180 lines
20 KiB
XML
180 lines
20 KiB
XML
<document id="8D13F4BB4FE1BA794577ACFE610DE2E8" ID-DOI="10.5252/cryptogamie-bryologie2021v42a2" ID-ISSN="1776-0992" ID-Zenodo-Dep="12215282" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719005151522" checkinUser="felipe" docAuthor="Thouvenot, Louis" docDate="2021" docId="03F1EA67FFA1FFC2E3CA27ECFBA10AE3" docLanguage="en" docName="CryptogamieBryologie.42.2.11-18.pdf" docOrigin="Cryptogamie, Bryologie 20 (2)" docSource="http://dx.doi.org/10.5252/cryptogamie-bryologie2021v42a2" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Schistochila integerrima Steph." docType="treatment" docVersion="2" lastPageNumber="13" masterDocId="FFC8921FFFA2FFC6E00C235FFFB00C0F" masterDocTitle="Schistochila gradsteinii sp. nov., a new species from New Caledonia related to S. vitreocincta (Schistochilaceae, Marchantiophyta), with a key to the local species and a description of the gynoecium of S. integerrima" masterLastPageNumber="18" masterPageNumber="11" pageNumber="12" updateTime="1719018869330" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="DC1A0E68125F5BFF9B3E62A48C32ECB0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5AD525CF4CBDEA570FF6C0B444BF6953">
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<mods:title id="CAFF205DD03082D3273630FD0480549D">Schistochila gradsteinii sp. nov., a new species from New Caledonia related to S. vitreocincta (Schistochilaceae, Marchantiophyta), with a key to the local species and a description of the gynoecium of S. integerrima</mods:title>
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</mods:titleInfo>
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<mods:name id="7429ACF37B65473992ACB132E4D97D11" type="personal">
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<mods:role id="D91045D57A0BABBB1486DF424077F41E">
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<mods:roleTerm id="6CBC9974A63118B312BD16C1C0D30719">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="09ACF54DA5E775814E96654358081204">Thouvenot, Louis</mods:namePart>
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<mods:affiliation id="4172AD6E6DECC3198BE72EDD59F9E243">Saint Léon, F- 66000 Perpignan (France) thouvenot. louis @ orange. fr</mods:affiliation>
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<mods:nameIdentifier id="3777826A6870663113FE972FA1DA2CDB" type="email">louis@orange.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="9DDFB6C89D202215E82626A67386D0DB">text</mods:typeOfResource>
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<mods:titleInfo id="B7C95FE22B06041FA99D2444D8AE4C59">
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<mods:title id="1023855D33E11485FBF0FE87787A78D9">Cryptogamie, Bryologie</mods:title>
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</mods:titleInfo>
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<mods:part id="49A9CCFE1F21A3BD1FC3B442E97EDC18">
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<mods:date id="0876BC268256A2C7056012087B94FEBA">2021</mods:date>
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<mods:detail id="75359B52C6733B9E80D37FDA6FACE033" type="pubDate">
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<mods:number id="A02905EA0948F8A4C117C378775DCBEE">2021-02-12</mods:number>
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</mods:detail>
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<mods:detail id="96EA16DCEB5E1287808371387C46FD6C" type="volume">
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<mods:number id="71F7944A9FB6190214C4084E9AFBF368">20</mods:number>
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</mods:detail>
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<mods:detail id="539BBBDA01279C103CF7AAAE4F41972B" type="issue">
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<mods:number id="4DCD385F676ED4610CAE7BEEE9CF0A95">2</mods:number>
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</mods:detail>
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<mods:start id="923DD92615E7AB322CF2ECAAFFB820DB">11</mods:start>
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<mods:end id="0A6E67758F50F3EF6AA98FD32D6F6D91">18</mods:end>
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<mods:url id="D22D6D40A928D806ECE0771E25C27D47">http://dx.doi.org/10.5252/cryptogamie-bryologie2021v42a2</mods:url>
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</mods:location>
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<mods:classification id="95CC5F4470497D4147FBC4B2A448220C">journal article</mods:classification>
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<mods:identifier id="91607FCA8750BD8F9398A0CC63165626" type="DOI">10.5252/cryptogamie-bryologie2021v42a2</mods:identifier>
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<mods:identifier id="ADA7BAD034F73719C61102C84F438B0B" type="ISSN">1776-0992</mods:identifier>
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<mods:identifier id="86823AC82B7C2A9B150AB96CC1BB756B" type="Zenodo-Dep">12215282</mods:identifier>
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</mods:mods>
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<treatment id="03F1EA67FFA1FFC2E3CA27ECFBA10AE3" LSID="urn:lsid:plazi:treatment:03F1EA67FFA1FFC2E3CA27ECFBA10AE3" httpUri="http://treatment.plazi.org/id/03F1EA67FFA1FFC2E3CA27ECFBA10AE3" lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12">
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<subSubSection id="C34208FAFFA1FFC5E3CA27ECFB29090A" pageId="3" pageNumber="12" type="nomenclature">
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<paragraph id="8BE75B71FFA1FFC5E3CA27ECFAA708C1" blockId="3.[966,1303,1203,1230]" box="[966,1303,1203,1230]" pageId="3" pageNumber="12">
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<heading id="D0AFEC1DFFA1FFC5E3CA27ECFAA708C1" box="[966,1303,1203,1230]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="12" reason="4">
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<taxonomicName id="4C5820F2FFA1FFC5E3CA27ECFAA708C1" ID-CoL="4V974" authority="Steph." authorityName="Steph." box="[966,1303,1203,1230]" class="Jungermanniopsida" family="Schistochilaceae" genus="Schistochila" kingdom="Plantae" order="Jungermanniales" pageId="3" pageNumber="12" phylum="Marchantiophyta" rank="species" species="integerrima">
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<emphasis id="B92C8763FFA1FFC5E3CA27ECFB7B08C1" bold="true" box="[966,1227,1203,1230]" italics="true" pageId="3" pageNumber="12">Schistochila integerrima</emphasis>
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Steph.
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE75B71FFA1FFC5E34F27B2FB29090A" blockId="3.[835,1177,1261,1285]" box="[835,1177,1261,1285]" pageId="3" pageNumber="12">
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<bibCitation id="5E72531DFFA1FFC5E34F27B2FB25090A" box="[835,1173,1261,1285]" journal="Species Hepaticarum" page="492" pageId="3" pageNumber="12" score="0.0033280253576670055" volume="6" year="1924">
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<emphasis id="B92C8763FFA1FFC5E34F27B2FBB4090B" box="[835,1028,1261,1284]" italics="true" pageId="3" pageNumber="12">Species Hepaticarum</emphasis>
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6: 492 (1924)
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</bibCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34208FAFFA1FFC5E3212643FA1F0966" pageId="3" pageNumber="12" type="description">
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<paragraph id="8BE75B71FFA1FFC5E3212643FA1F0966" blockId="3.[813,1456,1308,1385]" pageId="3" pageNumber="12">
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<smallCapsWord id="8D01CDADFFA1FFC5E3212643FC3E093C" baselines="1326,1327" box="[813,910,1308,1332]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Specimen" pageId="3" pageNumber="12">SPECIMEN</smallCapsWord>
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<smallCapsWord id="8D01CDADFFA1FFC5E399267DFC49093C" baselines="1327" box="[917,1017,1314,1331]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="examined" pageId="3" pageNumber="12">EXAMINED</smallCapsWord>
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. —
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<emphasis id="B92C8763FFA1FFC5E4262643FB750939" bold="true" box="[1066,1221,1308,1334]" pageId="3" pageNumber="12">
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<collectingCountry id="F34F1BE1FFA1FFC5E4262643FB750939" box="[1066,1221,1308,1334]" name="New Caledonia" pageId="3" pageNumber="12">New Caledonia</collectingCountry>
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</emphasis>
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.
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<collectingRegion id="499C9593FFA1FFC5E4DF2643FADF093B" box="[1235,1391,1308,1332]" country="New Caledonia" name="Province Nord" pageId="3" pageNumber="12">North Province</collectingRegion>
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, Poindimié, Amoa valley, trail to Goro Até Mèkébo, on trunks in cloud forest,
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<quantity id="4CA0F694FFA1FFC5E362260EFC000966" box="[878,944,1361,1385]" metricMagnitude="2" metricUnit="m" metricValue="7.24" pageId="3" pageNumber="12" unit="m" value="724.0">724 m</quantity>
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,
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<date id="FFE67DB1FFA1FFC5E3B6260EFB950966" box="[954,1061,1361,1385]" pageId="3" pageNumber="12" value="2019-10-12">12.X.2019</date>
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,
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<emphasis id="B92C8763FFA1FFC5E421260EFB5A0966" box="[1069,1258,1361,1385]" italics="true" pageId="3" pageNumber="12">Thouvenot NC2825</emphasis>
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(PC[PC0712101]).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34208FAFFA1FFC5E32126CCFC400BE4" pageId="3" pageNumber="12" type="discussion">
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<paragraph id="8BE75B71FFA1FFC5E32126CCFC1C09A3" blockId="3.[813,940,1427,1453]" box="[813,940,1427,1453]" pageId="3" pageNumber="12">
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<heading id="D0AFEC1DFFA1FFC5E32126CCFC1C09A3" box="[813,940,1427,1453]" fontSize="9" level="3" pageId="3" pageNumber="12" reason="5">
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<smallCapsWord id="8D01CDADFFA1FFC5E32126CCFC1C09A3" baselines="1446,1447" box="[813,940,1427,1453]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comments" pageId="3" pageNumber="12">COMMENTS</smallCapsWord>
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</heading>
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</paragraph>
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<paragraph id="8BE75B71FFA1FFC5E32126ECFAF30B44" blockId="3.[810,1457,1459,2027]" pageId="3" pageNumber="12">
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Among the
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<taxonomicName id="4C5820F2FFA1FFC5E3A426ECFBC009C2" box="[936,1136,1459,1485]" pageId="3" pageNumber="12">
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<emphasis id="B92C8763FFA1FFC5E3A426ECFB9009C2" box="[936,1056,1459,1485]" italics="true" pageId="3" pageNumber="12">Schistochila</emphasis>
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species
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</taxonomicName>
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lacking underleaves (formerly regarded as
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<taxonomicName id="4C5820F2FFA1FFC5E3A0268CFB0F09E2" authority="Nees ex Mont." authorityName="Mont." box="[940,1215,1491,1517]" class="Jungermanniopsida" family="Schistochilaceae" genus="Gottschea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Jungermanniales" pageId="3" pageNumber="12" phylum="Marchantiophyta" rank="genus">
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<emphasis id="B92C8763FFA1FFC5E3A0268CFBA009E2" box="[940,1040,1491,1517]" italics="true" pageId="3" pageNumber="12">Gottschea</emphasis>
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Nees ex Mont.
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</taxonomicName>
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) and with entire margined leaves,
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<taxonomicName id="4C5820F2FFA1FFC5E3B726ACFBE00A03" box="[955,1104,1523,1549]" class="Jungermanniopsida" family="Schistochilaceae" genus="Schistochila" kingdom="Plantae" order="Jungermanniales" pageId="3" pageNumber="12" phylum="Marchantiophyta" rank="species" species="integerrima">
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<emphasis id="B92C8763FFA1FFC5E3B726ACFBE00A03" box="[955,1104,1523,1549]" italics="true" pageId="3" pageNumber="12">S. integerrima</emphasis>
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</taxonomicName>
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is easily distinguished from species with subequal lobes and inflated keels (see below) by the insertion of the dorsal lobes on the dorsal face of the ventral ones. On the other hand, it is less easy to distinguish from
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<taxonomicName id="4C5820F2FFA1FFC5E321252CFBAF0A83" authority="(Mont.) Lindb." authorityName="Lindb." baseAuthorityName="Mont." box="[813,1055,1650,1677]" class="Jungermanniopsida" family="Schistochilaceae" genus="Schistochila" kingdom="Plantae" order="Jungermanniales" pageId="3" pageNumber="12" phylum="Marchantiophyta" rank="species" species="neesii">
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<emphasis id="B92C8763FFA1FFC5E321252CFCCC0A83" box="[813,892,1651,1677]" italics="true" pageId="3" pageNumber="12">S. neesii</emphasis>
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(Mont.) Lindb.
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</taxonomicName>
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(Mascaregnes) or
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<taxonomicName id="4C5820F2FFA1FFC5E4D8252CFADF0A83" authority="Horik." authorityName="Horik." box="[1236,1391,1650,1677]" class="Jungermanniopsida" family="Schistochilaceae" genus="Schistochila" kingdom="Plantae" order="Jungermanniales" pageId="3" pageNumber="12" phylum="Marchantiophyta" rank="species" species="nuda">
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<emphasis id="B92C8763FFA1FFC5E4D8252CFA900A83" box="[1236,1312,1650,1677]" italics="true" pageId="3" pageNumber="12">S. nuda</emphasis>
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Horik.
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</taxonomicName>
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(Asia) from which it differs only by larger leaves (leaves
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<quantity id="4CA0F694FFA1FFC5E55E25CDFA1D0AA2" box="[1362,1453,1682,1709]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="12" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5-6 mm</quantity>
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long vs
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<quantity id="4CA0F694FFA1FFC5E38825EDFC6E0AC2" box="[900,990,1714,1741]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="12" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
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long) and the total absence of teeth at the leaf apex which are unevenly present in the former species. As hypothesized by
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<bibRefCitation id="EFC92680FFA1FFC5E41C25ADFB3E0B03" author="SO M. L." box="[1040,1166,1778,1804]" pageId="3" pageNumber="12" pagination="255 - 275" refId="ref4787" refString="SO M. L. 2003 b. - Schistochila (Hepaticae) in Oceania. New Zealand Journal of Botany 41: 255 - 275. https: // doi. org / 10.1080 / 0 028825 X. 2003.9512845" type="journal article" year="2003">So (2003b)</bibRefCitation>
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, further studies involving molecular methods and more material would likely show these species and
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<taxonomicName id="4C5820F2FFA1FFC5E3E7246DFBCF0B44" box="[1003,1151,1842,1868]" class="Jungermanniopsida" family="Schistochilaceae" genus="Schistochila" kingdom="Plantae" order="Jungermanniales" pageId="3" pageNumber="12" phylum="Marchantiophyta" rank="species" species="integerrima">
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<emphasis id="B92C8763FFA1FFC5E3E7246DFBCF0B44" box="[1003,1151,1842,1868]" italics="true" pageId="3" pageNumber="12">S. integerrima</emphasis>
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</taxonomicName>
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to be conspecific.
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</paragraph>
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<paragraph id="8BE75B71FFA1FFC5E34F240DFC400BE4" blockId="3.[810,1457,1459,2027]" pageId="3" pageNumber="12">
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In the diagnosis, Stephani does not describe the gametangia of
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<taxonomicName id="4C5820F2FFA1FFC5E37E242DFBB70B84" box="[882,1031,1906,1932]" class="Jungermanniopsida" family="Schistochilaceae" genus="Schistochila" kingdom="Plantae" order="Jungermanniales" pageId="3" pageNumber="12" phylum="Marchantiophyta" rank="species" species="integerrima">
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<emphasis id="B92C8763FFA1FFC5E37E242DFBB70B84" box="[882,1031,1906,1932]" italics="true" pageId="3" pageNumber="12">S. integerrima</emphasis>
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</taxonomicName>
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and
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<bibRefCitation id="EFC92680FFA1FFC5E431242EFB080B84" author="SO M. L." box="[1085,1208,1905,1931]" pageId="3" pageNumber="12" pagination="255 - 275" refId="ref4787" refString="SO M. L. 2003 b. - Schistochila (Hepaticae) in Oceania. New Zealand Journal of Botany 41: 255 - 275. https: // doi. org / 10.1080 / 0 028825 X. 2003.9512845" type="journal article" year="2003">So (2003b)</bibRefCitation>
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states that the plant is known only in sterile condition. Here we provide a description of fertile shoots with unfertilized gynoecia. Androecia remain unknown.
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</paragraph>
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</subSubSection>
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<caption id="DF270BF9FFA6FFC2E088254FFB430A37" ID-DOI="http://doi.org/10.5281/zenodo.12215285" ID-Zenodo-Dep="12215285" httpUri="https://zenodo.org/record/12215285/files/figure.png" pageId="4" pageNumber="13" startId="4.[132,143,1552,1570]" targetBox="[132,1455,216,1508]" targetPageId="4" targetType="figure">
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<paragraph id="8BE75B71FFA6FFC2E088254FFB430A37" blockId="4.[132,1456,1552,1592]" pageId="4" pageNumber="13">
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<smallCapsWord id="8D01CDADFFA6FFC2E088254FFF2D0A2E" baselines="1565,1566" box="[132,157,1552,1570]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="4" pageNumber="13">FIG</smallCapsWord>
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. 1. —
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<taxonomicName id="4C5820F2FFA6FFC2E0D8254FFE6B0A2D" authority="Steph." authorityName="Steph." box="[212,475,1552,1570]" class="Jungermanniopsida" family="Schistochilaceae" genus="Schistochila" kingdom="Plantae" order="Jungermanniales" pageId="4" pageNumber="13" phylum="Marchantiophyta" rank="species" species="integerrima">
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<emphasis id="B92C8763FFA6FFC2E0D8254FFE2B0A2D" box="[212,411,1552,1570]" italics="true" pageId="4" pageNumber="13">Schistochila integerrima</emphasis>
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Steph.
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</taxonomicName>
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:
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<emphasis id="B92C8763FFA6FFC2E1ED254FFE5E0A2D" bold="true" box="[481,494,1552,1570]" pageId="4" pageNumber="13">A</emphasis>
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, cluster of archegonia surrounded by scales, bract involucre and bracts (longitudinal section);
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<emphasis id="B92C8763FFA6FFC2E4F0254FFAB90A2D" bold="true" box="[1276,1289,1552,1570]" pageId="4" pageNumber="13">B</emphasis>
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,
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<emphasis id="B92C8763FFA6FFC2E518254FFA920A2D" bold="true" box="[1300,1314,1552,1570]" pageId="4" pageNumber="13">C</emphasis>
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, bracts;
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<emphasis id="B92C8763FFA6FFC2E566254FFAC80A2D" bold="true" box="[1386,1400,1552,1570]" pageId="4" pageNumber="13">D</emphasis>
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, bracteole;
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<emphasis id="B92C8763FFA6FFC2E0BB2579FF730A37" bold="true" box="[183,195,1574,1592]" pageId="4" pageNumber="13">E</emphasis>
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, top of fertile shoot with gynoecium and fertile innovation;
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<emphasis id="B92C8763FFA6FFC2E2A02579FD070A37" bold="true" box="[684,695,1574,1592]" pageId="4" pageNumber="13">F</emphasis>
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, gynoecium (top view). From
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<emphasis id="B92C8763FFA6FFC2E3A22579FBFF0A37" box="[942,1103,1574,1592]" italics="true" pageId="4" pageNumber="13">Thouvenot NC2825</emphasis>
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. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<subSubSection id="C34208FAFFA6FFC2E088252CFBA10AE3" pageId="4" pageNumber="13" type="description">
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<paragraph id="8BE75B71FFA6FFC2E088252CFDE90A83" blockId="4.[132,601,1650,1677]" box="[132,601,1650,1677]" pageId="4" pageNumber="13">
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<heading id="D0AFEC1DFFA6FFC2E088252CFDE90A83" box="[132,601,1650,1677]" fontSize="9" level="3" pageId="4" pageNumber="13" reason="5">
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<smallCapsWord id="8D01CDADFFA6FFC2E088252CFEA50A84" baselines="1670,1670" box="[132,277,1651,1677]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="4" pageNumber="13">DESCRIPTION</smallCapsWord>
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<smallCapsWord id="8D01CDADFFA6FFC2E1112527FE890A84" baselines="1670" box="[285,313,1656,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="4" pageNumber="13">OF</smallCapsWord>
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<smallCapsWord id="8D01CDADFFA6FFC2E14C2527FEFD0A84" baselines="1670" box="[320,333,1656,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="a" pageId="4" pageNumber="13">A</smallCapsWord>
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<smallCapsWord id="8D01CDADFFA6FFC2E1582527FE140A84" baselines="1670" box="[340,420,1656,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="female" pageId="4" pageNumber="13">FEMALE</smallCapsWord>
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<smallCapsWord id="8D01CDADFFA6FFC2E1A62527FDB10A84" baselines="1670" box="[426,513,1656,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="branch" pageId="4" pageNumber="13">BRANCH</smallCapsWord>
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(
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<figureCitation id="136347F4FFA6FFC2E21E252DFDE00A83" box="[530,592,1650,1676]" captionStart="FIG" captionStartId="4.[132,143,1552,1570]" captionTargetBox="[132,1455,216,1508]" captionTargetId="figure-206@4.[698,1267,769,1325]" captionTargetPageId="4" captionText="FIG. 1. — Schistochila integerrima Steph.: A, cluster of archegonia surrounded by scales, bract involucre and bracts (longitudinal section); B, C, bracts; D, bracteole; E, top of fertile shoot with gynoecium and fertile innovation; F, gynoecium (top view). From Thouvenot NC2825. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12215285" httpUri="https://zenodo.org/record/12215285/files/figure.png" pageId="4" pageNumber="13">Fig. 1</figureCitation>
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)
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</heading>
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</paragraph>
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<paragraph id="8BE75B71FFA6FFC2E08825CDFCB70BE4" blockId="4.[130,777,1682,2028]" pageId="4" pageNumber="13">
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A series of underleaves present on fertile shoots just below the gynoecia, the underleaves all deeply and unevenly bilobate with acute lobes, the outermost underleaves divided half-way to the base with margins indistinctly toothed, the innermost underleaves more deeply divided, the lobes tube-like and laciniate. Fertile innovations frequent so that gynoecia are usually in several levels situated one above the other, separated by a few normal leaves with underleaves present or absent; leaves just below the gynoecium progressively modified into bracts, the outermost ciliate only at the base, the innermost leaves
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<quantity id="4CA0F694FFA6FFC2E0C7248EFEE00BE3" box="[203,336,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="4" pageNumber="13" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5-4.5 mm</quantity>
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long, heavily laciniate, connate at base,
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</paragraph>
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<paragraph id="8BE75B71FFA6FFC2E322252DFBA10AE3" blockId="4.[811,1457,1650,1772]" pageId="4" pageNumber="13">linked together in a fringed involucre surrounding linear to furcate scales and many archegonia, ventral lobes canaliculate to involute with margins undulate and apices rounded, dorsal lobes linear, involute.</paragraph>
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</subSubSection>
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</treatment>
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</document> |