treatments-xml/data/03/FD/87/03FD878BFFC0FFE1FCBEFDE35A1A3BEA.xml
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<document id="4BCF47AD09975A997084702FD2E4D0C1" ID-DOI="10.11646/zootaxa.5367.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10145106" ID-ZooBank="5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700167363318" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="03FD878BFFC0FFE1FCBEFDE35A1A3BEA" docLanguage="en" docName="zootaxa.5367.1.1.pdf" docOrigin="Zootaxa 5367 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena (Hydraenopsis) concatenata Perkins 2023, new species" docType="treatment" docVersion="2" lastPageNumber="17" masterDocId="FFC4FFF3FFD0FFF1FC29FFA5596F3C37" masterDocTitle="New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)" masterLastPageNumber="86" masterPageNumber="1" pageNumber="17" updateTime="1700168056660" updateUser="ExternalLinkService">
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<mods:title id="28D8AD35AB073B89598FBC5202C04F26">New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="3792F595FB5984ABBA2D92B82822D517">Perkins, Philip D.</mods:namePart>
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<mods:title id="1293E14C39788BD8172648CF88948070">Zootaxa</mods:title>
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<mods:date id="27293699FC6774E1A0DCA93B51DA331B">2023</mods:date>
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<mods:number id="DA86F7A217B24A37DE269BC5AC52C061">2023-11-07</mods:number>
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<mods:identifier id="EA2E62B6D73F88D477EE3F7FFBF6B770" type="DOI">10.11646/zootaxa.5367.1.1</mods:identifier>
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<treatment id="03FD878BFFC0FFE1FCBEFDE35A1A3BEA" LSID="urn:lsid:plazi:treatment:03FD878BFFC0FFE1FCBEFDE35A1A3BEA" httpUri="http://treatment.plazi.org/id/03FD878BFFC0FFE1FCBEFDE35A1A3BEA" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="C34E6516FFC0FFE1FCBEFDE35A663E57" box="[151,777,582,609]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BEB369DFFC0FFE1FCBEFDE35A663E57" blockId="16.[151,777,582,645]" box="[151,777,582,609]" pageId="16" pageNumber="17">
<heading id="D0A381F1FFC0FFE1FCBEFDE35A663E57" bold="true" box="[151,777,582,609]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="4">
<emphasis id="B920EA8FFFC0FFE1FCBEFDE35A663E57" bold="true" box="[151,777,582,609]" pageId="16" pageNumber="17">
<taxonomicName id="4C544D1EFFC0FFE1FCBEFDE35B043E56" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[151,619,582,609]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="concatenata" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B920EA8FFFC0FFE1FCBEFDE35B043E56" bold="true" box="[151,619,582,609]" italics="true" pageId="16" pageNumber="17">Hydraena (Hydraenopsis) concatenata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21357F4FFC0FFE1FE53FDE25A663E57" box="[634,777,582,609]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC0FFE1FCBEFDCE5B933EB2" box="[151,764,619,645]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BEB369DFFC0FFE1FCBEFDCE5B933EB2" blockId="16.[151,777,582,645]" box="[151,764,619,645]" pageId="16" pageNumber="17">
<figureCitation id="136F2A18FFC0FFE1FCBEFDCE598C3EB2" box="[151,227,619,645]" captionStart="FIGURE 3" captionStartId="7.[151,250,1639,1663]" captionTargetBox="[206,1381,181,1615]" captionTargetId="figure-18@7.[206,1381,181,1615]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of Hydraena species, H. leechi Group." figureDoi="http://doi.org/10.5281/zenodo.10145112" httpUri="https://zenodo.org/record/10145112/files/figure.png" pageId="16" pageNumber="17">Figs. 3</figureCitation>
(habitus comparison), 9 (habitus and aedeagus)
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC0FFE1FCBEFD165AB43EC6" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BEB369DFFC0FFE1FCBEFD165AB43EC6" blockId="16.[151,1437,690,2013]" pageId="16" pageNumber="17">
<emphasis id="B920EA8FFFC0FFE1FCBEFD1658233EFB" bold="true" box="[151,332,690,717]" pageId="16" pageNumber="17">Type Material.</emphasis>
<materialsCitation id="3B3C3CC0FFC0FFE1FD7EFD1658173EC6" collectingDate="1981-05-15" collectionCode="NMNH" collectorName="P. J. Spangler" country="Mexico" location="Balzapote" municipality="Los Tuxtlas area" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<emphasis id="B920EA8FFFC0FFE1FD7EFD1658AB3EFA" bold="true" box="[343,452,691,717]" pageId="16" pageNumber="17">
<typeStatus id="54EF883FFFC0FFE1FD7EFD1658AB3EFA" box="[343,452,691,717]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
<collectingCountry id="F343760DFFC0FFE1FE1CFD165BC53EFA" box="[565,682,691,717]" name="Mexico" pageId="16" pageNumber="17">MEXICO</collectingCountry>
:
<collectingRegion id="4990F87FFFC0FFE1FE90FD165A4F3EFA" box="[697,800,691,717]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="16" pageNumber="17">Veracruz</collectingRegion>
,
<collectingMunicipality id="6B8FACE7FFC0FFE1FF07FD165A9F3EFA" box="[814,1008,691,717]" pageId="16" pageNumber="17">Los Tuxtlas area</collectingMunicipality>
nr.
<location id="8E8B6046FFC0FFE1F80AFD165DF93EFA" LSID="urn:lsid:plazi:treatment:03FD878BFFC0FFE1FCBEFDE35A1A3BEA:8E8B6046FFC0FFE1F80AFD165DF93EFA" box="[1059,1174,691,717]" country="Mexico" municipality="Los Tuxtlas area" name="Balzapote" pageId="16" pageNumber="17" stateProvince="Veracruz de Ignacio de la Llave">Balzapote</location>
,
<date id="FFEA105DFFC0FFE1F88CFD175C703EFA" box="[1189,1311,690,717]" pageId="16" pageNumber="17" value="1981-05-15">
<collectingDate id="EFAEE9B5FFC0FFE1F88CFD175C703EFA" box="[1189,1311,690,717]" pageId="16" pageNumber="17" value="1981-05-15">15 v 1981</collectingDate>
</date>
, coll.
<collectorName id="26A1534BFFC0FFE1F945FD1659923EC6" pageId="16" pageNumber="17">P. J. Spangler</collectorName>
” (
<collectionCode id="ED45AE58FFC0FFE1FD3FFD7258023EC6" box="[278,365,727,753]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3C3CC0FFC0FFE1FDADFD725AB93EC6" box="[388,982,727,753]" collectingDate="1981-05-15" collectionCode="NMNH" collectorName="P. J. Spangler" country="Mexico" location="Balzapote" municipality="Los Tuxtlas area" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<emphasis id="B920EA8FFFC0FFE1FDADFD7258943EC6" bold="true" box="[388,507,727,753]" pageId="16" pageNumber="17">
<typeStatus id="54EF883FFFC0FFE1FDADFD72589A3EC6" box="[388,501,727,753]" pageId="16" pageNumber="17" type="paratype">Paratype</typeStatus>
:
</emphasis>
Same data as holotype (
<specimenCount id="9D52FD14FFC0FFE1FF26FD725A013EC6" box="[783,878,727,753]" count="1" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
<collectionCode id="ED45AE58FFC0FFE1FF5CFD725AA43EC6" box="[885,971,727,753]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC0FFE1FCEEFD5F5A51386E" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BEB369DFFC0FFE1FCEEFD5F5A51386E" blockId="16.[151,1437,690,2013]" pageId="16" pageNumber="17">
<emphasis id="B920EA8FFFC0FFE1FCEEFD5F58BC3F23" bold="true" box="[199,467,762,788]" pageId="16" pageNumber="17">Differential Diagnosis.</emphasis>
This new species is a member of the
<taxonomicName id="4C544D1EFFC0FFE1FF57FD5E5A8B3F22" box="[894,996,763,789]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="leechi">
<emphasis id="B920EA8FFFC0FFE1FF57FD5E5A8B3F22" box="[894,996,763,789]" italics="true" pageId="16" pageNumber="17">H. leechi</emphasis>
</taxonomicName>
Group,
<taxonomicName id="4C544D1EFFC0FFE1F868FD5E5C013F22" authority="Subgroup" authorityName="Subgroup" box="[1089,1390,763,789]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="scintillabella">
<emphasis id="B920EA8FFFC0FFE1F868FD5E5D993F22" box="[1089,1270,763,789]" italics="true" pageId="16" pageNumber="17">H. scintillabella</emphasis>
Subgroup
</taxonomicName>
, the group of species that have a pronotal scintilla, and the males lack metatibial brushes (
<bibRefCitation id="EFC54B6CFFC0FFE1F873FCBA5C703F0E" author="Perkins, P. D." box="[1114,1311,799,825]" pageId="16" pageNumber="17" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980: 36</bibRefCitation>
). Members will key to couplet
<quantity id="4CAC9B78FFC0FFE1FD5CFCE658C03F6A" box="[373,431,835,861]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="16" pageNumber="17" unit="in" value="23.0">23 in</quantity>
<bibRefCitation id="EFC54B6CFFC0FFE1FD9EFCE65BE13F6A" author="Perkins, P. D." box="[439,654,835,861]" pageId="16" pageNumber="17" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980: 67)</bibRefCitation>
. This species can be recognized by the combination of the pronotum with a scintilla, the pronotal foveae deep, the metaventral plaques very narrow, very indistinct or absent, and the unusual aedeagus (
<figureCitation id="136F2A18FFC0FFE1FD40FC2E58C23F92" box="[361,429,907,933]" captionStart="FIGURE 9" captionStartId="17.[151,250,1792,1816]" captionTargetBox="[293,1294,181,1768]" captionTargetId="figure-18@17.[293,1294,181,1768]" captionTargetPageId="17" captionText="FIGURE 9. Hydraena (Hydraenopsis) concatenata new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145124" httpUri="https://zenodo.org/record/10145124/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
). The aedeagus appears to be a mix of the characters found in most
<taxonomicName id="4C544D1EFFC0FFE1F8B0FC2E5C213F92" box="[1177,1358,907,933]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B920EA8FFFC0FFE1F8B0FC2E5C213F92" box="[1177,1358,907,933]" italics="true" pageId="16" pageNumber="17">H. marginicollis</emphasis>
</taxonomicName>
Group species (species which lack a pronotal scintilla) such as the paramere shape and paramere setation, but with the robustness of the lateral shape, and the distal shape of the main-piece indicating the
<taxonomicName id="4C544D1EFFC0FFE1F87CFC765DD33FDA" box="[1109,1212,979,1005]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="leechi">
<emphasis id="B920EA8FFFC0FFE1F87CFC765DD33FDA" box="[1109,1212,979,1005]" italics="true" pageId="16" pageNumber="17">H. leechi</emphasis>
</taxonomicName>
Group. Apparently, this species is a link between the two groups, reaffirming the position of the
<taxonomicName id="4C544D1EFFC0FFE1FFCFFC525D243826" box="[998,1099,1015,1041]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="leechi">
<emphasis id="B920EA8FFFC0FFE1FFCFFC525D243826" box="[998,1099,1015,1041]" italics="true" pageId="16" pageNumber="17">H. leechi</emphasis>
</taxonomicName>
Group as a derived branch off of the
<taxonomicName id="4C544D1EFFC0FFE1FCF6FBBE5B6B3802" box="[223,516,1051,1077]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Hydraenopsis">
<emphasis id="B920EA8FFFC0FFE1FCF6FBBE58203802" box="[223,335,1051,1077]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
(
<emphasis id="B920EA8FFFC0FFE1FD76FBBE58943802" box="[351,507,1051,1077]" italics="true" pageId="16" pageNumber="17">Hydraenopsis</emphasis>
)
</taxonomicName>
lineage. See also the diagnosis of
<taxonomicName id="4C544D1EFFC0FFE1FFAFFBBE5D4A3802" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[902,1061,1051,1077]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="multivilosa" status="n. sp.">
<emphasis id="B920EA8FFFC0FFE1FFAFFBBE5D4A3802" box="[902,1061,1051,1077]" italics="true" pageId="16" pageNumber="17">H. multivilosa</emphasis>
</taxonomicName>
<emphasis id="B920EA8FFFC0FFE1F805FBBE5D043802" bold="true" box="[1068,1131,1051,1077]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A21357F4FFC0FFE1F805FBBE5D043802" box="[1068,1131,1051,1077]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, another species indicating the links between the
<taxonomicName id="4C544D1EFFC0FFE1FDA2FB9A589E386E" box="[395,497,1087,1113]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="leechi">
<emphasis id="B920EA8FFFC0FFE1FDA2FB9A589E386E" box="[395,497,1087,1113]" italics="true" pageId="16" pageNumber="17">H. leechi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D1EFFC0FFE1FE0EFB9A5A51386E" authority="Groups." authorityName="Groups." box="[551,830,1087,1113]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B920EA8FFFC0FFE1FE0EFB9A5BB2386E" box="[551,733,1087,1113]" italics="true" pageId="16" pageNumber="17">H. marginicollis</emphasis>
Groups.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC0FFE1FCEEFBC75A373B46" pageId="16" pageNumber="17" type="description">
<paragraph id="8BEB369DFFC0FFE1FCEEFBC7587539F6" blockId="16.[151,1437,690,2013]" pageId="16" pageNumber="17">
<emphasis id="B920EA8FFFC0FFE1FCEEFBC7583E384B" bold="true" box="[199,337,1122,1148]" pageId="16" pageNumber="17">Description</emphasis>
. Size:
<typeStatus id="54EF883FFFC0FFE1FD8DFBC65B69384A" box="[420,518,1123,1149]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.67/0.71; head width 0.40; pronotum 0.43/0.58, PA 0.45, PB 0.49; elytra 1.00/0.71. Habitus as illustrated (
<figureCitation id="136F2A18FFC0FFE1F805FB225D1C3896" box="[1068,1139,1159,1185]" captionStart="FIGURE 9" captionStartId="17.[151,250,1792,1816]" captionTargetBox="[293,1294,181,1768]" captionTargetId="figure-18@17.[293,1294,181,1768]" captionTargetPageId="17" captionText="FIGURE 9. Hydraena (Hydraenopsis) concatenata new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145124" httpUri="https://zenodo.org/record/10145124/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
). Dorsum with head dark brown to black, pronotum with dark brown very diffusely margined disc surrounded by wide brown margins, elytra brown, maxillary palpi missing from specimen. Dorsum of head and pronotum moderately finely moderately densely punctulate, more sparsely punctulate on pronotal disc than on surrounding areas, interstices shining; punctures of frons disc ca. 12xef, interstices ca. 1xpd; punctures of clypeus very fine, lateral areas microreticulate. Punctures of pronotal disc ca. 23xef, interstices on pronotal disc quite variable, ca. 13xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin emarginate over middle ½, with scintilla; antero- and posterolateral pronotal fovea deep, posteromedial foveae moderately deep; disc anteriorly with shallow transverse impression.
</paragraph>
<paragraph id="8BEB369DFFC0FFE1FCEEFA6E599E3A42" blockId="16.[151,1437,690,2013]" pageId="16" pageNumber="17">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Each puncture with fine short recumbent seta. Intervals not raised, width ca. 1xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BEB369DFFC0FFE1FCEEF9DA5A373B46" blockId="16.[151,1437,690,2013]" pageId="16" pageNumber="17">Venter: Mentum sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate in median concavity, surrounding areas smooth, shining. Genae raised, shining, posterior margin slightly raised laterally. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, very indistinct or absent, located at sides of deep median depression. AIS width at straight posterior margin ca. 3x P2. Pro- and meso tibiae slender, straight. Metatibia very slightly arcuate, gradually slightly increasing in width from base to apex. Abdominal apex symmetrical; last tergite with very shallow apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC0FFE1FCEEF8DF5A323BA2" box="[199,861,1914,1941]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BEB369DFFC0FFE1FCEEF8DF5A323BA2" blockId="16.[151,1437,690,2013]" box="[199,861,1914,1941]" pageId="16" pageNumber="17">
<emphasis id="B920EA8FFFC0FFE1FCEEF8DF580E3BA3" bold="true" box="[199,353,1914,1940]" pageId="16" pageNumber="17">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54EF883FFFC0FFE1FEE5F8DE5B933BA2" box="[716,764,1915,1941]" pageId="16" pageNumber="17">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC0FFE1FCEEF83A5A1A3BEA" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BEB369DFFC0FFE1FCEEF83A5A1A3BEA" blockId="16.[151,1437,690,2013]" pageId="16" pageNumber="17">
<emphasis id="B920EA8FFFC0FFE1FCEEF83A58213B8E" bold="true" box="[199,334,1951,1977]" pageId="16" pageNumber="17">Etymology.</emphasis>
Named in reference to the aedeagal morphology, which appears to represent a linking form between the
<taxonomicName id="4C544D1EFFC0FFE1FCE9F86658493BEA" box="[192,294,1987,2013]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="leechi">
<emphasis id="B920EA8FFFC0FFE1FCE9F86658493BEA" box="[192,294,1987,2013]" italics="true" pageId="16" pageNumber="17">H. leechi</emphasis>
</taxonomicName>
Group and the
<taxonomicName id="4C544D1EFFC0FFE1FDFDF8665BE53BEA" box="[468,650,1987,2013]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B920EA8FFFC0FFE1FDFDF8665BE53BEA" box="[468,650,1987,2013]" italics="true" pageId="16" pageNumber="17">H. marginicollis</emphasis>
</taxonomicName>
Group of
<taxonomicName id="4C544D1EFFC0FFE1FED7F8665A013BEA" box="[766,878,1987,2013]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B920EA8FFFC0FFE1FED7F8665A013BEA" box="[766,878,1987,2013]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>