageId="4" pageNumber="217">
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22. Elytra with conspicuously coarse and dense punctation (
<figureCitation id="136F2A32F108FFF87665F87957F878F9" box="[832,908,1961,1985]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="4" pageNumber="217">Fig. 19</figureCitation>
). Pronotum strongly convex in cross-section and weakly transverse, approximately 1.1 times as broad as long (
<figureCitation id="136F2A32F108FFF87645F81957DE78D9" box="[864,938,1993,2017]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="4" pageNumber="217">Fig. 19</figureCitation>
). Head and pronotum dark-brown to blackishbrown; elytra dark-reddish with infuscate postero-lateral portions (
<figureCitation id="136F2A32F108FFF87684F839579E7739" box="[929,1002,2025,2049]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="4" pageNumber="217">Fig. 19</figureCitation>
). Large species; body length 6.5–8.0 mm; length of forebody 3.0–
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. Male sexual characters unknown.
<collectingRegion id="4990F855F108FFF8768CF7D957A37718" box="[937,983,2057,2080]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="217">Java</collectingRegion>
, Borneo. ................................ punctipennis
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F109FFF975BAFF29511B7E29" blockId="5.[159,1391,248,273]" box="[159,1391,248,273]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BAFF29511B7E29" box="[159,1391,248,273]" pageId="5" pageNumber="218">– Elytral punctation less coarse and less dense. Character combination different. ..................................................... 23</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BEB36B7F109FFF975BAFEF8511B7E87" blockId="5.[159,1392,295,495]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BAFEF8511B7E87" pageId="5" pageNumber="218">
23. Abdomen distinctly bicoloured; tergites III–V reddish and tergites VI–VII extensively black (
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: figure 112). Elytra distinctly bicoloured with the anterior portion reddish and the posterior portion sharply black (
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: figure 56). Punctation of elytra coarse (
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: figure 56). Antennomeres IX–X or VIII–X pale-yellowish, sharply contrasting with the black antennomeres IV–VII (
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: figure 1). Median lobe of aedeagus as in
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: figures 176–177).
<collectingCountry id="F3437627F109FFF977EFFE7857417EF8" box="[714,821,424,448]" name="Myanmar" pageId="5" pageNumber="218">Myanmar</collectingCountry>
;
<collectingCountry id="F3437627F109FFF9761AFE7757EA7E87" box="[831,926,423,447]" name="Vietnam" pageId="5" pageNumber="218">Vietnam</collectingCountry>
. ..........................................................
<taxonomicName id="4C544D34F109FFF97031FE77511B7E87" authorityName="SCHEERPELTZ" authorityYear="1965" box="[1300,1391,423,447]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B920EAA5F109FFF97031FE77511B7E87" bold="true" box="[1300,1391,423,447]" italics="true" pageId="5" pageNumber="218">malaisei</emphasis>
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</paragraph>
<paragraph id="8BEB36B7F109FFF975BAFE07511B7ED7" blockId="5.[159,1392,295,495]" box="[159,1391,471,495]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BAFE07511B7ED7" box="[159,1391,471,495]" pageId="5" pageNumber="218">– Abdomen not distinctly bicoloured. ............................................................................................................................... 24</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BEB36B7F109FFF975BAFDD6511B7D65" blockId="5.[159,1391,518,717]" pageId="5" pageNumber="218">
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24. Small species; length of forebody
<quantity id="4CAC9B52F109FFF9770DFDD656DD7D25" box="[552,681,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="5" pageNumber="218" unit="mm" value="2.25" valueMax="2.4" valueMin="2.1">2.1–2.4 mm</quantity>
. Antennomere XI elongate, approximately as long as the combined length of VIII–X (
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: figures 26, 39). Pubescence inconspicuous, moderately long, suberect, and moderately dense. Median lobe of aedeagus as in
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: figures 223–224, 234–235). Widespread. ...................
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<emphasis id="B920EAA5F109FFF97031FD95511B7D65" bold="true" box="[1300,1391,581,605]" italics="true" pageId="5" pageNumber="218">gratellus</emphasis>
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</keyLead>
</paragraph>
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– Larger species; length of forebody&gt;
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. Antennomere XI short and of conical shape, approximately as long as the combined length of IX and X at most. (
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: figure185). Pubescence longer, denser, and sub-erect to erect. Species of the
<taxonomicName id="4C544D34F109FFF97492FD6656657DF5" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[439,529,693,717]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B920EAA5F109FFF97492FD6656657DF5" box="[439,529,693,717]" italics="true" pageId="5" pageNumber="218">Z. hirtus</emphasis>
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group. .......................................................................................................................... 25
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<paragraph id="8BEB36B7F109FFF975BAFD34511A7C44" blockId="5.[159,1392,740,1099]" pageId="5" pageNumber="218">
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25. Pronotum with coarse and rather dense punctation (
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: figure 78). Antennae shorter and with more transverse antennomeres IV–X; antennomeres IV weakly transverse and X more than twice as broad as long (
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: figure 38). Abdominal tergites VI and VII with very dense and rather fine non-setiferous punctation anteriorly (
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: figure 16). Median lobe of aedeagus as in
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: figures 255–256).
<collectingRegion id="4990F855F109FFF97028FC9451187C64" box="[1293,1388,836,860]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="218">Sulawesi</collectingRegion>
. ........................................................................................................................................................................
<taxonomicName id="4C544D34F109FFF971DDFCB4511A7C44" authorityName="ASSING" authorityYear="2017" box="[1272,1390,868,892]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="nigrihirtus">
<emphasis id="B920EAA5F109FFF971DDFCB4511A7C44" bold="true" box="[1272,1390,868,892]" italics="true" pageId="5" pageNumber="218">nigrihirtus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BEB36B7F109FFF975BAFC44511B7B73" blockId="5.[159,1392,740,1099]" pageId="5" pageNumber="218">
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– Pronotum with less distinct and less coarse punctation (
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: figure 184). Antennae longer and with less transverse antennomeres IV–X; antennomeres IV approximately as long as broad and X less than twice as broad as long (
<bibRefCitation id="EFC54B46F109FFF97441FC03558B7CD3" author="ASSING, V." box="[356,511,979,1003]" pageId="5" pageNumber="218" pagination="13 - 111" refId="ref25807" refString="ASSING, V. 2016 b: On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae). - Contributions to Entomology 66 (1): 13 - 111 - DOI: 10.21248 / contrib. entomol. 66.1.13 - 111." type="journal article" year="2016">ASSING 2016b</bibRefCitation>
: figure185). Abdominal tergites VI and VII with distinctly less fine and less dense punctation (
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: figure 186). Pubescence of dorsal surface long, dense, and sub-erect to erect; all sternites with long, dense, and erect pubescence. Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F109FFF97127FBC350DC7B13" author="ASSING, V." box="[1026,1192,1043,1067]" pageId="5" pageNumber="218" pagination="13 - 111" refId="ref25807" refString="ASSING, V. 2016 b: On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae). - Contributions to Entomology 66 (1): 13 - 111 - DOI: 10.21248 / contrib. entomol. 66.1.13 - 111." type="journal article" year="2016">ASSING (2016b</bibRefCitation>
: figures 461–462).
<collectingRegion id="4990F855F109FFF975E1FBE354857B73" box="[196,241,1075,1099]" country="Indonesia" name="Bali" pageId="5" pageNumber="218">Bali</collectingRegion>
. ................................................................................................................................................................
<taxonomicName id="4C544D34F109FFF97024FBE3511B7B73" authorityName="ASSING" authorityYear="2016" box="[1281,1391,1075,1099]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="wunderlei">
<emphasis id="B920EAA5F109FFF97024FBE3511B7B73" bold="true" box="[1281,1391,1075,1099]" italics="true" pageId="5" pageNumber="218">wunderlei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F109FFF975BAFBB2511B7BF1" pageId="5" pageNumber="218">
<paragraph id="8BEB36B7F109FFF975BAFBB2511B7BA2" blockId="5.[159,1391,1122,1226]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BAFBB2511B7BA2" pageId="5" pageNumber="218">
26. Legs bicoloured; femora brown to black with the bases of the meso- and metafemora yellow. Forebody of uniformly dark-brown to black coloration. Species of the
<taxonomicName id="4C544D34F109FFF97796FB5357797BA2" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[691,781,1154,1178]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B920EAA5F109FFF97796FB5357797BA2" box="[691,781,1154,1178]" italics="true" pageId="5" pageNumber="218">Z. hirtus</emphasis>
</taxonomicName>
group. ................................................................................. 27
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F109FFF975BAFB62511B7BF1" blockId="5.[159,1391,1122,1226]" box="[159,1391,1201,1226]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BAFB62511B7BF1" box="[159,1391,1201,1226]" pageId="5" pageNumber="218">– Legs of uniformly yellowish coloration. ......................................................................................................................... 28</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F109FFF975BAFB31511B7AFF" pageId="5" pageNumber="218">
<paragraph id="8BEB36B7F109FFF975BAFB31511B7A00" blockId="5.[159,1391,1248,1479]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BAFB31511B7A00" pageId="5" pageNumber="218">
27. Smaller species; length of forebody
<quantity id="4CAC9B52F109FFF9771AFB3156E77BC0" box="[575,659,1249,1273]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="218" unit="mm" value="2.8">2.8 mm</quantity>
. Antennae shorter,
<quantity id="4CAC9B52F109FFF97646FB3157C07BC0" box="[867,948,1249,1273]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="218" unit="mm" value="1.7">1.7 mm</quantity>
long; antennomere IV weakly transverse; antennomeres IX–X more than twice as broad as long (
<bibRefCitation id="EFC54B46F109FFF97628FAD157E37A20" author="ASSING, V." box="[781,919,1280,1305]" pageId="5" pageNumber="218" pagination="1257 - 1282" refId="ref25690" refString="ASSING, V. 2015: On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 47 (2): 1257 - 1282 - http: // www. zobodat. at / pdf / LBB _ 0047 _ 2 _ 1257 - 1282. pdf." type="journal article" year="2015">ASSING 2015</bibRefCitation>
: figure 57). Abdomen partly brownish. Male sexual characters unknown.
<collectingCountry id="F3437627F109FFF974D6FAF056267A00" box="[499,594,1312,1336]" name="Vietnam" pageId="5" pageNumber="218">Vietnam</collectingCountry>
. .................................................................................................................
<taxonomicName id="4C544D34F109FFF97033FAF0511B7A00" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[1302,1391,1312,1336]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="B920EAA5F109FFF97033FAF0511B7A00" bold="true" box="[1302,1391,1312,1336]" italics="true" pageId="5" pageNumber="218">funestus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F109FFF975BAFA80511B7AFF" blockId="5.[159,1391,1248,1479]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BAFA80511B7AFF" pageId="5" pageNumber="218">
– Larger species; length of forebody
<quantity id="4CAC9B52F109FFF97713FA8056C37A5F" box="[566,695,1360,1384]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.7" metricValueMin="3.6" pageId="5" pageNumber="218" unit="mm" value="3.65" valueMax="3.7" valueMin="3.6">3.6–3.7 mm</quantity>
. Antennae
<quantity id="4CAC9B52F109FFF97614FA9F57C47A5F" box="[817,944,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="5" pageNumber="218" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
long; antennomere IV noticeably oblong; antennomeres IX–X less than twice as broad as long (
<figureCitation id="136F2A32F109FFF977D6FAA0571F7AB0" box="[755,875,1392,1416]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="5" pageNumber="218">Figs 11–12</figureCitation>
). Abdomen black, except for the brown posterior margins of tergites VII and VIII (
<figureCitation id="136F2A32F109FFF97700FA4056057A90" box="[549,625,1424,1448]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="5" pageNumber="218">Fig. 49</figureCitation>
). Median lobe of the aedeagus as in
<figureCitation id="136F2A32F109FFF976D5FA4050117A90" box="[1008,1125,1424,1448]" captionStart="Figs 87–102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87–102: Zyras parvilobatus (87–89), Z. hlavaci (90–95; 90–92: holotype; 93–95: paratype from Selangor), Z. hirtiventris, holotype (96–98), Z. lunatus (99–101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (87–88, 90–91, 93–94, 96–97, 99–100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87–101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="5" pageNumber="218">Figs 96–97</figureCitation>
.
<collectingCountry id="F3437627F109FFF9714BFA4050B87A90" box="[1134,1228,1424,1448]" name="Vietnam" pageId="5" pageNumber="218">Vietnam</collectingCountry>
,
<collectingCountry id="F3437627F109FFF971F2FA4051787A9F" box="[1239,1292,1424,1447]" name="Laos" pageId="5" pageNumber="218">Laos</collectingCountry>
. .............. ..............................................................................................................................................................................
<taxonomicName id="4C544D34F109FFF971D3FA7F511B7AFF" authorityName="Assing" authorityYear="2017" box="[1270,1391,1455,1479]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F109FFF971D3FA7F511B7AFF" bold="true" box="[1270,1391,1455,1479]" italics="true" pageId="5" pageNumber="218">hirtiventris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F109FFF975BAFA0F511B797E" pageId="5" pageNumber="218">
<paragraph id="8BEB36B7F109FFF975BAFA0F511B792E" blockId="5.[158,1391,1503,1606]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BAFA0F511B792E" pageId="5" pageNumber="218">
28. Pronotum and elytra with very sparse and fine punctation; interstices on elytra on average more than twice as broad as diameter of punctures (
<figureCitation id="136F2A32F109FFF97738FA2F561D792F" box="[541,617,1535,1559]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="5" pageNumber="218">Fig. 24</figureCitation>
). Median lobe of aedeagus as in
<figureCitation id="136F2A32F109FFF976E5FA2F5041792F" box="[960,1077,1535,1559]" captionStart="Figs 56–57" captionStartId="16.[189,233,233,257]" captionText="Figs 56–57; paramere (Fig. 58) longer than median lobe, 0.77 mm long, and with rather large and distinctly flattened apical lobe." pageId="5" pageNumber="218">Figs 56–57</figureCitation>
. Java. .......................
<taxonomicName id="4C544D34F109FFF97028FA2E511B792E" authorityName="Last" authorityYear="1969" box="[1293,1391,1534,1558]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="facundus">
<emphasis id="B920EAA5F109FFF97028FA2E511B792E" bold="true" box="[1293,1391,1534,1558]" italics="true" pageId="5" pageNumber="218">facundus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F109FFF975BBF9FE511B797E" blockId="5.[158,1391,1503,1606]" box="[158,1391,1582,1606]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BBF9FE511B797E" box="[158,1391,1582,1606]" pageId="5" pageNumber="218">– Pronotum and elytra with much denser and usually also coarser punctation. ......................................................... 29</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F109FFF975BBF98D511B783C" pageId="5" pageNumber="218">
<paragraph id="8BEB36B7F109FFF975BBF98D511B798D" blockId="5.[158,1391,1629,1796]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BBF98D511B798D" pageId="5" pageNumber="218">
29. Abdomen bicoloured with segments III–V or III–VI reddish and VI–VIII or VII–VIII extensively infuscate. For
<taxonomicName id="4C544D34F109FFF975E1F9AE555379AD" authorityName="DVORAK" authorityYear="1996" box="[196,295,1662,1685]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="218" phylum="Arthropoda" rank="species" species="quasar">
<emphasis id="B920EAA5F109FFF975E1F9AE555379AD" box="[196,295,1662,1685]" italics="true" pageId="5" pageNumber="218">Z. quasar</emphasis>
</taxonomicName>
from Vietam with a posteriorly somewhat infuscate tergite V (
<bibRefCitation id="EFC54B46F109FFF976F2F9AD500679AD" author="ASSING, V." box="[983,1138,1661,1685]" pageId="5" pageNumber="218" pagination="13 - 111" refId="ref25807" refString="ASSING, V. 2016 b: On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae). - Contributions to Entomology 66 (1): 13 - 111 - DOI: 10.21248 / contrib. entomol. 66.1.13 - 111." type="journal article" year="2016">ASSING 2016b</bibRefCitation>
: figure 180) follow this alternative. .......................................................................................................................................................................... 30
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F109FFF975BBF91D511B783C" blockId="5.[158,1391,1629,1796]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BBF91D511B783C" pageId="5" pageNumber="218">– Abdomen of uniformly reddish or blackish coloration, or reddish with tergite VI (and sometimes also the anterior portion of tergite VII weakly) infuscate. ........................................................................................................................ 34</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F109FFF975BBF8CB511B789A" pageId="5" pageNumber="218">
<paragraph id="8BEB36B7F109FFF975BBF8CB511B786B" blockId="5.[158,1391,1819,1954]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BBF8CB511B786B" pageId="5" pageNumber="218">30. Pronotum with fine and sparse punctation; dorsal surface smooth, not impressed, except for the usual posteromedian impression. ........................................................................................................................................................... 31</keyLead>
</paragraph>
<paragraph id="8BEB36B7F109FFF975BBF8BB511B789A" blockId="5.[158,1391,1819,1954]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BBF8BB511B789A" pageId="5" pageNumber="218">– Pronotum with coarse and rather dense, or coarse and very irregularly distributed punctation; in the latter case pronotum with uneven surface (punctation situated in more or less pronounced impressions). ......................... 32</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F109FFFA75BBF86A51E17E08" lastPageId="6" lastPageNumber="219" pageId="5" pageNumber="218">
<paragraph id="8BEB36B7F109FFF975BBF86A511B7729" blockId="5.[158,1391,1977,2065]" pageId="5" pageNumber="218">
<keyLead id="30A59582F109FFF975BBF86A511B7729" pageId="5" pageNumber="218">
31. Antennomere XI strongly elongate, at least as long as, usually longer than the combined length of antennomeres VIII–X (
<bibRefCitation id="EFC54B46F109FFF97404F80A55CC78C9" author="ASSING, V." box="[289,440,2009,2034]" pageId="5" pageNumber="218" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 43). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F109FFF9765BF809506C78C9" author="ASSING, V." box="[894,1048,2009,2033]" pageId="5" pageNumber="218" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">Assing (2016a</bibRefCitation>
: figures 272–273). Widespread. ............................................................................................................................................................................... castaneus
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E0FF2951E17E08" blockId="6.[197,1429,248,305]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E0FF2951E17E08" pageId="6" pageNumber="219">
– Antennomere XI shorter, shorter than the combined length of antennomeres VIII–X (
<bibRefCitation id="EFC54B46F10AFFFA715CFF2951657E28" author="ASSING, V." box="[1145,1297,248,273]" pageId="6" pageNumber="219" pagination="13 - 111" refId="ref25807" refString="ASSING, V. 2016 b: On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae). - Contributions to Entomology 66 (1): 13 - 111 - DOI: 10.21248 / contrib. entomol. 66.1.13 - 111." type="journal article" year="2016">ASSING 2016b</bibRefCitation>
: figure 179). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10AFFFA770AFEC956A47E08" author="ASSING, V." box="[559,720,280,305]" pageId="6" pageNumber="219" pagination="13 - 111" refId="ref25807" refString="ASSING, V. 2016 b: On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae). - Contributions to Entomology 66 (1): 13 - 111 - DOI: 10.21248 / contrib. entomol. 66.1.13 - 111." type="journal article" year="2016">ASSING (2016b</bibRefCitation>
: figures 457–458).
<collectingCountry id="F3437627F10AFFFA76BDFEC857837E08" box="[920,1015,280,304]" name="Vietnam" pageId="6" pageNumber="219">Vietnam</collectingCountry>
. ....................................................
<taxonomicName id="4C544D34F10AFFFA706EFEC951E17E08" authorityName="DVORAK" authorityYear="1996" box="[1355,1429,281,304]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="219" phylum="Arthropoda" rank="species" species="quasar">
<emphasis id="B920EAA5F10AFFFA706EFEC951E17E08" bold="true" box="[1355,1429,281,304]" italics="true" pageId="6" pageNumber="219">quasar</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10AFFFA75E0FE9851E17D37" pageId="6" pageNumber="219">
<paragraph id="8BEB36B7F10AFFFA75E0FE9851E17E87" blockId="6.[197,1430,327,527]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E0FE9851E17E87" pageId="6" pageNumber="219">
32. Pronotum strongly transverse, nearly 1.3 times as broad as long and 1.2 times as broad as head; surface without distinct impressions (except for the usual postero-median impression); punctation dense and rather coarse (
<bibRefCitation id="EFC54B46F10AFFFA702DFEB851E57EB8" author="ASSING, V." box="[1288,1425,360,384]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 68). Abdominal tergite VI reddish or brown with reddish margins (
<bibRefCitation id="EFC54B46F10AFFFA76CDFE5850047E98" author="ASSING, V." box="[1000,1136,392,416]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 102). Median lobe of aedeagus with conspicuously short ventral process (
<bibRefCitation id="EFC54B46F10AFFFA7627FE7857FE7EF8" author="ASSING, V." box="[770,906,424,448]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figures 203–204).
<collectingCountry id="F3437627F10AFFFA7168FE7750DA7E87" box="[1101,1198,423,447]" name="Thailand" pageId="6" pageNumber="219">Thailand</collectingCountry>
. ..............
<taxonomicName id="4C544D34F10AFFFA7036FE7751E17E87" authorityName="ASSING" authorityYear="2017" box="[1299,1429,423,447]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="219" phylum="Arthropoda" rank="species" species="brevilobatus">
<emphasis id="B920EAA5F10AFFFA7036FE7751E17E87" bold="true" box="[1299,1429,423,447]" italics="true" pageId="6" pageNumber="219">brevilobatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E0FE0751E17D37" blockId="6.[197,1430,327,527]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E0FE0751E17D37" pageId="6" pageNumber="219">– Pronotum less transverse, 1.15–1.20 times as broad as long; surface smooth or uneven. Abdominal tergites VI reddish and VII–VIII extensively infuscate. .................................................................................................................. 33</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10AFFFA75E1FDF651E07C34" pageId="6" pageNumber="219">
<paragraph id="8BEB36B7F10AFFFA75E1FDF651E07DA6" blockId="6.[196,1429,550,781]" pageId="6" pageNumber="219">
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33. Pronotum with uneven surface; punctation conspicuously irregularly distributed and situated in more or less pronounced impressions (
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: figure 67). Antennae more slender; antennomere XI elongate, approximately as long as the combined length of VIII–X (
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: figures 30–31). Median lobe of aedeagus as in
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: figures 198–199).
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;
<collectingCountry id="F3437627F10AFFFA7790FD56569E7DA5" box="[693,746,646,669]" name="Laos" pageId="6" pageNumber="219">Laos</collectingCountry>
. .......................................................................................
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<emphasis id="B920EAA5F10AFFFA7031FD5651E07DA6" bold="true" box="[1300,1428,646,670]" italics="true" pageId="6" pageNumber="219">novinversus</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E1FD6551E07C34" blockId="6.[196,1429,550,781]" pageId="6" pageNumber="219">
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– Pronotum with smooth surface; punctation moderately irregularly distributed (
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: figure 172). antennae less slender; antennomere XI shorter, barely as long as the combined length of IX and X (
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: figure 173) Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10AFFFA778AFD2557247C35" author="ASSING, V." box="[687,848,757,781]" pageId="6" pageNumber="219" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 231–232). Widespread. .................
<taxonomicName id="4C544D34F10AFFFA7007FD2451E07C34" authorityName="CAMERON" authorityYear="1930" box="[1314,1428,756,780]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="219" phylum="Arthropoda" rank="species" species="bettotanus">
<emphasis id="B920EAA5F10AFFFA7007FD2451E07C34" bold="true" box="[1314,1428,756,780]" italics="true" pageId="6" pageNumber="219">bettotanus</emphasis>
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</paragraph>
</keyStep>
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<paragraph id="8BEB36B7F10AFFFA75E1FCF451E17C64" blockId="6.[196,1429,804,907]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1FCF451E17C64" pageId="6" pageNumber="219">34. Abdomen predominantly reddish, sometimes with tergite VI and the anterior portion of tergite VII more or less distinctly infuscate. ........................................................................................................................................................... 35</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E1FCA451E17CB3" blockId="6.[196,1429,804,907]" box="[196,1429,883,907]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1FCA451E17CB3" box="[196,1429,883,907]" pageId="6" pageNumber="219">– Abdomen predominantly black. ...................................................................................................................................... 40</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10AFFFA75E1FC7251E17B71" pageId="6" pageNumber="219">
<paragraph id="8BEB36B7F10AFFFA75E1FC7251E17CC2" blockId="6.[196,1429,930,1098]" pageId="6" pageNumber="219">
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35. Small species of variable coloration; length of forebody
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. Antennomere XI elongate, at least as long as the combined length of antennomeres VIII–X (
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: figures 26, 39). Forebody with fine punctation. Median lobe of aedeagus as
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: figures 222–223, 234–235). Widespread. .................................
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<emphasis id="B920EAA5F10AFFFA701CFC3251E17CC2" bold="true" box="[1337,1429,994,1018]" italics="true" pageId="6" pageNumber="219">gratellus</emphasis>
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</keyLead>
</paragraph>
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– Larger species; length of forebody&gt;
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. Antennomere short and of conical shape, shorter than the combined length of IX and X. Body with long pubescence. Species of the
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<emphasis id="B920EAA5F10AFFFA76A4FBE257AF7B71" box="[897,987,1073,1097]" italics="true" pageId="6" pageNumber="219">Z. hirtus</emphasis>
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group. ..................................................... 36
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36. Forebody dark-brown to black (
<figureCitation id="136F2A32F10AFFFA7716FBB0560B7B41" box="[563,639,1120,1145]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="6" pageNumber="219">Fig. 33</figureCitation>
). Body of larger average size; length of forebody 3.0–
<quantity id="4CAC9B52F10AFFFA71B9FBB0509B7B40" box="[1180,1263,1120,1144]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="219" unit="mm" value="3.5">3.5 mm</quantity>
. Abdomen with more extensive and coarser non-setiferous punctation on tergites VI and VII (
<figureCitation id="136F2A32F10AFFFA7109FB50500C7BA1" box="[1068,1144,1152,1177]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="6" pageNumber="219">Fig. 48</figureCitation>
). Median lobe of aedeagus larger,
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long, and shaped as in
<figureCitation id="136F2A32F10AFFFA77E7FB7057F07B81" box="[706,900,1184,1209]" captionStart="Figs 87–102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87–102: Zyras parvilobatus (87–89), Z. hlavaci (90–95; 90–92: holotype; 93–95: paratype from Selangor), Z. hirtiventris, holotype (96–98), Z. lunatus (99–101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (87–88, 90–91, 93–94, 96–97, 99–100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87–101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="6" pageNumber="219">Figs 90–91, 93–94</figureCitation>
. Peninsular
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. .................................
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<emphasis id="B920EAA5F10AFFFA7062FB7051E17B80" bold="true" box="[1351,1429,1184,1208]" italics="true" pageId="6" pageNumber="219">hlavaci</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E1FB0051E17A10" blockId="6.[196,1429,1120,1320]" pageId="6" pageNumber="219">
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– Forebody reddish to dark-brown. Body of smaller average size,
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. Abdomen with finer and less extensive non-setiferous punctation on tergites VI and VII. Median lobe of aedeagus &lt;
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long and of different shape. .............................................................................................................................................................................................. 37
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37. Slightly larger species; length of forebody 3.0–
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. Abdominal tergite VI infuscate. Median lobe of aedeagus very broad in ventral view (
<bibRefCitation id="EFC54B46F10AFFFA7735FA8F56EE7A4F" author="ASSING, V." box="[528,666,1375,1399]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figures 229–230). Borneo. ........................................................
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<emphasis id="B920EAA5F10AFFFA7007FA8E51E07A4E" bold="true" box="[1314,1428,1374,1398]" italics="true" pageId="6" pageNumber="219">parahirtus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E1FA5E51E17AFE" blockId="6.[196,1429,1343,1478]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1FA5E51E17AFE" pageId="6" pageNumber="219">
– Smaller species; length of forebody
<quantity id="4CAC9B52F10AFFFA7748FA5E569B7A9D" box="[621,751,1422,1446]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="6" pageNumber="219" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 mm</quantity>
. Abdominal tergite VI not distinctly infuscate. Median lobe of aedeagus more slender in ventral view. .......................................................................................................................... 38
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10AFFFA75E1FA0D51E1797C" pageId="6" pageNumber="219">
<paragraph id="8BEB36B7F10AFFFA75E1FA0D51E0792D" blockId="6.[196,1429,1501,1604]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1FA0D51E0792D" pageId="6" pageNumber="219">
38. Ventral process of aedeagus distinctly angled subapically in lateral view (
<figureCitation id="136F2A32F10AFFFA76CFFA0D50047ACD" box="[1002,1136,1501,1525]" captionStart="Figs 87–102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87–102: Zyras parvilobatus (87–89), Z. hlavaci (90–95; 90–92: holotype; 93–95: paratype from Selangor), Z. hirtiventris, holotype (96–98), Z. lunatus (99–101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (87–88, 90–91, 93–94, 96–97, 99–100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87–101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="6" pageNumber="219">Figs 99–100</figureCitation>
). Peninsular
<collectingCountry id="F3437627F10AFFFA71DCFA0D512E7ACD" box="[1273,1370,1501,1525]" name="Malaysia" pageId="6" pageNumber="219">Malaysia</collectingCountry>
. ....... .....................................................................................................................................................................................
<taxonomicName id="4C544D34F10AFFFA7066FA2D51E0792D" authorityName="Assing" authorityYear="2017" box="[1347,1428,1533,1557]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="219" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B920EAA5F10AFFFA7066FA2D51E0792D" bold="true" box="[1347,1428,1533,1557]" italics="true" pageId="6" pageNumber="219">lunatus</emphasis>
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</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E1F9FD51E1797C" blockId="6.[196,1429,1501,1604]" box="[196,1429,1580,1604]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1F9FD51E1797C" box="[196,1429,1580,1604]" pageId="6" pageNumber="219">– Ventral process of aedeagus not angled subapically in lateral view. Sunda Islands. ................................................. 39</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10AFFFA75E1F98B51E079DA" pageId="6" pageNumber="219">
<paragraph id="8BEB36B7F10AFFFA75E1F98B51E079AB" blockId="6.[196,1429,1627,1762]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1F98B51E079AB" pageId="6" pageNumber="219">
39. Aedeagus with more slender ventral process both in lateral and in ventral view and with more prominent crista apicalis (
<figureCitation id="136F2A32F10AFFFA7463F9AB55CB79AB" box="[326,447,1659,1683]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="6" pageNumber="219">Figs 73–74</figureCitation>
).
<collectingRegion id="4990F855F10AFFFA74F5F9AC558A79AB" box="[464,510,1660,1683]" country="Indonesia" name="Jawa Timur" pageId="6" pageNumber="219">Java</collectingRegion>
. .................................................................................................................................
<taxonomicName id="4C544D34F10AFFFA7003F9AB51E079AB" authorityName="Cameron" authorityYear="1939" box="[1318,1428,1659,1683]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="219" phylum="Arthropoda" rank="species" species="flavorufus">
<emphasis id="B920EAA5F10AFFFA7003F9AB51E079AB" bold="true" box="[1318,1428,1659,1683]" italics="true" pageId="6" pageNumber="219">flavorufus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E1F97B51E079DA" blockId="6.[196,1429,1627,1762]" pageId="6" pageNumber="219">
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– Aedeagus with less slender ventral process and with broader, less prominent crista apcialis (
<figureCitation id="136F2A32F10AFFFA7190F97A515A79FA" box="[1205,1326,1706,1730]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="6" pageNumber="219">Figs 63–64</figureCitation>
). Borneo. ............................................................................................................................................................................
<taxonomicName id="4C544D34F10AFFFA702BF91A51E079DA" authorityName="Cameron" authorityYear="1943" box="[1294,1428,1738,1762]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="219" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B920EAA5F10AFFFA702BF91A51E079DA" bold="true" box="[1294,1428,1738,1762]" italics="true" pageId="6" pageNumber="219">matangensis</emphasis>
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</paragraph>
</keyStep>
<keyStep id="30A02E12F10AFFFA75E1F92A51E178B9" pageId="6" pageNumber="219">
<paragraph id="8BEB36B7F10AFFFA75E1F92A51E1780A" blockId="6.[196,1429,1785,1921]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1F92A51E1780A" pageId="6" pageNumber="219">40. Antennomere XI elongate, approximately as long as the combined length of antennomeres VIII–X. Pronotum rather small in relation to head, only slightly broader than head. .............................................................................. 41</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10AFFFA75E1F89951E178B9" blockId="6.[196,1429,1785,1921]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1F89951E178B9" pageId="6" pageNumber="219">– Antennomere XI shorter, approximately as long as the combined length of antennomeres IX–X. Pronotum relatively larger, much broader than head. ..................................................................................................................................... 42</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10AFFFB75E1F848511B7E48" lastPageId="7" lastPageNumber="220" pageId="6" pageNumber="219">
<paragraph id="8BEB36B7F10AFFFA75E1F84851E17728" blockId="6.[196,1429,1944,2064]" pageId="6" pageNumber="219">
<keyLead id="30A59582F10AFFFA75E1F84851E17728" pageId="6" pageNumber="219">
41. Antennae more slender (
<bibRefCitation id="EFC54B46F10AFFFA74D3F84856F57888" author="ASSING, V." box="[502,641,1944,1968]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figures 28–29). Pronotum with very uneven surface; punctation strongly irregular and situated in more or less pronounced impressions (
<bibRefCitation id="EFC54B46F10AFFFA76BEF868505378E8" author="ASSING, V." box="[923,1063,1976,2000]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 65). Male tergite VIII with four blunt projections posteriorly (
<bibRefCitation id="EFC54B46F10AFFFA777FF808569078C8" author="ASSING, V." box="[602,740,2008,2032]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 196). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10AFFFA718AF808513778C8" author="ASSING, V." box="[1199,1347,2008,2032]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 191–194). Widespread (
<bibRefCitation id="EFC54B46F10AFFFA74CCF82856077728" author="ASSING, V." box="[489,627,2040,2064]" pageId="6" pageNumber="219" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: map 6). .................................................................................................. proximus
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10BFFFB75BAFF29511B7E48" blockId="7.[159,1391,248,368]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFF29511B7E48" pageId="7" pageNumber="220">
– Antennae less slender (
<bibRefCitation id="EFC54B46F10BFFFB74F5FF2956177E29" author="ASSING, V." box="[464,611,249,273]" pageId="7" pageNumber="220" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 33). Pronotum without impressions and with less irregularly distributed punctation (
<bibRefCitation id="EFC54B46F10BFFFB74E3FEC956267E09" author="ASSING, V." box="[454,594,281,305]" pageId="7" pageNumber="220" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 69). Male tergite VIII only with a median pair of more acute projections posteriorly (
<bibRefCitation id="EFC54B46F10BFFFB74A9FEE956687E69" author="ASSING, V." box="[396,540,313,337]" pageId="7" pageNumber="220" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
: figure 206). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10BFFFB712FFEE950D07E69" author="ASSING, V." box="[1034,1188,313,337]" pageId="7" pageNumber="220" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017</bibRefCitation>
: figures 208–211).
<collectingCountry id="F3437627F10BFFFB75E0FE88555E7E48" box="[197,298,344,368]" name="Thailand" pageId="7" pageNumber="220">Thailand</collectingCountry>
. .......................................................................................................................................................
<taxonomicName id="4C544D34F10BFFFB71D9FE88511B7E48" authorityName="ASSING" authorityYear="2017" box="[1276,1391,344,368]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B920EAA5F10BFFFB71D9FE88511B7E48" bold="true" box="[1276,1391,344,368]" italics="true" pageId="7" pageNumber="220">parvicollis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyLead id="30A59582F10BFFFB75BAFE58511B7D57" pageId="7" pageNumber="220">
<keyStep id="30A02E12F10BFFFB75BAFE58511B7D57" pageId="7" pageNumber="220">
<paragraph id="8BEB36B7F10BFFFB75BAFE58511B7EC7" blockId="7.[159,1392,391,623]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFE58511B7EC7" pageId="7" pageNumber="220">
42. Body rather large (length of forebody
<quantity id="4CAC9B52F10BFFFB777CFE5856AD7E98" box="[601,729,392,416]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.7" metricValueMin="3.4" pageId="7" pageNumber="220" unit="mm" value="3.55" valueMax="3.7" valueMin="3.4">3.4–3.7 mm</quantity>
) and of robust shape (
<bibRefCitation id="EFC54B46F10BFFFB76EDFE5850277EA7" author="ASSING, V." box="[968,1107,391,416]" pageId="7" pageNumber="220" pagination="1257 - 1282" refId="ref25690" refString="ASSING, V. 2015: On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 47 (2): 1257 - 1282 - http: // www. zobodat. at / pdf / LBB _ 0047 _ 2 _ 1257 - 1282. pdf." type="journal article" year="2015">ASSING 2015</bibRefCitation>
: figure 15). Antennae long (
<quantity id="4CAC9B52F10BFFFB75E9FE78553A7EF8" box="[204,334,424,448]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="7" pageNumber="220" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7–2.9 mm</quantity>
) and massive (
<bibRefCitation id="EFC54B46F10BFFFB74C9FE7856027E87" author="ASSING, V." box="[492,630,423,448]" pageId="7" pageNumber="220" pagination="1257 - 1282" refId="ref25690" refString="ASSING, V. 2015: On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 47 (2): 1257 - 1282 - http: // www. zobodat. at / pdf / LBB _ 0047 _ 2 _ 1257 - 1282. pdf." type="journal article" year="2015">ASSING 2015</bibRefCitation>
: figure 16). Elytra bicoloured with the anterior third reddish-yellow and the posterior two-thirds black (
<bibRefCitation id="EFC54B46F10BFFFB7723FE1856FA7EE7" author="ASSING, V." box="[518,654,455,480]" pageId="7" pageNumber="220" pagination="1257 - 1282" refId="ref25690" refString="ASSING, V. 2015: On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 47 (2): 1257 - 1282 - http: // www. zobodat. at / pdf / LBB _ 0047 _ 2 _ 1257 - 1282. pdf." type="journal article" year="2015">ASSING 2015</bibRefCitation>
: figure 15). Median lobe of aedeagus as in
<bibRefCitation id="EFC54B46F10BFFFB711FFE1850BF7EE7" author="ASSING, V." box="[1082,1227,455,480]" pageId="7" pageNumber="220" pagination="1257 - 1282" refId="ref25690" refString="ASSING, V. 2015: On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 47 (2): 1257 - 1282 - http: // www. zobodat. at / pdf / LBB _ 0047 _ 2 _ 1257 - 1282. pdf." type="journal article" year="2015">ASSING (2015</bibRefCitation>
: figures 33–34). North
<collectingCountry id="F3437627F10BFFFB7429FE37551F7EC7" box="[268,363,487,511]" name="Vietnam" pageId="7" pageNumber="220">Vietnam</collectingCountry>
. ...............................................................................................................................................
<taxonomicName id="4C544D34F10BFFFB71CDFE37511B7EC7" authorityName="ASSING" authorityYear="2015" box="[1256,1391,487,511]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="fansipanicus">
<emphasis id="B920EAA5F10BFFFB71CDFE37511B7EC7" bold="true" box="[1256,1391,487,511]" italics="true" pageId="7" pageNumber="220">fansipanicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10BFFFB75BAFDC7511B7D57" blockId="7.[159,1392,391,623]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFDC7511B7D57" pageId="7" pageNumber="220">– Body less robust and mostly smaller. Antennae less massive, slender in species with antennae of similar length. Elytra often bicoloured, but either more extensively reddish-yellow with infuscate postero-lateral portions or blackish with only the humeral portions reddish or reddish-yellow. ......................................................................... 43</keyLead>
</paragraph>
</keyStep>
</keyLead>
<keyStep id="30A02E12F10BFFFB75BAFD56511B7C55" pageId="7" pageNumber="220">
<paragraph id="8BEB36B7F10BFFFB75BAFD56511B7DC6" blockId="7.[159,1392,646,877]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFD56511B7DC6" pageId="7" pageNumber="220">
43. Species of more slender habitus (
<bibRefCitation id="EFC54B46F10BFFFB7706FD5656CE7DA6" author="ASSING, V." box="[547,698,646,670]" pageId="7" pageNumber="220" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 81). Antennae more slender with antennomere IV oblong and antennomere X less than 1.5 times as broad as long (
<bibRefCitation id="EFC54B46F10BFFFB7600FD7657C87D86" author="ASSING, V." box="[805,956,678,702]" pageId="7" pageNumber="220" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 24). Punctation of pronotum and elytra moderately dense (
<bibRefCitation id="EFC54B46F10BFFFB74F5FD1656127DE6" author="ASSING, V." box="[464,614,710,734]" pageId="7" pageNumber="220" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 81). Median lobe of aedeagus as in
<figureCitation id="136F2A32F10BFFFB7103FD1650ED7DE6" box="[1062,1177,710,734]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="7" pageNumber="220">Figs 78–79</figureCitation>
and
<bibRefCitation id="EFC54B46F10BFFFB71E9FD1651187DE6" author="ASSING, V." box="[1228,1388,710,734]" pageId="7" pageNumber="220" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 221–222).
<collectingCountry id="F3437627F10BFFFB74A6FD3655B37DC6" box="[387,455,742,766]" name="China" pageId="7" pageNumber="220">China</collectingCountry>
,
<collectingCountry id="F3437627F10BFFFB74F4FD36563B7DC6" box="[465,591,742,766]" name="Hong Kong" pageId="7" pageNumber="220">Hong Kong</collectingCountry>
,
<collectingCountry id="F3437627F10BFFFB777DFD3656F97DC5" box="[600,653,742,765]" name="Laos" pageId="7" pageNumber="220">Laos</collectingCountry>
. .................................................................................................
<taxonomicName id="4C544D34F10BFFFB71CBFD36511B7DC6" authorityName="PACE" authorityYear="1998" box="[1262,1391,742,766]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="notaticornis">
<emphasis id="B920EAA5F10BFFFB71CBFD36511B7DC6" bold="true" box="[1262,1391,742,766]" italics="true" pageId="7" pageNumber="220">notaticornis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10BFFFB75BAFCC5511B7C55" blockId="7.[159,1392,646,877]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFCC5511B7C55" pageId="7" pageNumber="220">– Body less slender and with a relatively larger pronotum. Antennae less slender; antennomere X at least 1.5 times as broad as long and antennomere IV usually as long as broad or weakly transverse. Punctation of pronotum and elytra mostly denser. ......................................................................................................................................................... 44</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10BFFFB75BAFC54511B7B33" pageId="7" pageNumber="220">
<paragraph id="8BEB36B7F10BFFFB75BAFC54511B7C84" blockId="7.[159,1392,900,1035]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFC54511B7C84" pageId="7" pageNumber="220">
44. Ventral process of aedeagus very short in relation to basal capsule (
<figureCitation id="136F2A32F10BFFFB76B0FC54507B7CA4" box="[917,1039,900,924]" captionStart="Figs 87–102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87–102: Zyras parvilobatus (87–89), Z. hlavaci (90–95; 90–92: holotype; 93–95: paratype from Selangor), Z. hirtiventris, holotype (96–98), Z. lunatus (99–101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (87–88, 90–91, 93–94, 96–97, 99–100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87–101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="7" pageNumber="220">Figs 87–88</figureCitation>
). Pronotum with maximal width near anterior angles, slightly more strongly tapering posteriorly.
<collectingCountry id="F3437627F10BFFFB764BFC7557D77C84" box="[878,931,933,956]" name="Laos" pageId="7" pageNumber="220">Laos</collectingCountry>
. ..................................................
<taxonomicName id="4C544D34F10BFFFB71C2FC74511B7C84" authorityName="Assing" authorityYear="2017" box="[1255,1391,932,956]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="parvilobatus">
<emphasis id="B920EAA5F10BFFFB71C2FC74511B7C84" bold="true" box="[1255,1391,932,956]" italics="true" pageId="7" pageNumber="220">parvilobatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10BFFFB75BAFC04511B7B33" blockId="7.[159,1392,900,1035]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFC04511B7B33" pageId="7" pageNumber="220">– Aedeagus with distinctly longer ventral process. Pronotum with lateral margins more evenly convex in dorsal view. .............................................................................................................................................................................................. 45</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E12F10BFFFB75BAFBF2511B7B91" pageId="7" pageNumber="220">
<paragraph id="8BEB36B7F10BFFFB75BAFBF2511A7B62" blockId="7.[159,1391,1058,1193]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFBF2511A7B62" pageId="7" pageNumber="220">
45. Median lobe of aedeagus with apically longer and more acute ventral process and with longer crista apicalis in lateral view (
<bibRefCitation id="EFC54B46F10BFFFB7468FB9255977B62" author="ASSING, V." box="[333,483,1090,1114]" pageId="7" pageNumber="220" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figures 215–216).
<collectingCountry id="F3437627F10BFFFB778EFB92569B7B62" box="[683,751,1090,1114]" name="China" pageId="7" pageNumber="220">China</collectingCountry>
:
<collectingRegion id="4990F855F10BFFFB77DCFB93573B7B62" box="[761,847,1091,1114]" country="China" name="Yunnan" pageId="7" pageNumber="220">Yunnan</collectingRegion>
;
<collectingCountry id="F3437627F10BFFFB767CFB9257C87B62" box="[857,956,1090,1114]" name="Thailand" pageId="7" pageNumber="220">Thailand</collectingCountry>
. ..............................................
<taxonomicName id="4C544D34F10BFFFB71CCFB92511A7B62" authorityName="ASSING" authorityYear="2016" box="[1257,1390,1090,1114]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="caloderoides">
<emphasis id="B920EAA5F10BFFFB71CCFB92511A7B62" bold="true" box="[1257,1390,1090,1114]" italics="true" pageId="7" pageNumber="220">caloderoides</emphasis>
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</paragraph>
<paragraph id="8BEB36B7F10BFFFB75BAFBA2511B7B91" blockId="7.[159,1391,1058,1193]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFBA2511B7B91" pageId="7" pageNumber="220">– Median lobe of aedeagus with apically shorter and less acute ventral process and with shorter crista apicalis in lateral view. ......................................................................................................................................................................... 46</keyLead>
</paragraph>
</keyStep>
<keyLead id="30A59582F10BFFFB75BAFB10511B7A50" pageId="7" pageNumber="220">
<keyStep id="30A02E12F10BFFFB75BAFB10511B7A50" pageId="7" pageNumber="220">
<paragraph id="8BEB36B7F10BFFFB75BAFB10511B7A20" blockId="7.[159,1392,1216,1384]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFB10511B7A20" pageId="7" pageNumber="220">
46. Ventral process of aedeagus basally broader in ventral view (
<figureCitation id="136F2A32F10BFFFB760FFB1057D47BE1" box="[810,928,1216,1241]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="7" pageNumber="220">Figs 70–71</figureCitation>
). Elytra practically completely black; anterior margin only very narrowly reddish-yellow (
<figureCitation id="136F2A32F10BFFFB77AAFB3056AF7BC0" box="[655,731,1248,1272]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="7" pageNumber="220">Fig. 25</figureCitation>
). Tergite VIII anteriorly with longitudinal striae (
<figureCitation id="136F2A32F10BFFFB71C5FB3051587BC1" box="[1248,1324,1248,1273]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="7" pageNumber="220">Fig. 39</figureCitation>
). Java. ................................................................................................................................................................................
<taxonomicName id="4C544D34F10BFFFB7024FAD0511B7A20" authorityName="Assing" authorityYear="2017" box="[1281,1391,1280,1304]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="latibasalis">
<emphasis id="B920EAA5F10BFFFB7024FAD0511B7A20" bold="true" box="[1281,1391,1280,1304]" italics="true" pageId="7" pageNumber="220">latibasalis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36B7F10BFFFB75BAFAE0511B7A50" blockId="7.[159,1392,1216,1384]" pageId="7" pageNumber="220">
<keyLead id="30A59582F10BFFFB75BAFAE0511B7A50" pageId="7" pageNumber="220">
– Ventral process of aedeagus more slender in ventral view (
<figureCitation id="136F2A32F10BFFFB760CFAE057E97A70" box="[809,925,1328,1352]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="7" pageNumber="220">Figs 76–77</figureCitation>
and
<bibRefCitation id="EFC54B46F10BFFFB76F4FAE050137A70" author="ASSING, V." box="[977,1127,1328,1352]" pageId="7" pageNumber="220" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figures 193–194). Elytra mostly more extensively reddish-yellow (
<bibRefCitation id="EFC54B46F10BFFFB775DFA80577A7A50" author="ASSING, V." box="[632,782,1360,1384]" pageId="7" pageNumber="220" pagination="87 - 175" refId="ref25746" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 66).
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,
<collectingCountry id="F3437627F10BFFFB76FCFA8050797A5F" box="[985,1037,1360,1383]" name="Laos" pageId="7" pageNumber="220">Laos</collectingCountry>
,
<collectingCountry id="F3437627F10BFFFB7132FA8050027A50" box="[1047,1142,1360,1384]" name="Vietnam" pageId="7" pageNumber="220">Vietnam</collectingCountry>
. ............................... wei
</keyLead>
</paragraph>
</keyStep>
</keyLead>
</key>
</subSubSection>
</treatment>
</document>
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