treatments-xml/data/03/DB/C8/03DBC80ABB63FFE0C6EA0A0DFEB1FE6A.xml
2024-06-21 12:22:17 +02:00

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<mods:title id="AE092E0D593BBFAD974CF7D68A2BBD92">Taxonomic revision of the genus Cabassous McMurtrie, 1831 (Cingulata Chlamyphoridae), with revalidation of Cabassous squamicaudis (Lund, 1845)</mods:title>
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<mods:affiliation id="3F5893C4257DF1715149312B8645750F">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, 10001, China. andefeijo @ gmail. com; https: // orcid. org / 0000 - 0002 - 4643 - 2293;</mods:affiliation>
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<mods:affiliation id="2702D1FE33A293D6732C72BE41651110">Coordenação de Ciências Biológicas, Universidade do Estado de Mato Grosso, Caixa Postal 08, 78690 - 000, Nova Xavantina, Mato Grosso, Brazil.</mods:affiliation>
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<paragraph id="8BCD791CBB63FFE3C6EA0A0DFDEAFC27" blockId="13.[151,584,895,958]" box="[151,584,895,922]" pageId="13" pageNumber="60">
<heading id="D085CE70BB63FFE3C6EA0A0DFDEAFC27" bold="true" box="[151,584,895,922]" fontSize="11" level="1" pageId="13" pageNumber="60" reason="1">
<taxonomicName id="4C72029FBB63FFE3C6EA0A0DFDEAFC27" ID-CoL="P43M" authority="Wetzel, 1980" authorityName="Wetzel" authorityYear="1980" box="[151,584,895,922]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B906A50EBB63FFE3C6EA0A0DFDEAFC27" bold="true" box="[151,584,895,922]" pageId="13" pageNumber="60">
<emphasis id="B906A50EBB63FFE3C6EA0A0DFE01FC27" bold="true" box="[151,419,895,922]" italics="true" pageId="13" pageNumber="60">Cabassous chacoensis</emphasis>
<bibRefCitation id="EFE304EDBB63FFE3C7D70AF2FDEAFC27" author="Wetzel, R. M." box="[426,584,896,922]" pageId="13" pageNumber="60" pagination="323 - 357" refId="ref20214" refString="Wetzel, R. M. (1980) Revision of the naked-tailed armadillos, genus Cabassous McMurtrie. Annals of Carnegie Museum, 49, 323 - 357." type="journal article" year="1980">Wetzel, 1980</bibRefCitation>
</emphasis>
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<paragraph id="8BCD791CBB63FFE3C6EA0AD6FE5FFC03" blockId="13.[151,584,895,958]" box="[151,509,932,958]" pageId="13" pageNumber="60">Chacoan naked-tailed armadillo</paragraph>
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<subSubSection id="C3682A97BB63FFE3C6EA0A96FDBAFAD0" pageId="13" pageNumber="60" type="reference_group">
<paragraph id="8BCD791CBB63FFE3C6EA0A96FC08FC46" blockId="13.[151,1436,995,1172]" box="[151,938,995,1019]" pageId="13" pageNumber="60">
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<treatmentCitation id="0AD35F0DBB63FFE3C6EA0A96FDB9FC46" author="Lahille, F." box="[151,539,995,1019]" page="204" pageId="13" pageNumber="60" year="1899">
<taxonomicName id="4C72029FBB63FFE3C6EA0A96FDB9FC46" authority=": Lahille, 1899: 204" authorityName="Lahille" authorityPageNumber="204" authorityYear="1899" box="[151,539,995,1019]" class="Mammalia" family="Dasypodidae" genus="Xenurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="gymnurus">
<emphasis id="B906A50EBB63FFE3C6EA0A96FEF4FC46" box="[151,342,996,1019]" italics="true" pageId="13" pageNumber="60">Xenurus gymnurus</emphasis>
:
<bibRefCitation id="EFE304EDBB63FFE3C71E0A91FDB9FC46" author="Lahille, F." box="[355,539,995,1019]" pageId="13" pageNumber="60" pagination="165 - 206" refId="ref16642" refString="Lahille, F. (1899) Ensayo sobre la distribucion geografica de los mamiferos en la Republica Argentina. Primera reunion del Congreso Cientifico Latino Americo [1898], Buenos Aires, 3, 165 - 206." type="journal article" year="1899">Lahille, 1899:204</bibRefCitation>
</taxonomicName>
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. Not
<treatmentCitation id="0AD35F0DBB63FFE3C42D0A96FC04FC46" author="Olfers, I. von" box="[592,934,995,1019]" page="220" pageId="13" pageNumber="60" year="1818">
<taxonomicName id="4C72029FBB63FFE3C42D0A96FC04FC46" authority="Olfers 1818: 220" authorityName="Olfers" authorityPageNumber="220" authorityYear="1818" box="[592,934,995,1019]" class="Mammalia" family="Dasypodidae" genus="Tatus" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="gymnurus">
<emphasis id="B906A50EBB63FFE3C42D0A96FD53FC46" box="[592,753,996,1019]" italics="true" pageId="13" pageNumber="60">Tatus gymnurus</emphasis>
<bibRefCitation id="EFE304EDBB63FFE3C48A0A91FC04FC46" author="Olfers, I. von" box="[759,934,995,1019]" pageId="13" pageNumber="60" pagination="192 - 237" refId="ref17656" refString="Olfers, I. von (1818) Bemerkungen zu Illiger's Ueberblick der Saugthiere, nach ihrer Vertheilung uber die Welttheile, rucksichtlich der Sudamericanischen Arten (Species). In: Eschwege, W. L. (Ed.), Journal von Brasilien oder vermischte nachrichten aus Brasilien, auf wisenschaftlichen Reisen gesammelt. Verlage des Gr. H. S. Landes-Industrie-Comptoirs, Weimar, pp. 192 - 237." type="book chapter" year="1818">Olfers 1818: 220</bibRefCitation>
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</paragraph>
<paragraph id="8BCD791CBB63FFE3C6EA0D70FB20FBA7" blockId="13.[151,1436,995,1172]" box="[151,1154,1026,1050]" pageId="13" pageNumber="60">
<treatmentCitationGroup id="AB625E32BB63FFE3C6EA0D70FB20FBA7" box="[151,1154,1026,1050]" pageId="13" pageNumber="60">
<treatmentCitation id="0AD35F0DBB63FFE3C6EA0D70FDBCFBA7" author="Yepes, J." box="[151,542,1026,1050]" page="441" pageId="13" pageNumber="60" year="1935">
<taxonomicName id="4C72029FBB63FFE3C6EA0D70FDBCFBA7" authority=": Yepes, 1935: 441" authorityName="Yepes" authorityPageNumber="441" authorityYear="1935" box="[151,542,1026,1050]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="loricatus">
<emphasis id="B906A50EBB63FFE3C6EA0D70FEC7FBA7" box="[151,357,1026,1050]" italics="true" pageId="13" pageNumber="60">Cabassous loricatus</emphasis>
:
<bibRefCitation id="EFE304EDBB63FFE3C70F0D70FDBCFBA7" author="Yepes, J." box="[370,542,1026,1050]" pageId="13" pageNumber="60" pagination="438 - 444" refId="ref20614" refString="Yepes, J. (1935) Las especies argentinas del genero &quot; Cabassous &quot; (Dasypodidae). Physis, 11, 438 - 444." type="journal article" year="1935">Yepes, 1935:441</bibRefCitation>
</taxonomicName>
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. Part; not
<treatmentCitation id="0AD35F0DBB63FFE3C4F80D70FBDCFBA7" author="Wagner, J. A." box="[645,1150,1026,1050]" page="176" pageId="13" pageNumber="60" year="1855">
<taxonomicName id="4C72029FBB63FFE3C4F80D70FBDCFBA7" authority="J. A. Wagner, 1855: 174 - 176" authorityName="J. A. Wagner" authorityPageNumber="174 - 176" authorityYear="1855" box="[645,1150,1026,1050]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="loricatus">
<emphasis id="B906A50EBB63FFE3C4F80D70FCF1FBA7" box="[645,851,1026,1050]" italics="true" pageId="13" pageNumber="60">Cabassous loricatus</emphasis>
J. A.
<bibRefCitation id="EFE304EDBB63FFE3C5F00D70FBDCFBA7" author="Wagner, J. A." box="[909,1150,1026,1050]" pageId="13" pageNumber="60" refId="ref20141" refString="Wagner, J. A. (1855) Die Saugthiere in Abbildungen nach der Natur mit Beschreibungen von Dr. Johann Christian Daniel von Schreber. Supplementband. Funfte Abtheilung: Die Affen, Zahnlucker, Beutelthiere, Hufthiere, Insektenfresser und Handflugler. Supplementband 5. T. O. Weigel, Leipzig, 810 pp. https: // doi. org / 10.5962 / bhl. title. 67399" type="book" year="1855">Wagner, 1855: 174-176</bibRefCitation>
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</paragraph>
<paragraph id="8BCD791CBB63FFE3C6EA0D52FB8FFB85" blockId="13.[151,1436,995,1172]" box="[151,1069,1056,1081]" pageId="13" pageNumber="60">
<treatmentCitationGroup id="AB625E32BB63FFE3C6EA0D52FB8FFB85" box="[151,1069,1056,1081]" pageId="13" pageNumber="60">
<treatmentCitation id="0AD35F0DBB63FFE3C6EA0D52FD91FB85" author="Cabrera, A." box="[151,563,1056,1080]" page="219" pageId="13" pageNumber="60" year="1958">
<taxonomicName id="4C72029FBB63FFE3C6EA0D52FD91FB85" authority=": Cabrera, 1958: 219" authorityName="Cabrera" authorityPageNumber="219" authorityYear="1958" box="[151,563,1056,1080]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="loricatus">
<emphasis id="B906A50EBB63FFE3C6EA0D52FEC7FB85" box="[151,357,1056,1080]" italics="true" pageId="13" pageNumber="60">Cabassous loricatus</emphasis>
:
<bibRefCitation id="EFE304EDBB63FFE3C70E0D52FD91FB85" author="Cabrera, A." box="[371,563,1056,1080]" pageId="13" pageNumber="60" pagination="1 - 307" refId="ref14025" refString="Cabrera, A. (1958) Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot;, Ciencias Zoologicas, 4 (1), 1 - 307." type="journal article" year="1958">Cabrera, 1958:219</bibRefCitation>
</taxonomicName>
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. Part; not
<taxonomicName id="4C72029FBB63FFE3C4E40D52FB8BFB85" authority="J. A Wagner, 1855" authorityName="J. A. Wagner" authorityYear="1855" box="[665,1065,1056,1081]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="loricatus">
<emphasis id="B906A50EBB63FFE3C4E40D52FCC5FB85" box="[665,871,1056,1080]" italics="true" pageId="13" pageNumber="60">Cabassous loricatus</emphasis>
J.
<bibRefCitation id="EFE304EDBB63FFE3C5FE0D53FB8BFB85" author="Wagner, J. A." box="[899,1065,1056,1081]" pageId="13" pageNumber="60" refId="ref20141" refString="Wagner, J. A. (1855) Die Saugthiere in Abbildungen nach der Natur mit Beschreibungen von Dr. Johann Christian Daniel von Schreber. Supplementband. Funfte Abtheilung: Die Affen, Zahnlucker, Beutelthiere, Hufthiere, Insektenfresser und Handflugler. Supplementband 5. T. O. Weigel, Leipzig, 810 pp. https: // doi. org / 10.5962 / bhl. title. 67399" type="book" year="1855">A Wagner, 1855</bibRefCitation>
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</paragraph>
<paragraph id="8BCD791CBB63FFE3C6EA0D4DFB96FBEA" blockId="13.[151,1436,995,1172]" box="[151,1076,1087,1112]" pageId="13" pageNumber="60">
<treatmentCitationGroup id="AB625E32BB63FFE3C6EA0D4DFB96FBEA" box="[151,1076,1087,1112]" pageId="13" pageNumber="60">
<treatmentCitation id="0AD35F0DBB63FFE3C6EA0D4DFD93FBEA" author="Moeller, W." box="[151,561,1087,1111]" page="420" pageId="13" pageNumber="60" year="1968">
<taxonomicName id="4C72029FBB63FFE3C6EA0D4DFD93FBEA" authority=": Moeller, 1968: 420" authorityName="Moeller" authorityPageNumber="420" authorityYear="1968" box="[151,561,1087,1111]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="loricatus">
<emphasis id="B906A50EBB63FFE3C6EA0D4DFEC7FBEA" box="[151,357,1087,1111]" italics="true" pageId="13" pageNumber="60">Cabassous loricatus</emphasis>
:
<bibRefCitation id="EFE304EDBB63FFE3C70E0D4DFD93FBEA" author="Moeller, W." box="[371,561,1087,1111]" pageId="13" pageNumber="60" pagination="411 - 528" refId="ref17531" refString="Moeller, W. (1968) Allometrische Analyse der Gurteltierschadel, Ein Beitrag zur Phylogenie der Dasypodidae Bonaparte, 1838. Zoologische Jahrbuecher Anatomie, 85, 411 - 528." type="journal article" year="1968">Moeller, 1968:420</bibRefCitation>
</taxonomicName>
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. Part; not
<taxonomicName id="4C72029FBB63FFE3C4E50D4DFB92FBEA" authority="J. A. Wagner, 1855" authorityName="J. A. Wagner" authorityYear="1855" box="[664,1072,1087,1112]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="loricatus">
<emphasis id="B906A50EBB63FFE3C4E50D4DFCC4FBEA" box="[664,870,1087,1111]" italics="true" pageId="13" pageNumber="60">Cabassous loricatus</emphasis>
J. A.
<bibRefCitation id="EFE304EDBB63FFE3C5DD0D32FB92FBEA" author="Wagner, J. A." box="[928,1072,1087,1112]" pageId="13" pageNumber="60" refId="ref20141" refString="Wagner, J. A. (1855) Die Saugthiere in Abbildungen nach der Natur mit Beschreibungen von Dr. Johann Christian Daniel von Schreber. Supplementband. Funfte Abtheilung: Die Affen, Zahnlucker, Beutelthiere, Hufthiere, Insektenfresser und Handflugler. Supplementband 5. T. O. Weigel, Leipzig, 810 pp. https: // doi. org / 10.5962 / bhl. title. 67399" type="book" year="1855">Wagner, 1855</bibRefCitation>
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</paragraph>
<paragraph id="8BCD791CBB63FFE3C6EA0D2CFEA9FB29" blockId="13.[151,1436,995,1172]" pageId="13" pageNumber="60">
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<treatmentCitation id="0AD35F0DBB63FFE3C6EA0D2CFD9DFBCB" author="Wetzel, R. M." box="[151,575,1118,1142]" page="335" pageId="13" pageNumber="60" year="1980">
<taxonomicName id="4C72029FBB63FFE3C6EA0D2CFD9DFBCB" ID-CoL="P43M" authority="Wetzel, 1980: 335" authorityName="Wetzel" authorityPageNumber="335" authorityYear="1980" box="[151,575,1118,1142]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B906A50EBB63FFE3C6EA0D2CFED9FBCB" box="[151,379,1118,1142]" italics="true" pageId="13" pageNumber="60">Cabassous chacoensis</emphasis>
<bibRefCitation id="EFE304EDBB63FFE3C7FF0D2CFD9DFBCB" author="Wetzel, R. M." box="[386,575,1118,1142]" pageId="13" pageNumber="60" pagination="323 - 357" refId="ref20214" refString="Wetzel, R. M. (1980) Revision of the naked-tailed armadillos, genus Cabassous McMurtrie. Annals of Carnegie Museum, 49, 323 - 357." type="journal article" year="1980">Wetzel, 1980: 335</bibRefCitation>
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.
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locality: “
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, Depto.
<collectingRegion id="49B6B7FEBB63FFE3C5E10D2CFBEDFBCB" box="[924,1103,1118,1142]" country="Paraguay" name="Presidente Hayes" pageId="13" pageNumber="60">Presidente Hayes</collectingRegion>
,
<quantity id="4C8AD4F9BB63FFE3C2270D2CFB00FBCB" box="[1114,1186,1118,1142]" metricMagnitude="3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="13" pageNumber="60" unit="km" value="6.0" valueMax="7.0" valueMin="5.0">5-7 km</quantity>
W
<location id="8EAD2FC7BB63FFE3C2B50D2CFF5FFB29" LSID="urn:lsid:plazi:treatment:03DBC80ABB63FFE0C6EA0A0DFEB1FE6A:8EAD2FC7BB63FFE3C2B50D2CFF5FFB29" country="Paraguay" name="Estancia Juan de Zalazar" pageId="13" pageNumber="60" stateProvince="Presidente Hayes">Estancia Juan de Zalazar</location>
”.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCD791CBB63FFE3C6EA0DB1FDBAFAD0" blockId="13.[151,1437,1219,2037]" pageId="13" pageNumber="60">
<emphasis id="B906A50EBB63FFE3C6EA0DB1FF7EFB60" bold="true" box="[151,220,1219,1245]" pageId="13" pageNumber="60">Types</emphasis>
.
<materialsCitation id="3B1A7341BB63FFE3C6960DB1FDB6FAD0" ID-GBIF-Occurrence="3128119304" collectingDate="1974-07-28" collectionCode="UCM" country="Paraguay" location="It" municipality="Filadelfia" pageId="13" pageNumber="60" specimenCode="CM 67067, CONN 16982, USNM 531004" specimenCount="1" stateProvince="Boqueron" typeStatus="holotype">
The
<typeStatus id="54C9C7BEBB63FFE3C7630DB1FE22FB60" box="[286,384,1219,1245]" pageId="13" pageNumber="60" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBD4D167BB63FFE3C7F20DB1FDAEFB63" box="[399,524,1219,1246]" collectionCode="CM" country="China" name="Chongqing Museum" pageId="13" pageNumber="60">CM 67067</specimenCode>
) is an adult male collected on
<date id="FFCC5FDCBB63FFE3C5180DB1FC5BFB60" box="[869,1017,1219,1245]" pageId="13" pageNumber="60" value="1974-07-28">
<collectingDate id="EF88A634BB63FFE3C5180DB1FC5BFB60" box="[869,1017,1219,1245]" pageId="13" pageNumber="60" value="1974-07-28">28 July 1974</collectingDate>
</date>
, no collector listed, original numbers PWM 247 and
<collectionCode id="ED63E1D9BB63FFE3C73F0D95FE20FABC" box="[322,386,1255,1281]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="13" pageNumber="60" type="University or college">UCM</collectionCode>
[originally cited as CONN] 16891.
<location id="8EAD2FC7BB63FFE3C5680D95FC84FABC" LSID="urn:lsid:plazi:treatment:03DBC80ABB63FFE0C6EA0A0DFEB1FE6A:8EAD2FC7BB63FFE3C5680D95FC84FABC" box="[789,806,1255,1281]" country="Paraguay" municipality="Filadelfia" name="It" pageId="13" pageNumber="60" stateProvince="Boqueron">It</location>
consists of a stuffed skin, skull, and a carcass preserved in alcohol.
<bibRefCitation id="EFE304EDBB63FFE3C76A0C79FE1AFA98" author="Wetzel, R. M." box="[279,440,1291,1317]" pageId="13" pageNumber="60" pagination="323 - 357" refId="ref20214" refString="Wetzel, R. M. (1980) Revision of the naked-tailed armadillos, genus Cabassous McMurtrie. Annals of Carnegie Museum, 49, 323 - 357." type="journal article" year="1980">Wetzel (1980)</bibRefCitation>
listed
<specimenCount id="9D74B295BB63FFE3C47B0C79FD0BFA98" box="[518,681,1291,1317]" pageId="13" pageNumber="60" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
: an adult (
<specimenCode id="DBD4D167BB63FFE3C55D0C79FC60FA9B" box="[800,962,1291,1318]" collectionCode="CONN" country="USA" lsid="urn:lsid:biocol.org:col:15523" name="University of Connecticut" pageId="13" pageNumber="60" type="Herbarium">CONN 16982</specimenCode>
) female [erroneously cited as male in the original publication] collected in the type locality, catalog number PWM 246; and a male (
<specimenCode id="DBD4D167BB63FFE3C2060C5DFA88FAF4" box="[1147,1322,1326,1353]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="60" type="Museum">USNM 531004</specimenCode>
) collected in
<collectingMunicipality id="6BA9E366BB63FFE3C6C90C21FE80FAD0" box="[180,290,1363,1389]" pageId="13" pageNumber="60">Filadelfia</collectingMunicipality>
,
<collectingRegion id="49B6B7FEBB63FFE3C7500C21FE3CFAD0" box="[301,414,1363,1389]" country="Paraguay" name="Boqueron" pageId="13" pageNumber="60">Boquerón</collectingRegion>
,
<collectingCountry id="F365398CBB63FFE3C7D70C21FDB6FAD0" box="[426,532,1363,1389]" name="Paraguay" pageId="13" pageNumber="60">Paraguay</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3682A97BB63FFE3C6BA0C05FB66FA08" pageId="13" pageNumber="60" type="materials_examined">
<paragraph id="8BCD791CBB63FFE3C6BA0C05FB66FA08" blockId="13.[151,1437,1219,2037]" pageId="13" pageNumber="60">
<emphasis id="B906A50EBB63FFE3C6BA0C05FEC3FA2D" bold="true" box="[199,353,1398,1425]" pageId="13" pageNumber="60">Type locality</emphasis>
.
<materialsCitation id="3B1A7341BB63FFE3C7120C05FB62FA08" ID-GBIF-Occurrence="3128119305" country="Paraguay" location="Departament" municipality="Chaco" pageId="13" pageNumber="60" specimenCount="1" stateProvince="Presidente Hayes" typeStatus="holotype">
The
<typeStatus id="54C9C7BEBB63FFE3C7DE0C05FDA7FA2C" box="[419,517,1399,1425]" pageId="13" pageNumber="60" type="holotype">holotype</typeStatus>
was collected at “
<locationDeviation id="9A7F603EBB63FFE3C4AB0C04FB21FA2C" box="[726,1155,1398,1425]" location="Estancia Juan de Zalazar" pageId="13" pageNumber="60">
<quantity id="4C8AD4F9BB63FFE3C4AB0C04FC82FA2C" box="[726,800,1398,1425]" metricMagnitude="3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="13" pageNumber="60" unit="km" value="6.0" valueMax="7.0" valueMin="5.0">5-7km</quantity>
W of Estancia Juan de Zalazar
</locationDeviation>
,
<location id="8EAD2FC7BB63FFE3C2F20C05FA82FA2C" LSID="urn:lsid:plazi:treatment:03DBC80ABB63FFE0C6EA0A0DFEB1FE6A:8EAD2FC7BB63FFE3C2F20C05FA82FA2C" box="[1167,1312,1399,1425]" country="Paraguay" municipality="Chaco" name="Departament" pageId="13" pageNumber="60" stateProvince="Presidente Hayes">Departament</location>
<collectingRegion id="49B6B7FEBB63FFE3C3550C05FF42FA08" country="Paraguay" name="Presidente Hayes" pageId="13" pageNumber="60">Presidente Hayes</collectingRegion>
,
<collectingCountry id="F365398CBB63FFE3C6960CE9FEF4FA08" box="[235,342,1435,1461]" name="Paraguay" pageId="13" pageNumber="60">Paraguay</collectingCountry>
” (
<bibRefCitation id="EFE304EDBB63FFE3C7130CE9FDA2FA08" author="Wetzel, R. M." box="[366,512,1435,1461]" pageId="13" pageNumber="60" pagination="323 - 357" refId="ref20214" refString="Wetzel, R. M. (1980) Revision of the naked-tailed armadillos, genus Cabassous McMurtrie. Annals of Carnegie Museum, 49, 323 - 357." type="journal article" year="1980">Wetzel 1980</bibRefCitation>
), in an area of thorn forest and mixed grasses in the dry
<collectingMunicipality id="6BA9E366BB63FFE3C2080CE9FB62FA08" box="[1141,1216,1435,1461]" pageId="13" pageNumber="60">Chaco</collectingMunicipality>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3682A97BB63FFE0C6BA0CCCFA22FE9E" lastPageId="14" lastPageNumber="61" pageId="13" pageNumber="60" type="diagnosis">
<paragraph id="8BCD791CBB63FFE3C6BA0CCCFBC8F90F" blockId="13.[151,1437,1219,2037]" pageId="13" pageNumber="60">
<emphasis id="B906A50EBB63FFE3C6BA0CCCFE98FA65" bold="true" box="[199,314,1470,1496]" pageId="13" pageNumber="60">Diagnosis</emphasis>
. This species can be easily differentiated from other congeneric taxa by its much smaller ears (
<figureCitation id="13496599BB63FFE3C32C0CCDFF05FA43" captionStart="FIGURE 6" captionStartId="8.[151,250,1893,1917]" captionTargetBox="[173,1415,990,1863]" captionTargetId="figure-66@8.[168,1420,984,1870]" captionTargetPageId="8" captionText="FIGURE 6. Dorsal view of the head of Cabassous species. Note that C. chacoensis has much smaller ears in relation to its congeners and C. squamicaudis has a larger number of scutes on the cephalic shield. Images not in scale." figureDoi="http://doi.org/10.5281/zenodo.4773373" httpUri="https://zenodo.org/record/4773373/files/figure.png" pageId="13" pageNumber="60">Figure 6</figureCitation>
), limbs and sides of the body hairy, and its overall smaller size (
<tableCitation id="C6F04CA7BB63FFE3C5160C91FC60FA40" box="[875,962,1507,1533]" captionStart="TABLE 2" captionStartId="4.[151,239,947,971]" captionTargetBox="[163,1405,1016,1927]" captionTargetPageId="4" captionText="TABLE 2. Descriptive statistics (mean, standard deviation, minimum and maximum values and number of specimens) of the craniodental variables (in mm) for the Cabassous species. Mean ± SD/ (Min-Max) N." httpUri="http://table.plazi.org/id/DF0D2994BB6AFFEAC6EA0AC1FBCFFC55" pageId="13" pageNumber="60" tableUuid="DF0D2994BB6AFFEAC6EA0AC1FBCFFC55">Table 2</tableCitation>
). The scutes on the cephalic shield (45-52) exhibit a concentric organization with a large and central scute surrounded by smaller polygonal scutes (
<figureCitation id="13496599BB63FFE3C3510F75FA2CF99F" box="[1324,1422,1543,1570]" captionStart="FIGURE 6" captionStartId="8.[151,250,1893,1917]" captionTargetBox="[173,1415,990,1863]" captionTargetId="figure-66@8.[168,1420,984,1870]" captionTargetPageId="8" captionText="FIGURE 6. Dorsal view of the head of Cabassous species. Note that C. chacoensis has much smaller ears in relation to its congeners and C. squamicaudis has a larger number of scutes on the cephalic shield. Images not in scale." figureDoi="http://doi.org/10.5281/zenodo.4773373" httpUri="https://zenodo.org/record/4773373/files/figure.png" pageId="13" pageNumber="60">Figure 6</figureCitation>
). The cheek region is usually naked or with very small scutes. The carapace is brownish with 11-13 movable bands and hairy on its edges. The tail is overall naked with only sparse scutes and hairs.
<taxonomicName id="4C72029FBB63FFE3C24E0F3DFB6DF9D4" authorityName="Wetzel" authorityYear="1980" box="[1075,1231,1615,1641]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B906A50EBB63FFE3C24E0F3DFB6DF9D4" box="[1075,1231,1615,1641]" italics="true" pageId="13" pageNumber="60">C. chacoensis</emphasis>
</taxonomicName>
exhibits wide and marked curved zygomatic arcs, its teeth are anteroposteriorly constricted, making them wider than long. The body of the mandible is well curved and the ramus shows a more vertical profile (
<figureCitation id="13496599BB63FFE3C5960FE5FBFFF90F" box="[1003,1117,1687,1714]" captionStart="FIGURE 4" captionStartId="7.[151,250,1678,1702]" captionTargetBox="[178,1412,425,1642]" captionTargetId="figure-121@7.[163,1425,417,1654]" captionTargetPageId="7" captionText="FIGURE 4. Skulls and mandibles of Cabassous species. (a) C. squamicaudis (AMNH 133318). (b) C. unicinctus (AMNH 136256). (c) C. centralis (AMNH 10752). (d) C. chacoensis (skull: CM 67067, the holotype). (e) C. tatouay (skull: MCN-M 2109, mandible: CMNX 77)." figureDoi="http://doi.org/10.5281/zenodo.4773361" httpUri="https://zenodo.org/record/4773361/files/figure.png" pageId="13" pageNumber="60">Figure 4d</figureCitation>
).
</paragraph>
<paragraph id="8BCD791CBB63FFE3C6BA0FC8FB77F8FC" blockId="13.[151,1437,1219,2037]" pageId="13" pageNumber="60">
<emphasis id="B906A50EBB63FFE3C6BA0FC8FE4AF969" bold="true" box="[199,488,1722,1748]" pageId="13" pageNumber="60">Geographic distribution</emphasis>
.
<taxonomicName id="4C72029FBB63FFE3C7880FC9FD52F968" authorityName="Wetzel" authorityYear="1980" box="[501,752,1723,1749]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B906A50EBB63FFE3C7880FC9FD52F968" box="[501,752,1723,1749]" italics="true" pageId="13" pageNumber="60">Cabassous chacoensis</emphasis>
</taxonomicName>
occurs from western
<collectingCountry id="F365398CBB63FFE3C59D0FC9FBEAF968" box="[992,1096,1723,1749]" name="Paraguay" pageId="13" pageNumber="60">Paraguay</collectingCountry>
to central
<collectingCountry id="F365398CBB63FFE3C2C10FC9FA8CF968" box="[1212,1326,1723,1749]" name="Argentina" pageId="13" pageNumber="60">Argentina</collectingCountry>
, comprising mainly the Gran
<collectingRegion id="49B6B7FEBB63FFE3C7000FADFDB5F944" box="[381,535,1759,1785]" country="Argentina" name="Chaco" pageId="13" pageNumber="60">Chaco region</collectingRegion>
. However, given that the species is rarely recorded (
<bibRefCitation id="EFE304EDBB63FFE3C2200FADFB51F944" author="Chebez, J. C." box="[1117,1267,1759,1785]" pageId="13" pageNumber="60" refId="ref14065" refString="Chebez, J. C. (1994) Los que se van. Tomo 3. Editorial Albatros, Buenos Aires, 336 pp." type="book" year="1994">Chebez 1994</bibRefCitation>
), its range may be larger than it is currently known. For example, some isolated records in
<collectingCountry id="F365398CBB63FFE3C5A70E71FBEBF8A0" box="[986,1097,1795,1821]" name="Argentina" pageId="13" pageNumber="60">Argentina</collectingCountry>
also include the Yungas (
<bibRefCitation id="EFE304EDBB63FFE3C31B0E71FEF3F8FC" author="Cirignoli, S. &amp; Abba, A. M. &amp; Tamburini, D. M. &amp; Decarre, J. &amp; Castro, L. B. &amp; Superina, M. &amp; Rogel, T. G. &amp; Aguero, A. J. &amp; Albrecht, C. D." pageId="13" pageNumber="60" pagination="2019" refId="ref14200" refString="Cirignoli, S., Abba, A. M., Tamburini, D. M., Decarre, J., Castro, L. B., Superina, M., Rogel, T. G., Aguero, A. J. &amp; Albrecht, C. D. (2019) Cabassous chacoensis. Categorizacion 2019 de los mamiferos de Argentina segun su riesgo de extincion. In: SayDS-SAREM (Eds.), Lista Roja de los mamiferos de Argentina. https: // doi. org / 10.31687 / SaremLR. 19.044" type="book chapter" year="2019">
Cirignoli
<emphasis id="B906A50EBB63FFE3C6A50E55FEB2F8FC" box="[216,272,1831,1857]" italics="true" pageId="13" pageNumber="60">et al.</emphasis>
2019
</bibRefCitation>
) and montane grassland High Monte in
<collectingRegion id="49B6B7FEBB63FFE3C56E0E55FCD6F8FC" box="[787,884,1831,1857]" country="Argentina" name="La Rioja" pageId="13" pageNumber="60">La Rioja</collectingRegion>
(Monguillot &amp; Miatello 2010).
</paragraph>
<paragraph id="8BCD791CBB63FFE0C6BA0E39FA22FE9E" blockId="13.[151,1437,1219,2037]" lastBlockId="14.[151,1436,157,471]" lastPageId="14" lastPageNumber="61" pageId="13" pageNumber="60">
<bibRefCitation id="EFE304EDBB63FFE3C6BA0E39FEC5F8D8" author="Wetzel, R. M." box="[199,359,1867,1893]" pageId="13" pageNumber="60" pagination="323 - 357" refId="ref20214" refString="Wetzel, R. M. (1980) Revision of the naked-tailed armadillos, genus Cabassous McMurtrie. Annals of Carnegie Museum, 49, 323 - 357." type="journal article" year="1980">Wetzel (1980)</bibRefCitation>
listed
<collectingCountry id="F365398CBB63FFE3C7CE0E39FE55F8D8" box="[435,503,1867,1893]" name="Brazil" pageId="13" pageNumber="60">Brazil</collectingCountry>
(“
<collectingRegion id="49B6B7FEBB63FFE3C46C0E39FD09F8D8" box="[529,683,1867,1893]" country="Brazil" name="Mato Grosso" pageId="13" pageNumber="60">Mato Grosso</collectingRegion>
”) as part of the distribution of
<taxonomicName id="4C72029FBB63FFE3C2770E39FAA4F8D8" authorityName="Wetzel" authorityYear="1980" box="[1034,1286,1867,1893]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B906A50EBB63FFE3C2770E39FAA4F8D8" box="[1034,1286,1867,1893]" italics="true" pageId="13" pageNumber="60">Cabassous chacoensis</emphasis>
</taxonomicName>
, but this was based on a specimen obtained from the Buenos Aires Zoo in 1904. Since then, no additional record has been reported, leading several authors to not include it as part of the Brazilian fauna (
<bibRefCitation id="EFE304EDBB63FFE3C2770EE1FB7FF810" author="Wetzel, R. M. &amp; Gardner, A. L. &amp; Redford, K. H. &amp; Eisenberg, J. F." box="[1034,1245,1939,1965]" pageId="13" pageNumber="60" pagination="128 - 156" refId="ref20304" refString="Wetzel, R. M., Gardner, A. L., Redford, K. H. &amp; Eisenberg, J. F. (2008) Order Cingulata. In: Gardner, A. L. (Ed.), Mammals of South America. Marsupials, Xenarthrans, Shrews and Bats. Vol. 1. University of Chicago Press, Chicago, Illinois, pp. 128 - 156. https: // doi. org / 10.7208 / chicago / 9780226282428.001.0001" type="book chapter" year="2008">
Wetzel
<emphasis id="B906A50EBB63FFE3C21C0EE1FB38F810" box="[1121,1178,1939,1965]" italics="true" pageId="13" pageNumber="60">et al.</emphasis>
2008
</bibRefCitation>
,
<bibRefCitation id="EFE304EDBB63FFE3C2940EE1FA3AF810" author="Hayssen, V." box="[1257,1432,1939,1965]" pageId="13" pageNumber="60" pagination="24 - 27" refId="ref16226" refString="Hayssen, V. (2014 b) Cabassous chacoensis (Cingulata: Dasypodidae). Mammalian Species, 46 (908), 24 - 27. https: // doi. org / 10.1644 / 908" type="journal article" year="2014">Hayssen 2014b</bibRefCitation>
,
<bibRefCitation id="EFE304EDBB63FFE3C6EA0EC5FEDEF86C" author="Brandao, M. V. &amp; Garbino, G. S. T. &amp; Semedo, T. B. F. &amp; Feijo, A. &amp; Nascimento, F. O. &amp; Fernandes-Ferreira, H. &amp; Rossi, R. V. &amp; Dalponte, J. &amp; Carmignotto, A. P." box="[151,380,1975,2001]" pageId="13" pageNumber="60" pagination="263 - 307" refId="ref13759" refString="Brandao, M. V., Garbino, G. S. T., Semedo, T. B. F., Feijo, A., Nascimento, F. O., Fernandes-Ferreira, H., Rossi, R. V., Dalponte, J. &amp; Carmignotto, A. P. (2019) Mammals of Mato Grosso, Brazil: annotated species list and historical review. Mastozoologia Neotropical, 26 (2), 263 - 307. https: // doi. org / 10.31687 / saremMN. 19.26.2.0.03" type="journal article" year="2019">
Brandão
<emphasis id="B906A50EBB63FFE3C6820EC5FE9BF86C" box="[255,313,1975,2001]" italics="true" pageId="13" pageNumber="60">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation id="EFE304EDBB63FFE3C7F50EC5FDF7F86C" author="Abreu, E. F. &amp; Casali, D. M. &amp; Garbino, G. S. T. &amp; Loretto, D. &amp; Loss, A. C. &amp; Marmontel, M. &amp; Nascimento, M. C. &amp; Oliveira, M. L. &amp; Pavan, S. E. &amp; Tirelli, F. P." box="[392,597,1975,2001]" pageId="13" pageNumber="60" refId="ref13039" refString="Abreu, E. F., Casali, D. M., Garbino, G. S. T., Loretto, D., Loss, A. C., Marmontel, M., Nascimento, M. C., Oliveira, M. L., Pavan, S. E. &amp; Tirelli, F. P. (2020) Lista de Mamiferos do Brasil. Comite de Taxonomia da Sociedade Brasileira de Mastozoologia (CT-SBMz). Available from: https: // www. sbmz. org / mamiferos-do-brasil / (accessed 26 March 2021)" type="url" year="2020">
Abreu
<emphasis id="B906A50EBB63FFE3C7AA0EC5FDABF86C" box="[471,521,1975,2001]" italics="true" pageId="13" pageNumber="60">et al</emphasis>
. 2020
</bibRefCitation>
;
<bibRefCitation id="EFE304EDBB63FFE3C41C0EC5FCE8F86C" author="Quintela, F. M. &amp; Rosa, C. A. &amp; Feijo, A." box="[609,842,1975,2001]" pageId="13" pageNumber="60" pagination="20191004" refId="ref18027" refString="Quintela, F. M., Rosa, C. A. &amp; Feijo, A. (2020) Updated and annotated checklist of recent mammals from Brazil. Anais da Academia Brasileira de Ciencias, 92 (Supplement 2), e 20191004. https: // doi. org / 10.1590 / 0001 - 3765202020191004" type="journal article" year="2020">
Quintela
<emphasis id="B906A50EBB63FFE3C4B70EC5FCA6F86C" box="[714,772,1975,2001]" italics="true" pageId="13" pageNumber="60">et al.</emphasis>
2020
</bibRefCitation>
). Nevertheless,
<taxonomicName id="4C72029FBB63FFE3C5830EC5FB39F86C" authorityName="Wetzel" authorityYear="1980" box="[1022,1179,1975,2001]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B906A50EBB63FFE3C5830EC5FB39F86C" box="[1022,1179,1975,2001]" italics="true" pageId="13" pageNumber="60">C. chacoensis</emphasis>
</taxonomicName>
has been registered in
<collectingCountry id="F365398CBB63FFE3C6EA0EA9FF5DF848" box="[151,255,2011,2037]" name="Paraguay" pageId="13" pageNumber="60">Paraguay</collectingCountry>
about
<quantity id="4C8AD4F9BB63FFE3C7360EA9FE02F848" box="[331,416,2011,2037]" metricMagnitude="5" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="60" unit="km" value="220.0">220 km</quantity>
from the Brazilian border. It is therefore an open question whether the species went locally extinct, has gone unnoticed for over a century, or never occurred in
<collectingCountry id="F365398CBB60FFE0C5DF09EFFC48FF0A" box="[930,1002,157,183]" name="Brazil" pageId="14" pageNumber="61">Brazil</collectingCountry>
; in this latter case the
<collectingCountry id="F365398CBB60FFE0C29009EFFAF7FF0A" box="[1261,1365,157,183]" name="Paraguay" pageId="14" pageNumber="61">Paraguay</collectingCountry>
River might represent the barrier between the two countries. On the other hand, the absence of
<taxonomicName id="4C72029FBB60FFE0C2F509B3FA86FF66" authorityName="Wetzel" authorityYear="1980" box="[1160,1316,193,219]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B906A50EBB60FFE0C2F509B3FA86FF66" box="[1160,1316,193,219]" italics="true" pageId="14" pageNumber="61">C. chacoensis</emphasis>
</taxonomicName>
in
<collectingCountry id="F365398CBB60FFE0C33709B3FA3EFF66" box="[1354,1436,193,219]" name="Bolivia" pageId="14" pageNumber="61">Bolivia</collectingCountry>
might reflect the lack of studies on this genus given that the dry Chaco extends northward to the Bolivian east portion (
<figureCitation id="13496599BB60FFE0C6AC087BFE91FE9E" box="[209,307,265,291]" captionStart="FIGURE 8" captionStartId="11.[151,250,1282,1306]" captionTargetBox="[222,1356,181,1257]" captionTargetId="figure-17@11.[222,1366,181,1257]" captionTargetPageId="11" captionText="FIGURE 8. Distribution of the five living species of Cabassous in the Neotropical region. Question marks represent unconfirmed occurrence." figureDoi="http://doi.org/10.5281/zenodo.4773380" httpUri="https://zenodo.org/record/4773380/files/figure.png" pageId="14" pageNumber="61">Figure 8</figureCitation>
). Among all specimens of
<taxonomicName id="4C72029FBB60FFE0C422087BFD7AFE9E" authorityName="McMurtrie" authorityYear="1831" box="[607,728,265,291]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="14" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="B906A50EBB60FFE0C422087BFD7AFE9E" box="[607,728,265,291]" italics="true" pageId="14" pageNumber="61">Cabassous</emphasis>
</taxonomicName>
examined in this study, only three were collected in
<collectingCountry id="F365398CBB60FFE0C35B087BFADEFE9E" box="[1318,1404,265,291]" name="Bolivia" pageId="14" pageNumber="61">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3682A97BB60FFE0C6BA085FFEB1FE6A" pageId="14" pageNumber="61" type="discussion">
<paragraph id="8BCD791CBB60FFE0C6BA085FFEB1FE6A" blockId="14.[151,1436,157,471]" pageId="14" pageNumber="61">
<emphasis id="B906A50EBB60FFE0C6BA085FFE96FEFA" bold="true" box="[199,308,301,327]" pageId="14" pageNumber="61">Remarks</emphasis>
.
<bibRefCitation id="EFE304EDBB60FFE0C73F085FFDF5FEFA" author="Hutterer, R. &amp; Peters, G." box="[322,599,301,327]" pageId="14" pageNumber="61" pagination="3 - 27" refId="ref16422" refString="Hutterer, R. &amp; Peters, G. (2010) Type specimens of mammals (Mammalia) in the collections of the Zoologisches Forschungsmuseum Alexander Koenig, Bonn. Bonn Zoological Bulletin, 59 (1 - 2), 3 - 27." type="journal article" year="2010">Hutterer &amp; Peters (2010)</bibRefCitation>
listed an adult female (ZFMK 60.317, field number 448) housed at the Zoologisches Forschungsmuseum Alexander Koenig, Bonn,
<collectingCountry id="F365398CBB60FFE0C5590823FC2EFED6" box="[804,908,337,363]" name="Germany" pageId="14" pageNumber="61">Germany</collectingCountry>
as a
<typeStatus id="54C9C7BEBB60FFE0C5BA0823FB85FED6" box="[967,1063,337,363]" pageId="14" pageNumber="61" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C72029FBB60FFE0C2330823FB49FED6" authorityName="Wetzel" authorityYear="1980" box="[1102,1259,337,363]" class="Mammalia" family="Dasypodidae" genus="Cabassous" kingdom="Animalia" order="Cingulata" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="chacoensis">
<emphasis id="B906A50EBB60FFE0C2330823FB49FED6" box="[1102,1259,337,363]" italics="true" pageId="14" pageNumber="61">C. chacoensis</emphasis>
</taxonomicName>
. This specimen is represented by a skin and skull and was collected in Filadelfia,
<collectingRegion id="49B6B7FEBB60FFE0C50C0807FC40FE32" box="[881,994,373,399]" country="Paraguay" name="Boqueron" pageId="14" pageNumber="61">Boqueron</collectingRegion>
,
<collectingCountry id="F365398CBB60FFE0C5900807FBF7FE32" box="[1005,1109,373,399]" name="Paraguay" pageId="14" pageNumber="61">Paraguay</collectingCountry>
by J. Unger on
<date id="FFCC5FDCBB60FFE0C37B0807FF70FE09" pageId="14" pageNumber="61" value="1958-12-10">10 December 1958</date>
. However, although
<bibRefCitation id="EFE304EDBB60FFE0C7C508EBFDF4FE0E" author="Wetzel, R. M." box="[440,598,409,435]" pageId="14" pageNumber="61" pagination="323 - 357" refId="ref20214" refString="Wetzel, R. M. (1980) Revision of the naked-tailed armadillos, genus Cabassous McMurtrie. Annals of Carnegie Museum, 49, 323 - 357." type="journal article" year="1980">Wetzel (1980)</bibRefCitation>
listed this individual among the material examined, he did not include it as a
<typeStatus id="54C9C7BEBB60FFE0C6D608CFFEADFE6A" box="[171,271,445,471]" pageId="14" pageNumber="61" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>