163 lines
19 KiB
XML
163 lines
19 KiB
XML
<document id="DB2C50CE49BB21512BE7DFEE78071EF7" ID="10.11646/zootaxa.4144.4.5" ID-DOI="10.11646/zootaxa.4144.4.5" ID-GBIF-Dataset="9bbc5f70-a78f-4ef8-946e-11077c8bbb46" ID-ISSN="1175-5326" ID-Zenodo-Dep="266732" ID-ZooBank="DE8B262C-1465-4998-88AD-392880B06912" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469773920918" checkinUser="plazi" docAuthor="Couri, Márcia & Pont, Adrian" docDate="2016" docId="03DB87F3FFE0FFECFF39E3D50815F8F6" docLanguage="en" docName="zootaxa.4144.4.5.pdf" docOrigin="Zootaxa 4144 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenosia kilembana Emden 1940, syn. nov." docType="treatment" docVersion="7" lastPageNumber="538" masterDocId="FFE2FF8BFFE9FFE5FFAEE75C0C13FF86" masterDocTitle="Species of Coenosia Meigen (Diptera, Muscidae) described by Fritz van Emden from the British Museum Ruwenzori Expedition of 1934 – 1935" masterLastPageNumber="555" masterPageNumber="529" pageNumber="538" updateTime="1698366539694" updateUser="plazi">
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<mods:title id="A5FB01C3C4BCA36AAF8293F7424C2CB3">Species of Coenosia Meigen (Diptera, Muscidae) described by Fritz van Emden from the British Museum Ruwenzori Expedition of 1934 – 1935</mods:title>
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<mods:namePart id="C5D410047CEAE677490053CB228280F0">Couri, Márcia</mods:namePart>
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<mods:name id="0852C2BE7DC20FB41868D139CF601339" type="personal">
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<mods:namePart id="EA42594545D77CAE17A3FE1FB62F2CD4">Pont, Adrian</mods:namePart>
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<mods:title id="914AB2B2CAAF4FEAC218DAA73F3A06DE">Zootaxa</mods:title>
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<mods:part id="AD9B03493CDB02C24F134B05C86F0FEF">
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<mods:date id="0055421901E202DA4613F65EF3E88B2D">2016</mods:date>
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<mods:number id="37EDD9869296815932D465BC41D379CB">4144</mods:number>
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<mods:number id="7ABDC8D6FDBE3962ABBA3309480BB9AC">4</mods:number>
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<treatment id="03DB87F3FFE0FFECFF39E3D50815F8F6" ID-DOI="http://doi.org/10.5281/zenodo.6074783" ID-GBIF-Taxon="123514569" ID-Zenodo-Dep="6074783" LSID="urn:lsid:plazi:treatment:03DB87F3FFE0FFECFF39E3D50815F8F6" httpUri="http://treatment.plazi.org/id/03DB87F3FFE0FFECFF39E3D50815F8F6" lastPageNumber="538" pageId="9" pageNumber="538">
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<subSubSection id="C368656EFFE0FFECFF39E3D5089BFB42" pageId="9" pageNumber="538" type="nomenclature">
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<paragraph id="8BCD36E5FFE0FFECFF39E3D50E9BFB25" blockId="9.[151,1160,1161,1221]" box="[151,648,1161,1187]" pageId="9" pageNumber="538">
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<heading id="D0858189FFE0FFECFF39E3D50E9BFB25" bold="true" box="[151,648,1161,1187]" fontSize="11" level="1" pageId="9" pageNumber="538" reason="1">
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<treatmentCitationGroup id="AB6211CBFFE0FFECFF39E3D50E9BFB25" box="[151,648,1161,1187]" pageId="9" pageNumber="538">
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<taxonomicName id="4C724D66FFE0FFECFF39E3D50E9BFB25" ID-CoL="92P38" authority="Emden, 1940: 216" authorityName="Emden" authorityPageNumber="216" authorityYear="1940" box="[151,648,1161,1187]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="kilembana">
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<emphasis id="B906EAF7FFE0FFECFF39E3D50E9BFB25" bold="true" box="[151,648,1161,1187]" pageId="9" pageNumber="538">
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<emphasis id="B906EAF7FFE0FFECFF39E3D50D9EFB25" bold="true" box="[151,397,1161,1187]" italics="true" pageId="9" pageNumber="538">Coenosia kilembana</emphasis>
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(
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<treatmentCitation id="0AD310F4FFE0FFECFE33E3D50E93FB25" author="Emden" box="[413,640,1161,1187]" page="216" pageId="9" pageNumber="538" year="1940">
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<bibRefCitation id="EFE34B14FFE0FFECFE33E3D50E93FB25" author="Emden" box="[413,640,1161,1187]" pageId="9" pageNumber="538" refString="Emden, F. I. van (1940) Muscidae: B - Coenosiinae. In: Ruwenzori Expedition 1934 - 35. Vol. 2. British Museum (Natural History), London, pp. 91 - 255." type="book chapter" year="1940">Emden, 1940: 216</bibRefCitation>
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</treatmentCitation>
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)
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</emphasis>
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</taxonomicName>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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<paragraph id="8BCD36E5FFE0FFECFF39E3F0089BFB42" blockId="9.[151,1160,1161,1221]" box="[151,1160,1196,1221]" pageId="9" pageNumber="538">
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(
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<figureCitation id="13492A60FFE0FFECFF31E3F00D2FFB43" box="[159,316,1196,1221]" captionStart="FIGURES 101 – 112" captionStartId="18.[151,264,1912,1935]" captionTargetBox="[197,1391,206,1879]" captionTargetId="figure@18.[183,1404,193,1891]" captionTargetPageId="18" captionText="FIGURES 101 – 112. Sternite 5; cercal plate and surstyli in dorsal view; epandrium, cercal plate, surstylus and aedeagus in lateral view: 101 – 103. Coenosia niveifrons Stein. 104 – 106. Coenosia ruwenzorica Speiser. (syn: Coenosia kilembana syn. nov.). 107 – 109. Coenosia morio (Emden). 110 – 112. Coenosia somereni (Emden)." httpUri="https://zenodo.org/record/266741/files/figure.png" pageId="9" pageNumber="538">Figs 104–106</figureCitation>
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, as
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<taxonomicName id="4C724D66FFE0FFECFECBE3F00DE3FB42" box="[357,496,1196,1220]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="B906EAF7FFE0FFECFECBE3F00DE3FB42" box="[357,496,1196,1220]" italics="true" pageId="9" pageNumber="538">C. rebmanni</emphasis>
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</taxonomicName>
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) (junior synonym of
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<taxonomicName id="4C724D66FFE0FFECFD4EE3F0081FFB42" authority="Speiser, 1924" authorityName="Speiser" authorityYear="1924" box="[736,1036,1196,1221]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="B906EAF7FFE0FFECFD4EE3F00F78FB42" box="[736,875,1196,1220]" italics="true" pageId="9" pageNumber="538">C. rebmanni</emphasis>
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Speiser, 1924
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</taxonomicName>
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,
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<emphasis id="B906EAF7FFE0FFECFBB9E3F0086CFB42" bold="true" box="[1047,1151,1196,1220]" pageId="9" pageNumber="538">
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<taxonomicNameLabel id="A235578CFFE0FFECFBB9E3F0086CFB42" box="[1047,1151,1196,1220]" pageId="9" pageNumber="538">syn. nov.</taxonomicNameLabel>
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</emphasis>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C368656EFFE0FFECFF39E3A80E42FA8B" box="[151,593,1268,1293]" pageId="9" pageNumber="538" type="materials_examined">
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<paragraph id="8BCD36E5FFE0FFECFF39E3A80E42FA8B" blockId="9.[151,1436,1268,1905]" box="[151,593,1268,1293]" pageId="9" pageNumber="538">Holotype male seen; in good condition.</paragraph>
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</subSubSection>
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<subSubSection id="C368656EFFE0FFECFF69E24B0EC7F9F2" pageId="9" pageNumber="538" type="diagnosis">
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<paragraph id="8BCD36E5FFE0FFECFF69E24B0EC7F9F2" blockId="9.[151,1436,1268,1905]" pageId="9" pageNumber="538">
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<emphasis id="B906EAF7FFE0FFECFF69E24B0D52FAB6" bold="true" box="[199,321,1303,1328]" pageId="9" pageNumber="538">Diagnosis.</emphasis>
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General colour brown with grey pollinosity; scutum brown; postpronotum and pleura grey dusted; frons dark, fronto-orbital plate weakly grey pollinose; face, parafacial and gena silver from certain angles; palpus light brown; fore coxa yellow or partially infuscated, mid and hind coxae partially grey pollinose; femora yellow on basal two-thirds, remainder of legs brown; the narrowest point of the fronto-facial stripe below the base of the antenna; antennae inserted at dorsal third of eye; postpedicel long, about six times the length of pedicel; arista short-haired, bare on apical third; scutellum with both pairs of setae present and long, with setulae on disc; anterior and lower katepisternals about two-thirds the length of the long posterior seta; haltere yellow; lower calypter about 2.5 times as long as upper one; fore tibia with a very long posteroventral seta; hind tarsomeres 1–3 moderately to distinctly compressed; hind tibia with one anterior and one long anterodorsal inserted at almost the same level, the anterior one much shorter; sternite 5 as in
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<figureCitation id="13492A60FFE0FFECFDDEE1000EC3F9F2" box="[624,720,1628,1653]" captionStart="FIGURES 101 – 112" captionStartId="18.[151,264,1912,1935]" captionTargetBox="[197,1391,206,1879]" captionTargetId="figure@18.[183,1404,193,1891]" captionTargetPageId="18" captionText="FIGURES 101 – 112. Sternite 5; cercal plate and surstyli in dorsal view; epandrium, cercal plate, surstylus and aedeagus in lateral view: 101 – 103. Coenosia niveifrons Stein. 104 – 106. Coenosia ruwenzorica Speiser. (syn: Coenosia kilembana syn. nov.). 107 – 109. Coenosia morio (Emden). 110 – 112. Coenosia somereni (Emden)." httpUri="https://zenodo.org/record/266741/files/figure.png" pageId="9" pageNumber="538">Fig. 104</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C368656EFFE0FFECFF69E1DC0815F8F6" pageId="9" pageNumber="538" type="description">
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<paragraph id="8BCD36E5FFE0FFECFF69E1DC08C3F91F" blockId="9.[151,1436,1268,1905]" box="[199,1232,1664,1689]" pageId="9" pageNumber="538">
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<emphasis id="B906EAF7FFE0FFECFF69E1DC0D68F91E" box="[199,379,1664,1688]" italics="true" pageId="9" pageNumber="538">Male terminalia</emphasis>
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. Cercal plate and surstylus as in
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<figureCitation id="13492A60FFE0FFECFD43E1DC0F5EF91F" box="[749,845,1664,1689]" captionStart="FIGURES 101 – 112" captionStartId="18.[151,264,1912,1935]" captionTargetBox="[197,1391,206,1879]" captionTargetId="figure@18.[183,1404,193,1891]" captionTargetPageId="18" captionText="FIGURES 101 – 112. Sternite 5; cercal plate and surstyli in dorsal view; epandrium, cercal plate, surstylus and aedeagus in lateral view: 101 – 103. Coenosia niveifrons Stein. 104 – 106. Coenosia ruwenzorica Speiser. (syn: Coenosia kilembana syn. nov.). 107 – 109. Coenosia morio (Emden). 110 – 112. Coenosia somereni (Emden)." httpUri="https://zenodo.org/record/266741/files/figure.png" pageId="9" pageNumber="538">Fig. 105</figureCitation>
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. Aedeagal complex as in
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<figureCitation id="13492A60FFE0FFECFBC5E1DC08DFF91F" box="[1131,1228,1664,1689]" captionStart="FIGURES 101 – 112" captionStartId="18.[151,264,1912,1935]" captionTargetBox="[197,1391,206,1879]" captionTargetId="figure@18.[183,1404,193,1891]" captionTargetPageId="18" captionText="FIGURES 101 – 112. Sternite 5; cercal plate and surstyli in dorsal view; epandrium, cercal plate, surstylus and aedeagus in lateral view: 101 – 103. Coenosia niveifrons Stein. 104 – 106. Coenosia ruwenzorica Speiser. (syn: Coenosia kilembana syn. nov.). 107 – 109. Coenosia morio (Emden). 110 – 112. Coenosia somereni (Emden)." httpUri="https://zenodo.org/record/266741/files/figure.png" pageId="9" pageNumber="538">Fig. 106</figureCitation>
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.
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</paragraph>
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<paragraph id="8BCD36E5FFE0FFECFF69E1F80815F8F6" blockId="9.[151,1436,1268,1905]" pageId="9" pageNumber="538">
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<emphasis id="B906EAF7FFE0FFECFF69E1F80D02F93A" bold="true" box="[199,273,1700,1724]" pageId="9" pageNumber="538">Notes.</emphasis>
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Included in the
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<taxonomicName id="4C724D66FFE0FFECFE67E1F80E20F93A" box="[457,563,1700,1724]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="B906EAF7FFE0FFECFE67E1F80E20F93A" box="[457,563,1700,1724]" italics="true" pageId="9" pageNumber="538">rebmanni</emphasis>
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</taxonomicName>
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group.
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<bibRefCitation id="EFE34B14FFE0FFECFD25E1F80F3FF93A" author="Emden" box="[651,812,1700,1725]" pageId="9" pageNumber="538" refString="Emden, F. I. van (1940) Muscidae: B - Coenosiinae. In: Ruwenzori Expedition 1934 - 35. Vol. 2. British Museum (Natural History), London, pp. 91 - 255." type="book chapter" year="1940">Emden (1940)</bibRefCitation>
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stated that there are probably only two valid species in this “group”—
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<taxonomicName id="4C724D66FFE0FFECFE93E1940E9EF966" authority="Emden, 1940" authorityName="Emden" authorityYear="1940" box="[317,653,1736,1761]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="translucida">
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<emphasis id="B906EAF7FFE0FFECFE93E1940DCDF966" box="[317,478,1736,1760]" italics="true" pageId="9" pageNumber="538">C. translucida</emphasis>
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(
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<bibRefCitation id="EFE34B14FFE0FFECFE42E1940E97F966" author="Emden" box="[492,644,1736,1761]" pageId="9" pageNumber="538" refString="Emden, F. I. van (1940) Muscidae: B - Coenosiinae. In: Ruwenzori Expedition 1934 - 35. Vol. 2. British Museum (Natural History), London, pp. 91 - 255." type="book chapter" year="1940">Emden, 1940</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="4C724D66FFE0FFECFD6AE1940F5CF966" box="[708,847,1736,1760]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="B906EAF7FFE0FFECFD6AE1940F5CF966" box="[708,847,1736,1760]" italics="true" pageId="9" pageNumber="538">C. rebmanni</emphasis>
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</taxonomicName>
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– and that the other species are probably subspecies of
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<taxonomicName id="4C724D66FFE0FFECFF1BE1B00D52F882" box="[181,321,1772,1796]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="B906EAF7FFE0FFECFF1BE1B00D52F882" box="[181,321,1772,1796]" italics="true" pageId="9" pageNumber="538">C. rebmanni</emphasis>
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</taxonomicName>
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.
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<bibRefCitation id="EFE34B14FFE0FFECFEE1E1B10DC6F882" author="Pont" box="[335,469,1772,1796]" pageId="9" pageNumber="538" refString="Pont, A. C. (1980) Family Muscidae. In: Crosskey, R. W. (Ed.), Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, pp. 721 - 761." type="book chapter" year="1980">Pont (1980)</bibRefCitation>
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considered this species as a subspecies of
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<taxonomicName id="4C724D66FFE0FFECFC18E1B00852F882" box="[950,1089,1772,1796]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="B906EAF7FFE0FFECFC18E1B00852F882" box="[950,1089,1772,1796]" italics="true" pageId="9" pageNumber="538">C. rebmanni</emphasis>
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</taxonomicName>
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. We dissected a male
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<typeStatus id="54C98847FFE0FFECFA95E1B1098FF883" box="[1339,1436,1773,1797]" pageId="9" pageNumber="538" type="paratype">paratype</typeStatus>
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of
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<taxonomicName id="4C724D66FFE0FFECFF17E04D0D41F8AE" box="[185,338,1808,1832]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="kilembana">
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<emphasis id="B906EAF7FFE0FFECFF17E04D0D41F8AE" box="[185,338,1808,1832]" italics="true" pageId="9" pageNumber="538">C. kilembana</emphasis>
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</taxonomicName>
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and a male of
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<taxonomicName id="4C724D66FFE0FFECFDA7E04D0E84F8AE" box="[521,663,1808,1832]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="B906EAF7FFE0FFECFDA7E04D0E84F8AE" box="[521,663,1808,1832]" italics="true" pageId="9" pageNumber="538">C. rebmanni</emphasis>
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</taxonomicName>
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and the terminalia are exactly the same. On the other hand, other species of this group that we dissected (
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<taxonomicName id="4C724D66FFE0FFECFDCEE0680EE0F8CA" box="[608,755,1844,1868]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="ciliventris">
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<emphasis id="B906EAF7FFE0FFECFDCEE0680EE0F8CA" box="[608,755,1844,1868]" italics="true" pageId="9" pageNumber="538">C. ciliventris</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C724D66FFE0FFECFC82E0680FDDF8CA" box="[812,974,1844,1868]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="translucida">
|
||
<emphasis id="B906EAF7FFE0FFECFC82E0680FDDF8CA" box="[812,974,1844,1868]" italics="true" pageId="9" pageNumber="538">C. translucida</emphasis>
|
||
</taxonomicName>
|
||
) have very characteristic terminalia. We are proposing the new synonymy of
|
||
<taxonomicName id="4C724D66FFE0FFECFD9EE0050ED6F8F6" box="[560,709,1880,1904]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="kilembana">
|
||
<emphasis id="B906EAF7FFE0FFECFD9EE0050ED6F8F6" box="[560,709,1880,1904]" italics="true" pageId="9" pageNumber="538">C. kilembana</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C724D66FFE0FFECFCABE0050F9CF8F6" box="[773,911,1880,1904]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="rebmanni">
|
||
<emphasis id="B906EAF7FFE0FFECFCABE0050F9CF8F6" box="[773,911,1880,1904]" italics="true" pageId="9" pageNumber="538">C. rebmanni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B906EAF7FFE0FFECFC30E0040815F8F6" bold="true" box="[926,1030,1880,1904]" pageId="9" pageNumber="538">
|
||
<taxonomicNameLabel id="A235578CFFE0FFECFC30E0040815F8F6" box="[926,1030,1880,1904]" pageId="9" pageNumber="538">syn. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |